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<document id="5B7064D7F8FEF6F130AE85860DFABEA0" ID-DOI="10.5281/zenodo.191317" ID-GBIF-Dataset="94e38b1e-c437-4315-8538-daaf32535555" ID-ISSN="1175-5326" ID-Zenodo-Dep="191317" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461126606189" checkinUser="plazi" docAuthor="Elberg, Kaupo, Rozkošný, Rudolf &amp; Knutson, Lloyd" docDate="2009" docId="039F8602D56BD0035F930E5196E2FE5E" docLanguage="en" docName="zt02288p060.pdf" docOrigin="Zootaxa 2288" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sepedon spinipes Scopoli 1763" docType="treatment" docVersion="9" lastPageNumber="59" masterDocId="FFA6FE7AD56CD00B5F040B349741FF9E" masterDocTitle="A review of of the Holarctic Sepedon fuscipennis and S. spinipes groups with description of a new species (Diptera: Sciomyzidae)" masterLastPageNumber="60" masterPageNumber="51" pageNumber="58" updateTime="1698596924154" updateUser="plazi">
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<mods:title id="18C96BBF9165492DA24347A27295F0AA">A review of of the Holarctic Sepedon fuscipennis and S. spinipes groups with description of a new species (Diptera: Sciomyzidae)</mods:title>
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<mods:namePart id="C0F95FC88B4F75DCEC99E5C767291AB5">Elberg, Kaupo</mods:namePart>
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<mods:namePart id="8AD17066FC6D3BA81C1E6B906EDFE296">Rozkošný, Rudolf</mods:namePart>
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<mods:namePart id="37A4FEFF2DDF0AA73F3C702F338AE99D">Knutson, Lloyd</mods:namePart>
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<mods:date id="3AB2A5CCB93C244FB7B6A0A569F93435">2009</mods:date>
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<treatment id="039F8602D56BD0035F930E5196E2FE5E" ID-DOI="http://doi.org/10.5281/zenodo.6216450" ID-GBIF-Taxon="119626352" ID-Zenodo-Dep="6216450" LSID="urn:lsid:plazi:treatment:039F8602D56BD0035F930E5196E2FE5E" httpUri="http://treatment.plazi.org/id/039F8602D56BD0035F930E5196E2FE5E" lastPageId="8" lastPageNumber="59" pageId="7" pageNumber="58">
<subSubSection id="C32C649FD56BD00C5F930E519618FA3C" pageId="7" pageNumber="58" type="nomenclature">
<paragraph id="8B893714D56BD00C5F930E519564FAE1" blockId="7.[151,549,1381,1442]" box="[151,549,1381,1407]" pageId="7" pageNumber="58">
<heading id="D0C18078D56BD00C5F930E519564FAE1" bold="true" box="[151,549,1381,1407]" fontSize="11" level="1" pageId="7" pageNumber="58" reason="1">
<taxonomicName id="4C364C97D56BD00C5F930E519564FAE1" ID-CoL="4WPX2" authority="Scopoli, 1763" authorityName="Scopoli" authorityYear="1763" box="[151,549,1381,1407]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B942EB06D56BD00C5F930E519564FAE1" bold="true" box="[151,549,1381,1407]" pageId="7" pageNumber="58">
<emphasis id="B942EB06D56BD00C5F930E519627FAE1" bold="true" box="[151,358,1381,1407]" italics="true" pageId="7" pageNumber="58">Sepedon spinipes</emphasis>
(
<bibRefCitation id="EFA74AE5D56BD00C5E720E51955CFAE1" author="Scopoli" box="[374,541,1381,1407]" pageId="7" pageNumber="60" refString="Scopoli, G. A. (1763) Entomologia Carniolica exhibens Insecta Carnioliae indigene et distribute in ordines, genera, species, varietates. Methodo Linnaeana. I. T. Trattner, Vindobonae, (30) + 420 pp." type="book" year="1763">Scopoli, 1763</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B893714D56BD00C5F930EBC9618FA3C" blockId="7.[151,549,1381,1442]" box="[151,345,1416,1442]" pageId="7" pageNumber="58">
(
