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<document id="625C39093B3C671613C4E9F49B7BCBE9" ID-CLB-Dataset="55466" ID-DOI="10.5281/zenodo.5169525" ID-GBIF-Dataset="4910761a-706e-4f53-a5a8-6c72379bdbc6" ID-ISSN="1942-1354" ID-Zenodo-Dep="5169525" ID-ZooBank="DA6CBFE5-927E-45B6-9D05-69AC97AF7B76" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628277966057" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2017" docId="03DA133F7517B1334BBCFA9789586D51" docLanguage="en" docName="InsectaMundi.0592.1-16.pdf" docOrigin="Insecta Mundi 2017 (592)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Agnelius nageli Paulsen 2017, new combination" docType="treatment" docVersion="7" lastPageNumber="13" masterDocId="FFE36B47751BB13D4B02FFA38A0B6E57" masterDocTitle="Revision of the endemic Madagascan stag beetle genus Ganelius Benesh, and description of a new, related genus (Coleoptera: Lucanidae: Lucaninae: Figulini)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="11" updateTime="1699056540890" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="6CE1FD7EB3D5FBE55FB54743B1C33AEA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DD32A5141A7AD8FB819D5E3CC435C3CE">
<mods:title id="4EA18653A26D1D487B98E18019BB7BC3">Revision of the endemic Madagascan stag beetle genus Ganelius Benesh, and description of a new, related genus (Coleoptera: Lucanidae: Lucaninae: Figulini)</mods:title>
</mods:titleInfo>
<mods:name id="EF83AFCD91A03B6C024A8E5E2FE5CA50" type="personal">
<mods:role id="EB62B5F384CAD2FDBE485808B50A9826">
<mods:roleTerm id="E860AE20B91A36C0EB5792DF446F1FF2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="19DAB116159E38FBAE2D9C32E62A395D">Paulsen, M. J.</mods:namePart>
<mods:affiliation id="E19C858017B9272B2490E50411B10AC1">Systematic Research Collections University of Nebraska State Museum W 436 Nebraska Hall Lincoln, NE 68588 - 0546</mods:affiliation>
<mods:nameIdentifier id="1822E1271743F6B10713B8DA17606A22" type="email">mjpaulsen@unl.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="DBBE05EC902F844BAEC5938E6262DA82">text</mods:typeOfResource>
<mods:relatedItem id="35A35DC140C8705F748B5BACF9BEFEA9" type="host">
<mods:titleInfo id="FB261F21F4EC9283AB0960DC04838EDD">
<mods:title id="10B7F1D4C7D336AEE7429058DA455B39">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="9B6701F07F465713E760EB71566CA895">
<mods:date id="BCFCED6B6456CD892A47EEA9BA14423A">2017</mods:date>
<mods:detail id="2DDB7EB4A6F807F835E241DC6564EEBD" type="pubDate">
<mods:number id="675F3515EB1723429D4DBC8016B192D3">2017-12-29</mods:number>
</mods:detail>
<mods:detail id="185B31EC16F6708D135283042AD130F2" type="volume">
<mods:number id="4CD53FF0DC0B50F338D6C4B58F622106">2017</mods:number>
</mods:detail>
<mods:detail id="09415003BC9103F2622A6BDBB4FE4FA0" type="issue">
<mods:number id="18CEF562818B7D2A4161E3D6D4981054">592</mods:number>
</mods:detail>
<mods:extent id="758721292DF696393F1CF37348721340" unit="page">
<mods:start id="00579B697D57F9D24E798BD08E7559DE">1</mods:start>
<mods:end id="36307E64E4F8F55134015675C6DA9BA3">16</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2EF44699E06BF6688ACDBF7E275F6712">journal article</mods:classification>
<mods:identifier id="79D0E0B80B37DAB94E453302E88A703B" type="CLB-Dataset">55466</mods:identifier>
<mods:identifier id="5A2B7CB52AB45F6952284710A7A9E9D7" type="DOI">10.5281/zenodo.5169525</mods:identifier>
<mods:identifier id="7F5CD379D495EEFBDDBE95ED15917A6E" type="GBIF-Dataset">4910761a-706e-4f53-a5a8-6c72379bdbc6</mods:identifier>
