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<document id="972B745E52B058E46C7DFFB051FD16D1" ID-CLB-Dataset="5666" ID-DOI="10.11646/zootaxa.3785.3.7" ID-GBIF-Dataset="8eddca4b-08a5-433a-8e3a-c89fb483818e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4913363" ID-ZooBank="D6E3D055-4390-4DC2-A9AC-0DBC40C2740F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623170138633" checkinUser="felipe" docAuthor="Neubauer, Thomas A., Harzhauser, Mathias, Georgopoulou, Elisavet, Mandic, Oleg &amp; Kroh, Andreas" docDate="2014" docId="03A7D746F074FFA6FF3DFADDFAE0F81C" docLanguage="en" docName="zt03785p468.pdf" docOrigin="Zootaxa 3785 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrobia alpha Jekelius 1944" docType="treatment" docVersion="6" lastPageNumber="459" masterDocId="FF9EAF3EF072FFA0FFAAFFE8FFBBFF9A" masterDocTitle="Replacement names and nomenclatural comments for problematic species-group names in Europe's Neogene freshwater Gastropoda" masterLastPageNumber="468" masterPageNumber="453" pageNumber="459" updateTime="1699005021158" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A0BBA13309746491CECA935DEB10C920">Replacement names and nomenclatural comments for problematic species-group names in Europe's Neogene freshwater Gastropoda</mods:title>
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<mods:namePart id="F4646C2C45791E85CD6EBB8A1432FB06">Neubauer, Thomas A.</mods:namePart>
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<mods:namePart id="D830FE4E7F5817985996A3093E465367">Harzhauser, Mathias</mods:namePart>
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<mods:namePart id="3D2B562F2FAEC57EB7978B31DF5C6E36">Georgopoulou, Elisavet</mods:namePart>
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<mods:namePart id="0F2C0ED014A777DFE4462298821B9E14">Mandic, Oleg</mods:namePart>
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<mods:namePart id="CF1E8D1E5E5D16AB49DEC8B21DBBFDC8">Kroh, Andreas</mods:namePart>
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<mods:title id="1A4C1678102F716931EE4A75F2FC22AA">Zootaxa</mods:title>
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<treatment id="03A7D746F074FFA6FF3DFADDFAE0F81C" ID-DOI="http://doi.org/10.5281/zenodo.4913381" ID-GBIF-Taxon="183399406" ID-Zenodo-Dep="4913381" LSID="urn:lsid:plazi:treatment:03A7D746F074FFA6FF3DFADDFAE0F81C" httpUri="http://treatment.plazi.org/id/03A7D746F074FFA6FF3DFADDFAE0F81C" lastPageNumber="459" pageId="6" pageNumber="459">
<subSubSection id="C31435DBF074FFA6FF3DFADDFDB1FAD5" box="[151,522,1333,1359]" pageId="6" pageNumber="459" type="nomenclature">
<paragraph id="8BB16650F074FFA6FF3DFADDFDB1FAD5" blockId="6.[151,522,1333,1359]" box="[151,522,1333,1359]" pageId="6" pageNumber="459">
<heading id="D0F9D13CF074FFA6FF3DFADDFDB1FAD5" bold="true" box="[151,522,1333,1359]" fontSize="11" level="1" pageId="6" pageNumber="459" reason="1">
<taxonomicName id="4C0E1DD3F074FFA6FF3DFADDFDB1FAD5" ID-CoL="3N5CY" authority="Jekelius, 1944" authorityName="Jekelius" authorityYear="1944" box="[151,522,1333,1359]" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="459" phylum="Mollusca" rank="species" species="alpha">