<figureCitation id="130D2B91D56BD00C5F9B0EBC9666FA3C" box="[159,295,1416,1442]" captionStart="FIGURES 20 27" captionStartId="7.[151,269,1208,1232]" captionTargetBox="[186,1404,454,1147]" captionTargetId="figure@7.[151,1436,395,1184]" captionTargetPageId="7" captionText="FIGURES 20 27. Species of Sepedon spinipes group, male terminalia. 20 23. S. americana. 24 27. S. spinipes. 20, 24. Lateral view. 21, 25. Hypandrium in lateral view. 22, 26. Aedeagus in lateral view. 23, 27. Epiphallus in caudal view. Scales: 0.5 mm (Figs 20, 24), 0.3 mm (Figs 21 23, 24 27)." httpUri="https://zenodo.org/record/191321/files/figure.png" pageId="7" pageNumber="58">Figs 2427</figureCitation>
,
<figureCitation id="130D2B91D56BD00C5E360EBC9610FA3C" box="[306,337,1416,1442]" captionStart="FIGURES 28 29" captionStartId="8.[151,269,686,710]" captionTargetBox="[432,1143,514,655]" captionTargetId="figure@8.[430,1146,514,658]" captionTargetPageId="8" captionText="FIGURES 28 29. Wings of Sepedon species. 28. S. hecate sp. nov. 29. S. spinipes." httpUri="https://zenodo.org/record/191322/files/figure.png" pageId="7" pageNumber="58">29</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32C649FD56BD00C5F930EE194F2F8B6" pageId="7" pageNumber="58" type="diagnosis">
<paragraph id="8B893714D56BD00C5F930EE19485F9A3" blockId="7.[151,1437,1493,2025]" pageId="7" pageNumber="58">
<emphasis id="B942EB06D56BD00C5F930EE19656FA71" bold="true" box="[151,279,1493,1519]" pageId="7" pageNumber="58">Diagnosis.</emphasis>
Cerci usually reaching beyond lower margin of epandrium in lateral view; surstylus much longer than distiphallus (
<figureCitation id="130D2B91D56BD00C5E6D0EC89680F988" box="[361,449,1532,1558]" captionStart="FIGURES 20 27" captionStartId="7.[151,269,1208,1232]" captionTargetBox="[186,1404,454,1147]" captionTargetId="figure@7.[151,1436,395,1184]" captionTargetPageId="7" captionText="FIGURES 20 27. Species of Sepedon spinipes group, male terminalia. 20 23. S. americana. 24 27. S. spinipes. 20, 24. Lateral view. 21, 25. Hypandrium in lateral view. 22, 26. Aedeagus in lateral view. 23, 27. Epiphallus in caudal view. Scales: 0.5 mm (Figs 20, 24), 0.3 mm (Figs 21 23, 24 27)." httpUri="https://zenodo.org/record/191321/files/figure.png" pageId="7" pageNumber="58">Fig. 24</figureCitation>
); emargination of basal part of hypandrium conspicuous (
<figureCitation id="130D2B91D56BD00C5B6E0EC89382F988" box="[1130,1219,1532,1558]" captionStart="FIGURES 20 27" captionStartId="7.[151,269,1208,1232]" captionTargetBox="[186,1404,454,1147]" captionTargetId="figure@7.[151,1436,395,1184]" captionTargetPageId="7" captionText="FIGURES 20 27. Species of Sepedon spinipes group, male terminalia. 20 23. S. americana. 24 27. S. spinipes. 20, 24. Lateral view. 21, 25. Hypandrium in lateral view. 22, 26. Aedeagus in lateral view. 23, 27. Epiphallus in caudal view. Scales: 0.5 mm (Figs 20, 24), 0.3 mm (Figs 21 23, 24 27)." httpUri="https://zenodo.org/record/191321/files/figure.png" pageId="7" pageNumber="58">Fig. 25</figureCitation>
); distiphallus only short, barely longer than epiphallus and not dilated apically (
<figureCitation id="130D2B91D56BD00C5C5A0D1794F7F9A3" box="[862,950,1571,1597]" captionStart="FIGURES 20 27" captionStartId="7.[151,269,1208,1232]" captionTargetBox="[186,1404,454,1147]" captionTargetId="figure@7.[151,1436,395,1184]" captionTargetPageId="7" captionText="FIGURES 20 27. Species of Sepedon spinipes group, male terminalia. 20 23. S. americana. 24 27. S. spinipes. 20, 24. Lateral view. 21, 25. Hypandrium in lateral view. 22, 26. Aedeagus in lateral view. 23, 27. Epiphallus in caudal view. Scales: 0.5 mm (Figs 20, 24), 0.3 mm (Figs 21 23, 24 27)." httpUri="https://zenodo.org/record/191321/files/figure.png" pageId="7" pageNumber="58">Fig. 26</figureCitation>
).
