treatments-xml/data/03/9F/87/039F87C4FA13FFD0CBAFBDD4FC9CF985.xml

<document id="BBCA8670379CA9F489619E1354D3BD84" ID-DOI="10.11646/zootaxa.4090.1.1" ID-GBIF-Dataset="e6b35f5a-435e-4473-b29e-1e4d842f84b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="263299" ID-ZooBank="203BAC25-B542-48FE-B5AD-EBA8C0285833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457964633549" checkinUser="plazi" docAuthor="Fallon, Phillip J." docDate="2016" docId="039F87C4FA13FFD0CBAFBDD4FC9CF985" docLanguage="en" docName="15612-5136-1-PB.pdf.imf" docOrigin="Zootaxa 4090 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Decoradrillia harlequina Fallon, 2016, new species" docType="treatment" docVersion="12" lastPageNumber="121" masterDocId="FFA6FFBCFA65FFA8CB38B91BFFE8FFAA" masterDocTitle="Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species" masterLastPageNumber="363" masterPageNumber="1" pageNumber="119" updateTime="1698184378282" updateUser="plazi">
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<mods:title id="73F2A9ABA286C30859B7FDBB5E730DC6">Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species</mods:title>
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<mods:namePart id="04F24B8D52FB47B4F982EE1079B51943">Fallon, Phillip J.</mods:namePart>
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<mods:date id="F29EA64738429EBE16ABE26713C888CE">2016</mods:date>
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<treatment id="039F87C4FA13FFD0CBAFBDD4FC9CF985" ID-DOI="http://doi.org/10.5281/zenodo.6076376" ID-GBIF-Taxon="117911748" ID-Zenodo-Dep="6076376" LSID="urn:lsid:plazi:treatment:039F87C4FA13FFD0CBAFBDD4FC9CF985" httpUri="http://treatment.plazi.org/id/039F87C4FA13FFD0CBAFBDD4FC9CF985" lastPageId="120" lastPageNumber="121" pageId="118" pageNumber="119">
<subSubSection id="C32C6559FA13FFDECBAFBDD4FD05FA24" pageId="118" pageNumber="119" type="nomenclature">
<paragraph id="8B8936D2FA13FFDECBAFBDD4FD8BFB40" blockId="118.[151,611,1231,1291]" box="[151,611,1231,1258]" pageId="118" pageNumber="119">
<heading id="D0C181BEFA13FFDECBAFBDD4FD8BFB40" bold="true" box="[151,611,1231,1258]" fontSize="11" level="1" pageId="118" pageNumber="119" reason="1">
<emphasis id="B942EAC0FA13FFDECBAFBDD4FD8BFB40" bold="true" box="[151,611,1231,1258]" pageId="118" pageNumber="119">
<taxonomicName id="4C364D51FA13FFDECBAFBDD4FE2DFB43" ID-CoL="6CFQG" box="[151,453,1231,1257]" class="Gastropoda" family="Drilliidae" genus="Decoradrillia" kingdom="Animalia" order="Neogastropoda" pageId="118" pageNumber="119" phylum="Mollusca" rank="species" species="harlequina" status="sp. nov.">
<emphasis id="B942EAC0FA13FFDECBAFBDD4FE2DFB43" bold="true" box="[151,453,1231,1257]" italics="true" pageId="118" pageNumber="119">Decoradrillia harlequina</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A27157BBFA13FFDECAECBDCBFD8BFB40" box="[468,611,1232,1258]" pageId="118" pageNumber="119" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8936D2FA13FFDECBAFBDE9FE2DFAA1" blockId="118.[151,611,1231,1291]" box="[151,453,1266,1291]" pageId="118" pageNumber="119">
(
<figureCitation id="130D2A57FA13FFDECBA7BDE9FEE0FAA1" box="[159,264,1266,1291]" captionStart="PLATE 51" captionStartId="114.[151,234,1892,1914]" captionTargetBox="[213,1370,179,1874]" captionTargetId="figure@114.[213,1370,173,1874]" captionTargetPageId="114" captionText="PLATE 51. Decoradrillia spp. shell surface SEM images. Figs. 1 – 2: Decoradrillia pulchella (Reeve, 1845), Anse Bateau Bay, Tobago I., Trinidad &amp; Tobago (ANSP 338495), 240 x &amp; 930 x, respectively. Figs. 3 – 4: Decoradrillia harlequina, new species: Fig. 3: Indian Cay, Grand Bahama I. (ANSP 366927), 240 x; Fig. 4: holotype, Devil’s Bay, Grenada (USNM 1291335), 1,000 x. Figs. 5 – 6: Decoradrillia interstincta, new species, &quot; Big Rocks &quot;, Utila I., Bay Is., Honduras (ANSP 464986), 250 x &amp; 1,000 x, respectively." httpUri="https://zenodo.org/record/263351/files/figure.png" pageId="118" pageNumber="119">Plates 51</figureCitation>
, Figs. 3, 4 &amp; 54)
</paragraph>
<paragraph id="8B8936D2FA13FFDECBAFBC21FD4DFAFA" blockId="118.[151,749,1338,1422]" box="[151,677,1338,1360]" pageId="118" pageNumber="119">
<taxonomicName id="4C364D51FA13FFDECBAFBC21FEE5FAFA" box="[151,269,1338,1360]" class="Gastropoda" family="Pseudomelatomidae" genus="Crassispira" kingdom="Animalia" order="Neogastropoda" pageId="118" pageNumber="119" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA13FFDECBAFBC21FEE5FAFA" box="[151,269,1338,1360]" italics="true" pageId="118" pageNumber="119">Crassispira</emphasis>
</taxonomicName>
sp.: Humfrey (1975: 182, pl. 22. fig. 7).