<mods:identifier id="90C8FAF9CF89C412926C1E660F545B40" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="CF1EDABBA500D7DFE1001A6AEB6372E4" type="Zenodo-Dep">5169525</mods:identifier>
<mods:identifier id="9ACBD0F81C035B5694785E42DAE0C9C2" type="ZooBank">DA6CBFE5-927E-45B6-9D05-69AC97AF7B76</mods:identifier>
</mods:mods>
<treatment id="03DA133F7517B1334BBCFA9789586D51" ID-DOI="http://doi.org/10.5281/zenodo.6567182" ID-GBIF-Taxon="195537731" ID-Zenodo-Dep="6567182" LSID="urn:lsid:plazi:treatment:03DA133F7517B1334BBCFA9789586D51" httpUri="http://treatment.plazi.org/id/03DA133F7517B1334BBCFA9789586D51" lastPageId="14" lastPageNumber="13" pageId="12" pageNumber="11">
<subSubSection id="C369F1A27517B1314BBCFA9789276B1B" box="[190,812,1332,1357]" pageId="12" pageNumber="11" type="nomenclature">
<paragraph id="8BCCA2297517B1314BBCFA9789276B1B" blockId="12.[190,812,1332,1357]" box="[190,812,1332,1357]" pageId="12" pageNumber="11">
<heading id="D08415457517B1314BBCFA9789276B1B" bold="true" box="[190,812,1332,1357]" fontSize="10" level="2" pageId="12" pageNumber="11" reason="3">
<emphasis id="B9077E3B7517B1314BBCFA9789276B1B" bold="true" box="[190,812,1332,1357]" pageId="12" pageNumber="11">
<taxonomicName id="4C73D9AA7517B1314BBCFA9788256B1B" ID-CoL="65T6Y" authority="(Kriesche)" authorityName="Paulsen" authorityYear="2017" baseAuthorityName="Kriesche" baseAuthorityYear="2017" box="[190,558,1332,1356]" class="Insecta" family="Lucanidae" genus="Agnelius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="nageli" status="comb. nov.">
<emphasis id="B9077E3B7517B1314BBCFA978B9C6B1B" bold="true" box="[190,407,1332,1356]" italics="true" pageId="12" pageNumber="11">Agnelius nageli</emphasis>
(Kriesche)
</taxonomicName>
,
<taxonomicNameLabel id="A234C3407517B131493FFA9689276B1B" box="[573,812,1332,1357]" pageId="12" pageNumber="11" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C369F1A27517B1314BC2FAD489836BD8" box="[192,904,1399,1423]" pageId="12" pageNumber="11" type="reference_group">
<paragraph id="8BCCA2297517B1314BC2FAD489836BD8" blockId="12.[192,904,1399,1423]" box="[192,904,1399,1423]" pageId="12" pageNumber="11">
<taxonomicName id="4C73D9AA7517B1314BC2FAD488646BD8" ID-CoL="73XVQ" authority="Kriesche 1926: 385" authorityName="Kriesche" authorityPageNumber="385" authorityYear="1926" box="[192,623,1399,1423]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="nageli">
<emphasis id="B9077E3B7517B1314BC2FAD48B746BD8" box="[192,383,1399,1423]" italics="true" pageId="12" pageNumber="11">Nigidius nageli</emphasis>
Kriesche 1926: 385
</taxonomicName>
(original combination)
</paragraph>
</subSubSection>
<subSubSection id="C369F1A27517B1314BC2FA18886E6957" pageId="12" pageNumber="11" type="materials_examined">
<paragraph id="8BCCA2297517B1314BC2FA1888926860" blockId="12.[192,1440,1466,1591]" pageId="12" pageNumber="11">
<emphasis id="B9077E3B7517B1314BC2FA188B836B85" bold="true" box="[192,392,1466,1491]" pageId="12" pageNumber="11">Type material.</emphasis>
<materialsCitation id="3B1BA8747517B1314A8EFA1888926860" ID-GBIF-Occurrence="3776282309" collectionCode="MNHN" collectorName="R. Oberthur" country="Madagascar" determinerName="M. J. Paulsen" location="Perrot Freres" pageId="12" pageNumber="11" specimenCount="1" specimenCount-male="1" stateProvince="Antsianaka" typeStatus="neotype">
<typeStatus id="54C81C8B7517B1314A8EFA188BF96B84" box="[396,498,1467,1491]" pageId="12" pageNumber="11" type="neotype">Neotype</typeStatus>
male,