<emphasis id="B97ABA42F074FFA6FF3DFADDFDB1FAD5" bold="true" box="[151,522,1333,1359]" pageId="6" pageNumber="459">
<emphasis id="B97ABA42F074FFA6FF3DFADDFEEFFAD5" bold="true" box="[151,340,1333,1359]" italics="true" pageId="6" pageNumber="459">Hydrobia alpha</emphasis>
<bibRefCitation id="EF9F1BA1F074FFA6FEF6FADDFDB1FAD5" author="Jekelius, E." box="[348,522,1333,1359]" pageId="6" pageNumber="459" pagination="1 - 167" refId="ref10036" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului geologic al Romaniei, 5, 1 - 167." type="journal article" year="1944">Jekelius, 1944</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C31435DBF074FFA6FF3DFA94FCEDFA2A" pageId="6" pageNumber="459" type="reference_group">
<paragraph id="8BB16650F074FFA6FF3DFA94FB63FA08" blockId="6.[151,1240,1404,1456]" box="[151,1240,1404,1426]" pageId="6" pageNumber="459">
<treatmentCitationGroup id="AB1E417EF074FFA6FF3DFA94FB63FA08" box="[151,1240,1404,1426]" pageId="6" pageNumber="459">
<treatmentCitation id="0AAF4041F074FFA6FF3DFA94FD82FA08" author="Jekelius, E." box="[151,569,1404,1426]" page="59" pageId="6" pageNumber="459" year="1944">
<taxonomicName id="4C0E1DD3F074FFA6FF3DFA94FD82FA08" ID-CoL="3N5MB" authority="Jekelius 1944: 59" authorityName="Jekelius" authorityPageNumber="59" authorityYear="1944" box="[151,569,1404,1426]" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="459" phylum="Mollusca" rank="species" species="subprotracta">
<emphasis id="B97ABA42F074FFA6FF3DFA94FEC4FA08" box="[151,383,1404,1426]" italics="true" pageId="6" pageNumber="459">Hydrobia subprotracta</emphasis>
<bibRefCitation id="EF9F1BA1F074FFA6FE2FFA94FD82FA08" author="Jekelius, E." box="[389,569,1404,1426]" pageId="6" pageNumber="459" pagination="1 - 167" refId="ref10036" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului geologic al Romaniei, 5, 1 - 167." type="journal article" year="1944">Jekelius 1944: 59</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 10, figs 1–4 [non
<taxonomicName id="4C0E1DD3F074FFA6FCBDFA94FB75FA08" ID-CoL="3N5MB" authority="Zhizhchenko, 1936" authorityName="Zhizhchenko" authorityYear="1936" box="[791,1230,1404,1426]" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="459" phylum="Mollusca" rank="species" species="subprotracta">
<emphasis id="B97ABA42F074FFA6FCBDFA94FC44FA08" box="[791,1023,1404,1426]" italics="true" pageId="6" pageNumber="459">Hydrobia subprotracta</emphasis>
<bibRefCitation id="EF9F1BA1F074FFA6FBAFFA94FB75FA08" author="Zhizhchenko, B." box="[1029,1230,1404,1426]" pageId="6" pageNumber="459" refId="ref12435" refString="Zhizhchenko, B. (1936) Paleontologija SSSR. Tom X, Chast' 3: Chokrakskie Mollyuski. Izdatel'stvo Akademii Nauk SSSR, Moskva, 355 pp." type="book" year="1936">Zhizhchenko, 1936</bibRefCitation>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB16650F074FFA6FF3DFA72FCEDFA2A" blockId="6.[151,1240,1404,1456]" box="[151,854,1434,1456]" pageId="6" pageNumber="459">
