151 lines
20 KiB
XML
151 lines
20 KiB
XML
<document id="7B8A0A5DEF8E20E774C86F1A66EE438F" ID-DOI="10.11646/zootaxa.4090.1.1" ID-GBIF-Dataset="e6b35f5a-435e-4473-b29e-1e4d842f84b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="263299" ID-ZooBank="203BAC25-B542-48FE-B5AD-EBA8C0285833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457964633549" checkinUser="plazi" docAuthor="Fallon, Phillip J." docDate="2016" docId="039F87C4FA69FFA5CBAFB8E5FA73FDA7" docLanguage="en" docName="15612-5136-1-PB.pdf.imf" docOrigin="Zootaxa 4090 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Drilliidae Olsson 1964" docType="treatment" docVersion="12" lastPageNumber="13" masterDocId="FFA6FFBCFA65FFA8CB38B91BFFE8FFAA" masterDocTitle="Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species" masterLastPageNumber="363" masterPageNumber="1" pageNumber="13" updateTime="1698184378282" updateUser="plazi">
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<mods:titleInfo id="23ADB86B10B8FC88D14FEFD130CE2867">
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<mods:title id="D0E260F628001F6D86BD25F6C78F51AD">Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species</mods:title>
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<mods:name id="D3CA47364D293B20C1E6D49FC15ACEBA" type="personal">
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<mods:namePart id="8BF69ACADE89F2A2D124FB26F3216FA9">Fallon, Phillip J.</mods:namePart>
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<mods:typeOfResource id="B30C2B244AF29032FC528E576C92669A">text</mods:typeOfResource>
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<mods:title id="8C44C754F6290F9098EC6B5430CE10B6">Zootaxa</mods:title>
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<mods:part id="6D37FAD1B1CEC2CABEC8F2B20E70687B">
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<mods:date id="5CD04A74CE9E707E6D1A8897ED21CD0C">2016</mods:date>
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<mods:detail id="7817A71BDB61396CD7107C180693B21B" type="volume">
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<mods:number id="CC91BD5008F4DE37BB61FA8AB22C7E97">4090</mods:number>
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<mods:detail id="F9B0ADA5ED2ABEB01D50E10E7B66370F" type="issue">
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<mods:number id="2084D1BF347CAF87C619414558B4F9E9">1</mods:number>
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<mods:classification id="24766681BADC897D962113F206995676">journal article</mods:classification>
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<mods:identifier id="A0D820988F7B675EE1D002E3648C9F09" type="DOI">10.11646/zootaxa.4090.1.1</mods:identifier>
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<mods:identifier id="D69516CCB2FC5B9F1CC24364A030C961" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8D2467DD1DD810B5A4868582421EA340" type="Zenodo-Dep">263299</mods:identifier>
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<treatment id="039F87C4FA69FFA5CBAFB8E5FA73FDA7" ID-DOI="http://doi.org/10.5281/zenodo.6076235" ID-GBIF-Taxon="117911506" ID-Zenodo-Dep="6076235" LSID="urn:lsid:plazi:treatment:039F87C4FA69FFA5CBAFB8E5FA73FDA7" httpUri="http://treatment.plazi.org/id/039F87C4FA69FFA5CBAFB8E5FA73FDA7" lastPageId="13" lastPageNumber="13" pageId="12" pageNumber="13">
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<subSubSection id="C32C6559FA69FFA4CBAFB8E5FE06FDB2" box="[151,494,510,536]" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="8B8936D2FA69FFA4CBAFB8E5FE06FDB2" blockId="12.[151,494,510,536]" box="[151,494,510,536]" pageId="12" pageNumber="13">
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<heading id="D0C181BEFA69FFA4CBAFB8E5FE06FDB2" bold="true" box="[151,494,510,536]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
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<taxonomicName id="4C364D51FA69FFA4CBAFB8E5FE06FDB2" ID-CoL="7NHDF" authority="Olsson, 1964" authorityName="Olsson" authorityYear="1964" box="[151,494,510,536]" class="Gastropoda" family="Drilliidae" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="6" phylum="Mollusca" rank="family">
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<emphasis id="B942EAC0FA69FFA4CBAFB8E5FE06FDB2" bold="true" box="[151,494,510,536]" pageId="12" pageNumber="13">DRILLIIDAE Olsson, 1964</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FA69FFA4CBAFBB5EFCC4FCB8" pageId="12" pageNumber="13" type="diagnosis">
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<paragraph id="8B8936D2FA69FFA4CBAFBB5EFCC4FCB8" blockId="12.[151,1436,581,966]" pageId="12" pageNumber="13">
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<emphasis id="B942EAC0FA69FFA4CBAFBB5EFEF9FDF4" bold="true" box="[151,273,581,606]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
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According to Kilburn (1988: 172): species with a claviform shell facies (tall spire, truncated base, and parietal lobe) that possess a drilliine radula, formula 1-1-R-1-1 with rachidians small to large, laterals broad arched and pectinate, marginals long, sharply pointed, with axial ribs as the dominant sculptural element, and operculum oblanceolate with a terminal nucleus. Taylor
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<emphasis id="B942EAC0FA69FFA4C9A2BBA9FD3DFD60" box="[666,725,689,714]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
