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<document id="52111EFEA830F0DAD39952C4D86462B2" ID-CLB-Dataset="42381" ID-DOI="10.11646/zootaxa.3873.3.1" ID-GBIF-Dataset="815053ff-47ad-4d35-bd4f-c45e262a1663" ID-ISSN="1175-5326" ID-Zenodo-Dep="228641" ID-ZooBank="022136BF-394F-4959-8CE7-1F4971923223" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460935070357" checkinUser="plazi" docAuthor="Katsiaras, Nikolaos, Simboura, Nomiki &amp; Koutsoubas, Drosos" docDate="2014" docId="03A75B379108FFF007DBFDEDFA7F7D01" docLanguage="en" docName="zt03873p217.pdf" docOrigin="Zootaxa 3873 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Marphysa adenensis Gravier 1900" docType="treatment" docVersion="9" lastPageNumber="206" masterDocId="FF9E234F910AFFF6074CFFAAFFFD7F3C" masterDocTitle="The rare subgroup C 1 of Marphysa (Polychaeta, Eunicidae): re-description of species and first records in the Mediterranean Sea" masterLastPageNumber="217" masterPageNumber="201" pageNumber="203" updateTime="1698582671682" updateUser="ExternalLinkService">
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<mods:title id="C1A249E7427A1A558249AF45A8F30B43">The rare subgroup C 1 of Marphysa (Polychaeta, Eunicidae): re-description of species and first records in the Mediterranean Sea</mods:title>
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<mods:namePart id="36409C5CDC2AFFE6C694F4FE6E963398">Katsiaras, Nikolaos</mods:namePart>
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<mods:namePart id="A21D00890090026A652065CB9373530D">Simboura, Nomiki</mods:namePart>
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<treatment id="03A75B379108FFF007DBFDEDFA7F7D01" ID-DOI="http://doi.org/10.5281/zenodo.5613403" ID-GBIF-Taxon="119590487" ID-Zenodo-Dep="5613403" LSID="urn:lsid:plazi:treatment:03A75B379108FFF007DBFDEDFA7F7D01" httpUri="http://treatment.plazi.org/id/03A75B379108FFF007DBFDEDFA7F7D01" lastPageId="6" lastPageNumber="206" pageId="2" pageNumber="203">
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<paragraph id="8BB1EA219108FFF407DBFDEDFDBD7D5E" blockId="2.[151,587,583,643]" box="[151,576,583,610]" pageId="2" pageNumber="203">
<heading id="D0F95D4D9108FFF407DBFDEDFDBD7D5E" bold="true" box="[151,576,583,610]" fontSize="11" level="1" pageId="2" pageNumber="203" reason="1">
<taxonomicName id="4C0E91A29108FFF407DBFDEDFDBD7D5E" ID-CoL="3Y88M" authority="Gravier, 1900" authorityName="Gravier" authorityYear="1900" box="[151,576,583,610]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="2" pageNumber="203" phylum="Annelida" rank="species" species="adenensis">
<emphasis id="B97A36339108FFF407DBFDEDFDBD7D5E" bold="true" box="[151,576,583,610]" pageId="2" pageNumber="203">
<emphasis id="B97A36339108FFF407DBFDEDFE727D5D" bold="true" box="[151,399,583,609]" italics="true" pageId="2" pageNumber="203">Marphysa adenensis</emphasis>
<bibRefCitation id="EF9F97D09108FFF406DAFDE2FDBD7D5E" author="Gravier" box="[406,576,584,610]" pageId="2" pageNumber="203" refString="Gravier, C. (1900) Contribution a l' etude des Annelides Polychaetes de la Mer Rouge. Premiere partie. Nouvelles Archives du Museum d' Historie Naturelle Paris, Series 4, 2 (2), 137 - 282." type="journal article" year="1900">Gravier, 1900</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB1EA219108FFF407DBFDC0FDB67DBF" blockId="2.[151,587,583,643]" box="[151,587,618,643]" pageId="2" pageNumber="203">
<figureCitation id="1335F6A49108FFF407DBFDC0FF297DBF" box="[151,212,618,643]" captionStart="FIGURE 1" captionStartId="3.[151,250,1903,1925]" captionTargetBox="[188,1379,213,1853]" captionTargetId="figure@3.[180,1390,195,1864]" captionTargetPageId="3" captionText="FIGURE 1. Marphysa adenensis (GP- 4 B) a) Anterior long capillary seta (segment 10); b) anterior composite falcigers (segment 10); c) branchial-body composite falciger; d) posterior composite falcigers (segment 74); e) acicular seta (segment 73); f h) pectinate setae" httpUri="https://zenodo.org/record/228642/files/figure.png" pageId="2" pageNumber="203">Fig. 1</figureCitation>
ah, 2ad, 3, 4ad, 5ad,
<tableCitation id="C68CDF9A9108FFF406B4FDC0FDB67DBF" box="[504,587,618,643]" captionStart="TABLE 2" captionStartId="13.[151,239,260,283]" captionTargetBox="[159,1427,446,963]" captionTargetPageId="13" captionText="TABLE 2. Comparison of important diagnostic characters of all species belonging to Marphysa C 1 Group. Species marked with asterisk (*) were not personally examined and source of the information are published descriptions. The composite falcigers blades gradation is referred to anterior segments." httpUri="http://table.plazi.org/id/DF71BAA99107FFFB07DBFEAEFC8B7E67" pageId="2" pageNumber="203" tableUuid="DF71BAA99107FFFB07DBFEAEFC8B7E67">Table 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C314B9AA9108FFF407DBFD18FEEB7DDB" pageId="2" pageNumber="203" type="reference_group">
<paragraph id="8BB1EA219108FFF407DBFD18FEEB7DDB" blockId="2.[151,1436,690,743]" pageId="2" pageNumber="203">
<treatmentCitationGroup id="AB1ECD0F9108FFF407DBFD18FEEB7DDB" pageId="2" pageNumber="203">
<taxonomicName id="4C0E91A29108FFF407DBFD18FDD57DF4" ID-CoL="3Y88M" authority="Gravier 1900: 270" authorityName="Gravier" authorityPageNumber="270" authorityYear="1900" box="[151,552,690,712]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="2" pageNumber="203" phylum="Annelida" rank="species" species="adenensis">
<emphasis id="B97A36339108FFF407DBFD18FE9A7DF4" box="[151,359,690,712]" italics="true" pageId="2" pageNumber="203">Marphysa adenensis</emphasis>
<treatmentCitation id="0AAFCC309108FFF40621FD18FDD57DF4" author="Gravier" box="[365,552,690,712]" page="270" pageId="2" pageNumber="203" year="1900">