<collectionCode id="ED623AEC7517B1314943FA1888946B85" box="[577,671,1467,1490]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="11">MNHN</collectionCode>
,
<emphasis id="B9077E3B7517B13149ABFA18898E6B85" bold="true" box="[681,901,1466,1491]" pageId="12" pageNumber="11">here designated</emphasis>
, labeled: a) “
<collectingCountry id="F364E2B97517B1314F20FA198EB36B85" box="[1058,1208,1466,1490]" pageId="12" pageNumber="11">Madagascar</collectingCountry>
/
<collectingRegion id="49B76CCB7517B1314FC8FA198F526B85" box="[1226,1369,1466,1490]" pageId="12" pageNumber="11">Antsianaka</collectingRegion>
/
<location id="8EACF4F27517B1314E6EFA188B306BA3" LSID="urn:lsid:plazi:treatment:03DA133F7517B1334BBCFA9789586D51:8EACF4F27517B1314E6EFA188B306BA3" country="Madagascar" name="Perrot Freres" pageId="12" pageNumber="11" stateProvince="Antsianaka">Perrot Frères</location>
/ 2º Semestre 1890”; b) on light blue paper, “Museum Paris / 1952 / Coll.
<collectorName id="2686C7FF7517B1314FBAFA7F8F5B6BA3" box="[1208,1360,1500,1524]" pageId="12" pageNumber="11">R. Oberthur</collectorName>
”; c) on red paper, “
<taxonomicName id="4C73D9AA7517B1314A50FA5E88196842" authorityName="Kriesche" authorityYear="1926" box="[338,530,1533,1557]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="nageli">
<emphasis id="B9077E3B7517B1314A50FA5E88196842" box="[338,530,1533,1557]" italics="true" pageId="12" pageNumber="11">Nigidius nageli</emphasis>
</taxonomicName>
/
<specimenCount id="9D7569A07517B1314929FA5D88346842" box="[555,575,1534,1557]" pageId="12" pageNumber="11" type="male">♂</specimenCount>
Kriesche /
<typeStatus id="54C81C8B7517B13149CEFA5D89586841" box="[716,851,1534,1558]" pageId="12" pageNumber="11" type="neotype">NEOTYPE</typeStatus>
/ des. MJ Paulsen”; d) “
<taxonomicName id="4C73D9AA7517B1314F7FFA5E8EE36842" authorityName="Paulsen" authorityYear="2017" box="[1149,1256,1533,1557]" class="Insecta" family="Lucanidae" genus="Agnelius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7517B1314F7FFA5E8EE36842" box="[1149,1256,1533,1557]" italics="true" pageId="12" pageNumber="11">Agnelius</emphasis>
</taxonomicName>
/
<taxonomicName id="4C73D9AA7517B1314E03FA5E8F466842" authorityName="Kriesche" authorityYear="1926" box="[1281,1357,1533,1557]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="nageli">
<emphasis id="B9077E3B7517B1314E03FA5E8F466842" box="[1281,1357,1533,1557]" italics="true" pageId="12" pageNumber="11">nageli</emphasis>
</taxonomicName>
/ (Kriesche, 1926) / det.
<determinerName id="7E104EED7517B1314AA0F9BC88486860" box="[418,579,1567,1591]" pageId="12" pageNumber="11">M.J. Paulsen</determinerName>
2017”.
</materialsCitation>
</paragraph>
<paragraph id="8BCCA2297517B1314BC2F9C1886E6957" blockId="12.[192,1440,1634,1792]" pageId="12" pageNumber="11">
<emphasis id="B9077E3B7517B1314BC2F9C18BBD682D" bold="true" box="[192,438,1634,1658]" pageId="12" pageNumber="11">Non-type material</emphasis>
(
<figureCitation id="1348BEAC7517B1314AC6F9C1881E682D" box="[452,533,1634,1658]" captionStart="Figures 33–37" captionStartId="17.[192,298,1697,1721]" captionTargetBox="[233,1386,399,1638]" captionTargetId="figure-36@17.[192,1440,364,1666]" captionTargetPageId="17" captionText="Figures 33–37. Species distributions. 33) Ganelius madagascariensis. 34) G. oberndorferi. 35) G. gnamptus. 36) G. zombi. 37) Agnelius nageli." figureDoi="http://doi.org/10.5281/zenodo.5169541" httpUri="https://zenodo.org/record/5169541/files/figure.png" pageId="12" pageNumber="11">Fig. 37</figureCitation>
).