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(1993) included some additional morphological detail in their diagnosis: sculpture usually well developed, anal sinus deep, located on the shoulder, sometimes made sub-tubular by a parietal lobe; protoconch smooth, sometimes carinate.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FA69FFA4CBFFBA05FBC1FA75" pageId="12" pageNumber="13" type="discussion">
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<paragraph id="8B8936D2FA69FFA4CBFFBA05FCE0FC6F" blockId="12.[151,1436,581,966]" pageId="12" pageNumber="13">
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<emphasis id="B942EAC0FA69FFA4CBFFBA05FED3FC9F" bold="true" box="[199,315,798,821]" pageId="12" pageNumber="13">Remarks.</emphasis>
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The most distinguishing characteristic of the
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<taxonomicName id="4C364D51FA69FFA4C87BBA06FC46FC9C" box="[835,942,797,822]" class="Gastropoda" family="Drilliidae" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="family">Drilliidae</taxonomicName>
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is the structure of the radula. However, the radula has been examined for only a few of the TWA drilliid taxa (Taylor
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<emphasis id="B942EAC0FA69FFA4C8D4BA59FBCFFCF0" box="[1004,1063,833,858]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
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, 1993; Maes, 1984; and Tippett, 1995), so genera have been included in the family on a more convenient combination of shell morphology and anatomical examination in the most recent published works (Bouchet
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<emphasis id="B942EAC0FA69FFA4C8FBBA91FC17FC08" box="[963,1023,905,930]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
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, 2011, and Puillandre
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<emphasis id="B942EAC0FA69FFA4CE30BA91FAACFC08" box="[1288,1348,905,930]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
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, 2011). Shell characteristics common to all TWA
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<taxonomicName id="4C364D51FA69FFA4C953BAB6FD3EFC6C" box="[619,726,941,966]" class="Gastropoda" family="Drilliidae" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="family">Drilliidae</taxonomicName>
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are:
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</paragraph>
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<paragraph id="8B8936D2FA69FFA4CBAFBAEDFD57FB98" blockId="12.[151,1436,1013,1362]" pageId="12" pageNumber="13">1. A fusiform shell, truncated anteriorly in most genera, usually glossy, with a tall spire, and a large last whorl usually 50% or more of the total shell length;</paragraph>
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<paragraph id="8B8936D2FA69FFA4CBAFBD25FC37FBFF" blockId="12.[151,1436,1013,1362]" box="[151,991,1085,1110]" pageId="12" pageNumber="13">2. Smooth protoconch of 1½–3 whorls. A median carina present in some;</paragraph>
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<paragraph id="8B8936D2FA69FFA4CBAFBD79FD0EFB34" blockId="12.[151,1436,1013,1362]" pageId="12" pageNumber="13">3. Sculpture primarily of axial ribs; some genera with spiral grooves, incised lines, or threads that may be between or override ribs; subsutural cord absent;</paragraph>
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<paragraph id="8B8936D2FA69FFA4CBAFBDB1FB5FFB4F" blockId="12.[151,1436,1013,1362]" pageId="12" pageNumber="13">
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4. Surface microsculpture of spiral grooves or threads present on at least part of the shell surface in all but a single genus (
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<taxonomicName id="4C364D51FA69FFA4CA58BDD6FE09FB4C" box="[352,481,1229,1254]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA69FFA4CA58BDD6FE09FB4C" box="[352,481,1229,1254]" italics="true" pageId="12" pageNumber="13">Lissodrillia</emphasis>
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</taxonomicName>
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); incremental growth lines can be absent, microscopic or strong;
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</paragraph>
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<paragraph id="8B8936D2FA69FFA4CBAFBDE9FB11FAA0" blockId="12.[151,1436,1013,1362]" box="[151,1273,1265,1290]" pageId="12" pageNumber="13">5. Anal sinus on whorl shoulder adjacent to suture, shallow to moderately deep, usually U-shaped;</paragraph>
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<paragraph id="8B8936D2FA69FFA4CBAFBC0DFB81FA84" blockId="12.[151,1436,1013,1362]" box="[151,1129,1301,1326]" pageId="12" pageNumber="13">6. Parietal callus or lobe at the posterior end of the aperture beside the anal sinus; and</paragraph>
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<paragraph id="8B8936D2FA69FFA4CBAFBC21FD52FAF8" blockId="12.[151,1436,1013,1362]" box="[151,698,1337,1362]" pageId="12" pageNumber="13">7. Varix present in fully developed individuals.</paragraph>