</paragraph>
<paragraph id="8B8936D2FA13FFDECBAFBC42FD44FAC5" blockId="118.[151,749,1338,1422]" box="[151,684,1369,1391]" pageId="118" pageNumber="119">“? Unknown”: Williams (2005; 2009: species 9006).</paragraph>
<paragraph id="8B8936D2FA13FFDECBAFBC62FD05FA24" blockId="118.[151,749,1338,1422]" box="[151,749,1400,1422]" pageId="118" pageNumber="119">
<taxonomicName id="4C364D51FA13FFDECBAFBC62FEEDFA24" box="[151,261,1401,1422]" class="Gastropoda" family="Drilliidae" genus="Fenimorea" kingdom="Animalia" order="Neogastropoda" pageId="118" pageNumber="119" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA13FFDECBAFBC62FEEDFA24" box="[151,261,1401,1422]" italics="true" pageId="118" pageNumber="119">Fenimorea</emphasis>
</taxonomicName>
sp. 1: Daccarett &amp; Bossio (2011: 134, fig. 832).
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA13FFDECBAFBCA0FD3DF9C9" pageId="118" pageNumber="119" type="description">
<paragraph id="8B8936D2FA13FFDECBAFBCA0FD3DF9C9" blockId="118.[151,1436,1466,1962]" pageId="118" pageNumber="119">
<emphasis id="B942EAC0FA13FFDECBAFBCA0FF0BFA78" bold="true" box="[151,227,1467,1490]" pageId="118" pageNumber="119">
<typeStatus id="548D8870FA13FFDECBAFBCA0FF37FA78" box="[151,223,1467,1490]" pageId="118" pageNumber="119">Types</typeStatus>
.
</emphasis>
<typeStatus id="548D8870FA13FFDECBD3BCA1FEBBFA79" box="[235,339,1466,1491]" pageId="118" pageNumber="119" type="holotype">Holotype</typeStatus>
16.1 x 
<quantity id="4CCE9B37FA13FFDECA92BCA0FDE9FA78" box="[426,513,1467,1491]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="118" pageNumber="119" unit="mm" value="6.2">6.2 mm</quantity>
(USNM 1291335); 9 
<typeStatus id="548D8870FA13FFDEC9C3BCA0FC83FA78" box="[763,875,1467,1490]" pageId="118" pageNumber="119" type="paratype">paratypes</typeStatus>
: 4 spec, 17.2 x 
<quantity id="4CCE9B37FA13FFDECF23BCA0FB9AFA78" box="[1051,1138,1467,1491]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="118" pageNumber="119" unit="mm" value="6.2">6.2 mm</quantity>
(USNM 1291336), 16.9 x 6.3, 14.4 x 6.0 mm (ANSP 464967), and 15.2 x 
<quantity id="4CCE9B37FA13FFDEC98EBCC4FCE5FA5C" box="[694,781,1503,1527]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="118" pageNumber="119" unit="mm" value="5.3">5.3 mm</quantity>
(MZSP 
<date id="FF881012FA13FFDEC856BCC5FC2DFA5D" box="[878,965,1502,1527]" pageId="118" pageNumber="119" value="63-12-20">122063</date>
), from the 
<typeStatus id="548D8870FA13FFDECF79BCC4FB99FA5C" box="[1089,1137,1503,1526]" pageId="118" pageNumber="119">type</typeStatus>
locality, G. Mackintosh! 
<date id="FF881012FA13FFDECEB6BCC4FEE1F9B1" pageId="118" pageNumber="119" value="1996-08-04">4 Aug 1996</date>
. 2 spec., 15.0 x 6.0 &amp; 16.4 x 
<quantity id="4CCE9B37FA13FFDEC973BF18FD4AF9B0" box="[587,674,1539,1563]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="118" pageNumber="119" unit="mm" value="6.2">6.2 mm</quantity>
, in 
<quantity id="4CCE9B37FA13FFDEC9F2BF18FCECF9B0" box="[714,772,1539,1563]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="118" pageNumber="119" unit="m" value="14.0">14 m</quantity>
, Annas Shoal, St. Georges, 
<collectingCountry id="F3217642FA13FFDECF01BF19FB73F9B1" box="[1081,1179,1538,1563]" name="Grenada" pageId="118" pageNumber="119">Grenada</collectingCountry>
, G. Mackintosh! 