<bibRefCitation id="EF9F97D09108FFF40621FD18FE0B7DF4" author="Gravier" box="[365,502,690,712]" pageId="2" pageNumber="203" refString="Gravier, C. (1900) Contribution a l' etude des Annelides Polychaetes de la Mer Rouge. Premiere partie. Nouvelles Archives du Museum d' Historie Naturelle Paris, Series 4, 2 (2), 137 - 282." type="journal article" year="1900">Gravier 1900</bibRefCitation>
: 270
</treatmentCitation>
</taxonomicName>
, Plate 14 Figures 9192; Text-figures 140143;
<treatmentCitation id="0AAFCC309108FFF40340FD19FB587DF4" author="Day" box="[1036,1189,690,712]" page="644" pageId="2" pageNumber="203" year="1962">
<bibRefCitation id="EF9F97D09108FFF40340FD19FB8F7DF4" author="Day" box="[1036,1138,690,712]" pageId="2" pageNumber="203" refString="Day, J. H. (1962) Polychaeta from several localities in the Western Indian Ocean Proceeding of the Zoological Society of London, 139 (4), 627 - 656. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1962. tb 01597. x" type="journal article" year="1962">Day 1962</bibRefCitation>
: 644
</treatmentCitation>
;
<treatmentCitation id="0AAFCC309108FFF403E3FD19FABB7DF4" author="Day" box="[1199,1350,690,712]" page="399" pageId="2" pageNumber="203" year="1967">
<bibRefCitation id="EF9F97D09108FFF403E3FD19FAE87DF4" author="Day" box="[1199,1301,690,712]" pageId="2" pageNumber="203" refString="Day, J. H. (1967) A monograph on the polychaeta of southern Africa. Part I. Errantia. British Museum (Natural History), London, 458 pp." type="book" year="1967">Day 1967</bibRefCitation>
: 399
</treatmentCitation>
, Figures 17.6ps
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</paragraph>
</subSubSection>
<subSubSection id="C314B9AA9108FFF407DBFCBEFAAB7B39" pageId="2" pageNumber="203" type="materials_examined">
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<emphasis id="B97A36339108FFF407DBFCBEFEBA7C11" bold="true" box="[151,327,788,813]" pageId="2" pageNumber="203">
<typeStatus id="54B554839108FFF407DBFCBEFF2F7C11" box="[151,210,788,813]" pageId="2" pageNumber="203">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54B554839108FFF4061CFCBEFE457C11" box="[336,440,788,813]" pageId="2" pageNumber="203" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED1F72E49108FFF40685FCBEFDDD7C10" LSID="urn:lsid:biocol.org:col:34988" box="[457,544,788,812]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="203">MNHN</collectionCode>
<collectionCode id="ED1F72E49108FFF40566FCBEFD8D7C10" box="[554,624,788,812]" pageId="2" pageNumber="203">POLY</collectionCode>
<typeStatus id="54B554839108FFF40536FCBFFD3D7C10" box="[634,704,789,812]" pageId="2" pageNumber="203">
<collectionCode id="ED1F72E49108FFF40536FCBFFD3D7C10" box="[634,704,789,812]" pageId="2" pageNumber="203">TYPE</collectionCode>
</typeStatus>
0531) from Aden Sea, 1900,
<specimenCount id="9D0821A89108FFF40357FCBEFB5D7C11" box="[1051,1184,788,813]" pageId="2" pageNumber="203" type="generic">1 specimen</specimenCount>
,
<quantity id="4CF647C49108FFF403E1FCBEFB087C11" box="[1197,1269,788,813]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="203" unit="mm" value="45.0">45mm</quantity>
long and 0.75 width, 200 segments, posterior end missing, jaws removed.
</paragraph>
<paragraph id="8BB1EA219108FFF4078BFCF6FAAB7B39" blockId="2.[151,1437,788,1928]" pageId="2" pageNumber="203">
<emphasis id="B97A36339108FFF4078BFCF6FE547C49" bold="true" box="[199,425,860,885]" pageId="2" pageNumber="203">Non-type material.</emphasis>
Lesvos Island, Gera Gulf
<date id="FFB0CCE19108FFF4059DFCF6FCBA7C49" box="[721,839,860,885]" pageId="2" pageNumber="203" value="2009-09-15">15/9/2009</date>
:
<specimenCount id="9D0821A89108FFF4041EFCF6FC217C49" box="[850,988,860,885]" pageId="2" pageNumber="203" type="generic">3 specimens</specimenCount>
(GP-4B): 76 segments
<quantity id="4CF647C49108FFF403A9FCF6FAC97C48" box="[1253,1332,860,884]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="203" unit="mm" value="30.0">30 mm</quantity>
long and
<quantity id="4CF647C49108FFF407DBFCD5FF137CAB" box="[151,238,895,920]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="203" unit="mm" value="2.5">2.5 mm</quantity>
wide, 62 segments
<quantity id="4CF647C49108FFF4069DFC2AFDDF7CAB" box="[465,546,896,920]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="203" unit="mm" value="24.0">24 mm</quantity>
long and
<quantity id="4CF647C49108FFF405D9FCD5FD107CAB" box="[661,749,895,920]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="203" unit="mm" value="2.5">2.5 mm</quantity>
wide, 74 segments
<quantity id="4CF647C49108FFF4049CFCD5FBDD7CAB" box="[976,1056,895,920]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="203" unit="mm" value="25.0">25 mm</quantity>
long and
<quantity id="4CF647C49108FFF403DFFC2AFB307CA4" box="[1171,1229,896,920]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="203" unit="mm" value="1.0">1mm</quantity>
wide;
<specimenCount id="9D0821A89108FFF40250FC2AFA607CA4" box="[1308,1437,895,920]" pageId="2" pageNumber="203" type="generic">1 specimen</specimenCount>
(GP-2B): in the 25 segments
<quantity id="4CF647C49108FFF406A1FC0EFDC27C80" box="[493,575,932,956]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="203" unit="mm" value="30.0">30 mm</quantity>
long and
<quantity id="4CF647C49108FFF405FBFC0EFD067C80" box="[695,763,932,956]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="203" unit="mm" value="4.0">4 mm</quantity>
wide; Methoni
<date id="FFB0CCE19108FFF404F9FC0EFBD67C80" box="[949,1067,932,956]" pageId="2" pageNumber="203" value="2012-04-24">24/4/2012</date>
:
<specimenCount id="9D0821A89108FFF40375FC0EFB3B7C81" box="[1081,1222,932,957]" pageId="2" pageNumber="203" type="generic">3 specimens</specimenCount>
(1/WFD 3R1): 54 segments