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<paragraph id="8B8936D2FA69FFA4CBFFBC65FBC1FA75" blockId="12.[151,1436,1406,1683]" pageId="12" pageNumber="13">Characters are identified that are diagnostic for each genus so that species may be properly placed. Most TWA genera have been fairly well defined by past workers; however, additional characteristics that distinguish one from another, such as shell surface microsculpture, have been added to their diagnoses.</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FA69FFA5CBFFBCF0FA73FDA7" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="key">
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<paragraph id="8B8936D2FA69FFA4CBFFBCF0FB4FF938" blockId="12.[151,1436,1406,1683]" pageId="12" pageNumber="13">
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<emphasis id="B942EAC0FA69FFA4CBFFBCF0FE9CF9A8" bold="true" box="[199,372,1515,1538]" pageId="12" pageNumber="13">Key to genera.</emphasis>
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The following key to shallow water western Atlantic drilliids is based on some aspects of shell form and detail of sculpture: axials, spirals, varix form and position, and microsculpture (growth increments or very fine striae and spirals, which may be grooves and/or threads). Some species may only marginally fit for the described character in this key. Confirmation of a species placement or clarification of characteristics of a genus can be made by reviewing the key characteristics paragraph in the introduction to that genus.
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</paragraph>
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<paragraph id="8B8936D2FA69FFA5CBAFBFA7FA73FDA7" blockId="12.[151,1436,1724,2038]" lastBlockId="13.[151,1436,151,525]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
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1. Sculpture includes strong spiral elements.................................................................. 2. - Shell mostly smooth with just axial ribs, or spirals, if present, much less prominent than ribs......................... 5. 2. Protoconch whorls smooth............................................................................. 3. - One or all protoconch whorls carinate..............................................................
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<taxonomicName id="4C364D51FA69FFA4CE0BBE08FA73F883" ID-CoL="7NVM5" box="[1331,1435,1811,1833]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA69FFA4CE0BBE08FA73F883" box="[1331,1435,1811,1833]" italics="true" pageId="12" pageNumber="13">Calliclava</emphasis>
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</taxonomicName>
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3. Varix ½-turn from edge of outer lip, dorsum of last whorl lacking ribs....................................
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<taxonomicName id="4C364D51FA69FFA4CE0ABE2BFA73F8EC" ID-CoL="7NNBL" box="[1330,1435,1840,1862]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA69FFA4CE0ABE2BFA73F8EC" box="[1330,1435,1840,1862]" italics="true" pageId="12" pageNumber="13">Agladrillia</emphasis>
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</taxonomicName>
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- Varix about ⅓-turn from edge of outer lip................................................................. 4. 4. Shoulders tabulate or angular..................................................................
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<taxonomicName id="4C364D51FA69FFA4CE2EBE70FA74F82B" ID-CoL="7NZQQ" box="[1302,1436,1899,1921]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA69FFA4CE2EBE70FA74F82B" box="[1302,1436,1899,1921]" italics="true" pageId="12" pageNumber="13">Clathrodrillia</emphasis>
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</taxonomicName>
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- Shoulders round, spirals usually very fine...........................................................
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<taxonomicName id="4C364D51FA69FFA4CE0BBE93FA73F834" ID-CoL="7NQMK" box="[1331,1435,1928,1950]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA69FFA4CE0BBE93FA73F834" box="[1331,1435,1928,1950]" italics="true" pageId="12" pageNumber="13">Bellaspira</emphasis>
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</taxonomicName>
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5. Fine spiral grooves present over most of the teleoconch surface................................................ 6. - Microscopic spirals mostly restricted to last whorl or anterior fasciole........................................... 8. 6. Spirals and striae subequal and closely spaced, but not packed..........................................
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<taxonomicName id="4C364D51FA69FFA4CE0ABEFAFA72F85C" ID-CoL="7P2D7" box="[1330,1434,2017,2038]" class="Gastropoda" family="Drilliidae" genus="Fenimorea" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA69FFA4CE0ABEFAFA72F85C" box="[1330,1434,2017,2038]" italics="true" pageId="12" pageNumber="13">Fenimorea</emphasis>
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</taxonomicName>
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- Spirals larger than growth striae, either packed, or widely spread............................................... 7. 7. Spirals are widely spaced incised lines. Very shallow anal sinus..........................................