<materialsCitation id="3B1BA8747517B1314928F9C18F30682D" ID-GBIF-Occurrence="3776282329" box="[554,1339,1634,1658]" collectionCode="LBC" collectorName="Ambatofitorahana" location="FIANARANTSOA" pageId="12" pageNumber="11" specimenCount="1" stateProvince="Fianarantsoa">
<emphasis id="B9077E3B7517B1314928F9C18908682D" bold="true" box="[554,771,1634,1658]" pageId="12" pageNumber="11">MADAGASCAR:</emphasis>
<collectingRegion id="49B76CCB7517B131480AF9C189E2682D" box="[776,1001,1634,1658]" country="Madagascar" name="Fianarantsoa" pageId="12" pageNumber="11">FIANARANTSOA</collectingRegion>
:
<collectorName id="2686C7FF7517B13148F3F9C18EC4682D" box="[1009,1231,1634,1658]" pageId="12" pageNumber="11">Ambatofitorahana</collectorName>
(1;
<collectionCode id="ED623AEC7517B1314FFAF9C18F38682D" box="[1272,1331,1634,1658]" pageId="12" pageNumber="11">LBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1BA8747517B1314E47F9C18B9468CC" ID-GBIF-Occurrence="3776282315" collectionCode="MNHN" pageId="12" pageNumber="11" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa">
Ambositra (
<specimenCount id="9D7569A07517B1314A03F9208B2068CC" box="[257,299,1667,1691]" pageId="12" pageNumber="11" type="male">1 ♂</specimenCount>
;
<collectionCode id="ED623AEC7517B1314A3BF9208B9368CC" box="[313,408,1667,1691]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="11">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1BA8747517B1314AAEF92088D868CC" ID-GBIF-Occurrence="3776282306" box="[428,723,1667,1691]" collectionCode="CASC" location="Talakately" pageId="12" pageNumber="11" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa">
<location id="8EACF4F27517B1314AAEF920883A68CC" LSID="urn:lsid:plazi:treatment:03DA133F7517B1334BBCFA9789586D51:8EACF4F27517B1314AAEF920883A68CC" box="[428,561,1667,1691]" name="Talakately" pageId="12" pageNumber="11" stateProvince="Fianarantsoa">Talakately</location>
(
<specimenCount id="9D7569A07517B1314946F920886468CC" box="[580,623,1667,1691]" pageId="12" pageNumber="11" type="male">1 ♂</specimenCount>
;
<collectionCode id="ED623AEC7517B131497EF92088C068CC" box="[636,715,1667,1691]" pageId="12" pageNumber="11">CASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1BA8747517B13149DDF920880468EA" ID-GBIF-Occurrence="3776282304" collectionCode="MNHN" county="Vondrozo" location="Lake Alaotra" municipality="Antsianaka" pageId="12" pageNumber="11" specimenCount="19" specimenCount-female="6" specimenCount-male="13" stateProvince="Toamasina">
<collectingCounty id="62ADDAA57517B13149DDF920895868CC" box="[735,851,1667,1691]" pageId="12" pageNumber="11">Vondrozo</collectingCounty>
(
<specimenCount id="9D7569A07517B1314861F920899A68CC" box="[867,913,1667,1691]" pageId="12" pageNumber="11" type="male">2 ♂</specimenCount>
;
<collectionCode id="ED623AEC7517B131489CF92089F568CC" box="[926,1022,1667,1691]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="11">MNHN</collectionCode>
).