</paragraph>
<paragraph id="8B893714D56BD00C5FC10D7E94F2F8B6" blockId="7.[151,1437,1493,2025]" pageId="7" pageNumber="58">
At first glance,
<taxonomicName id="4C364C97D56BD00C5E860D7E9542F9FD" box="[386,515,1610,1636]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B942EB06D56BD00C5E860D7E96D0F9FD" box="[386,401,1610,1635]" italics="true" pageId="7" pageNumber="58">S</emphasis>
.
<emphasis id="B942EB06D56BD00C5EA50D7E9542F9FD" box="[417,515,1610,1635]" italics="true" pageId="7" pageNumber="58">spinipes</emphasis>
</taxonomicName>
might be mistaken for the new Palaearctic species,
<taxonomicName id="4C364C97D56BD00C5B730D7E9272F9FD" box="[1143,1331,1610,1635]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="hecate" status="sp. nov.">
<emphasis id="B942EB06D56BD00C5B730D7E9272F9FD" box="[1143,1331,1610,1635]" italics="true" pageId="7" pageNumber="58">Sepedon hecate</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A271567DD56BD00C5A380D7D92DDF9FD" box="[1340,1436,1609,1635]" pageId="7" pageNumber="58" rank="species">
<emphasis id="B942EB06D56BD00C5A380D7D92DDF9FD" bold="true" box="[1340,1436,1609,1635]" pageId="7" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
However, detailed comparison of both species shows reliable differences.
<taxonomicName id="4C364C97D56BD00C5CFA0D45933DF914" box="[1022,1148,1648,1674]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B942EB06D56BD00C5CFA0D45934CF914" box="[1022,1037,1649,1674]" italics="true" pageId="7" pageNumber="58">S</emphasis>
.
<emphasis id="B942EB06D56BD00C5B180D45933DF914" box="[1052,1148,1649,1674]" italics="true" pageId="7" pageNumber="58">spinipes</emphasis>
</taxonomicName>
is generally smaller and paler, the black orbital spot is of a different shape, the pedicel is stouter, the dark longitudinal stripe above the postpronotal callus and notopleuron is almost indistinct, the katatergite is setulose in the posterior part, cell r4+5 is parallel-sided, and the posterior crossvein is much more bowed than sinuate (
<figureCitation id="130D2B91D56BD00C5B420DD393DEF89F" box="[1094,1183,1767,1793]" captionStart="FIGURES 28 29" captionStartId="8.[151,269,686,710]" captionTargetBox="[432,1143,514,655]" captionTargetId="figure@8.[430,1146,514,658]" captionTargetPageId="8" captionText="FIGURES 28 29. Wings of Sepedon species. 28. S. hecate sp. nov. 29. S. spinipes." httpUri="https://zenodo.org/record/191322/files/figure.png" pageId="7" pageNumber="58">Fig. 28</figureCitation>
). Distinct differences were also found in the shape of the male terminalia, as noted above.