<date id="FF881012FA13FFDECE60BF18FF27F995" pageId="118" pageNumber="119" value="1996-08-04">4 Aug 1996</date>
(M. Williams coll.); 3 spec., 15.8 x 6.5, 14.8 x 5.4, &amp; 16.0 x 5.9 (apex gone) mm, in 
<quantity id="4CCE9B37FA13FFDECF89BF3CFB06F994" box="[1201,1262,1575,1599]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="118" pageNumber="119" unit="m" value="14.0">14 m</quantity>
, Chatham Bay, Union I., G. Mackintosh! 
<date id="FF881012FA13FFDECA8EBF50FDDDF9C9" box="[438,565,1611,1635]" pageId="118" pageNumber="119" value="2007-04-08">8 Apr 2007</date>
(UF 496635).
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA13FFD0CBFFBF74FA75FE2D" lastPageId="120" lastPageNumber="121" pageId="118" pageNumber="119" type="materials_examined">
<paragraph id="8B8936D2FA13FFDECBFFBF74FD2FF92C" blockId="118.[151,1436,1466,1962]" box="[199,711,1646,1671]" pageId="118" pageNumber="119">
<emphasis id="B942EAC0FA13FFDECBFFBF74FE8DF92D" bold="true" box="[199,357,1646,1671]" pageId="118" pageNumber="119">
<typeStatus id="548D8870FA13FFDECBFFBF74FEEAF92C" box="[199,258,1647,1670]" pageId="118" pageNumber="119">Type</typeStatus>
locality.
</emphasis>
Devil’s Bay, 
<collectingCountry id="F3217642FA13FFDECAC5BF75FDB7F92D" box="[509,607,1646,1671]" name="Grenada" pageId="118" pageNumber="119">Grenada</collectingCountry>
, in 
<specimenCount id="9D30FD5BFA13FFDEC9BFBF74FD2FF92C" box="[647,711,1647,1671]" pageId="118" pageNumber="119" type="generic">
<quantity id="4CCE9B37FA13FFDEC9BFBF74FD2BF92C" box="[647,707,1647,1671]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="118" pageNumber="119" unit="m" value="14.0">14 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="8B8936D2FA13FFDECBFFBF89FD87F800" blockId="118.[151,1436,1466,1962]" pageId="118" pageNumber="119">
<emphasis id="B942EAC0FA13FFDECBFFBF89FE14F901" bold="true" box="[199,508,1682,1707]" pageId="118" pageNumber="119">Other material examined.</emphasis>
An additional 
<specimenCount id="9D30FD5BFA13FFDEC99DBF88FCD5F901" box="[677,829,1682,1707]" pageId="118" pageNumber="119" type="generic">12 specimens</specimenCount>
have been examined: 
<emphasis id="B942EAC0FA13FFDECF00BF89FB2FF900" bold="true" box="[1080,1223,1682,1707]" italics="true" pageId="118" pageNumber="119">
<collectingCountry id="F3217642FA13FFDECF00BF89FB72F901" box="[1080,1178,1682,1707]" name="Bahamas" pageId="118" pageNumber="119">Bahama</collectingCountry>
Is.:
</emphasis>
1 spec., 14.5 x 
<quantity id="4CCE9B37FA13FFDECE41BF88FF2DF964" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="118" pageNumber="119" unit="mm" value="5.8">5.8 mm</quantity>
, in 
<quantity id="4CCE9B37FA13FFDECBC1BFACFED2F964" box="[249,314,1719,1743]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="118" pageNumber="119" unit="m" value="18.0">18 m</quantity>
, Indian Cay, Grand 
<collectingCountry id="F3217642FA13FFDEC909BFADFD65F965" box="[561,653,1718,1743]" name="Bahamas" pageId="118" pageNumber="119">Bahama</collectingCountry>
I., 
<geoCoordinate id="EE025015FA13FFDEC986BFACFCABF965" box="[702,835,1719,1743]" direction="north" orientation="latitude" pageId="118" pageNumber="119" precision="15" value="26.712502">26°42'45''N</geoCoordinate>
, 
<geoCoordinate id="EE025015FA13FFDEC86DBFACFC03F965" box="[853,1003,1719,1743]" direction="west" orientation="longitude" pageId="118" pageNumber="119" precision="15" value="-78.65417">078°39'15''W</geoCoordinate>
, J. Worsfold! 
<date id="FF881012FA13FFDECFA1BFACFADBF965" box="[1177,1331,1719,1743]" pageId="118" pageNumber="119" value="1985-04-16">16 Apr 1985</date>
? (ANSP 366927). 
<emphasis id="B942EAC0FA13FFDECA3BBFC1FE69F959" bold="true" box="[259,385,1754,1779]" italics="true" pageId="118" pageNumber="119">
<collectingCountry id="F3217642FA13FFDECA3BBFC1FE92F959" box="[259,378,1754,1779]" name="Sint Martin" pageId="118" pageNumber="119">St. Martin</collectingCountry>
:
</emphasis>
1 spec., 19.5 x 
<quantity id="4CCE9B37FA13FFDEC90DBFC0FD63F958" box="[565,651,1755,1779]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="118" pageNumber="119" unit="mm" value="6.8">6.8 mm</quantity>
(MNHN, ex J. Colomb coll.). 
<emphasis id="B942EAC0FA13FFDEC8DEBFC1FB25F959" bold="true" box="[998,1229,1754,1779]" italics="true" pageId="118" pageNumber="119">
<collectingCountry id="F3217642FA13FFDEC8DEBFC1FB2EF959" box="[998,1222,1754,1779]" name="Trinidad and Tobago" pageId="118" pageNumber="119">Trinidad &amp; Tobago</collectingCountry>
:
</emphasis>
1 spec. 13.8 x 
<quantity id="4CCE9B37FA13FFDECE41BFC0FF2DF8BC" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="118" pageNumber="119" unit="mm" value="5.3">5.3 mm</quantity>
, in 
<quantity id="4CCE9B37FA13FFDECBD6BFE4FEC2F8BC" box="[238,298,1791,1815]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="118" pageNumber="119" unit="m" value="29.0">29 m</quantity>
, entrance to 
<collectingCountry id="F3217642FA13FFDECA82BFE4FE05F8BC" box="[442,493,1791,1814]" name="Isle of Man" pageId="118" pageNumber="119">Man</collectingCountry>
of War Bay, 
<collectingCountry id="F3217642FA13FFDEC9BBBFE5FD3FF8BD" box="[643,727,1790,1815]" name="Trinidad and Tobago" pageId="118" pageNumber="119">Tobago</collectingCountry>
I., R. Masino! (author’s coll.); 2 spec., 15.0 x 5.9 &amp; 16.7 x 
<quantity id="4CCE9B37FA13FFDECE41BFE4FF2DF890" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="118" pageNumber="119" unit="mm" value="6.5">6.5 mm</quantity>
, in 
<quantity id="4CCE9B37FA13FFDECBD7BE38FEC3F890" box="[239,299,1827,1851]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="118" pageNumber="119" unit="m" value="20.0">20 m</quantity>
, Rocky Pt., Mt. Irving Bay, 
<collectingCountry id="F3217642FA13FFDEC955BE39FD29F891" box="[621,705,1826,1851]" name="Trinidad and Tobago" pageId="118" pageNumber="119">Tobago</collectingCountry>
I., G. Mackintosh! 
<date id="FF881012FA13FFDEC8A6BE38FBC4F891" box="[926,1068,1827,1851]" pageId="118" pageNumber="119" value="1998-09-20">20 Sep 1998</date>
(M. Williams coll.). 
<emphasis id="B942EAC0FA13FFDECE24BE38FA6EF890" bold="true" box="[1308,1414,1827,1850]" italics="true" pageId="118" pageNumber="119">
<collectingCountry id="F3217642FA13FFDECE24BE38FA96F890" box="[1308,1406,1827,1850]" name="Panama" pageId="118" pageNumber="119">Panama</collectingCountry>
:
</emphasis>
3 spec., 14.4 x 5.6, 11.6 x 4.3 &amp; 18.1 x 
<quantity id="4CCE9B37FA13FFDEC971BE51FD4AF8CB" box="[585,674,1866,1890]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="118" pageNumber="119" unit="mm" value="6.2">6.2 mm</quantity>
, in 
<quantity id="4CCE9B37FA13FFDEC9F6BE51FD14F8CB" box="[718,764,1866,1890]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="118" pageNumber="119" unit="m" value="4.0">4 m</quantity>
, 5th Cay, Hollandes Cays, G. Mackintosh! 
<date id="FF881012FA13FFDECFE5BE51FA82F8C8" box="[1245,1386,1866,1890]" pageId="118" pageNumber="119" value="1992-10-19">19 Oct 1992</date>
(M. Williams coll.). 
<emphasis id="B942EAC0FA13FFDECA6CBE76FE4CF82C" bold="true" box="[340,420,1901,1926]" italics="true" pageId="118" pageNumber="119">
<collectingCountry id="F3217642FA13FFDECA6CBE76FE74F82C" box="[340,412,1901,1926]" name="Belize" pageId="118" pageNumber="119">Belize</collectingCountry>
:
</emphasis>
1 spec., 15.2 x 
<quantity id="4CCE9B37FA13FFDEC952BE75FD2CF82F" box="[618,708,1902,1926]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="118" pageNumber="119" unit="mm" value="6.6">6.6 mm</quantity>
(apex gone), on shallow reef, 
<collectingCountry id="F3217642FA13FFDECF0BBE75FB44F82C" box="[1075,1196,1901,1926]" name="Saint Pierre and Miquelon" pageId="118" pageNumber="119">San Pedro</collectingCountry>
, Ambergris Cay, M. Williams! 
<date id="FF881012FA13FFDECA35BE89FE61F800" box="[269,393,1938,1962]" pageId="118" pageNumber="119" value="1990-10-08">8 Oct 1990</date>
(M. Williams coll.).
</paragraph>
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<paragraph id="8B8936D2FA12FFDFCBAFBE07FC44F840" blockId="119.[151,1436,1820,2026]" pageId="119" pageNumber="120">
<emphasis id="B942EAC0FA12FFDFCBAFBE07FEFFF898" bold="true" box="[151,279,1820,1842]" pageId="119" pageNumber="120">PLATE 54.</emphasis>
<taxonomicName id="4C364D51FA12FFDFCA1BBE07FDC3F898" box="[291,555,1820,1842]" class="Gastropoda" family="Drilliidae" genus="Decoradrillia" kingdom="Animalia" order="Neogastropoda" pageId="119" pageNumber="120" phylum="Mollusca" rank="species" species="harlequina" status="sp. nov.">
<emphasis id="B942EAC0FA12FFDFCA1BBE07FDC3F898" box="[291,555,1820,1842]" italics="true" pageId="119" pageNumber="120">Decoradrillia harlequina</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A27157BBFA12FFDFC906BE06FD2BF898" box="[574,707,1820,1842]" pageId="119" pageNumber="120" rank="species">new species</taxonomicNameLabel>
. 
<emphasis id="B942EAC0FA12FFDFC9EBBE06FC88F898" bold="true" box="[723,864,1821,1842]" pageId="119" pageNumber="120">Upper panel</emphasis>
. Figs. 1–3: holotype, Devil's Bay, Grenada (USNM 1291335). Fig. 1: ventral, lateral &amp; dorsal views; Fig. 2: enlarged view of protoconch (not to scale); Fig. 3: apical view, V = varix, L = edge of outer lip. 
<emphasis id="B942EAC0FA12FFDFCA80BE41FDD7F8C5" bold="true" box="[440,575,1882,1903]" pageId="119" pageNumber="120">Lower panel</emphasis>
. Fig. 4: paratype, from the type locality (USNM 1291336); Fig. 5: Man of War Bay, Tobago I., Trinidad &amp; Tobago (ANSP 464967); Figs. 6–7: paratypes, Annas Shoal, St. Georges, Grenada (P. Williams coll.); Figs. 8–9: Hollandes Cays, San Blas Is., Panama (P. Williams coll.); Fig. 10: San Pedro, Ambergris Cay, Belize (P. Williams coll.); Figs. 11–12: Mt. Irving Bay, Tobago I., Trinidad &amp; Tobago (P. Williams coll.); Fig. 13: paratype, Union I., St. Vincent &amp; the Grenadines (UF 496635); Fig. 14: St Martin (MNHN, ex J. Colomb coll.).
</paragraph>
</caption>
<paragraph id="8B8936D2FA1DFFD0CBFFB983FA75FE2D" blockId="120.[151,1437,151,1583]" pageId="120" pageNumber="121">
<emphasis id="B942EAC0FA1DFFD0CBFFB983FE47FF1A" bold="true" box="[199,431,151,176]" pageId="120" pageNumber="121">Range and habitat.</emphasis>
<collectingCountry id="F3217642FA1DFFD0CA80B98CFDFCFF1A" box="[440,532,151,176]" name="Bahamas" pageId="120" pageNumber="121">Bahama</collectingCountry>
Is. (Grand 
<collectingCountry id="F3217642FA1DFFD0C9A1B98CFD1DFF1A" box="[665,757,151,176]" name="Bahamas" pageId="120" pageNumber="121">Bahama</collectingCountry>
I.); Greater Antilles (
<collectingCountry id="F3217642FA1DFFD0C8D7B98CFBA6FF1A" box="[1007,1102,151,176]" name="Jamaica" pageId="120" pageNumber="121">Jamaica</collectingCountry>
), Lesser Antilles (
<collectingCountry id="F3217642FA1DFFD0CE19B98CFA70FF1A" box="[1313,1432,151,176]" name="Sint Martin" pageId="120" pageNumber="121">St. Martin</collectingCountry>
; SVG; 
<collectingCountry id="F3217642FA1DFFD0CBD8B9A7FEABFF7F" box="[224,323,188,213]" name="Grenada" pageId="120" pageNumber="121">Grenada</collectingCountry>
); South 
<collectingCountry id="F3217642FA1DFFD0CA9BB9A7FDEBFF7F" box="[419,515,188,213]" name="United States of America" pageId="120" pageNumber="121">America</collectingCountry>
(
<collectingCountry id="F3217642FA1DFFD0C92CB9A7FD1DFF7F" box="[532,757,188,213]" name="Trinidad and Tobago" pageId="120" pageNumber="121">Trinidad &amp; Tobago</collectingCountry>
: 
<collectingCountry id="F3217642FA1DFFD0C839B9A7FCBDFF7F" box="[769,853,188,213]" name="Trinidad and Tobago" pageId="120" pageNumber="121">Tobago</collectingCountry>
I.; 
<collectingCountry id="F3217642FA1DFFD0C846B9A7FC19FF7F" box="[894,1009,188,213]" name="Colombia" pageId="120" pageNumber="121">Colombia</collectingCountry>
: San Andreas Is.); 
<collectingCountry id="F3217642FA1DFFD0CFF4B9A6FACDFF7E" box="[1228,1317,189,212]" name="Panama" pageId="120" pageNumber="121">Panama</collectingCountry>
(San Blas Is.: Hollandes Cays); and 
<collectingCountry id="F3217642FA1DFFD0CA87B9C4FDEFFF52" box="[447,519,223,248]" name="Belize" pageId="120" pageNumber="121">Belize</collectingCountry>
(Ambergris Cay). Published photographs of specimens from 
<collectingCountry id="F3217642FA1DFFD0CFFEB9C4FAC8FF52" box="[1222,1312,223,248]" name="Jamaica" pageId="120" pageNumber="121">Jamaica</collectingCountry>
(Humfrey, 1975) and 
<collectingCountry id="F3217642FA1DFFD0CA28B81FFE96FEB7" box="[272,382,260,285]" name="Colombia" pageId="120" pageNumber="121">Colombia</collectingCountry>
(San Andrea Is.) (Daccarett &amp; Bossio, 2011) are identifiable as this species. This species has been collected in coral reefs in mostly a rather narrow depth range of 
<quantity id="4CCE9B37FA1DFFD0C8A4B833FBE9FE95" box="[924,1025,296,320]" metricMagnitude="1" metricUnit="m" metricValue="1.95" metricValueMax="2.9" metricValueMin="1.0" pageId="120" pageNumber="121" unit="m" value="19.5" valueMax="29.0" valueMin="10.0">
10– 
<specimenCount id="9D30FD5BFA1DFFD0C8FDB833FBE9FE95" box="[965,1025,296,320]" pageId="120" pageNumber="121" type="generic">29 m</specimenCount>
</quantity>
. Most specimens exhibit evidence of predator attacks with severe repairs on the spire and body whorls, not counting missing lips, post-varix (which may have been broken in handling). The shell is also very solid, probably a defense against this high predatory pressure.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA1DFFD0CBFFB88FFB0DFBD5" pageId="120" pageNumber="121" type="description">
<paragraph id="8B8936D2FA1DFFD0CBFFB88FFB0DFBD5" blockId="120.[151,1437,151,1583]" pageId="120" pageNumber="121">
<emphasis id="B942EAC0FA1DFFD0CBFFB88FFE73FE07" bold="true" box="[199,411,404,429]" pageId="120" pageNumber="121">
Description. 
<emphasis id="B942EAC0FA1DFFD0CA58B88FFE73FE07" bold="true" box="[352,411,404,429]" italics="true" pageId="120" pageNumber="121">Shell</emphasis>
</emphasis>
small, to 
<quantity id="4CCE9B37FA1DFFD0C932B88FFD99FE06" box="[522,625,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="120" pageNumber="121" unit="mm" value="19.5">19.5 mm</quantity>
, fusiform, truncated anteriorly, up to 11¼ convex, bulging whorls; body whorl an approximately 57% of shell height; aperture moderately wide, columella straight. 
<emphasis id="B942EAC0FA1DFFD0CE2EB8ACFA73FE7A" bold="true" box="[1302,1435,439,464]" italics="true" pageId="120" pageNumber="121">Protoconch</emphasis>
approximately 2½ smooth whorls, well rounded, the first not immersed in the second. 
<emphasis id="B942EAC0FA1DFFD0CF49B8C7FACBFE5F" bold="true" box="[1137,1315,476,501]" italics="true" pageId="120" pageNumber="121">Axial sculpture</emphasis>
of arcuate ribs on the first teleoconch whorl, which transition into straight ribs by 2nd broad, and the remaining ribs, which are bulging, round-crested and extend from suture-to-suture, broader than intercostal spaces; reduced, narrower, and slightly recurved on shoulder. Ribs number 9–10 on the penultimate, and 7–10 to the varix on the body whorl. Growth striae cover entire shell surface; striae on the trailing sides of ribs run parallel with the ribs; striae on the leading side of the ribs cross over ribs obliquely at rib shoulders; striae in the sulcus are recurved, echoing the shape of the anal sinus. 
<emphasis id="B942EAC0FA1DFFD0CA9BBBADFE37FD65" bold="true" box="[419,479,694,719]" italics="true" pageId="120" pageNumber="121">Varix</emphasis>
wider and more prominent than the adjacent ribs, located about ¼–⅓-turn back from the edge of the outer lip. 
<emphasis id="B942EAC0FA1DFFD0CAFDBBC1FD6AFD59" bold="true" box="[453,642,730,755]" italics="true" pageId="120" pageNumber="121">Spiral sculpture</emphasis>
of 2–3 raised threads on anterior portion of teleoconch whorls, and approximately 17 below the periphery of the body whorl, fairly evenly spaced up to the anterior fasciole. Between the threads are 5 or more finer microscopic threads or grooves, approximately 40/mm on the 
<typeStatus id="548D8870FA1DFFD0CFDCBA39FAA1FC91" box="[1252,1353,802,827]" pageId="120" pageNumber="121" type="holotype">holotype</typeStatus>
, which impart a matte finish to shell surface. 
<emphasis id="B942EAC0FA1DFFD0C972BA5DFD70FCF5" bold="true" box="[586,664,838,863]" italics="true" pageId="120" pageNumber="121">Sulcus</emphasis>
narrow and concave with curved extensions of the ribs and growth striae whose shape reflect the round outline of the anal sinus. 
<emphasis id="B942EAC0FA1DFFD0C851BA71FC3BFC29" bold="true" box="[873,979,874,899]" italics="true" pageId="120" pageNumber="121">Outer lip</emphasis>
thin, edge forming a low arc from and congruent with the anal sinus to the stromboid notch. 
<emphasis id="B942EAC0FA1DFFD0C818BA95FC74FC0D" bold="true" box="[800,924,910,935]" italics="true" pageId="120" pageNumber="121">Anal sinus</emphasis>
deep and round, constricted by parietal lobe, almost closed in older specimens, edges flared, angled away from shell axis. 
<emphasis id="B942EAC0FA1DFFD0CF39BAA8FB81FC61" bold="true" box="[1025,1129,946,971]" italics="true" pageId="120" pageNumber="121">Inner lip</emphasis>
margined, thick at anterior end, thinner on parietal wall, and ending posteriorly in a large tubercle. 
<emphasis id="B942EAC0FA1DFFD0C8EDBACDFB69FC45" bold="true" box="[981,1153,982,1007]" italics="true" pageId="120" pageNumber="121">Anterior canal</emphasis>
deep, open, and slightly twisted to the right when viewed ventrally, end of canal flared up dorsally, notched; anterior fasciole slightly swollen; with about 8 spiral threads. 
<emphasis id="B942EAC0FA1DFFD0C90EBD05FD90FB9D" bold="true" box="[566,632,1054,1079]" italics="true" pageId="120" pageNumber="121">Color</emphasis>
yellowish white to white base with 3 narrow brown bands, about equal in width, the posterior-most one broken by the crests of the ribs forming a row of brown spots. Only two bands are visible on the spire whorls, one at the suture and the row of spots just above the whorl’s periphery.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA1DFFD0CBFFBD90FE52FA4D" pageId="120" pageNumber="121" type="discussion">
<paragraph id="8B8936D2FA1DFFD0CBFFBD90FE52FA4D" blockId="120.[151,1437,151,1583]" pageId="120" pageNumber="121">
<emphasis id="B942EAC0FA1DFFD0CBFFBD90FE57FB08" bold="true" box="[199,447,1163,1186]" pageId="120" pageNumber="121">
Remarks. 
<emphasis id="B942EAC0FA1DFFD0CA7ABD90FE57FB08" box="[322,447,1163,1186]" italics="true" pageId="120" pageNumber="121">Taxonomy.</emphasis>
</emphasis>
<taxonomicName id="4C364D51FA1DFFD0CAFFBD91FD0AFB09" box="[455,738,1162,1187]" class="Gastropoda" family="Drilliidae" genus="Decoradrillia" kingdom="Animalia" order="Neogastropoda" pageId="120" pageNumber="121" phylum="Mollusca" rank="species" species="harlequina">Decoradrillia harlequina</taxonomicName>
is an appropriate representative of the new genus because it is widely distributed throughout the Caribbean and possesses strong generic characteristics as the 
<typeStatus id="548D8870FA1DFFD0CFEFBDABFAEFFB6D" box="[1239,1287,1200,1223]" pageId="120" pageNumber="121">type</typeStatus>
of the genus should: broad bulging ribs, enlarged, rib-like varix (not hump-like), elongate anterior canal, and presence of closely-packed wavy microscopic spiral threads. 
<emphasis id="B942EAC0FA1DFFD0C986BDEDFCD5FAA5" bold="true" box="[702,829,1270,1295]" italics="true" pageId="120" pageNumber="121">Variability.</emphasis>
The average total length of 16 unbroken specimens is 
<quantity id="4CCE9B37FA1DFFD0CBAFBC00FEE9FA98" box="[151,257,1307,1331]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" pageId="120" pageNumber="121" unit="mm" value="15.7">15.7 mm</quantity>
(
<quantity id="4CCE9B37FA1DFFD0CA2DBC01FE28FA99" box="[277,448,1306,1331]" metricMagnitude="-2" metricUnit="m" metricValue="1.555" metricValueMax="1.95" metricValueMin="1.16" pageId="120" pageNumber="121" unit="mm" value="15.55" valueMax="19.5" valueMin="11.6">11.6–19.5 mm</quantity>
) and their W/ 
<taxonomicName id="4C364D51FA1DFFD0C955BC01FD55FA99" box="[621,701,1306,1331]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="120" pageNumber="121" phylum="Mollusca" rank="species" species="ratio">L ratio</taxonomicName>
is 0.375. The most visible characteristic, its color pattern, is surprisingly constant among specimens examined so far. Those from 