<collectingRegion id="49B76CCB7517B1314F13F9208ECB68CC" box="[1041,1216,1667,1691]" country="Madagascar" name="Toamasina" pageId="12" pageNumber="11">TOAMASINA</collectingRegion>
:
<collectingMunicipality id="6BA838537517B1314FCEF9208F5268CC" box="[1228,1369,1667,1691]" pageId="12" pageNumber="11">Antsianaka</collectingMunicipality>
/
<location id="8EACF4F27517B1314E63F9208B1668EA" LSID="urn:lsid:plazi:treatment:03DA133F7517B1334BBCFA9789586D51:8EACF4F27517B1314E63F9208B1668EA" county="Vondrozo" municipality="Antsianaka" name="Lake Alaotra" pageId="12" pageNumber="11" stateProvince="Toamasina">Lake Alaotra</location>
(
<specimenCount id="9D7569A07517B1314A2FF9068B6D68EB" box="[301,358,1701,1725]" pageId="12" pageNumber="11" type="male">11 ♂</specimenCount>
,
<specimenCount id="9D7569A07517B1314A70F9068B9768EB" box="[370,412,1701,1725]" pageId="12" pageNumber="11" type="female">6 ♀</specimenCount>
,
<collectionCode id="ED623AEC7517B1314AAAF906880C68EA" box="[424,519,1701,1725]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="11">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1BA8747517B1314918F90688F568EA" ID-GBIF-Occurrence="3776282325" box="[538,766,1701,1725]" collectionCode="LBC" collectorName="Beparasy" pageId="12" pageNumber="11" specimenCount="1" stateProvince="Toamasina">
<collectorName id="2686C7FF7517B1314918F906888568EA" box="[538,654,1701,1725]" pageId="12" pageNumber="11">Beparasy</collectorName>
(1;
<collectionCode id="ED623AEC7517B13149B8F90688FD68EA" box="[698,758,1701,1725]" pageId="12" pageNumber="11">LBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1BA8747517B131480BF9068E5168EA" ID-GBIF-Occurrence="3776282330" box="[777,1114,1701,1725]" collectionCode="MNHN" location="Fanovana" pageId="12" pageNumber="11" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Toamasina">
<location id="8EACF4F27517B131480BF906898F68EB" LSID="urn:lsid:plazi:treatment:03DA133F7517B1334BBCFA9789586D51:8EACF4F27517B131480BF906898F68EB" box="[777,900,1701,1724]" name="Fanovana" pageId="12" pageNumber="11" stateProvince="Toamasina">Fanovana</location>
(
<specimenCount id="9D7569A07517B1314891F90689BD68EA" box="[915,950,1701,1725]" pageId="12" pageNumber="11" type="male">1♂</specimenCount>
,
<specimenCount id="9D7569A07517B13148C1F90689ED68EA" box="[963,998,1701,1725]" pageId="12" pageNumber="11" type="female">1♀</specimenCount>
;
<collectionCode id="ED623AEC7517B13148F1F9068E5968EA" box="[1011,1106,1701,1725]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="11">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1BA8747517B1314F67F9068B2C6889" ID-GBIF-Occurrence="3776282343" collectionCode="CASC, MNHN" location="Fenerive" pageId="12" pageNumber="11" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Toamasina">
<location id="8EACF4F27517B1314F67F9068ED968EA" LSID="urn:lsid:plazi:treatment:03DA133F7517B1334BBCFA9789586D51:8EACF4F27517B1314F67F9068ED968EA" box="[1125,1234,1701,1725]" name="Fenerive" pageId="12" pageNumber="11" stateProvince="Toamasina">Fenerive</location>
(
<specimenCount id="9D7569A07517B1314FE2F9068F0768EB" box="[1248,1292,1701,1725]" pageId="12" pageNumber="11" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D7569A07517B1314E15F9068F4A68EB" box="[1303,1345,1701,1725]" pageId="12" pageNumber="11" type="female">1 ♀</specimenCount>
;
<collectionCode id="ED623AEC7517B1314E4CF9068F9768EA" box="[1358,1436,1701,1725]" pageId="12" pageNumber="11">CASC</collectionCode>
,
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)
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;
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<location id="8EACF4F27517B1314A37F9648BEA6889" LSID="urn:lsid:plazi:treatment:03DA133F7517B1334BBCFA9789586D51:8EACF4F27517B1314A37F9648BEA6889" box="[309,481,1735,1758]" name="Maroantsetra" pageId="12" pageNumber="11" stateProvince="Toamasina">Maroantsetra</location>