</paragraph>
</subSubSection>
<subSubSection id="C32C649FD56BD0035FC20C009579FF2C" lastPageId="8" lastPageNumber="59" pageId="7" pageNumber="58" type="distribution">
<paragraph id="8B893714D56BD0035FC20C009579FF2C" blockId="7.[151,1437,1493,2025]" lastBlockId="8.[151,1437,152,448]" lastPageId="8" lastPageNumber="59" pageId="7" pageNumber="58">
<emphasis id="B942EB06D56BD00C5FC20C009626F8D0" bold="true" box="[198,359,1844,1870]" pageId="7" pageNumber="58">Distribution.</emphasis>
Details of the male genitalia of
<taxonomicName id="4C364C97D56BD00C5DF80C01943FF8D0" box="[764,894,1844,1870]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B942EB06D56BD00C5DF80C01944AF8D0" box="[764,779,1845,1870]" italics="true" pageId="7" pageNumber="58">S</emphasis>
.
<emphasis id="B942EB06D56BD00C5C1A0C01943FF8D0" box="[798,894,1845,1870]" italics="true" pageId="7" pageNumber="58">spinipes</emphasis>
</taxonomicName>
were recently re-examined in representative material from widespread parts of the distribution area and also compared with earlier published data from the Palaearctic, if they were accompanied by sufficiently detailed figures of the male terminalia. It is thus confirmed that
<taxonomicName id="4C364C97D56BD00C5E4A0C9D968EF85C" box="[334,463,1961,1987]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B942EB06D56BD00C5E4A0C9D961CF85C" box="[334,349,1961,1986]" italics="true" pageId="7" pageNumber="58">S</emphasis>
.
<emphasis id="B942EB06D56BD00C5E690C9D968EF85C" box="[365,463,1961,1986]" italics="true" pageId="7" pageNumber="58">spinipes</emphasis>
</taxonomicName>
occurs over the large territory from
<collectingCountry id="F3217784D56BD00C5C830C9D949EF85D" box="[903,991,1961,1987]" name="Ireland" pageId="7" pageNumber="58">Ireland</collectingCountry>
, the British Isles, and Scandinavia to North Africa (
<collectingCountry id="F3217784D56BD00C5E440CFB96EEF877" box="[320,431,1999,2025]" name="Morocco" pageId="7" pageNumber="58">Morocco</collectingCountry>
) and from
<collectingCountry id="F3217784D56BD00C5D370CFB95D9F877" box="[563,664,1999,2025]" name="Portugal" pageId="7" pageNumber="58">Portugal</collectingCountry>
to
<collectingCountry id="F3217784D56BD00C5DBB0CFB9473F877" box="[703,818,1999,2025]" name="Mongolia" pageId="7" pageNumber="58">Mongolia</collectingCountry>
and the Russian Far East. However, it has not been found in
<collectingCountry id="F3217784D564D0035E050BAC9605FF2C" box="[257,324,152,178]" name="Japan" pageId="8" pageNumber="59">Japan</collectingCountry>
(cf.
<bibRefCitation id="EFA74AE5D564D0035E7E0BAC956AFF2C" author="Sueyoshi" box="[378,555,152,178]" pageId="8" pageNumber="60" refString="Sueyoshi, M. (2001) A revision of Japanese Sciomyzidae (Diptera) with descriptions of three new species. Entomological Science 4, 485 - 506." type="journal article" year="2001">Sueyoshi 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32C649FD564D0035FC10B8A96E2FE5E" pageId="8" pageNumber="59" type="materials_examined">
<paragraph id="8B893714D564D0035FC10B8A96E2FE5E" blockId="8.[151,1437,152,448]" pageId="8" pageNumber="59">
<emphasis id="B942EB06D564D0035FC10B8A9687FF46" bold="true" box="[197,454,190,216]" pageId="8" pageNumber="59">Material examined.</emphasis>
<collectingCountry id="F3217784D564D0035ECB0B8A9534FF46" box="[463,629,190,216]" name="Portugal" pageId="8" pageNumber="59">PORTUGAL</collectingCountry>
: Evora,
<date id="FF8811D4D564D0035DDF0B8A9418FF46" box="[731,857,190,216]" pageId="8" pageNumber="59" value="1998-10-17">17.x.1998</date>
, 1 3, 1 Ƥ, J. Olejníček.;
<collectingCountry id="F3217784D564D0035B900B8A92D8FF46" box="[1172,1433,190,216]" name="Czech Republic" pageId="8" pageNumber="59">CZECH REPUBLIC</collectingCountry>
: Lednice,
<date id="FF8811D4D564D0035E0C0BD196C3FF61" box="[264,386,229,255]" pageId="8" pageNumber="59" value="1998-05-30">30.v.1998</date>
, 1 3, R. Rozkošný;
<collectingCountry id="F3217784D564D0035D7F0BD19454FF61" box="[635,789,229,255]" name="Slovakia" pageId="8" pageNumber="59">SLOVAKIA</collectingCountry>
: Čoltovo, 18.5.