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<taxonomicName id="4C364D51FA68FFA5CE0CB9AFFA73FF60" ID-CoL="7NQMK" box="[1332,1435,180,202]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA68FFA5CE0CB9AFFA73FF60" box="[1332,1435,180,202]" italics="true" pageId="13" pageNumber="14">Bellaspira</emphasis>
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</taxonomicName>
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- Spirals packed together; anal sinus a deep notch...................................................
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<taxonomicName id="4C364D51FA68FFA5CE2FB9CAFA73FF4D" ID-CoL="7P2TC" box="[1303,1435,209,231]" class="Gastropoda" family="Drilliidae" genus="Decoradrillia" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="277" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="B942EAC0FA68FFA5CE2FB9CAFA73FF4D" box="[1303,1435,209,231]" italics="true" pageId="13" pageNumber="14">Decoradrillia</emphasis>
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</taxonomicName>
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- Spirals of raised threads, coarse, shell feels sandpapery.................................................
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<taxonomicName id="4C364D51FA68FFA5CE0CB9F4FA73FEAF" ID-CoL="7P9MR" box="[1332,1435,239,261]" class="Gastropoda" family="Drilliidae" genus="Neodrillia" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA68FFA5CE0CB9F4FA73FEAF" box="[1332,1435,239,261]" italics="true" pageId="13" pageNumber="14">Neodrillia</emphasis>
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</taxonomicName>
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8. Ribs run suture-to-suture with only a slight change in sulcal area............................................... 9. - Ribs end at sulcus, or evanesce near suture on later whorls................................................... 10. 9. Microscopic spirals between ribs of last whorl, may be very faint.....................................
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<taxonomicName id="4C364D51FA68FFA5CE34B85CFA73FEF7" ID-CoL="7PHWK" box="[1292,1435,327,349]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA68FFA5CE34B85CFA73FEF7" box="[1292,1435,327,349]" italics="true" pageId="13" pageNumber="14">Syntomodrillia</emphasis>
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</taxonomicName>
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- Spirals on anterior fasciole only.................................................................
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<taxonomicName id="4C364D51FA68FFA5CE11B87FFA74FED0" ID-CoL="7P77Q" box="[1321,1436,356,378]" class="Gastropoda" family="Drilliidae" genus="Leptadrillia" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA68FFA5CE11B87FFA74FED0" box="[1321,1436,356,378]" italics="true" pageId="13" pageNumber="14">Leptadrillia</emphasis>
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</taxonomicName>
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- Spirals absent altogether........................................................................
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<taxonomicName id="4C364D51FA68FFA5CE10B89AFA74FE3D" ID-CoL="7P7PM" box="[1320,1436,385,407]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA68FFA5CE10B89AFA74FE3D" box="[1320,1436,385,407]" italics="true" pageId="13" pageNumber="14">Lissodrillia</emphasis>
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</taxonomicName>
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10. Varix ⅓-turn from edge of outer lip, not visible in ventral view.........................................
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<taxonomicName id="4C364D51FA68FFA5CE12B884FA73FE1F" ID-CoL="7PHCB" box="[1322,1435,415,437]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA68FFA5CE12B884FA73FE1F" box="[1322,1435,415,437]" italics="true" pageId="13" pageNumber="14">Splendrillia</emphasis>
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</taxonomicName>
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- Varix prominent, visible when shell is viewed ventrally, ¼-turn or less.......................................... 11. 11. Ribs usually to suture of later whorls; protoconch 1¾–2 whorls.........................................
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<taxonomicName id="4C364D51FA68FFA5CE08B8C2FA73FE45" ID-CoL="7NSCQ" box="[1328,1435,473,495]" class="Gastropoda" family="Drilliidae" genus="Cerodrillia" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FA68FFA5CE08B8C2FA73FE45" box="[1328,1435,473,495]" italics="true" pageId="13" pageNumber="14">Cerodrillia</emphasis>
|
||
</taxonomicName>
|
||
- Ribs usually absent in sulcus of later whorls; protoconch 2½–3 whorls...................................
|
||
<taxonomicName id="4C364D51FA68FFA5CE16B8ECFA73FDA7" box="[1326,1435,503,525]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FA68FFA5CE16B8ECFA73FDA7" box="[1326,1435,503,525]" italics="true" pageId="13" pageNumber="14">Douglassia</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |