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<document id="1532BC40069078F38F6689F4F8B79EB9" ID-DOI="10.11646/zootaxa.4090.1.1" ID-GBIF-Dataset="e6b35f5a-435e-4473-b29e-1e4d842f84b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="263299" ID-ZooBank="203BAC25-B542-48FE-B5AD-EBA8C0285833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457964633549" checkinUser="plazi" docAuthor="Fallon, Phillip J." docDate="2016" docId="039F87C4FA3AFFCACBAFBC2AFB1EFB4D" docLanguage="en" docName="15612-5136-1-PB.pdf.imf" docOrigin="Zootaxa 4090 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Clathrodrillia gibbosa Born 1778" docType="treatment" docVersion="13" lastPageNumber="99" masterDocId="FFA6FFBCFA65FFA8CB38B91BFFE8FFAA" masterDocTitle="Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species" masterLastPageNumber="363" masterPageNumber="1" pageNumber="96" updateTime="1698184378282" updateUser="plazi">
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<mods:title id="571DCBFB2E977393E5FAFFE4B05A8361">Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species</mods:title>
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<mods:namePart id="B551DC20BE338CC4EA5F93FEDC581E09">Fallon, Phillip J.</mods:namePart>
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<mods:date id="1E298798E61B2C57F405CDA57D484A44">2016</mods:date>
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<mods:identifier id="20137D6E545F101629F6461054991151" type="DOI">10.11646/zootaxa.4090.1.1</mods:identifier>
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<treatment id="039F87C4FA3AFFCACBAFBC2AFB1EFB4D" ID-DOI="http://doi.org/10.5281/zenodo.6076344" ID-GBIF-Taxon="117911730" ID-Zenodo-Dep="6076344" LSID="urn:lsid:plazi:treatment:039F87C4FA3AFFCACBAFBC2AFB1EFB4D" httpUri="http://treatment.plazi.org/id/039F87C4FA3AFFCACBAFBC2AFB1EFB4D" lastPageId="98" lastPageNumber="99" pageId="95" pageNumber="96">
<subSubSection id="C32C6559FA3AFFF7CBAFBC2AFBFBFA61" pageId="95" pageNumber="96" type="nomenclature">
<paragraph id="8B8936D2FA3AFFF7CBAFBC2AFDADFAE1" blockId="95.[151,581,1329,1389]" box="[151,581,1329,1355]" pageId="95" pageNumber="96">
<heading id="D0C181BEFA3AFFF7CBAFBC2AFDADFAE1" bold="true" box="[151,581,1329,1355]" fontSize="11" level="1" pageId="95" pageNumber="96" reason="1">
<taxonomicName id="4C364D51FA3AFFF7CBAFBC2AFDADFAE1" ID-CoL="VSYK" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,581,1329,1355]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA3AFFF7CBAFBC2AFDADFAE1" bold="true" box="[151,581,1329,1355]" pageId="95" pageNumber="96">
<emphasis id="B942EAC0FA3AFFF7CBAFBC2AFE49FAE1" bold="true" box="[151,417,1329,1355]" italics="true" pageId="95" pageNumber="96">Clathrodrillia gibbosa</emphasis>
(Born, 1778)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBC4FFF10FAC7" blockId="95.[151,581,1329,1389]" box="[151,248,1364,1389]" pageId="95" pageNumber="96">(Fig. 42)</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBC8CFBFBFA61" blockId="95.[151,1436,1430,2035]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBC8CFE02FA06" authority="Born, 1778: 325" authorityName="Born" authorityPageNumber="325" authorityYear="1778" box="[151,490,1430,1452]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosus">
<emphasis id="B942EAC0FA3AFFF7CBAFBC8CFED1FA06" box="[151,313,1430,1452]" italics="true" pageId="95" pageNumber="96">Murex gibbosus</emphasis>
Born, 1778: 325
</taxonomicName>
; Born (1780: 321, pl. 11, figs. 12, 13); Chemnitz (1795: 112–113, pl. 190, figs.
<date id="FF881012FA3AFFF7CE22BC8DFA78FA06" box="[1306,1424,1430,1452]" pageId="95" pageNumber="96" value="1829" valueMax="1830">1829–1830</date>
); Grant &amp; Gale (1931: 579, pl. 26, figs. 3a, b = Born’s figs.); E. H. Vokes (1971: 53).
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA3AFFC8CBAFBCCFFAFBFEED" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96" type="description">
<paragraph id="8B8936D2FA3AFFF7CBAFBCCFFC82FA40" blockId="95.[151,1436,1430,2035]" box="[151,874,1492,1514]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBCCFFE18FA40" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,496,1492,1514]" class="Gastropoda" family="Drilliidae" genus="Pleurotoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA3AFFF7CBAFBCCFFE8CFA40" box="[151,356,1492,1514]" italics="true" pageId="95" pageNumber="96">Pleurotoma gibbosa</emphasis>
(Born, 1778)
</taxonomicName>
: Sowerby I (1825: 58, cat. no. 1500).
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBCE9FEA5F98D" blockId="95.[151,1436,1430,2035]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBCE9FDF4F9A2" authority="Kiener 1840b: 35" authorityName="Kiener" authorityPageNumber="35" authorityYear="1840" box="[151,540,1522,1544]" class="Gastropoda" family="Drilliidae" genus="Pleurotoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA3AFFF7CBAFBCE9FE8BF9A2" box="[151,355,1522,1544]" italics="true" pageId="95" pageNumber="96">Pleurotoma gibbosa</emphasis>
Kiener 1840b: 35
</taxonomicName>
, pl. 12 [sic, 16], fig. 2. Kiener redescribed Born’s species and took authorship of the new combination.
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBF2BFB9DF9EC" blockId="95.[151,1436,1430,2035]" box="[151,1141,1584,1606]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBF2BFE8CF9EC" authority="Chemnitz" authorityName="Chemnitz" box="[151,356,1584,1606]" class="Gastropoda" family="Drilliidae" genus="Pleurotoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA3AFFF7CBAFBF2BFE8CF9EC" box="[151,356,1584,1606]" italics="true" pageId="95" pageNumber="96">Pleurotoma gibbosa</emphasis>
</taxonomicName>
“Chemnitz”: Reeve (1843b: pl. 5, sp. 30), an erroneous attribution by Reeve.
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBF55FB7EF9CE" blockId="95.[151,1436,1430,2035]" box="[151,1174,1614,1636]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBF55FEDAF9CE" authority="Chem" authorityName="Chem" box="[151,306,1614,1636]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA3AFFF7CBAFBF55FEDAF9CE" box="[151,306,1614,1636]" italics="true" pageId="95" pageNumber="96">Drillia gibbosa</emphasis>
</taxonomicName>
“Chem.”: H. &amp; A. Adams (1853: 90 [list]), an erroneous attribution by these authors.
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBF76FA7AF929" blockId="95.[151,1436,1430,2035]" box="[151,1426,1645,1667]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBF76FDA5F929" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,589,1645,1667]" class="Gastropoda" family="Drilliidae" genus="Pleurotoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa" subGenus="Drillia">
<emphasis id="B942EAC0FA3AFFF7CBAFBF76FE28F929" box="[151,448,1645,1667]" italics="true" pageId="95" pageNumber="96">Pleurotoma (Drillia) gibbosa</emphasis>
(Born, 1778)
</taxonomicName>
: Chenu (1859: 146, fig. 650 [as an example of the genus]); Paetel (1888: 63 [list]).
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBF97FBBBF908" blockId="95.[151,1436,1430,2035]" box="[151,1107,1676,1698]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBF97FDB4F908" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,604,1676,1698]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa" subGenus="Brachystoma">
<emphasis id="B942EAC0FA3AFFF7CBAFBF97FE27F908" box="[151,463,1676,1698]" italics="true" pageId="95" pageNumber="96">Drillia (Brachystoma) gibbosa</emphasis>
(Born, 1778)
</taxonomicName>
: Tryon (1884: 179, pl. 9, fig. 54 [= Reeve’s fig.]).
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBFB1FBDAF96A" blockId="95.[151,1436,1430,2035]" box="[151,1074,1706,1728]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBFB1FE92F96A" authority="Reeve" authorityName="Reeve" box="[151,378,1706,1728]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA3AFFF7CBAFBFB1FE92F96A" box="[151,378,1706,1728]" italics="true" pageId="95" pageNumber="96">Clathrodrillia gibbosa</emphasis>
</taxonomicName>
“Reeve”: Dall (1918a: 317, 323) is an erroneous attribution by Dall.
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBFD1FD11F975" blockId="95.[151,1436,1430,2035]" box="[151,761,1737,1759]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBFD1FE50F975" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,440,1737,1759]" class="Gastropoda" family="Turridae" genus="Turris" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA3AFFF7CBAFBFD1FEC3F975" box="[151,299,1737,1759]" italics="true" pageId="95" pageNumber="96">Turris gibbosa</emphasis>
(Born, 1778)
</taxonomicName>
: M. Smith (1940: 80, sp. 1110).
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBFF3FC53F954" blockId="95.[151,1436,1430,2035]" box="[151,955,1768,1790]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBFF3FD8DF954" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,613,1768,1790]" class="Gastropoda" family="Drilliidae" genus="Clavus" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa" subGenus="Clathrodrillia">
<emphasis id="B942EAC0FA3AFFF7CBAFBFF3FE30F954" box="[151,472,1768,1790]" italics="true" pageId="95" pageNumber="96">Clavus (Clathrodrillia) gibbosa</emphasis>
(Born, 1778)
</taxonomicName>
: Korobkov (1955: pl. 104, fig. 6).
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBE1DFA9DF8B6" blockId="95.[151,1436,1430,2035]" box="[151,1397,1798,1820]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBE1DFDB7F8B6" authority="Weisbord, 1962: 437" authorityName="Weisbord" authorityPageNumber="437" authorityYear="1962" box="[151,607,1798,1820]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="mareana">
<emphasis id="B942EAC0FA3AFFF7CBAFBE1DFE6BF8B6" box="[151,387,1798,1820]" italics="true" pageId="95" pageNumber="96">Clathrodrillia mareana</emphasis>
Weisbord, 1962: 437
</taxonomicName>
, pl. 41, figs. 16–17 [fossil], is a synonym according to Woodring (1970: 383).
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBE3EFB32F83D" blockId="95.[151,1436,1430,2035]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBE3EFDE2F891" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,522,1829,1851]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA3AFFF7CBAFBE3EFE94F891" box="[151,380,1829,1851]" italics="true" pageId="95" pageNumber="96">Clathrodrillia gibbosa</emphasis>
(Born, 1778)
</taxonomicName>
: Powell (1966: pl. 11, fig. 2); Kaicher (1984c: card 3944); K. Sunderland (1991: 14, text photo); Díaz (1994: 40 [incorrectly gives authorship date as 1780]); Diaz &amp; Puyana (1994: 226, pl. 68, fig. 896); Rolán &amp; Espinosa (1995); C.-K. Chang (2001a: 18, figs. 3–8; 215, figs. 19–22); Macsotay &amp; Campos (2001: 108, pl. 9, figs. 1, 8); Williams (2005: species 1542); Massemin
<emphasis id="B942EAC0FA3AFFF7C94FBE99FD43F83D" box="[631,683,1921,1943]" italics="true" pageId="95" pageNumber="96">et al.</emphasis>
(2009: 205); Daccarett &amp; Bossio (2011: 129, fig. 785).
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBEBBFC48F81C" blockId="95.[151,1436,1430,2035]" box="[151,928,1952,1974]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBEBBFD8BF81C" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,611,1952,1974]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa" subGenus="Clathrodrillia">
<emphasis id="B942EAC0FA3AFFF7CBAFBEBBFE3EF81C" box="[151,470,1952,1974]" italics="true" pageId="95" pageNumber="96">Drillia (Clathrodrillia) gibbosa</emphasis>
(Born, 1778)
</taxonomicName>
: Abbott (1974: 270, fig. 2999).
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBEA5FCE7F87E" blockId="95.[151,1436,1430,2035]" box="[151,783,1982,2004]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBEA5FE18F87E" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,496,1982,2004]" class="Gastropoda" family="Pseudomelatomidae" genus="Crassispira" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA3AFFF7CBAFBEA5FE8BF87E" box="[151,355,1982,2004]" italics="true" pageId="95" pageNumber="96">Crassispira gibbosa</emphasis>
(Born, 1778)
</taxonomicName>
: Sutty (1986: 125, fig. 139).
</paragraph>
<paragraph id="8B8936D2FA3AFFF7CBAFBEC6FB4BF859" blockId="95.[151,1436,1430,2035]" box="[151,1187,2013,2035]" pageId="95" pageNumber="96">
<taxonomicName id="4C364D51FA3AFFF7CBAFBEC6FE57F859" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,447,2013,2035]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA3AFFF7CBAFBEC6FEDAF859" box="[151,306,2013,2035]" italics="true" pageId="95" pageNumber="96">Drillia gibbosa</emphasis>
(Born, 1778)
</taxonomicName>
: Vermeij (1978: 274); Harasewych &amp; Moretzsohn (2010: 573, text figs.).
</paragraph>
<paragraph id="8B8936D2FA05FFC8CBAFB982FCFEFF04" blockId="96.[151,1437,152,327]" box="[151,790,152,174]" pageId="96" pageNumber="97">
<emphasis id="B942EAC0FA05FFC8CBAFB982FCFEFF04" bold="true" box="[151,790,152,174]" pageId="96" pageNumber="97">Not this species, fossil, or unconfirmed extralimital reports.</emphasis>
</paragraph>
<paragraph id="8B8936D2FA05FFC8CBAFB9ADFAFDFF66" blockId="96.[151,1437,152,327]" box="[151,1301,182,204]" pageId="96" pageNumber="97">
<taxonomicName id="4C364D51FA05FFC8CBAFB9ADFE18FF66" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,496,182,204]" class="Gastropoda" family="Drilliidae" genus="Pleurotoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA05FFC8CBAFB9ADFE8CFF66" box="[151,356,182,204]" italics="true" pageId="96" pageNumber="97">Pleurotoma gibbosa</emphasis>
(Born, 1778)
</taxonomicName>
: Simpson (1887c: 54 [Anna Maria Key, W Florida: “one slightly worn shell”]).
</paragraph>
<paragraph id="8B8936D2FA05FFC8CBAFB9CEFEA6FEA0" blockId="96.[151,1437,152,327]" pageId="96" pageNumber="97">
<taxonomicName id="4C364D51FA05FFC8CBAFB9CEFD4BFF41" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,675,213,235]" class="Gastropoda" family="Pseudomelatomidae" genus="Crassispira" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="gibbosa" subGenus="Clathrodrillia">
<emphasis id="B942EAC0FA05FFC8CBAFB9CEFDE7FF41" box="[151,527,213,235]" italics="true" pageId="96" pageNumber="97">Crassispira (Clathrodrillia) gibbosa</emphasis>
(Born, 1778)
</taxonomicName>
: Wenz (1943: 1408, fig. 3978 [fossil occurrence]); Glibert (1960 [fossil occurrence]).
</paragraph>
<paragraph id="8B8936D2FA05FFC8CBAFB809FAFBFEED" blockId="96.[151,1437,152,327]" pageId="96" pageNumber="97">
<taxonomicName id="4C364D51FA05FFC8CBAFB809FDEEFE82" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,518,274,296]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA05FFC8CBAFB809FE92FE82" box="[151,378,274,296]" italics="true" pageId="96" pageNumber="97">Clathrodrillia gibbosa</emphasis>
(Born, 1778)
</taxonomicName>
: Fargo (1953: 368; pl. 16, figs. 1, 1a [fossil occurrence]); Weisbord (1962: 435, pl. 41, figs. 13–15 [fossil occurrence]); Olsson (1964: 96 [fossil occurrence]); Woodring (1970: 383 [fossil occurrence]).
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA05FFC8CBAFB86EFB89F937" pageId="96" pageNumber="97" type="materials_examined">
<paragraph id="8B8936D2FA05FFC8CBAFB86EFECEFE7E" blockId="96.[151,1437,372,2017]" pageId="96" pageNumber="97">
<emphasis id="B942EAC0FA05FFC8CBAFB86EFEADFE27" bold="true" box="[151,325,372,397]" pageId="96" pageNumber="97">
<typeStatus id="548D8870FA05FFC8CBAFB86EFF3AFE26" box="[151,210,373,396]" pageId="96" pageNumber="97">Type</typeStatus>
material.
</emphasis>
Unknown. Born’s drawing is taken as representative of the
<typeStatus id="548D8870FA05FFC8C8DBB86EFBFFFE26" box="[995,1047,373,396]" pageId="96" pageNumber="97">type</typeStatus>
. The drawing is somewhat artistic, but since there is little confusion about the identity of this species the designation of a
<typeStatus id="548D8870FA05FFC8CFA0B883FB19FE05" box="[1176,1265,408,431]" pageId="96" pageNumber="97" type="neotype">neotype</typeStatus>
at this time is unnecessary.
</paragraph>
<paragraph id="8B8936D2FA05FFC8CBFFB8FBFF33FD22" blockId="96.[151,1437,372,2017]" pageId="96" pageNumber="97">
<emphasis id="B942EAC0FA05FFC8CBFFB8FBFE81FE52" bold="true" box="[199,361,479,504]" pageId="96" pageNumber="97">
<typeStatus id="548D8870FA05FFC8CBFFB8FBFEEAFE5D" box="[199,258,480,503]" pageId="96" pageNumber="97">Type</typeStatus>
locality.