, (
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;
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<location id="8EACF4F27517B13149A9F965890D6889" LSID="urn:lsid:plazi:treatment:03DA133F7517B1334BBCFA9789586D51:8EACF4F27517B13149A9F965890D6889" box="[683,774,1734,1758]" name="Perinet" pageId="12" pageNumber="11" stateProvince="Toamasina">Perinet</location>
(
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,
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;
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)
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;
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<collectorName id="2686C7FF7517B1314F02F9648E426888" box="[1024,1097,1735,1759]" pageId="12" pageNumber="11">Rogez</collectorName>
(
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,
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,
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;
<materialsCitation id="3B1BA8747517B1314BC2F94B886A6957" ID-GBIF-Occurrence="3776282301" box="[192,609,1768,1792]" collectionCode="MNHN" collectorName="Tamatave &amp; Toamasina" pageId="12" pageNumber="11" specimenCount="1" stateProvince="Toamasina">
<collectorName id="2686C7FF7517B1314BC2F94B8B316957" box="[192,314,1768,1792]" pageId="12" pageNumber="11">Tamatave</collectorName>
/
<collectorName id="2686C7FF7517B1314A46F94B8BC76957" box="[324,460,1768,1792]" pageId="12" pageNumber="11">Toamasina</collectorName>
(3;
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="C369F1A27517B1314BC2F88888C169D1" pageId="12" pageNumber="11" type="diagnosis">
<paragraph id="8BCCA2297517B1314BC2F88888C169D1" blockId="12.[191,1440,1835,1926]" pageId="12" pageNumber="11">
<emphasis id="B9077E3B7517B1314BC2F8888B5B6914" bold="true" box="[192,336,1835,1859]" pageId="12" pageNumber="11">Diagnosis.</emphasis>
This species is immediately separable from other Madagascan stag beetles by the characters discussed above. It is the only species with mandibles with a bifurcate median tooth (
<figureCitation id="1348BEAC7517B1314E04F8EF8F566933" box="[1286,1373,1868,1892]" captionStart="Figures 28–32" captionStartId="16.[192,302,1675,1699]" captionTargetBox="[230,1426,1030,1622]" captionTargetId="figure-48@16.[192,1440,1020,1634]" captionTargetPageId="16" captionText="Figures 28–32. Comparison of ocular canthi, females. 28) Ganelius madagascariensis. 29) G. oberndorferi. 30) G. gnamptus. 31) G. zombi. 32) Agnelius nageli." figureDoi="http://doi.org/10.5281/zenodo.5169539" httpUri="https://zenodo.org/record/5169539/files/figure.png" pageId="12" pageNumber="11">Fig. 32</figureCitation>
). The flattened, elongate form is also distinctive.
</paragraph>
</subSubSection>
<caption id="DF0CF2A17516B1304BC2F8F0894669D2" ID-DOI="http://doi.org/10.5281/zenodo.5169531" ID-Zenodo-Dep="5169531" httpUri="https://zenodo.org/record/5169531/files/figure.png" pageId="13" pageNumber="12" startId="13.[192,288,1875,1897]" targetBox="[207,1409,327,1742]" targetPageId="13">
<paragraph id="8BCCA2297516B1304BC2F8F0894669D2" blockId="13.[192,1441,1875,1926]" pageId="13" pageNumber="12">
<emphasis id="B9077E3B7516B1304BC2F8F08B7A693F" bold="true" box="[192,369,1875,1897]" pageId="13" pageNumber="12">Figures 17–20.</emphasis>
<taxonomicName id="4C73D9AA7516B1304A7AF8F088EF693E" authority="(Kriesche, 1926)" authorityName="Paulsen" authorityYear="2017" baseAuthorityName="Kriesche" baseAuthorityYear="2017" box="[376,740,1875,1897]" class="Insecta" family="Lucanidae" genus="Agnelius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="nageli">
<emphasis id="B9077E3B7516B1304A7AF8F0882F693F" box="[376,548,1875,1896]" italics="true" pageId="13" pageNumber="12">Agnelius nageli</emphasis>
(Kriesche, 1926)
</taxonomicName>
.
<emphasis id="B9077E3B7516B13049EDF8F0891F693E" bold="true" box="[751,788,1875,1897]" pageId="13" pageNumber="12">17)</emphasis>
Male neotype, dorsal habitus.
<emphasis id="B9077E3B7516B1304F6FF8F08E9A693E" bold="true" box="[1133,1169,1875,1897]" pageId="13" pageNumber="12">18)</emphasis>
Female, dorsal habitus.
<emphasis id="B9077E3B7516B1304BC2F8CC8AEE69D2" bold="true" box="[192,229,1903,1925]" pageId="13" pageNumber="12">19)</emphasis>
Neotype labels.
<emphasis id="B9077E3B7516B1304A9EF8CC8BCA69D2" bold="true" box="[412,449,1903,1925]" pageId="13" pageNumber="12">20)</emphasis>
Male genitalia, flagellum, &lt;20 mm.
</paragraph>
</caption>
<subSubSection id="C369F1A27515B1334BC2FF6189586D51" pageId="14" pageNumber="13" type="discussion">
<paragraph id="8BCCA2297515B1334BC2FF61895A6F6B" blockId="14.[190,1442,194,774]" pageId="14" pageNumber="13">
<emphasis id="B9077E3B7515B1334BC2FF618B4F6E8D" bold="true" box="[192,324,194,218]" pageId="14" pageNumber="13">Remarks.</emphasis>
<bibRefCitation id="EFE2DFD87515B1334A4FFF6188CB6E8D" author="Bartolozzi, L. &amp; K. Werner" box="[333,704,194,218]" pageId="14" pageNumber="13" refId="ref7286" refString="Bartolozzi, L., and K. Werner. 2004. Illustrated catalogue of the Lucanidae from Africa and Madagascar (Coleoptera: Lucanidae). Taita Publishers; Hradec Kralove, Czechia. 191 p." type="book" year="2004">Bartolozzi and Werner (2004)</bibRefCitation>
indicated that the
<typeStatus id="54C81C8B7515B13348B6FF6189E26E8D" box="[948,1001,194,218]" pageId="14" pageNumber="13">type</typeStatus>
of
<taxonomicName id="4C73D9AA7515B1334F17FF618F1A6E8D" authority="(Kriesche)" authorityName="Paulsen" authorityYear="2017" baseAuthorityName="Kriesche" baseAuthorityYear="2017" box="[1045,1297,194,218]" class="Insecta" family="Lucanidae" genus="Agnelius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="nageli">
<emphasis id="B9077E3B7515B1334F17FF618E8C6E8D" box="[1045,1159,194,218]" italics="true" pageId="14" pageNumber="13">G. nageli</emphasis>
(Kriesche)
</taxonomicName>
, which was originally deposited in Hannover,
<collectingCountry id="F364E2B97515B133496CFF4188EF6EAD" box="[622,740,226,250]" name="Germany" pageId="14" pageNumber="13">Germany</collectingCountry>
, is presumably lost. The stag beetle primary
<typeStatus id="54C81C8B7515B1334E1FFF408F546EAC" box="[1309,1375,227,251]" pageId="14" pageNumber="13">types</typeStatus>
from Hannover were taken to
<collectingRegion id="49B76CCB7515B1334AF9FEA0887A6F4C" box="[507,625,259,283]" country="Germany" name="Hamburg" pageId="14" pageNumber="13">Hamburg</collectingRegion>
for study by Paul Nagel (A. Eichler, NLMH, pers. comm.) where they were destroyed during WWII (
<bibRefCitation id="EFE2DFD87515B1334977FE87894F6F6B" author="Klapperich, H." box="[629,836,292,316]" pageId="14" pageNumber="13" pagination="67" refId="ref7497" refString="Klapperich, H. 1948. Notes about German and Austrian Coleopterists and collections. Coleopterists Bulletin 2: 67." type="journal article" year="1948">Klapperich 1948</bibRefCitation>
).
</paragraph>
<paragraph id="8BCCA2297515B1334BF2FEE78EDE6CD4" blockId="14.[190,1442,194,774]" pageId="14" pageNumber="13">
Although Kriesche’s
<typeStatus id="54C81C8B7515B1334AFAFEE7886A6F0B" box="[504,609,324,348]" pageId="14" pageNumber="13" type="holotype">holotype</typeStatus>
is lost, when comparing the available material with his original description, his species
<taxonomicName id="4C73D9AA7515B1334ADDFEC588446F2A" authorityName="Paulsen" authorityYear="2017" baseAuthorityName="Kriesche" baseAuthorityYear="2017" box="[479,591,357,381]" class="Insecta" family="Lucanidae" genus="Agnelius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="nageli">
<emphasis id="B9077E3B7515B1334ADDFEC588446F2A" box="[479,591,357,381]" italics="true" pageId="14" pageNumber="13">N. nageli</emphasis>
</taxonomicName>
can be distinguished. I had originally considered the species in question to be an undescribed species of
<taxonomicName id="4C73D9AA7515B133497DFE2588E66FC9" authorityName="Benesh" authorityYear="1955" box="[639,749,390,414]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7515B133497DFE2588E66FC9" box="[639,749,390,414]" italics="true" pageId="14" pageNumber="13">Ganelius</emphasis>
</taxonomicName>
from the eastern Madagascan forests that oddly lacked visible sexual dimorphism. The species was illustrated by Benesh (1955) as
<emphasis id="B9077E3B7515B1334F6CFE048F8B6FE8" box="[1134,1408,423,447]" italics="true" pageId="14" pageNumber="13">
<taxonomicName id="4C73D9AA7515B1334F6CFE048F706FE8" authorityName="Nonfried" authorityYear="1892" box="[1134,1403,423,447]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="oberndorferi">Nigidius oberndorferi</taxonomicName>
,
</emphasis>
in part (Benesh’s fig. 15), and in
<bibRefCitation id="EFE2DFD87515B1334927FE6488C86FB7" author="Fujita, H." box="[549,707,455,480]" pageId="14" pageNumber="13" refId="ref7468" refString="Fujita, H. 2010. The lucanid beetles of the world. Mushi-Sha's iconographic series of insects, 6. Mushi- Sha; Tokyo. 472 p." type="book" year="2010">Fujita (2010)</bibRefCitation>
as “
<taxonomicName id="4C73D9AA7515B13349F1FE64896A6F88" authorityName="Benesh" authorityYear="1955" box="[755,865,455,479]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7515B13349F1FE64896A6F88" box="[755,865,455,479]" italics="true" pageId="14" pageNumber="13">Ganelius</emphasis>
</taxonomicName>
? spp.” (species 1279). Characteristics described by Kriesche that cannot be ignored are the fitting description of the canthus, mentum with a buttonlike elevation, shiny and finely punctate frons, and especially the more elongate form. Most examined specimens are identical to Kriesche’s values of
<quantity id="4C8B0FCC7515B133480CFD89896F6C16" box="[782,868,554,578]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="13" unit="mm" value="19.0">19 mm</quantity>
in length (including mandibles) and
<quantity id="4C8B0FCC7515B1334E35FD898F756C16" box="[1335,1406,554,578]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="13" unit="mm" value="6.0">6 mm</quantity>
in width. Kriesche indicated that the specimen was female, which would be expected for a species that lacks sexual dimorphism in a time when genitalic dissections were rarely performed.
</paragraph>
<paragraph id="8BCCA2297515B1334BF2FD2F89586D51" blockId="14.[190,1442,194,774]" pageId="14" pageNumber="13">
A
<typeStatus id="54C81C8B7515B1334A0BFD2F8B626CF3" box="[265,361,652,676]" pageId="14" pageNumber="13" type="neotype">neotype</typeStatus>
is designated to tie the name to this species and eliminate confusion about its previously unknown identity. Kriesche’s original locality was simply ‘Madagaskar’ and a female indicated, however without genitalic dissection the lack of sexual dimorphism makes the sex equivocal; a
<typeStatus id="54C81C8B7515B1334FFFFD6D8F566CB1" box="[1277,1373,718,742]" pageId="14" pageNumber="13" type="neotype">neotype</typeStatus>
male is chosen from the MNHN material from Antsianaka.
</paragraph>
</subSubSection>
</treatment>
</document>
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