<treatmentCitationGroup id="AB1E417EF074FFA6FF3DFA72FCEDFA2A" box="[151,854,1434,1456]" pageId="6" pageNumber="459">
<taxonomicName id="4C0E1DD3F074FFA6FF3DFA72FE04FA2A" ID-CoL="3N5CY" authorityName="Jekelius" authorityYear="1944" box="[151,447,1434,1456]" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="459" phylum="Mollusca" rank="subSpecies" species="subprotracta" subSpecies="alpha">
<emphasis id="B97ABA42F074FFA6FF3DFA72FE04FA2A" box="[151,447,1434,1456]" italics="true" pageId="6" pageNumber="459">Hydrobia subprotracta alpha</emphasis>
</taxonomicName>
[sic]
<treatmentCitation id="0AAF4041F074FFA6FE53FA72FD16FA2A" author="Jekelius, E." box="[505,685,1434,1456]" page="59" pageId="6" pageNumber="459" year="1944">
<bibRefCitation id="EF9F1BA1F074FFA6FE53FA72FD16FA2A" author="Jekelius, E." box="[505,685,1434,1456]" pageId="6" pageNumber="459" pagination="1 - 167" refId="ref10036" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului geologic al Romaniei, 5, 1 - 167." type="journal article" year="1944">Jekelius 1944: 59</bibRefCitation>
</treatmentCitation>
, pl. 10, figs 5–7.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31435DBF074FFA6FF3DFA37FA9DF9D8" pageId="6" pageNumber="459" type="materials_examined">
<paragraph id="8BB16650F074FFA6FF3DFA37FC17FA62" blockId="6.[151,1436,1501,1926]" box="[151,940,1501,1529]" pageId="6" pageNumber="459">
<emphasis id="B97ABA42F074FFA6FF3DFA37FE8EFA62" bold="true" box="[151,309,1503,1528]" inLineHeading="true" pageId="6" pageNumber="459" reason="1">Type locality.</emphasis>
Poliţioană valley near Soceni,
<collectingRegion id="49CAA8B2F074FFA6FD38FA37FC8FFA62" box="[658,820,1503,1528]" country="Romania" name="Caras-Severin" pageId="6" pageNumber="459">Caras-Severin</collectingRegion>
,
<collectingCountry id="F31926C0F074FFA6FC95FA37FC13FA62" box="[831,936,1503,1528]" name="Romania" pageId="6" pageNumber="459">Romania</collectingCountry>
.
</paragraph>
<paragraph id="8BB16650F074FFA6FF6DF9EDFD1DF984" blockId="6.[151,1436,1501,1926]" box="[199,678,1541,1566]" pageId="6" pageNumber="459">
<emphasis id="B97ABA42F074FFA6FF6DF9EDFF47F984" bold="true" box="[199,252,1541,1566]" pageId="6" pageNumber="459">Age.</emphasis>
Sarmatian s.s. (= upper Serravallian).
</paragraph>
<paragraph id="8BB16650F074FFA6FF6DF9C1FA9DF9D8" blockId="6.[151,1436,1501,1926]" box="[199,1318,1577,1602]" pageId="6" pageNumber="459">
<emphasis id="B97ABA42F074FFA6FF6DF9C1FE80F9D8" bold="true" box="[199,315,1577,1602]" inLineHeading="true" pageId="6" pageNumber="459" reason="1">
<typeStatus id="54B5D8F2F074FFA6FF6DF9C1FE8DF9D8" box="[199,310,1577,1602]" pageId="6" pageNumber="459" type="holotype">Holotype</typeStatus>
.
</emphasis>
<bibRefCitation id="EF9F1BA1F074FFA6FEE8F9C1FE5CF9D8" author="Jekelius, E." box="[322,487,1577,1602]" pageId="6" pageNumber="459" pagination="1 - 167" refId="ref10036" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului geologic al Romaniei, 5, 1 - 167." type="journal article" year="1944">Jekelius (1944</bibRefCitation>
, pl. 10, fig. 2); no information about storage and collection number given.