</emphasis>
Unknown. Stated as Cumaná,
<collectingCountry id="F3217642FA05FFC8C9EAB8C4FCA0FE52" box="[722,840,479,504]" name="Venezuela" pageId="96" pageNumber="97">Venezuela</collectingCountry>
, by Weisbord (1962: 437) erroneously citing Fargo (1953: 368; pl. 16, figs. 1, 1a) who did not give the
<typeStatus id="548D8870FA05FFC8C9DABB1EFCFAFDB6" box="[738,786,517,540]" pageId="96" pageNumber="97">type</typeStatus>
locality. The
<typeStatus id="548D8870FA05FFC8C896BB1EFC36FDB6" box="[942,990,517,540]" pageId="96" pageNumber="97">type</typeStatus>
locality was unknown to Born (“
<emphasis id="B942EAC0FA05FFC8CE6FBB1EFF37FD95" italics="true" pageId="96" pageNumber="97">patria ignota</emphasis>
”). The
<typeStatus id="548D8870FA05FFC8CA0FBB33FE8FFD95" box="[311,359,552,575]" pageId="96" pageNumber="97">type</typeStatus>
locality of
<taxonomicName id="4C364D51FA05FFC8CAD4BB3CFC45FDEA" authority="Weisbord, 1962" authorityName="Weisbord" authorityYear="1962" box="[492,941,551,576]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="mareana">
<emphasis id="B942EAC0FA05FFC8CAD4BB3CFD1CFD95" box="[492,756,551,576]" italics="true" pageId="96" pageNumber="97">Clathrodrillia mareana</emphasis>
Weisbord, 1962
</taxonomicName>
is lower Mare Formation on hillside above west bank of Quebrada Mare Abajo, northern
<collectingCountry id="F3217642FA05FFC8C984BB57FCC7FDCF" box="[700,815,588,613]" name="Venezuela" pageId="96" pageNumber="97">Venezuela</collectingCountry>
(Weisbord, 1962: 438) [=Pliocene (Woodring, 1970: 383)].
</paragraph>
<paragraph id="8B8936D2FA05FFC8CBFFBB8FFB9DFAF2" blockId="96.[151,1437,372,2017]" pageId="96" pageNumber="97">
<emphasis id="B942EAC0FA05FFC8CBFFBB8FFE5AFD07" bold="true" box="[199,434,660,685]" pageId="96" pageNumber="97">Material examined.</emphasis>
A total of
<specimenCount id="9D30FD5BFA05FFC8C917BB8FFD3EFD07" box="[559,726,660,685]" pageId="96" pageNumber="97" type="generic">100 specimens</specimenCount>
were examined.
<emphasis id="B942EAC0FA05FFC8C8A0BB8FFBE4FD07" bold="true" box="[920,1036,660,685]" italics="true" pageId="96" pageNumber="97">
<collectingCountry id="F3217642FA05FFC8C8A0BB8FFBE4FD07" box="[920,1036,660,685]" name="Venezuela" pageId="96" pageNumber="97">Venezuela</collectingCountry>
</emphasis>
: 1 spec., 48.9 x
<quantity id="4CCE9B37FA05FFC8CFF2BB8FFAD9FD06" box="[1226,1329,660,684]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="96" pageNumber="97" unit="mm" value="17.3">17.3 mm</quantity>
, in
<quantity id="4CCE9B37FA05FFC8CE63BB8FFA7FFD06" box="[1371,1431,660,684]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="96" pageNumber="97" unit="m" value="15.0">15 m</quantity>
, Coche I., Nueva Esparta, G. Mackintosh!
<date id="FF881012FA05FFC8C954BBA3FD3DFD7A" box="[620,725,696,720]" pageId="96" pageNumber="97" value="1999-09">Sep 1999</date>
(author’s coll.), 70 spec., 48.1 x 18.3, 48.5 x 18.0, 47.3 x 16.7, 49.4 x 17.6, 48.4 x 18.0, 46.9 x 17.6, 48.6 x 17.2, 47.3 x 18.3, 47.1 x 17.0, 49.2 x 19.0, 47.8 x 17.6, 53.8 x 19.0, 49.0 x 17.3, 48.0 x 17.6, 50.9 x 18.2, 42.0 x 15.4, 42.5 x 16.0, 41.8 x 15.4, 44.1 x 16.3, 47.1 x 16.7, 42.6 x 16.2, 46.8 x 17.5, 41.7 x 16.1, 44.6 x 16.2, 42.7 x 15.6, 41.2 x 15.4, 47.1 x 16.6, 43.1 x 15.8, 44.6 x 16.1, 46.6 x 17.2, 42.8 x 15.5, 41.0 x 15.6, 47.3 x 18.0, 45.2 x 17.2, 44.4 x 15.7, 40.6 x 15.2, 47.8 x 17.8, 48.4 x 16.8, 49.4 x 17.9, 52.1 x 19.5, 46.6 x 16.7, 46.5 x 16.9, 47.5 x 17.3, 52.5 x 19.2, 47.5 x 16.7, 49.5 x 18.6, 51.3 x 18.2, 49.0 x 17.1, 48.9 x 18.1, 46.2 x 16.6, 49.1 x 17.7, 47.9 x 16.5, 50.4 x 18.4, 53.6 x 19.7, 44.7 x 17.8, 44.5 x 16.7, 44.1 x 16.0, 44.4 x 17.3, 44.0 x 16.5, 44.5 x 17.4, 44.9 x 17.9, 46.1 x 15.9, 45.9 x 16.5, 46.3 x 16.8, 43.7 x 16.1, 44.1 x 16.6, 44.3 x 16.4, 46.9 x 17.2, 44.1 x 16.7 &amp; 44.4 x
<quantity id="4CCE9B37FA05FFC8C9EABAC3FCD5FC45" box="[722,829,984,1008]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="96" pageNumber="97" unit="mm" value="16.5">16.5 mm</quantity>
, in
<quantity id="4CCE9B37FA05FFC8C856BAC3FC21FC45" box="[878,969,984,1008]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" pageId="96" pageNumber="97" unit="m" value="11.0" valueMax="14.0" valueMin="8.0">8–14 m</quantity>
, N side of Coche I., Nueva Esparta, G. Mackintosh!
<date id="FF881012FA05FFC8CA13BAE7FE09FBBE" box="[299,481,1020,1044]" pageId="96" pageNumber="97" value="1999-09-21" valueMax="1999-09-25" valueMin="1999-09-21">21–25 Sep 1999</date>
(author’s coll.); 18 spec., 25.9 x 9.9, 24.7 x 10.1, 32.7 x 12.7, 28.5 x 11.3, 23.2 x 9.2, 24.8 x 9.8, 26.0 x 9.8, 26.9 x 10.3, 26.2 x 10.3, 27.9 x 10.8, 28.6 x 10.7, 28.4 x 11.0, 27.7 x 10.6, 32.5 x 12.4, 32.1 x 12.3, 32.6 x 11.6, 45.3 x 16.3, 50.0 x
<quantity id="4CCE9B37FA05FFC8C96CBD5FFD52FBF6" box="[596,698,1092,1116]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="96" pageNumber="97" unit="mm" value="18.5">18.5 mm</quantity>
, in 5 &amp;
<quantity id="4CCE9B37FA05FFC8C82EBD5FFCB9FBF6" box="[790,849,1092,1116]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="96" pageNumber="97" unit="m" value="12.0">12 m</quantity>
, W end of Coche I., Nueva Esparta, G. Mackintosh!
<date id="FF881012FA05FFC8CBAFBD73FE69FB2A" box="[151,385,1128,1152]" pageId="96" pageNumber="97" value="1999-10-31" valueMax="1999-11-02" valueMin="1999-10-31">31 Oct–2 Nov 1999</date>
(author’s coll.); 1 spec. 44.2 x
<quantity id="4CCE9B37FA05FFC8C9C4BD73FCB3FB2A" box="[764,859,1128,1152]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="96" pageNumber="97" unit="mm" value="16.7">16.7mm</quantity>
, in
<quantity id="4CCE9B37FA05FFC8C8B3BD73FC22FBD5" box="[907,970,1128,1152]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="96" pageNumber="97" unit="m" value="12.0">12 m</quantity>
, N side of Coche I., Nueva Esparta, G. Mackintosh!
<date id="FF881012FA05FFC8CA15BD97FE53FB0E" box="[301,443,1164,1188]" pageId="96" pageNumber="97" value="1999-09-25">25 Sep 1999</date>
(UF 470284); 1 spec., 49.6 x
<quantity id="4CCE9B37FA05FFC8C82CBD97FC93FB0E" box="[788,891,1164,1188]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="96" pageNumber="97" unit="mm" value="18.7">18.7 mm</quantity>
, in
<quantity id="4CCE9B37FA05FFC8C89EBD97FC0AFB0E" box="[934,994,1164,1188]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="96" pageNumber="97" unit="m" value="14.0">14 m</quantity>
, N side of Coche I., Nueva Esparta, G. Mackintosh! 4–
<date id="FF881012FA05FFC8CA72BDABFE69FB62" box="[330,385,1199,1224]" pageId="96" pageNumber="97">5 Jul</date>
, 1993 (UF 470282); 1 spec., 50.5 x
<quantity id="4CCE9B37FA05FFC8C822BDABFC69FB6D" box="[794,897,1200,1224]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="96" pageNumber="97" unit="mm" value="19.1">19.1 mm</quantity>
, in
<quantity id="4CCE9B37FA05FFC8C892BDABFC30FB6D" box="[938,984,1200,1224]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="96" pageNumber="97" unit="m" value="6.0">6 m</quantity>
, Pampatar, Margarita I., Nueva Esparta, G. Mackintosh! (UF 470281); 1 spec., 36.9 x
<quantity id="4CCE9B37FA05FFC8C982BDCFFCCDFB46" box="[698,805,1236,1260]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="96" pageNumber="97" unit="mm" value="14.2">14.2 mm</quantity>
, in
<quantity id="4CCE9B37FA05FFC8C86EBDCFFC7EFB46" box="[854,918,1236,1260]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="96" pageNumber="97" unit="m" value="37.0">37 m</quantity>
, off Margarita I., Nueva Esparta, 1996 (UF 470283); 5 spec., 29.3 x 10.6, 25.0 x 9.3, 29.7 x 10.8, 30.1 x 11.4 &amp; 32.9 x
<quantity id="4CCE9B37FA05FFC8CF3BBDE3FB83FAA5" box="[1027,1131,1272,1296]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="96" pageNumber="97" unit="mm" value="12.5">12.5 mm</quantity>
, in
<quantity id="4CCE9B37FA05FFC8CFAFBDE3FB2EFAA5" box="[1175,1222,1272,1295]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="96" pageNumber="97" unit="m" value="7.0">7 m</quantity>
, Bahia Guamache, Margarita I., Nueva Esparta, G. Mackintosh!
<date id="FF881012FA05FFC8C9B6BC07FCF9FA9E" box="[654,785,1308,1333]" pageId="96" pageNumber="97" value="2001-07-29">29 Jul 2001</date>
(author’s coll.); 2 spec., 29.1 x 10.4 &amp; 30.6 x
<quantity id="4CCE9B37FA05FFC8CE2DBC07FA93FA9E" box="[1301,1403,1308,1332]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="96" pageNumber="97" unit="mm" value="10.6">10.6 mm</quantity>
, in
<quantity id="4CCE9B37FA05FFC8CBAFBC5BFF55FAF2" box="[151,189,1344,1368]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="96" pageNumber="97" unit="m" value="6.0">6m</quantity>
, Pampatar, Margarita I., Nueva Esparta, G. Mackintosh!
<date id="FF881012FA05FFC8C800BC5BFC29FAF2" box="[824,961,1344,1368]" pageId="96" pageNumber="97" value="1993-06-24">24 Jun 1993</date>
(author’s coll.).
</paragraph>
<paragraph id="8B8936D2FA05FFC8CBFFBC7EFB0FFA4D" blockId="96.[151,1437,372,2017]" pageId="96" pageNumber="97">
<emphasis id="B942EAC0FA05FFC8CBFFBC7EFE5AFAD7" bold="true" box="[199,434,1380,1405]" pageId="96" pageNumber="97">Range and habitat.</emphasis>
<collectingCountry id="F3217642FA05FFC8CA84BC7FFDC2FAD7" box="[444,554,1380,1405]" name="Colombia" pageId="96" pageNumber="97">Colombia</collectingCountry>
(Magdalena Dept.; Guajira Dept.);
<collectingCountry id="F3217642FA05FFC8C8F6BC7FFBA9FAD7" box="[974,1089,1380,1405]" name="Venezuela" pageId="96" pageNumber="97">Venezuela</collectingCountry>
(Zulia; Falcon; Sucre; Nueva Esparta: Margarita I.; Coche I.; Cubagua I.);
<collectingCountry id="F3217642FA05FFC8C993BC93FCEDFA0A" box="[683,773,1416,1440]" name="Guyana" pageId="96" pageNumber="97">Guyana</collectingCountry>
;
<collectingCountry id="F3217642FA05FFC8C82CBC9CFC6BFA0A" box="[788,899,1415,1440]" name="Suriname" pageId="96" pageNumber="97">Suriname</collectingCountry>
. It has been taken live by divers in relatively shallow water especially around Margarita I. (and the nearby uninhabited islets of Coche and Cubagua). Divercollected material has been reported from
<quantity id="4CCE9B37FA05FFC8C957BCCBFD2DFA4D" box="[623,709,1488,1512]" metricMagnitude="1" metricUnit="m" metricValue="1.3" metricValueMax="2.0" metricValueMin="0.6" pageId="96" pageNumber="97" unit="m" value="13.0" valueMax="20.0" valueMin="6.0">6–20 m</quantity>
depths, and from towed gear in as deep as
<specimenCount id="9D30FD5BFA05FFC8CF9FBCCBFB0FFA4D" box="[1191,1255,1488,1512]" pageId="96" pageNumber="97" type="generic">
<quantity id="4CCE9B37FA05FFC8CF9FBCCBFB0BFA4D" box="[1191,1251,1488,1512]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="96" pageNumber="97" unit="m" value="60.0">60 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="8B8936D2FA05FFC8CBFFBCEFFB89F937" blockId="96.[151,1437,372,2017]" pageId="96" pageNumber="97">
This taxon has been applied to fossil specimens in some of the works listed in the synonymy above. Fossils have been reported from
<collectingCountry id="F3217642FA05FFC8CA81BF0CFDC4F99A" box="[441,556,1559,1584]" name="Venezuela" pageId="96" pageNumber="97">Venezuela</collectingCountry>
(Lower Mare Formation, Weisbord, 1962) and from as far north as Sarasota Co., W Florida (Caloosahatchee Formation [YPM IP 559859]), which indicates a much wider distribution of this species during Pliocene-Pleistocene Epochs (5.3 million–11,700 years BP) than at present. Fossil specimens were not examined in this work to determine their degree of similarity to Recent specimens.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA05FFCACBFFBFBCFBD3FCBF" lastPageId="98" lastPageNumber="99" pageId="96" pageNumber="97" type="description">
<paragraph id="8B8936D2FA05FFCACBFFBFBCFBD3FCBF" blockId="96.[151,1437,372,2017]" lastBlockId="98.[151,1437,151,1256]" lastPageId="98" lastPageNumber="99" pageId="96" pageNumber="97">
<emphasis id="B942EAC0FA05FFC8CBFFBFBCFE73F96A" bold="true" box="[199,411,1703,1728]" pageId="96" pageNumber="97">
Description.
<emphasis id="B942EAC0FA05FFC8CA58BFBCFE73F96A" bold="true" box="[352,411,1703,1728]" italics="true" pageId="96" pageNumber="97">Shell</emphasis>
</emphasis>
large (to
<quantity id="4CCE9B37FA05FFC8C930BFB3FD85F915" box="[520,621,1704,1728]" metricMagnitude="-2" metricUnit="m" metricValue="5.38" pageId="96" pageNumber="97" unit="mm" value="53.8">53.8 mm</quantity>
in total length), fusiform, truncated anteriorly, with a tall, turreted spire, whorls straight to slightly convex and total about 14½; last whorl approximately 52% of the total length, constricted anteriorly by a furrow created by the stromboid notch. Sutures impressed, with a collar adjoining below. Sculpture of numerous ribs cut by spiral grooves. Aperture oval with a short but distinct anterior canal.
<emphasis id="B942EAC0FA05FFC8CE2EBE0FFA73F887" bold="true" box="[1302,1435,1812,1837]" italics="true" pageId="96" pageNumber="97">Protoconch</emphasis>
of approximately 1¾–2 smooth round whorls. Beginning of teleoconch usually obscure but marked by the first axial.
<emphasis id="B942EAC0FA05FFC8CBE4BE47FE7AF8DF" bold="true" box="[220,402,1884,1909]" italics="true" pageId="96" pageNumber="97">Axial Sculpture</emphasis>
of numerous thin axial ribs that extend from suture-to-suture, very slightly opisthocline on the spire whorls, more so on the last; narrower than their interspaces, 21–23 on the penultimate, 13–17 on the last whorl to the varix, the last 4–5 ribs larger, more widely spaced and acutely hooked at the shoulder. Ribs reduced to narrow crescents in sulcus of early whorls, finer crescents in later whorls that override collar to suture but obscured by coarse growth striae.
<emphasis id="B942EAC0FA07FFCACA90B98CFE0BFF1A" bold="true" box="[424,483,151,176]" italics="true" pageId="98" pageNumber="99">Varix</emphasis>
large, broad, hump-like, approximately ¼- to ⅓-whorl from the edge of the outer lip.
<emphasis id="B942EAC0FA07FFCACBAFB9A7FEB0FF7F" bold="true" box="[151,344,188,213]" italics="true" pageId="98" pageNumber="99">Spiral Sculpture</emphasis>
of incised grooves that cut the axial ribs creating a somewhat cancellate appearance, 6–8 on penultimate and 9–13 more on shell base to ridge demarcating anterior fasciole.
<emphasis id="B942EAC0FA07FFCACF73B9C4FB71FF52" bold="true" box="[1099,1177,223,248]" italics="true" pageId="98" pageNumber="99">Sulcus</emphasis>
with convex collar at suture, then a furrow at shoulder, which is the trace left but the apex of the anal sinus. Curved traces of ribs and spiral grooves appear in the sulcus of early whorls, but are entirely obscured by the heavy collar and growth striae on later whorls.
<emphasis id="B942EAC0FA07FFCACA77B857FE5EFECF" bold="true" box="[335,438,332,357]" italics="true" pageId="98" pageNumber="99">Outer lip</emphasis>
flattened from varix to edge of lip, jutting out beyond the margin of the previous whorl, and bearing several low axial folds, and a large reinforcing rib near edge; edge of lip forms a low arc along its length, heavily scalloped by tooth extensions of the spiral grooves. Stromboid notch deep, creating a spiral furrow that marks the shell base constriction.
<emphasis id="B942EAC0FA07FFCAC978B8ACFD53FE7A" bold="true" box="[576,699,439,464]" italics="true" pageId="98" pageNumber="99">Anal sinus</emphasis>
deeply notched, sides asymmetrically U-shaped, the outer lip side projects beyond the parietal wall side; sinus directed away from the main axis of the shell by heavy parietal lobe, flared inner edge of sinus, and outward flex of outer lip gives sinus a spout-like appearance.
<emphasis id="B942EAC0FA07FFCACF97BB1BFAFEFDB2" bold="true" box="[1199,1302,511,536]" italics="true" pageId="98" pageNumber="99">Inner lip</emphasis>
moderately wide, heavily enameled, emarginated, erect anteriorly along columella, recumbent and widest on parietal wall, developed into a heavy parietal lobe posteriorly.
<emphasis id="B942EAC0FA07FFCAC9E8BB5CFC94FDCA" bold="true" box="[720,892,583,608]" italics="true" pageId="98" pageNumber="99">Anterior canal</emphasis>
short but distinctly formed, open, with a deep apical notch, U-shaped, with an upturned end. Fasciole short, emerging from underneath current outer lip about halfway up the columella, bounded from shell base ridge along its margin; surface with dense incremental crescentshaped growth striae only.
<emphasis id="B942EAC0FA07FFCACAF6BBAFFDF8FD67" bold="true" box="[462,528,692,717]" italics="true" pageId="98" pageNumber="99">Color</emphasis>
dirty white with a reddish-brown band, confined mostly to rib crests, white to grayish between ribs, extending from shoulder to mid-whorl (to near suture on spire whorls) and with an additional band on shell base; varix with a pale to dark brown blotch; aperture reddish brown.
</paragraph>
</subSubSection>
<caption id="DF49665AFA04FFC9CBAFBFB6FD4DF85F" httpUri="https://zenodo.org/record/263342/files/figure.png" pageId="97" pageNumber="98" targetBox="[288,1302,193,1684]" targetPageId="97">
<paragraph id="8B8936D2FA04FFC9CBAFBFB6FD4DF85F" blockId="97.[151,1436,1709,2037]" pageId="97" pageNumber="98">
<emphasis id="B942EAC0FA04FFC9CBAFBFB6FEF9F969" bold="true" box="[151,273,1709,1731]" pageId="97" pageNumber="98">PLATE 42.</emphasis>
<taxonomicName id="4C364D51FA04FFC9CA20BFB6FD63F969" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[280,651,1709,1731]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA04FFC9CA20BFB6FE14F969" box="[280,508,1709,1731]" italics="true" pageId="97" pageNumber="98">Clathrodrillia gibbosa</emphasis>
(Born, 1778)
</taxonomicName>
.
<emphasis id="B942EAC0FA04FFC9C9ADBFB5FCA5F969" bold="true" box="[661,845,1709,1731]" pageId="97" pageNumber="98">Upper left panel.</emphasis>
Fig. 1: a fully developed representative specimen, 48.9 x 17.3 mm, from Coche I., Nueva Esparta, Venezuela (author’s coll.). Fig. 2: enlarged view (not to scale) of protoconch of a 29.3 x 10.6 mm specimen from Margarita, I., Nueva Esparta, Venezuela (author’s coll.).
<emphasis id="B942EAC0FA04FFC9CF23BFF0FB0EF8AA" bold="true" box="[1051,1254,1770,1792]" pageId="97" pageNumber="98">Upper right panel</emphasis>
. Fig. 3: drawing reproduced from Born (1780: pl. 11, figs. 12, 13) of
<taxonomicName id="4C364D51FA04FFC9C992BE11FCA4F8B5" box="[682,844,1801,1823]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="gibbosus">
<emphasis id="B942EAC0FA04FFC9C992BE11FCA4F8B5" box="[682,844,1801,1823]" italics="true" pageId="97" pageNumber="98">Murex gibbosus</emphasis>
</taxonomicName>
made available on the worldwide web by the AnimalBase Project Group (2005–2015). Fig. 4: enlarged view of outer lip showing marginal teeth, marginal axial fold, and stromboid notch of a 49.4 x 17.6 mm specimen from N side of Coche I., Nueva Esparta, Venezuela (author’s coll.). Fig. 5: enlarged view of area of anterior fasciole showing ridges left by advance of stromboid notch and edge of anterior canal notch on the end of the anterior canal during shell growth of a 42.0 x 15.4 mm specimen from N side of Coche I., Nueva Esparta, Venezuela, (author’s coll.). Fig. 6: same specimen as in Fig. 1, apical view, V=varix, L=edge of outer lip.
<emphasis id="B942EAC0FA04FFC9CF09BEB8FB55F812" bold="true" box="[1073,1213,1955,1976]" pageId="97" pageNumber="98">Lower panel</emphasis>
. Specimens showing variability in size and color. Figs. 7–12: W end of Coche I., Nueva Esparta, Venezuela (author’s coll.); Figs. 13–18: N side of Coche I., Nueva Esparta, Venezuela (author’s coll.).
</paragraph>
</caption>
<subSubSection id="C32C6559FA07FFCACBFFBA3BFB1EFB4D" pageId="98" pageNumber="99" type="discussion">
<paragraph id="8B8936D2FA07FFCACBFFBA3BFB1EFB4D" blockId="98.[151,1437,151,1256]" pageId="98" pageNumber="99">
<emphasis id="B942EAC0FA07FFCACBFFBA3BFE2AFC9D" bold="true" box="[199,450,800,823]" pageId="98" pageNumber="99">
Remarks.
<emphasis id="B942EAC0FA07FFCACA7CBA3BFE2AFC9D" bold="true" box="[324,450,800,823]" italics="true" pageId="98" pageNumber="99">Taxonomy.</emphasis>
</emphasis>
Born’s Latin description of the species was very brief.
<taxonomicName id="4C364D51FA07FFCACF7ABA04FAA8FC92" box="[1090,1344,799,824]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="98" pageNumber="99" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA07FFCACF7ABA04FAA8FC92" box="[1090,1344,799,824]" italics="true" pageId="98" pageNumber="99">Clathrodrillia gibbosa</emphasis>
</taxonomicName>
was redescribed and placed in
<taxonomicName id="4C364D51FA07FFCACA97BA5FFDDAFCF7" box="[431,562,836,861]" class="Gastropoda" family="Drilliidae" genus="Pleurotoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="98" pageNumber="99" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA07FFCACA97BA5FFDDAFCF7" box="[431,562,836,861]" italics="true" pageId="98" pageNumber="99">Pleurotoma</emphasis>
</taxonomicName>
by Kiener in his 11 volume work (Kiener, 1840). Dall (1918) subsequently selected this species as the
<typeStatus id="548D8870FA07FFCACAFEBA73FE1EFCD5" box="[454,502,872,895]" pageId="98" pageNumber="99">type</typeStatus>
of
<taxonomicName id="4C364D51FA07FFCAC923BA7CFD5FFC2A" box="[539,695,871,896]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="98" pageNumber="99" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA07FFCAC923BA7CFD5FFC2A" box="[539,695,871,896]" italics="true" pageId="98" pageNumber="99">Clathrodrillia</emphasis>
</taxonomicName>
, but did not describe it further. A more complete description of
<taxonomicName id="4C364D51FA07FFCACEBABA73FF18FC0F" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="98" pageNumber="99" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA07FFCACEBABA73FF18FC0F" italics="true" pageId="98" pageNumber="99">C. gibbosa</emphasis>
</taxonomicName>
was published by Fargo (1953: 368; pl. 16, figs. 1, 1a).
<emphasis id="B942EAC0FA07FFCAC867BA97FC3DFC0F" bold="true" box="[863,981,908,933]" italics="true" pageId="98" pageNumber="99">Variability</emphasis>
. Mature specimens can be quite variable in size, probably due to environmental factors. The smallest completely developed specimen, with varix and projecting scalloped lip, encountered in this work is
<quantity id="4CCE9B37FA07FFCAC9D0BACFFCA4FC46" box="[744,844,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="98" pageNumber="99" unit="mm" value="23.2">23.2 mm</quantity>
in total length (see Pl. 42, Fig. 7). The average total length of
<specimenCount id="9D30FD5BFA07FFCACA3BBAE3FE41FBBA" box="[259,425,1015,1040]" pageId="98" pageNumber="99" type="generic">100 specimens</specimenCount>
is
<quantity id="4CCE9B37FA07FFCACAF2BAE3FDD5FBA5" box="[458,573,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="4.232" pageId="98" pageNumber="99" unit="mm" value="42.32">42.32 mm</quantity>
(
<quantity id="4CCE9B37FA07FFCAC974BAECFD1BFBBA" box="[588,755,1015,1040]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="5.38" metricValueMin="2.32" pageId="98" pageNumber="99" unit="mm" value="38.5" valueMax="53.8" valueMin="23.2">23.2–53.8 mm</quantity>
), their average W/L is 0.371. The greatest length reported in the literature is
<quantity id="4CCE9B37FA07FFCACA7CBD06FE7BFB9E" box="[324,403,1053,1076]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="98" pageNumber="99" unit="mm" value="75.0">75 mm</quantity>
(Kaicher, 1984, card 3944); however, this may be a typographical error. The largest reported elsewhere is
<quantity id="4CCE9B37FA07FFCACA10BD5BFE90FBFD" box="[296,376,1088,1112]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="98" pageNumber="99" unit="mm" value="60.0">60 mm</quantity>
(M. Smith, 1940: 80), which is still quite large compared to the
<quantity id="4CCE9B37FA07FFCACF76BD5BFB5AFBFD" box="[1102,1202,1088,1112]" metricMagnitude="-2" metricUnit="m" metricValue="5.38" pageId="98" pageNumber="99" unit="mm" value="53.8">53.8 mm</quantity>
maximum specimen size examined for this work.
<emphasis id="B942EAC0FA07FFCACAE6BD7FFD6BFBD7" bold="true" box="[478,643,1124,1149]" italics="true" pageId="98" pageNumber="99">Identification.</emphasis>
This is a distinctive species; with the possible exception of Simpson’s 1887c report, miss-identifications have not been found in the published literature. The closest congener,
<taxonomicName id="4C364D51FA07FFCACE11BD93FEACFB6F" authority="Tippett, 1995" authorityName="Tippett" authorityYear="1995" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="98" pageNumber="99" phylum="Mollusca" rank="species" species="petuchi">
<emphasis id="B942EAC0FA07FFCACE11BD93FA75FB0A" box="[1321,1437,1159,1184]" italics="true" pageId="98" pageNumber="99">C. petuchi</emphasis>
(Tippett, 1995)
</taxonomicName>
, differs in being stockier, having wider and fewer ribs, and a different coloration, pale yellow to orange bands rather than brown. The varix also has a much lighter colored patch than in
<taxonomicName id="4C364D51FA07FFCACF4EBDCBFB18FB42" box="[1142,1264,1231,1256]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="98" pageNumber="99" phylum="Mollusca" rank="species" species="gibbosa">
<emphasis id="B942EAC0FA07FFCACF4EBDCBFB18FB42" box="[1142,1264,1231,1256]" italics="true" pageId="98" pageNumber="99">C. gibbosa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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