<quantity id="4CF647C49108FFF4064AFC62FE977CE3" box="[262,362,968,992]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="2" pageNumber="203" unit="mm" value="10.3">10.3 mm</quantity>
long and
<quantity id="4CF647C49108FFF40696FC62FDE67CE3" box="[474,539,968,992]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="203" unit="mm" value="1.0">1 mm</quantity>
wide, 43 segments
<quantity id="4CF647C49108FFF405B5FC6DFCC77CE3" box="[761,826,967,992]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="203" unit="mm" value="5.0">5 mm</quantity>
long and
<quantity id="4CF647C49108FFF404E6FC62FC167CE3" box="[938,1003,968,992]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="203" unit="mm" value="1.0">1 mm</quantity>
wide, 35 segments
<quantity id="4CF647C49108FFF40385FC62FAF77CE3" box="[1225,1290,968,992]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="203" unit="mm" value="9.0">9 mm</quantity>
long and
<quantity id="4CF647C49108FFF40235FC6DFF3E7B38" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="203" unit="mm" value="1.5">1.5 mm</quantity>
wide; Lesvos Strait
<date id="FFB0CCE19108FFF406E4FC46FDE37B38" box="[424,542,1004,1028]" pageId="2" pageNumber="203" value="2013-03-29">29/3/2013</date>
:
<specimenCount id="9D0821A89108FFF40564FC46FD4E7B39" box="[552,691,1004,1029]" pageId="2" pageNumber="203" type="generic">1 specimens</specimenCount>
(3/WFD 70R1): 42 segments
<quantity id="4CF647C49108FFF40348FC46FBA97B38" box="[1028,1108,1004,1028]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="203" unit="mm" value="20.0">20 mm</quantity>
long and
<quantity id="4CF647C49108FFF4038EFC46FAEC7B38" box="[1218,1297,1004,1028]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="203" unit="mm" value="1.7">1.7mm</quantity>
wide.
</paragraph>
</subSubSection>
<subSubSection id="C314B9AA9108FFF2078BFBA5FCD278F4" lastPageId="4" lastPageNumber="205" pageId="2" pageNumber="203" type="description">
<paragraph id="8BB1EA219108FFF4078BFBA5FD3D7A3C" blockId="2.[151,1437,788,1928]" pageId="2" pageNumber="203">
<emphasis id="B97A36339108FFF4078BFBA5FEA47B14" bold="true" box="[199,345,1039,1064]" pageId="2" pageNumber="203">Description.</emphasis>
<typeStatus id="54B554839108FFF4062CFBA5FE357B14" box="[352,456,1039,1064]" pageId="2" pageNumber="203" type="holotype">Holotype</typeStatus>
an anterior fragment of about 200 segments (
<figureCitation id="1335F6A49108FFF4049DFBA5FBEB7B14" box="[977,1046,1039,1064]" captionStart="FIGURE 5" captionStartId="7.[151,250,1430,1452]" captionTargetBox="[151,1436,193,1407]" captionTargetId="figure@7.[151,1436,193,1408]" captionTargetPageId="7" captionText="FIGURE 5. Holotype of Marphysa adenensis (MNHN POLY TYPE 0531) a) anterior body; b) anterior composite falcigers; c) pectinate seta; d) acicular seta; cf: composite falcigers; ps: pectinate seta; as: acicular seta; dc: dorsal cirri; vc: ventral cirri; ac: acicula." httpUri="https://zenodo.org/record/228646/files/figure.png" pageId="2" pageNumber="203">Fig. 5</figureCitation>
a). All 8 Mediterranean specimens anterior fragments; longest
<quantity id="4CF647C49108FFF40687FB9EFDE77B70" box="[459,538,1076,1100]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="203" unit="mm" value="30.0">30 mm</quantity>
long and 2.5 wide with 76 segments; shortest
<quantity id="4CF647C49108FFF4036DFB9EFB9F7B70" box="[1057,1122,1076,1101]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="203" unit="mm" value="5.0">5 mm</quantity>
long and
<quantity id="4CF647C49108FFF4039DFB9EFADA7B70" box="[1233,1319,1076,1100]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="203" unit="mm" value="0.9">0.9 mm</quantity>
wide with 43 segments. Prostomium rounded anteriorly, as long as first peristomial ring, which is longer than second peristomial ring. Three antennae present and a pair of dorso-lateral palps. Antennae slightly longer than prostomium; median longer. Antennae arranged in slightly curved line and slightly articulated. Eyes present, positioned behind the lateral antennae. Jaws removed from
<typeStatus id="54B554839108FFF40461FB6EFC6E7BE1" box="[813,915,1220,1245]" pageId="2" pageNumber="203" type="holotype">holotype</typeStatus>
. Maxillary formula of Mediterranean material: I(1+1), II(5-7+7-8), III(6-7+0), IV (4+8) (
<figureCitation id="1335F6A49108FFF40520FB4DFD4C7A3C" box="[620,689,1255,1280]" captionStart="FIGURE 3" captionStartId="5.[151,250,1305,1327]" captionTargetBox="[310,1277,199,1280]" captionTargetId="figure@5.[302,1285,193,1284]" captionTargetPageId="5" captionText="FIGURE 3. Marphysa adenensis (GP- 4 B). Maxillary apparatus; Mx. I IV: Maxillary I IV." httpUri="https://zenodo.org/record/228644/files/figure.png" pageId="2" pageNumber="203">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BB1EA219108FFF4078BFAA6FB0A7AC1" blockId="2.[151,1437,788,1928]" pageId="2" pageNumber="203">Dorsal cirri cirriform in anterior body, more thin and slender after mid-body; slightly longer than the parapodium in anterior region, increasing in size up to twice as long after the first branchiae. Ventral cirri tongueshaped with rounded tips in anterior body, tips with a distal papilla in branchial segments, which gradually becomes less distinct after mid-body. Ventral cirri slightly shorter than the parapodium lobe. Post-chaetal lobes elongated and tongue-shaped, much longer than acicular lobe, gradually decreasing down to a similar length in posterior body. Pre-chaetal lobes always truncated and straight, shorter than acicular lobes. Branchiae present from chaetiger 1017 until 1632, depending on specimen size; all pectinate and have up to 10 filaments.</paragraph>
<paragraph id="8BB1EA219108FFF4078BF9ADFE7978B4" blockId="2.[151,1437,788,1928]" pageId="2" pageNumber="203">