</paragraph>
</subSubSection>
<subSubSection id="C31435DBF074FFA6FF6DF9A5FAE0F81C" pageId="6" pageNumber="459" type="discussion">
<paragraph id="8BB16650F074FFA6FF6DF9A5FAE0F81C" blockId="6.[151,1436,1501,1926]" pageId="6" pageNumber="459">
<emphasis id="B97ABA42F074FFA6FF6DF9A5FEF7F9FC" bold="true" box="[199,332,1613,1638]" inLineHeading="true" pageId="6" pageNumber="459" reason="1">Discussion.</emphasis>
The species name
<taxonomicName id="4C0E1DD3F074FFA6FD9EF9A5FC51F9FC" authority="Jekelius, 1944" authorityName="Jekelius" authorityYear="1944" box="[564,1002,1613,1638]" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="459" phylum="Mollusca" rank="species" species="subprotracta">
<emphasis id="B97ABA42F074FFA6FD9EF9A5FC81F9FC" box="[564,826,1613,1638]" italics="true" pageId="6" pageNumber="459">Hydrobia subprotracta</emphasis>
<bibRefCitation id="EF9F1BA1F074FFA6FCEFF9A5FC51F9FC" author="Jekelius, E." box="[837,1002,1613,1638]" pageId="6" pageNumber="459" pagination="1 - 167" refId="ref10036" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului geologic al Romaniei, 5, 1 - 167." type="journal article" year="1944">Jekelius, 1944</bibRefCitation>
</taxonomicName>
is a primary homonym of
<taxonomicName id="4C0E1DD3F074FFA6FA98F9A5FDB5F910" authority="Zhizhchenko, 1936" authorityName="Zhizhchenko" authorityYear="1936" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="459" phylum="Mollusca" rank="species" species="subprotracta">
<emphasis id="B97ABA42F074FFA6FA98F9A5FE9CF910" italics="true" pageId="6" pageNumber="459">Hydrobia subprotracta</emphasis>
<bibRefCitation id="EF9F1BA1F074FFA6FE98F999FDB5F910" author="Zhizhchenko, B." box="[306,526,1649,1674]" pageId="6" pageNumber="459" refId="ref12435" refString="Zhizhchenko, B. (1936) Paleontologija SSSR. Tom X, Chast' 3: Chokrakskie Mollyuski. Izdatel'stvo Akademii Nauk SSSR, Moskva, 355 pp." type="book" year="1936">Zhizhchenko, 1936</bibRefCitation>
</taxonomicName>
and therefore unavailable. Formally, a replacement name is needed here. We refrain from such a nomenclatural act because of following reason:
<taxonomicName id="4C0E1DD3F074FFA6FC6BF97EFA90F934" authority="Jekelius, 1944" authorityName="Jekelius" authorityYear="1944" box="[961,1323,1685,1710]" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="459" phylum="Mollusca" rank="species" species="subprotracta">
<emphasis id="B97ABA42F074FFA6FC6BF97EFBC2F934" box="[961,1145,1685,1710]" italics="true" pageId="6" pageNumber="459">H. subprotracta</emphasis>
<bibRefCitation id="EF9F1BA1F074FFA6FB2FF97DFA90F934" author="Jekelius, E." box="[1157,1323,1685,1710]" pageId="6" pageNumber="459" pagination="1 - 167" refId="ref10036" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului geologic al Romaniei, 5, 1 - 167." type="journal article" year="1944">Jekelius, 1944</bibRefCitation>
</taxonomicName>
is herein considered synonymous with
<taxonomicName id="4C0E1DD3F074FFA6FE47F952FC27F948" authority="Jekelius, 1944" authorityName="Jekelius" authorityYear="1944" box="[493,924,1721,1746]" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="459" phylum="Mollusca" rank="subSpecies" species="subprotracta" subSpecies="alpha">
<emphasis id="B97ABA42F074FFA6FE47F952FD50F948" box="[493,747,1721,1746]" italics="true" pageId="6" pageNumber="459">H. subprotracta alpha</emphasis>
<bibRefCitation id="EF9F1BA1F074FFA6FD5FF951FC27F948" author="Jekelius, E." box="[757,924,1721,1746]" pageId="6" pageNumber="459" pagination="1 - 167" refId="ref10036" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului geologic al Romaniei, 5, 1 - 167." type="journal article" year="1944">Jekelius, 1944</bibRefCitation>
</taxonomicName>
. The name &quot;
