309 lines
55 KiB
XML
309 lines
55 KiB
XML
<document id="9EC210B677A7FA2C574C6137B69816A8" ID-DOI="10.1016/j.rbe.2017.04.004" ID-ISSN="1806-9665" ID-Zenodo-Dep="13195489" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722635027235" checkinUser="felipe" docAuthor="Ferreira, André da Silva, Almeida, Lúcia M. & Bravo, Freddy" docDate="2017" docId="6B7287A3FFE3FF9FF6ACFB70FD10F8E7" docLanguage="en" docName="RevBrasEntomol.61.3.208-223.pdf" docOrigin="Revista Brasileira de Entomologia 61 (3)" docSource="http://dx.doi.org/10.1016/j.rbe.2017.04.004" docStyle="DocumentStyle:0303D37C566E4E2A42C6591146119864.3:RevBrasEntomol.2015-.journal_article" docStyleId="0303D37C566E4E2A42C6591146119864" docStyleName="RevBrasEntomol.2015-.journal_article" docStyleVersion="3" docTitle="Pelidnota nordestina Ferreira, Almeida & Bravo, 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="212" masterDocId="974BFFDBFFE0FF9BF63EFFF3FF99FFD4" masterDocTitle="Three new species of Pelidnota MacLeay (Coleoptera, Scarabaeidae, Rutelinae) and new distributional records from northeast Brazil" masterLastPageNumber="223" masterPageNumber="208" pageNumber="211" updateTime="1722682458335" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5D1B312AF90E23FFE3D9F596F79F1D49">Three new species of Pelidnota MacLeay (Coleoptera, Scarabaeidae, Rutelinae) and new distributional records from northeast Brazil</mods:title>
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<mods:name id="1F31EF9F0FE3D4C08C18B88587DBB398" type="personal">
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<mods:roleTerm id="FC0CAE1A449691EEB7174460D87D5639">Author</mods:roleTerm>
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<mods:namePart id="DA038CC6B6B787E692CEDA64DB5E0C2C">Ferreira, André da Silva</mods:namePart>
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<mods:affiliation id="E2C99690180C280A30353ECBC9F6C4E7">Universidade Federal da Bahia, Instituto de Biologia, Programa de Pós-Graduação em Diversidade Animal, Bahia, BA, Brazil</mods:affiliation>
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</mods:name>
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<mods:name id="95D28019BA1DD26C433AD9CEA186F42C" type="personal">
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<mods:roleTerm id="1110C110AEEC2626236EBCB8336F7CC2">Author</mods:roleTerm>
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<mods:namePart id="CA5FADE8BF67623B80A9FD4A0FF360EF">Almeida, Lúcia M.</mods:namePart>
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<mods:affiliation id="2E06C832F141D817C279F4067E8E40F3">Universidade Federal do Paraná, Departamento de Zoologia, Laboratório de Sistemática e Bioecologia de Coleoptera, Curitiba, PR, Brazil</mods:affiliation>
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<mods:namePart id="10E9ABCFC27862E48A259EC5D3A43342">Bravo, Freddy</mods:namePart>
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<mods:affiliation id="ABF5B45299917590990BD79E3E324DCA">Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Laboratório de Sistemática de Insetos, Feira de Santana, BA, Brazil</mods:affiliation>
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<mods:title id="281C0C8A589D52FD206A9C5862640CF6">Revista Brasileira de Entomologia</mods:title>
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<mods:part id="43B3E47732D7BF91BBCD4A6459D3EB9F">
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<mods:date id="19BC309E3723929D8790DD66E8906D88">2017</mods:date>
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<mods:number id="A7F4C39D2543686FECF0C173E3B80757">2017-05-10</mods:number>
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<mods:number id="4C781B58A9F1D59EC8474541ACDD9ADC">61</mods:number>
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<mods:identifier id="E423430153C1E7EC15CAB243BD156C96" type="DOI">10.1016/j.rbe.2017.04.004</mods:identifier>
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<treatment id="6B7287A3FFE3FF9FF6ACFB70FD10F8E7" ID-DOI="http://doi.org/10.5281/zenodo.13194513" ID-Zenodo-Dep="13194513" LSID="urn:lsid:plazi:treatment:6B7287A3FFE3FF9FF6ACFB70FD10F8E7" httpUri="http://treatment.plazi.org/id/6B7287A3FFE3FF9FF6ACFB70FD10F8E7" lastPageId="4" lastPageNumber="212" pageId="3" pageNumber="211">
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<subSubSection id="ABC1653EFFE3FF98F6ACFB70FE5AFB43" box="[146,451,1155,1175]" pageId="3" pageNumber="211" type="nomenclature">
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<paragraph id="E36436B5FFE3FF98F6ACFB70FE5AFB43" blockId="3.[146,451,1155,1175]" box="[146,451,1155,1175]" pageId="3" pageNumber="211">
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<heading id="B82C81D9FFE3FF98F6ACFB70FE5AFB43" bold="true" box="[146,451,1155,1175]" fontSize="8" level="2" pageId="3" pageNumber="211" reason="2">
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<emphasis id="D1AFEAA7FFE3FF98F6ACFB70FE5AFB43" bold="true" box="[146,451,1155,1175]" pageId="3" pageNumber="211">
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<taxonomicName id="24DB4D36FFE3FF98F6ACFB70FEF4FB43" ID-CoL="75ZWV" authorityName="Ferreira & Almeida & Bravo" authorityYear="2017" box="[146,365,1155,1175]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="211" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
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<emphasis id="D1AFEAA7FFE3FF98F6ACFB70FEF4FB43" bold="true" box="[146,365,1155,1175]" italics="true" pageId="3" pageNumber="211">Pelidnota nordestina</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CA9C57DCFFE3FF98F74CFB77FE5AFB43" box="[370,451,1156,1175]" pageId="3" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="ABC1653EFFE3FF9FF6ACFB53FD10F8E7" lastPageId="4" lastPageNumber="212" pageId="3" pageNumber="211" type="description">
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<paragraph id="E36436B5FFE3FF98F6ACFB53FE8DFB67" blockId="3.[114,786,1184,1985]" box="[146,276,1184,1203]" pageId="3" pageNumber="211">
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(
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<figureCitation id="7BE02A30FFE3FF98F6A4FB53FE95FB67" box="[154,268,1184,1203]" captionStart="Figs" captionStartId="4.[85,120,1355,1369]" captionTargetBox="[250,1305,180,1309]" captionTargetId="figure-13@4.[249,1309,149,1324]" captionTargetPageId="4" captionText="Figs. 11–16. Pelidnota. 11–13, Pelidnota nordestina sp. nov., male holotype (dorsal, ventral, lateral); 14–16, P. pernambucana sp. nov., male holotype (dorsal, ventral, lateral). Scale bars: Figs. 11–13 =5.5 mm, Figs.14–16= 5.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195493" httpUri="https://zenodo.org/record/13195493/files/figure.png" pageId="3" pageNumber="211">Figs. 11–13</figureCitation>
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)
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</paragraph>
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<paragraph id="E36436B5FFE3FF98F6ACFB48FED3FA1E" blockId="3.[114,786,1184,1985]" pageId="3" pageNumber="211">
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<emphasis id="D1AFEAA7FFE3FF98F6ACFB48FF69FB1B" box="[146,240,1211,1231]" italics="true" pageId="3" pageNumber="211">Diagnosis</emphasis>
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. Labrum triangular (
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<figureCitation id="7BE02A30FFE3FF98F7EDFB4FFD85FB1B" box="[467,540,1212,1231]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 17–24. Mouthparts. 17–20, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 21–24, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.17–24 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195495" httpUri="https://zenodo.org/record/13195495/files/figure.png" pageId="3" pageNumber="211">Fig. 19</figureCitation>
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). Second teeth on the inner margin of mandible larger than first and third (
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<figureCitation id="7BE02A30FFE3FF98F482FB2BFC9DFB3F" box="[700,772,1240,1259]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 17–24. Mouthparts. 17–20, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 21–24, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.17–24 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195495" httpUri="https://zenodo.org/record/13195495/files/figure.png" pageId="3" pageNumber="211">Fig. 23</figureCitation>
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). Second teeth of maxillary galea sawed (
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<figureCitation id="7BE02A30FFE3FF98F427FB07FDC6FAD3" box="[537,607,1268,1287]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 25–28. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 27–28, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 25–28 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195497" httpUri="https://zenodo.org/record/13195497/files/figure.png" pageId="3" pageNumber="211">Fig. 27</figureCitation>
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). Surface area of pre-mentum excavated and lateral margin of post-mentum slight rounded (
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<figureCitation id="7BE02A30FFE3FF98F6EBFAD8FE8FFAEB" box="[213,278,1323,1343]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 29–32. Mouthparts. 29–32, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 29–32 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195499" httpUri="https://zenodo.org/record/13195499/files/figure.png" pageId="3" pageNumber="211">Fig. 31</figureCitation>
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). Anteroexternal coxal angle of metacoxa rounded and little acute (
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<figureCitation id="7BE02A30FFE3FF98F72AFAB4FECFFA8F" box="[276,342,1351,1371]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 33–40. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 33–36, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 37–40, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 33–36 =0.500 mm, Figs. 37–40 =1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13195501" httpUri="https://zenodo.org/record/13195501/files/figure.png" pageId="3" pageNumber="211">Fig. 39</figureCitation>
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). Pygidium dark brown, with reddish reflections on disk, with slight metallic green reflections only on lateral portion of basal margin (
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<figureCitation id="7BE02A30FFE3FF98F75BFA8CFE3FFA46" box="[357,422,1407,1426]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 41–46. Protibia and pygidium in dorsal view. 41–44, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 45–46, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 41–42 = 1.0mm, Figs. 45–76 =2.7 mm." figureDoi="http://doi.org/10.5281/zenodo.13195503" httpUri="https://zenodo.org/record/13195503/files/figure.png" pageId="3" pageNumber="211">Fig.45</figureCitation>
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). Six spine-like setae on the external longitudinal carinae and six setae on the inner longitudinal carinae of metatibia (
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<figureCitation id="7BE02A30FFE3FF98F6C5FA44FEA4FA1E" box="[251,317,1463,1482]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 53–56. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.53–56= 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195507" httpUri="https://zenodo.org/record/13195507/files/figure.png" pageId="3" pageNumber="211">Fig. 55</figureCitation>
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).