<typeStatus id="54B554839108FFF4078BF9ADFEC9791C" box="[199,308,1543,1568]" pageId="2" pageNumber="203" type="holotype">Holotype</typeStatus>
: Superior setae in each fascicle consist of: (1) 35 long capillaries, (2) 49 relatively shorter capillaries, (3) 12 heterodont pectinate setae, marginal teeth longer and unequal to each other (twice as long) throughout the body, with 45 teeth in anterior region (
<figureCitation id="1335F6A49108FFF4044FF9E5FCB57954" box="[771,840,1615,1640]" captionStart="FIGURE 4" captionStartId="6.[151,250,1826,1848]" captionTargetBox="[178,1384,632,1752]" captionTargetId="figure@6.[167,1397,625,1762]" captionTargetPageId="6" captionText="FIGURE 4. Holotype of Marphysa adenensis (MNHN POLY TYPE 0531) a) acicular seta; b) anterior composite falcigers; c) anterior pectinate seta (segment 13); d) posterior pectinate seta (segment 195)." httpUri="https://zenodo.org/record/228645/files/figure.png" pageId="2" pageNumber="203">Fig. 4</figureCitation>
c, 5c), increasing up to 78 towards the posterior end (
<figureCitation id="1335F6A49108FFF407D3F9DEFF1979B1" box="[159,228,1652,1677]" captionStart="FIGURE 4" captionStartId="6.[151,250,1826,1848]" captionTargetBox="[178,1384,632,1752]" captionTargetId="figure@6.[167,1397,625,1762]" captionTargetPageId="6" captionText="FIGURE 4. Holotype of Marphysa adenensis (MNHN POLY TYPE 0531) a) acicular seta; b) anterior composite falcigers; c) anterior pectinate seta (segment 13); d) posterior pectinate seta (segment 195)." httpUri="https://zenodo.org/record/228645/files/figure.png" pageId="2" pageNumber="203">Fig. 4</figureCitation>
d). Inferior setae consist only of composite falcigers, bidentate and hooded (
<figureCitation id="1335F6A49108FFF40376F9DEFB8379B1" box="[1082,1150,1652,1677]" captionStart="FIGURE 4" captionStartId="6.[151,250,1826,1848]" captionTargetBox="[178,1384,632,1752]" captionTargetId="figure@6.[167,1397,625,1762]" captionTargetPageId="6" captionText="FIGURE 4. Holotype of Marphysa adenensis (MNHN POLY TYPE 0531) a) acicular seta; b) anterior composite falcigers; c) anterior pectinate seta (segment 13); d) posterior pectinate seta (segment 195)." httpUri="https://zenodo.org/record/228645/files/figure.png" pageId="2" pageNumber="203">Fig. 4</figureCitation>
b, 5b). There are up to
<quantity id="4CF647C49108FFF402CCF9DEFF50798C" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="203" unit="in" value="20.0">20 in</quantity>
anterior body, but only up to 10 at the posterior part. No composite spinigers present. Towards the posterior body, the number and length of all setae is decreasing, except of pectinate which remains the same. The length of the falciger blades is relatively short (3240 µm in anterior chaetigers; decreasing down to 1220 µm in posterior) and about equal between falcigers of the same fascicle, but with a slight length gradation (from 90µm up to 140µm) in the branchial region (one and half times longer). Aciculae blunt and pale. Acicular setae, beginning from chaetiger 35; pale yellow, hooded and bidentate; teeth have almost 90° angle between them; subdistal almost twice as wide (
<figureCitation id="1335F6A49108FFF407B7F8C5FEC378B4" box="[251,318,1903,1928]" captionStart="FIGURE 4" captionStartId="6.[151,250,1826,1848]" captionTargetBox="[178,1384,632,1752]" captionTargetId="figure@6.[167,1397,625,1762]" captionTargetPageId="6" captionText="FIGURE 4. Holotype of Marphysa adenensis (MNHN POLY TYPE 0531) a) acicular seta; b) anterior composite falcigers; c) anterior pectinate seta (segment 13); d) posterior pectinate seta (segment 195)." httpUri="https://zenodo.org/record/228645/files/figure.png" pageId="2" pageNumber="203">Fig. 4</figureCitation>
a, 5d).
</paragraph>
<caption id="DF71BAA99109FFF507DBF8C5FE7678FE" httpUri="https://zenodo.org/record/228642/files/figure.png" pageId="3" pageNumber="204" targetBox="[188,1379,213,1853]" targetPageId="3">
<paragraph id="8BB1EA219109FFF507DBF8C5FE7678FE" blockId="3.[151,1436,1903,1986]" pageId="3" pageNumber="204">
<emphasis id="B97A36339109FFF507DBF8C5FEE578B8" bold="true" box="[151,280,1903,1925]" pageId="3" pageNumber="204">FIGURE 1.</emphasis>
<taxonomicName id="4C0E91A29109FFF50668F8C5FE0778B9" box="[292,506,1903,1925]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="204" phylum="Annelida" rank="species" species="adenensis">
<emphasis id="B97A36339109FFF50668F8C5FE0778B9" box="[292,506,1903,1925]" italics="true" pageId="3" pageNumber="204">Marphysa adenensis</emphasis>
</taxonomicName>
(GP-4B) a) Anterior long capillary seta (segment 10); b) anterior composite falcigers (segment 10); c) branchial-body composite falciger; d) posterior composite falcigers (segment 74); e) acicular seta (segment 73); fh) pectinate setae
</paragraph>
</caption>
<caption id="DF71BAA9910EFFF207DBFA09FEE57ACA" httpUri="https://zenodo.org/record/228643/files/figure.png" pageId="4" pageNumber="205" targetBox="[151,1436,193,1420]" targetPageId="4">
<paragraph id="8BB1EA21910EFFF207DBFA09FEE57ACA" blockId="4.[151,1436,1443,1526]" pageId="4" pageNumber="205">
<emphasis id="B97A3633910EFFF207DBFA09FEE87A84" bold="true" box="[151,277,1443,1465]" pageId="4" pageNumber="205">FIGURE 2.</emphasis>
<taxonomicName id="4C0E91A2910EFFF20652FA09FE0C7A85" box="[286,497,1443,1465]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="4" pageNumber="205" phylum="Annelida" rank="species" species="adenensis">
<emphasis id="B97A3633910EFFF20652FA09FE0C7A85" box="[286,497,1443,1465]" italics="true" pageId="4" pageNumber="205">Marphysa adenensis</emphasis>
</taxonomicName>
(1/WFD 3R1) a) anterior body; b) anterior composite falcigers (segment 10); c) posterior pectinate seta (segment 39); d) acicular seta (segment 35); cf: composite falcigers; ps: pectinate seta; as: acicular seta; pc: postchaetal lobe.