<taxonomicName id="4C0E1DD3F074FFA6FB9AF951FBCBF948" authorityName="Jekelius" authorityYear="1944" box="[1072,1136,1721,1746]" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="459" phylum="Mollusca" rank="species" species="alpha">
<emphasis id="B97ABA42F074FFA6FB9AF951FBCBF948" box="[1072,1136,1721,1746]" italics="true" pageId="6" pageNumber="459">alpha</emphasis>
</taxonomicName>
&quot; was fully spelled out by
<bibRefCitation id="EF9F1BA1F074FFA6FF3DF935FEF9F96C" author="Jekelius, E." box="[151,322,1757,1782]" pageId="6" pageNumber="459" pagination="1 - 167" refId="ref10036" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului geologic al Romaniei, 5, 1 - 167." type="journal article" year="1944">Jekelius (1944)</bibRefCitation>
and does not denote an infrasubspecific taxon; it is thus available as a species-group name (see also
<bibRefCitation id="EF9F1BA1F074FFA6FF3DF8E9FEA1F880" author="ICZN" box="[151,282,1793,1818]" pageId="6" pageNumber="459" refId="ref9970" refString="ICZN (1999) International Code of Zoological Nomenclature. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
, Article 45.6.4). Apart from stronger convex whorls as indicated by the description and illustrations of
<bibRefCitation id="EF9F1BA1F074FFA6FF3DF8CDFEFCF8A4" author="Jekelius, E." box="[151,327,1829,1854]" pageId="6" pageNumber="459" pagination="1 - 167" refId="ref10036" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului geologic al Romaniei, 5, 1 - 167." type="journal article" year="1944">Jekelius (1944)</bibRefCitation>
, it fully corresponds to &quot;
<taxonomicName id="4C0E1DD3F074FFA6FDCCF8CEFC70F8A4" authority="Jekelius, 1944" authorityName="Jekelius" authorityYear="1944" box="[614,971,1829,1854]" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="459" phylum="Mollusca" rank="species" species="subprotracta">
<emphasis id="B97ABA42F074FFA6FDCCF8CEFCA0F8A4" box="[614,795,1829,1854]" italics="true" pageId="6" pageNumber="459">H. subprotracta</emphasis>
<bibRefCitation id="EF9F1BA1F074FFA6FC8EF8CDFC70F8A4" author="Jekelius, E." box="[804,971,1829,1854]" pageId="6" pageNumber="459" pagination="1 - 167" refId="ref10036" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului geologic al Romaniei, 5, 1 - 167." type="journal article" year="1944">Jekelius, 1944</bibRefCitation>
</taxonomicName>
&quot;. Following this, &quot;
<taxonomicName id="4C0E1DD3F074FFA6FB08F8CDFB59F8A4" authorityName="Jekelius" authorityYear="1944" box="[1186,1250,1829,1854]" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="459" phylum="Mollusca" rank="species" species="alpha">
<emphasis id="B97ABA42F074FFA6FB08F8CDFB59F8A4" box="[1186,1250,1829,1854]" italics="true" pageId="6" pageNumber="459">alpha</emphasis>
</taxonomicName>
&quot; is the first and only available name for the taxon. We simply avoid introducing a replacement name for
<taxonomicName id="4C0E1DD3F074FFA6FB33F8A1FE81F81C" authority="Jekelius, 1944" authorityName="Jekelius" authorityYear="1944" class="Gastropoda" family="Hydrobiidae" genus="Hydrobia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="459" phylum="Mollusca" rank="species" species="subprotracta">
<emphasis id="B97ABA42F074FFA6FB33F8A1FA27F8F8" box="[1177,1436,1865,1890]" italics="true" pageId="6" pageNumber="459">Hydrobia subprotracta</emphasis>
<bibRefCitation id="EF9F1BA1F074FFA6FF3DF885FE81F81C" author="Jekelius, E." box="[151,314,1901,1926]" pageId="6" pageNumber="459" pagination="1 - 167" refId="ref10036" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului geologic al Romaniei, 5, 1 - 167." type="journal article" year="1944">Jekelius, 1944</bibRefCitation>
</taxonomicName>
, not to add another (redundant) name to the already existing bulk and confusing later workers.
</paragraph>
</subSubSection>
</treatment>
</document>
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