<collectingCountry id="F3217642FA1DFFD0C888BC25FBECFAFD" box="[944,1028,1342,1367]" name="Trinidad and Tobago" pageId="120" pageNumber="121">Tobago</collectingCountry>
I. have a brown, not white anterior fasciole (
<figureCitation id="130D2A57FA1DFFD0CBC5BC79FEB3FAD1" box="[253,347,1378,1403]" captionStart="PLATE 54" captionStartId="119.[151,234,1820,1842]" captionTargetBox="[248,1337,195,1796]" captionTargetId="figure@119.[248,1337,195,1796]" captionTargetPageId="119" captionText="PLATE 54. Decoradrillia harlequina, new species. Upper panel. Figs. 1 – 3: holotype, Devil's Bay, Grenada (USNM 1291335). Fig. 1: ventral, lateral &amp; dorsal views; Fig. 2: enlarged view of protoconch (not to scale); Fig. 3: apical view, V = varix, L = edge of outer lip. Lower panel. Fig. 4: paratype, from the type locality (USNM 1291336); Fig. 5: Man of War Bay, Tobago I., Trinidad &amp; Tobago (ANSP 464967); Figs. 6 – 7: paratypes, Annas Shoal, St. Georges, Grenada (P. Williams coll.); Figs. 8 – 9: Hollandes Cays, San Blas Is., Panama (P. Williams coll.); Fig. 10: San Pedro, Ambergris Cay, Belize (P. Williams coll.); Figs. 11 – 12: Mt. Irving Bay, Tobago I., Trinidad &amp; Tobago (P. Williams coll.); Fig. 13: paratype, Union I., St. Vincent &amp; the Grenadines (UF 496635); Fig. 14: St Martin (MNHN, ex J. Colomb coll.)." httpUri="https://zenodo.org/record/263354/files/figure.png" pageId="120" pageNumber="121">Plate 54</figureCitation>
, Figs. 11, dorsal view). 
<emphasis id="B942EAC0FA1DFFD0C95FBC79FBC6FAD1" box="[615,1070,1378,1403]" italics="true" pageId="120" pageNumber="121">
<emphasis id="B942EAC0FA1DFFD0C95FBC79FCE4FAD1" bold="true" box="[615,780,1378,1403]" italics="true" pageId="120" pageNumber="121">Identification.</emphasis>
<taxonomicName id="4C364D51FA1DFFD0C82ABC79FBC6FAD1" box="[786,1070,1378,1403]" class="Gastropoda" family="Drilliidae" genus="Decoradrillia" kingdom="Animalia" order="Neogastropoda" pageId="120" pageNumber="121" phylum="Mollusca" rank="species" species="harlequina">Decoradrillia harlequina</taxonomicName>
</emphasis>
is not likely to be confused with its congeners on account of its color pattern. It is most similar to 
<taxonomicName id="4C364D51FA1DFFD0C8A2BC93FB13FA35" authority="Tippett, 1995" authorityName="Tippett" authorityYear="1995" box="[922,1275,1414,1439]" class="Gastropoda" family="Drilliidae" genus="Fenimorea" kingdom="Animalia" order="Neogastropoda" pageId="120" pageNumber="121" phylum="Mollusca" rank="species" species="petiti">
<emphasis id="B942EAC0FA1DFFD0C8A2BC93FBB0FA35" box="[922,1112,1416,1439]" italics="true" pageId="120" pageNumber="121">Fenimorea petiti</emphasis>
Tippett, 1995
</taxonomicName>
but differs in having more swollen ribs, closely packed spiral threads, not the rows of “pits” formed by spiral grooves in 
<taxonomicName id="4C364D51FA1DFFD0CE78BCB0FA7DFA68" box="[1344,1429,1451,1474]" class="Gastropoda" family="Drilliidae" genus="Fenimorea" kingdom="Animalia" order="Neogastropoda" pageId="120" pageNumber="121" phylum="Mollusca" rank="species" species="petiti">
<emphasis id="B942EAC0FA1DFFD0CE78BCB0FA7DFA68" box="[1344,1429,1451,1474]" italics="true" pageId="120" pageNumber="121">F. petiti</emphasis>
</taxonomicName>
, and a different coloration.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA1DFFD0CBFFBCE8FC9CF985" pageId="120" pageNumber="121" type="etymology">
<paragraph id="8B8936D2FA1DFFD0CBFFBCE8FC9CF985" blockId="120.[151,1437,151,1583]" pageId="120" pageNumber="121">
<emphasis id="B942EAC0FA1DFFD0CBFFBCE8FEA6F9A0" bold="true" box="[199,334,1523,1546]" pageId="120" pageNumber="121">Etymology.</emphasis>
The Harlequin 
<taxonomicName id="4C364D51FA1DFFD0C93DBCE9FD77F9A1" box="[517,671,1522,1547]" class="Gastropoda" family="Drilliidae" genus="Decoradrillia" kingdom="Animalia" order="Neogastropoda" pageId="120" pageNumber="277" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B942EAC0FA1DFFD0C93DBCE9FD77F9A1" box="[517,671,1522,1547]" italics="true" pageId="120" pageNumber="121">Decoradrillia</emphasis>
</taxonomicName>
. The classic harlequin, in a light and dark checkerboard-patterned suit, and this gaily garbed gastropod have a similar comic appeal.
</paragraph>
</subSubSection>
</treatment>
</document>