<date id="FF8811D4D564D0035CCA0BD1930BFF60" box="[974,1098,228,255]" pageId="8" pageNumber="59" value="1980-01-03">1980, 1 3</date>
, R. Rozkošný (all
<collectionCode id="ED27AFD1D564D0035A3D0BD192D1FF61" box="[1337,1424,229,255]" pageId="8" pageNumber="59">FSMU</collectionCode>
).
<collectingCountry id="F3217784D564D0035F930A3897ADFEB8" box="[151,236,268,294]" name="Italy" pageId="8" pageNumber="59">ITALY</collectingCountry>
: Lazio, L. Fibreno,
<quantity id="4CCE9AF1D564D0035ED60A38954FFEB8" box="[466,526,268,294]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="59" unit="km" value="7.0">7 km</quantity>
SE of Sora,
<date id="FF8811D4D564D0035DA70A38944AFEB8" box="[675,779,268,294]" pageId="8" pageNumber="59" value="1990-05-07">7.v.1990</date>
, 1 Ƥ, L. Fornasari &amp; L.V. Knutson (
<collectionCode id="ED27AFD1D564D0035BB70A38924FFEB8" LSID="urn:lsid:biocol.org:col:34871" box="[1203,1294,268,294]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="59">USNM</collectionCode>
).
<collectingCountry id="F3217784D564D0035A260A3892D8FEB8" box="[1314,1433,268,294]" name="Greece" pageId="8" pageNumber="59">GREECE</collectingCountry>
: Crete, Gerani,
<quantity id="4CCE9AF1D564D0035E400A0696CEFED2" box="[324,399,306,332]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="59" unit="km" value="15.0">15 km</quantity>
W of Canea,
<date id="FF8811D4D564D0035D350A0695F2FED2" box="[561,691,306,332]" pageId="8" pageNumber="59" value="1963-03-21">21.iii.1963</date>
, 9 3, 2 Ƥ, L. Knutson (CU).
<collectingCountry id="F3217784D564D0035B080A069319FED2" box="[1036,1112,306,332]" name="Iran" pageId="8" pageNumber="59">IRAN</collectingCountry>
: Boneh Javaz,
<quantity id="4CCE9AF1D564D0035A0C0A069210FED2" box="[1288,1361,306,332]" metricMagnitude="4" metricUnit="m" metricValue="4.1" pageId="8" pageNumber="59" unit="km" value="41.0">41 km</quantity>
SE of Dezful,
<date id="FF8811D4D564D0035FFD0A6D962EFEED" box="[249,367,345,371]" pageId="8" pageNumber="59" value="1971-11-05">5.xi.1971</date>
, 5 3, 6 Ƥ, L. Knutson (
<collectionCode id="ED27AFD1D564D0035D970A6D95B0FEED" LSID="urn:lsid:biocol.org:col:34871" box="[659,753,345,371]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="59">USNM</collectionCode>
).
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: Bulgan Aimag, Teshig Soum, pond and marsh, wetland area near confluence Tariakhtain Gol &amp; Egiin Gol,
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,
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,
<date id="FF8811D4D564D0035BFE0AB492C3FE04" box="[1274,1410,384,410]" pageId="8" pageNumber="59" value="2005-07-10">10.vii.2005</date>
, 1 3, J. Gelhaus (
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).
</paragraph>
</subSubSection>
</treatment>
</document>