</paragraph>
</caption>
<paragraph id="8BB1EA21910EFFF2078BF989FCD278F4" blockId="4.[151,1437,1571,1992]" pageId="4" pageNumber="205">
Mediterranean material: Superior setae in each fascicle consist of: (1) 35 long serrated capillaries (
<figureCitation id="1335F6A4910EFFF20277F989FA817900" box="[1339,1404,1571,1596]" captionStart="FIGURE 1" captionStartId="3.[151,250,1903,1925]" captionTargetBox="[188,1379,213,1853]" captionTargetId="figure@3.[180,1390,195,1864]" captionTargetPageId="3" captionText="FIGURE 1. Marphysa adenensis (GP- 4 B) a) Anterior long capillary seta (segment 10); b) anterior composite falcigers (segment 10); c) branchial-body composite falciger; d) posterior composite falcigers (segment 74); e) acicular seta (segment 73); f h) pectinate setae" httpUri="https://zenodo.org/record/228642/files/figure.png" pageId="4" pageNumber="205">Fig. 1</figureCitation>
a), (2) 49 short serrated capillaries, (3) 12 heterodont pectinate setae, marginal teeth longer and unequal to each other (twice as long) throughout the body, with 48 teeth in total (
<figureCitation id="1335F6A4910EFFF2049CF9C1FBE979B8" box="[976,1044,1643,1668]" captionStart="FIGURE 1" captionStartId="3.[151,250,1903,1925]" captionTargetBox="[188,1379,213,1853]" captionTargetId="figure@3.[180,1390,195,1864]" captionTargetPageId="3" captionText="FIGURE 1. Marphysa adenensis (GP- 4 B) a) Anterior long capillary seta (segment 10); b) anterior composite falcigers (segment 10); c) branchial-body composite falciger; d) posterior composite falcigers (segment 74); e) acicular seta (segment 73); f h) pectinate setae" httpUri="https://zenodo.org/record/228642/files/figure.png" pageId="4" pageNumber="205">Fig. 1</figureCitation>
fh, 2c), regardless of specimen dimensions. Inferior setae consist only of serrated, bidentated and hooded composite falcigers. No composite spiniger present. In each fascicle of anterior body (
<figureCitation id="1335F6A4910EFFF20582F919FCF379F0" box="[718,782,1715,1740]" captionStart="FIGURE 1" captionStartId="3.[151,250,1903,1925]" captionTargetBox="[188,1379,213,1853]" captionTargetId="figure@3.[180,1390,195,1864]" captionTargetPageId="3" captionText="FIGURE 1. Marphysa adenensis (GP- 4 B) a) Anterior long capillary seta (segment 10); b) anterior composite falcigers (segment 10); c) branchial-body composite falciger; d) posterior composite falcigers (segment 74); e) acicular seta (segment 73); f h) pectinate setae" httpUri="https://zenodo.org/record/228642/files/figure.png" pageId="4" pageNumber="205">Fig. 1</figureCitation>
b, 2b), or until the mid-body in older individuals (
<figureCitation id="1335F6A4910EFFF20271F919FA8079F0" box="[1341,1405,1715,1740]" captionStart="FIGURE 1" captionStartId="3.[151,250,1903,1925]" captionTargetBox="[188,1379,213,1853]" captionTargetId="figure@3.[180,1390,195,1864]" captionTargetPageId="3" captionText="FIGURE 1. Marphysa adenensis (GP- 4 B) a) Anterior long capillary seta (segment 10); b) anterior composite falcigers (segment 10); c) branchial-body composite falciger; d) posterior composite falcigers (segment 74); e) acicular seta (segment 73); f h) pectinate setae" httpUri="https://zenodo.org/record/228642/files/figure.png" pageId="4" pageNumber="205">Fig. 1</figureCitation>
c), there are 34 falcigers with notably longer and thinner blades (up to 160 µm; 115.3 µm average in older, 68 µm average in younger specimens), more than twice as long as the other (up to 76 µm; 55.3 µm average in older, 34.6 µm average in younger specimens). The length and the number of all setae decrease close to the posterior end (
<figureCitation id="1335F6A4910EFFF20223F8B5FF5F7860" captionStart="FIGURE 1" captionStartId="3.[151,250,1903,1925]" captionTargetBox="[188,1379,213,1853]" captionTargetId="figure@3.[180,1390,195,1864]" captionTargetPageId="3" captionText="FIGURE 1. Marphysa adenensis (GP- 4 B) a) Anterior long capillary seta (segment 10); b) anterior composite falcigers (segment 10); c) branchial-body composite falciger; d) posterior composite falcigers (segment 74); e) acicular seta (segment 73); f h) pectinate setae" httpUri="https://zenodo.org/record/228642/files/figure.png" pageId="4" pageNumber="205">Fig. 1</figureCitation>
d), except the pectinate which remains the same. Aciculae blunt, pale yellow at the edge and dark brown at base; 23 per parapodium in anterior body, only 1 after the first branchiae. Acicular setae, beginning from chaetiger 2235 (depending on specimen dimensions) are pale yellow, hooded and bidentate; teeth have almost 90° angle between them; subdistal almost twice as wide (
<figureCitation id="1335F6A4910EFFF205EAF805FD1878FB" box="[678,741,1967,1992]" captionStart="FIGURE 1" captionStartId="3.[151,250,1903,1925]" captionTargetBox="[188,1379,213,1853]" captionTargetId="figure@3.[180,1390,195,1864]" captionTargetPageId="3" captionText="FIGURE 1. Marphysa adenensis (GP- 4 B) a) Anterior long capillary seta (segment 10); b) anterior composite falcigers (segment 10); c) branchial-body composite falciger; d) posterior composite falcigers (segment 74); e) acicular seta (segment 73); f h) pectinate setae" httpUri="https://zenodo.org/record/228642/files/figure.png" pageId="4" pageNumber="205">Fig. 1</figureCitation>
e, 2d).
</paragraph>
</subSubSection>
<caption id="DF71BAA9910FFFF307DBFAB3FBC87A13" box="[151,1077,1305,1327]" httpUri="https://zenodo.org/record/228644/files/figure.png" pageId="5" pageNumber="206" targetBox="[310,1277,199,1280]" targetPageId="5">
<paragraph id="8BB1EA21910FFFF307DBFAB3FBC87A13" blockId="5.[151,1077,1305,1327]" box="[151,1077,1305,1327]" pageId="5" pageNumber="206">
<emphasis id="B97A3633910FFFF307DBFAB3FEEF7A12" bold="true" box="[151,274,1305,1327]" pageId="5" pageNumber="206">FIGURE 3.</emphasis>
<taxonomicName id="4C0E91A2910FFFF30655FAB3FE177A13" box="[281,490,1305,1327]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="5" pageNumber="206" phylum="Annelida" rank="species" species="adenensis">
<emphasis id="B97A3633910FFFF30655FAB3FE177A13" box="[281,490,1305,1327]" italics="true" pageId="5" pageNumber="206">Marphysa adenensis</emphasis>
</taxonomicName>
(GP-4B). Maxillary apparatus; Mx. IIV: Maxillary IIV.
</paragraph>
</caption>
<subSubSection id="C314B9AA910FFFF3078BFAFDFA777AC3" pageId="5" pageNumber="206" type="distribution">
<paragraph id="8BB1EA21910FFFF3078BFAFDFB777A8B" blockId="5.[151,1436,1367,2004]" pageId="5" pageNumber="206">
<emphasis id="B97A3633910FFFF3078BFAFDFE9D7A52" bold="true" box="[199,352,1367,1392]" pageId="5" pageNumber="206">Distribution.</emphasis>
<emphasis id="B97A3633910FFFF3062AFAFDFE3C7A52" box="[358,449,1367,1391]" italics="true" pageId="5" pageNumber="206">Red Sea</emphasis>
: Aden (
<bibRefCitation id="EF9F97D0910FFFF30556FAFDFD497A53" author="Gravier" box="[538,692,1367,1392]" pageId="5" pageNumber="206" refString="Gravier, C. (1900) Contribution a l' etude des Annelides Polychaetes de la Mer Rouge. Premiere partie. Nouvelles Archives du Museum d' Historie Naturelle Paris, Series 4, 2 (2), 137 - 282." type="journal article" year="1900">Gravier 1900</bibRefCitation>
),
<emphasis id="B97A3633910FFFF3058BFAFDFCA27A53" box="[711,863,1367,1391]" italics="true" pageId="5" pageNumber="206">Indian Ocean</emphasis>
:
<collectingCountry id="F319AAB1910FFFF30422FAFDFC087A4C" box="[878,1013,1367,1392]" name="Madagascar" pageId="5" pageNumber="206">Madagascar</collectingCountry>
(
<bibRefCitation id="EF9F97D0910FFFF30348FAFDFB8B7A4C" author="Day" box="[1028,1142,1367,1392]" pageId="5" pageNumber="206" refString="Day, J. H. (1962) Polychaeta from several localities in the Western Indian Ocean Proceeding of the Zoological Society of London, 139 (4), 627 - 656. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1962. tb 01597. x" type="journal article" year="1962">Day 1962</bibRefCitation>
),
<collectingCountry id="F319AAB1910FFFF303C5FAFDFB157A4C" box="[1161,1256,1367,1392]" name="Réunion" pageId="5" pageNumber="206">Reunion</collectingCountry>
Island (
<bibRefCitation id="EF9F97D0910FFFF3020DFAFDFF047AAF" author="Bigot" pageId="5" pageNumber="206" refString="Bigot, L., Gremare, A., Amoroux, J. M., Frouin, P., Maire, O. &amp; Gaertner, J. C. (2008) Assessment of the ecological quality status of soft-bottoms in Reunion Island (tropical southwest Indian Ocean) using AZTI marine biotic indices. Marine Pollution Bulletin, 56, 704 - 722. http: // dx. doi. org / 10.1016 / j. marpolbul. 2007.12.020" type="journal article" year="2008">
Bigot
<emphasis id="B97A3633910FFFF302C4FAF2FF507AAF" italics="true" pageId="5" pageNumber="206">et al</emphasis>
. 2008
</bibRefCitation>
),
<emphasis id="B97A3633910FFFF30643FAD1FE4C7AAE" box="[271,433,1403,1427]" italics="true" pageId="5" pageNumber="206">Pacific Ocean</emphasis>
:
<collectingCountry id="F319AAB1910FFFF3068FFAD6FDB47AA8" box="[451,585,1404,1428]" name="Hong Kong" pageId="5" pageNumber="206">Hong Kong</collectingCountry>
(
<bibRefCitation id="EF9F97D0910FFFF30516FAD1FD287AAF" author="Mak" box="[602,725,1403,1428]" pageId="5" pageNumber="206" refString="Mak, P. M. S. (1980) The coral associated polychaetes of Hong Kong, with special references to the serpulids. In: Morton, B. &amp; Tseng, C. K. (Eds.), Proceedings of the First International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China, Hong Kong. Hong Kong University Press, Hong Kong, pp. 595 - 617." type="book chapter" year="1980">Mak 1980</bibRefCitation>
), Andaman Sea (
<bibRefCitation id="EF9F97D0910FFFF304D7FAD1FB587AAF" author="Aungtonya" box="[923,1189,1403,1427]" pageId="5" pageNumber="206" refString="Aungtonya, C., Thaipal, S. &amp; Bussarawit, S. (2002) A list of Polychaetes (Annelida) in the reference collection database of the phuket marine biological center, Thailand. Phuket Marine Biological Center Special Publication, 24, 21 - 32." type="journal article" year="2002">
Aungtonya
<emphasis id="B97A3633910FFFF30368FAD6FBAA7AAF" box="[1060,1111,1403,1427]" italics="true" pageId="5" pageNumber="206">et al</emphasis>
. 2002
</bibRefCitation>
).
<emphasis id="B97A3633910FFFF303F6FAD1FA687AAE" box="[1210,1429,1403,1427]" italics="true" pageId="5" pageNumber="206">Mediterranean Sea</emphasis>
: Gera Gulf, Lesvos Island; Mytilene Strait (East Aegean Sea), Methoni (South Ionian Sea).
</paragraph>
<paragraph id="8BB1EA21910FFFF3078BFA69FA777AC3" blockId="5.[151,1436,1367,2004]" pageId="5" pageNumber="206">
<emphasis id="B97A3633910FFFF3078BFA69FED67AE6" bold="true" box="[199,299,1475,1500]" pageId="5" pageNumber="206">Ecology.</emphasis>
Soft mud (
<collectingCountry id="F319AAB1910FFFF306FBFA69FDB97AE0" box="[439,580,1475,1500]" name="Madagascar" pageId="5" pageNumber="206">Madagascar</collectingCountry>
), Muddy sand with sea-shells (Andaman Sea), coral communities (
<collectingCountry id="F319AAB1910FFFF30210FA69FF277AC3" name="Hong Kong" pageId="5" pageNumber="206">Hong Kong</collectingCountry>
).
<taxonomicName id="4C0E91A2910FFFF307A0FA42FE907AC3" box="[236,365,1512,1535]" class="Liliopsida" family="Posidoniaceae" genus="Posidonia" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="oceanica">
<emphasis id="B97A3633910FFFF307A0FA42FE907AC3" box="[236,365,1512,1535]" italics="true" pageId="5" pageNumber="206">P. oceanica</emphasis>
</taxonomicName>
beds and two individuals have been found especially in shoot sheaths micro-habitat (
<collectingCountry id="F319AAB1910FFFF30265FA4DFA827AC3" box="[1321,1407,1511,1535]" name="Greece" pageId="5" pageNumber="206">Greece</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C314B9AA910FFFF0078BF9A1FA7F7D01" lastPageId="6" lastPageNumber="207" pageId="5" pageNumber="206" type="discussion">
<paragraph id="8BB1EA21910FFFF0078BF9A1FA7F7D01" blockId="5.[151,1436,1367,2004]" lastBlockId="6.[151,1436,151,573]" lastPageId="6" lastPageNumber="207" pageId="5" pageNumber="206">
<emphasis id="B97A3633910FFFF3078BF9A1FEC6791E" bold="true" box="[199,315,1547,1572]" pageId="5" pageNumber="206">Remarks.</emphasis>
<taxonomicName id="4C0E91A2910FFFF30608F9A1FDD0791F" box="[324,557,1547,1571]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="5" pageNumber="206" phylum="Annelida" rank="species" species="adenensis">
<emphasis id="B97A3633910FFFF30608F9A1FDD0791F" box="[324,557,1547,1571]" italics="true" pageId="5" pageNumber="206">Marphysa adenensis</emphasis>
</taxonomicName>
is the first species of the genus described as having present only composite falcigers in inferior setae and branchiae limited to the anterior body. Although the blades of the falcigers are referred as “long and narrow” both in the original description and the subsequent description by
<bibRefCitation id="EF9F97D0910FFFF303A2F9FEFA8F7950" author="Day" box="[1262,1394,1619,1644]" pageId="5" pageNumber="206" refString="Day, J. H. (1963 a) The polychaete fauna of South Africa. Part 8: New species and records from grab samples and dredgings. Bulletin of the British Museum of Natural History Zoology, 10 (7), 383 - 445." type="journal article" year="1963">Day (1963)</bibRefCitation>
, no remarks are made for the presence of length gradation among the blades. The re-examination of the
<typeStatus id="54B55483910FFFF303B8F9DDFAAB79AC" box="[1268,1366,1655,1680]" pageId="5" pageNumber="206" type="holotype">holotype</typeStatus>
in the present study showed that there is a length gradation in the branchial region. The Mediterranean specimens differ from the
<typeStatus id="54B55483910FFFF307B0F915FEA379E4" box="[252,350,1727,1752]" pageId="5" pageNumber="206" type="holotype">holotype</typeStatus>
by having a stronger gradation of falciger blades-length (more than twice as long, rather than one and half). However, there are a lot of falcigers missing blades in the
<typeStatus id="54B55483910FFFF3048EF949FBDA79C0" box="[962,1063,1763,1788]" pageId="5" pageNumber="206" type="holotype">holotype</typeStatus>
, after a century of preservation, especially in the first segments. Maxillary apparatus has been dissected out from the
<typeStatus id="54B55483910FFFF3032BF8ADFB31781C" box="[1127,1228,1799,1824]" pageId="5" pageNumber="206" type="holotype">holotype</typeStatus>
, but the maxillary formula of the Mediterranean specimens differ from the published formula by
<bibRefCitation id="EF9F97D0910FFFF3035CF886FB697878" author="Day" box="[1040,1172,1835,1860]" pageId="5" pageNumber="206" refString="Day, J. H. (1967) A monograph on the polychaeta of southern Africa. Part I. Errantia. British Museum (Natural History), London, 458 pp." type="book" year="1967">Day (1967)</bibRefCitation>
. In particular, the latter have more teeth in maxillary IV (4+
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the Mediterranean specimens, than 7+
<quantity id="4CF647C4910FFFF30365F8E5FB9E7854" box="[1065,1123,1871,1896]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="5" pageNumber="206" unit="in" value="11.0">11 in</quantity>
the
<typeStatus id="54B55483910FFFF303D4F8E5FB037854" box="[1176,1278,1871,1896]" pageId="5" pageNumber="206" type="holotype">holotype</typeStatus>
), as well as a slight variation in MxII (5-7+
<quantity id="4CF647C4910FFFF306AEF8D9FDDE78B0" box="[482,547,1907,1932]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="5" pageNumber="206" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">7-8 in</quantity>
the Mediterranean specimens, than 7+
<quantity id="4CF647C4910FFFF30498F8D9FC0078B0" box="[980,1021,1907,1932]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="5" pageNumber="206" unit="in" value="8.0">8 in</quantity>
the
<typeStatus id="54B55483910FFFF30360F8D9FB6D78B0" box="[1068,1168,1907,1932]" pageId="5" pageNumber="206" type="holotype">holotype</typeStatus>
) and in MxIII (6-7+0 in the Mediterranean specimens, than 7+0 in the
<typeStatus id="54B55483910FFFF305EBF83DFCF0788C" box="[679,781,1943,1968]" pageId="5" pageNumber="206" type="holotype">holotype</typeStatus>
), varying according to body size. However, the published formula could be considered inadequate, since it seems to be based only on 3 individuals in total (the
<typeStatus id="54B55483910FFFF303B1F811FAA278E8" box="[1277,1375,1979,2004]" pageId="5" pageNumber="206" type="holotype">holotype</typeStatus>
from Aden;
<bibRefCitation id="EF9F97D0910CFFF007A9FF3DFE807F8C" author="Gravier" box="[229,381,151,176]" pageId="6" pageNumber="207" refString="Gravier, C. (1900) Contribution a l' etude des Annelides Polychaetes de la Mer Rouge. Premiere partie. Nouvelles Archives du Museum d' Historie Naturelle Paris, Series 4, 2 (2), 137 - 282." type="journal article" year="1900">Gravier 1900</bibRefCitation>
and two individual collected from
<collectingCountry id="F319AAB1910CFFF00457FF3DFC5B7F8C" box="[795,934,151,176]" name="Madagascar" pageId="6" pageNumber="207">Madagascar</collectingCountry>
;
<bibRefCitation id="EF9F97D0910CFFF004F8FF32FBDB7F8C" author="Day" box="[948,1062,151,176]" pageId="6" pageNumber="207" refString="Day, J. H. (1962) Polychaeta from several localities in the Western Indian Ocean Proceeding of the Zoological Society of London, 139 (4), 627 - 656. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1962. tb 01597. x" type="journal article" year="1962">Day 1962</bibRefCitation>
) and this number should not be enough to cover the interspecies variation. Besides the above differences, the Mediterranean specimens show important similarities with the
<typeStatus id="54B55483910CFFF006B4FF75FDA07FC4" box="[504,605,223,248]" pageId="6" pageNumber="207" type="holotype">holotype</typeStatus>
, namely in having: a) 12 pectinate setae all along the body, up to eight teeth, b) pale yellow acicular setae, hooded and bidentate with teeth that have almost 90° angle between them and the subdistal only slightly longer than the distal, but twice as wide.
<taxonomicName id="4C0E91A2910CFFF004C5FE82FB8D7E7C" box="[905,1136,296,320]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="6" pageNumber="207" phylum="Annelida" rank="species" species="adenensis">
<emphasis id="B97A3633910CFFF004C5FE82FB8D7E7C" box="[905,1136,296,320]" italics="true" pageId="6" pageNumber="207">Marphysa adenensis</emphasis>
</taxonomicName>
is very similar to
<emphasis id="B97A3633910CFFF00277FE83FAAF7E7C" box="[1339,1362,297,320]" italics="true" pageId="6" pageNumber="207">M</emphasis>
. sp. A from the Gulf of
<collectingCountry id="F319AAB1910CFFF00629FEE6FE437E59" box="[357,446,332,357]" name="Mexico" pageId="6" pageNumber="207">Mexico</collectingCountry>
. Although the personal examination of this species
<typeStatus id="54B55483910CFFF00364FEE7FBA57E59" box="[1064,1112,333,357]" pageId="6" pageNumber="207">type</typeStatus>
materials was not possible, according to the published description (
<bibRefCitation id="EF9F97D0910CFFF00516FEC5FD117EB4" author="Gathof" box="[602,748,367,392]" pageId="6" pageNumber="207" refString="Gathof, J. M. (1984) Chapter 40 Eunicidae. In: Uebelacker, J. M. &amp; Johnson, P. G. (Eds.), Taxonomic guide to the polychaetes of the northern Gulf of Mexico. Vol. 6. Final Report to the Minerals Management Service, contract 14 - 12 - 001 - 29091. Barry A. Vittor &amp; Assoc. Inc. Mobile, Alabama, pp. 1 - 31. [pp. 40.1 - 40.30]" type="book chapter" year="1984">Gathof 1984</bibRefCitation>
) the two species are very similar in terms of strong falciger blades-length gradation (twice as long in figure 40-14ef), acicular shape (bidentate, hooded, 90° angle between the teeth and subdistal is wider than distal in figure 40-14h) and maxillary formula close to the interspecies variation (I:1+1, II:5-6+7, III:5-7+0, IV:2-4+5-12). The relation of
<emphasis id="B97A3633910CFFF004C6FE77FC5C7EC8" box="[906,929,477,500]" italics="true" pageId="6" pageNumber="207">Μ</emphasis>
. sp. A to
<taxonomicName id="4C0E91A2910CFFF00343FE77FB5E7EC8" box="[1039,1187,476,500]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="6" pageNumber="207" phylum="Annelida" rank="species" species="adenensis">
<emphasis id="B97A3633910CFFF00343FE77FBDB7EC8" box="[1039,1062,477,500]" italics="true" pageId="6" pageNumber="207">M</emphasis>
.
<emphasis id="B97A3633910CFFF00379FE76FB5E7EC8" box="[1077,1187,476,500]" italics="true" pageId="6" pageNumber="207">adenensis</emphasis>
</taxonomicName>
has to be defined in a future study, since there is a possibility to be a synonym species. Finally, it differs from the other species of Group C1 from
<collectingCountry id="F319AAB1910CFFF007B0FD8EFEA27D01" box="[252,351,548,573]" name="United States of America" pageId="6" pageNumber="207">America</collectingCountry>
,
<taxonomicName id="4C0E91A2910CFFF00626FD8FFE107D00" box="[362,493,548,572]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="6" pageNumber="207" phylum="Annelida" rank="species" species="conferta">
<emphasis id="B97A3633910CFFF00626FD8FFE107D00" box="[362,493,548,572]" italics="true" pageId="6" pageNumber="207">M. conferta</emphasis>
</taxonomicName>
, mainly in having pectinate setae of fewer teeth (based on published description).
</paragraph>
</subSubSection>
</treatment>
</document>