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</paragraph>
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<paragraph id="E36436B5FFE3FF98F6ACFA21FABAF9CA" blockId="3.[114,786,1184,1985]" lastBlockId="3.[833,1504,152,1984]" pageId="3" pageNumber="211">
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<emphasis id="D1AFEAA7FFE3FF98F6ACFA21FE14FA32" box="[146,397,1490,1510]" italics="true" pageId="3" pageNumber="211">
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<typeStatus id="3C608817FFE3FF98F6ACFA21FF73FA32" box="[146,234,1490,1510]" pageId="3" pageNumber="211" type="holotype">Holotype</typeStatus>
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male description
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</emphasis>
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. Total length
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<quantity id="24239B50FFE3FF98F42CFA20FDF2FA32" box="[530,619,1491,1510]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="3" pageNumber="211" unit="mm" value="25.7">25.7 mm</quantity>
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, width
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<quantity id="24239B50FFE3FF98F48DFA20FC97FA32" box="[691,782,1491,1510]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="3" pageNumber="211" unit="mm" value="13.1">13.1 mm</quantity>
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. Body elongated-oval, slightly convex.
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<emphasis id="D1AFEAA7FFE3FF98F7C5FA1DFDB7F9D6" box="[507,558,1518,1538]" italics="true" pageId="3" pageNumber="211">Color</emphasis>
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: Head golden-brown, with black margin. Pronotum, scutellum, and elytra golden-brown and with metallic green reflections. Anterior margin of pronotum and scutellum dark. Pronotum with one and elytra with two lateral black spots. Pygidium dark brown, with reddish reflections in middle, with slight metallic green reflections only on lateral portion of basal margin. Ventral surface and legs of same color as pygidium, and with metallic green reflections.
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<emphasis id="D1AFEAA7FFE3FF98F479F942FDE0F911" box="[583,633,1713,1733]" italics="true" pageId="3" pageNumber="211">Head</emphasis>
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: Antenna with 10 antennomeres; club composed by three antennomeres; scape elongated, shorter than antennomeres II–VII. Clypeus trapezoidal, puncture dense and large to moderately large; anterior margin concave. Frontoclypeal suture not evident. Frons width at middle region 2–3 times length of clypeus; punctures moderate to dense and moderately wide. Ocular canthus not reaching the middle of the eye. Interocular distance 3 times the transverse eye diameters. Labrum triangular and transversely elongated (
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<figureCitation id="7BE02A30FFE3FF98F466F862FD05F870" box="[600,668,1937,1956]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 17–24. Mouthparts. 17–20, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 21–24, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.17–24 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195495" httpUri="https://zenodo.org/record/13195495/files/figure.png" pageId="3" pageNumber="211">Fig. 19</figureCitation>
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), not fused to clypeus, with medial compression; acuminate anterior margin and posterior convex. Mandibles partially visible in dorsal view, outer margin black, with 2 exposed, recurved, outer teeth; 3 smaller teeth on inner margin, II larger than others; molar robust (
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<figureCitation id="7BE02A30FFE3FF98F3AEFF23FA4BFF37" box="[1424,1490,208,227]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 17–24. Mouthparts. 17–20, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 21–24, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.17–24 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195495" httpUri="https://zenodo.org/record/13195495/files/figure.png" pageId="3" pageNumber="211">Fig. 23</figureCitation>
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). Maxilla with galea bearing 6 teeth, first and second sawed; palp with 4 palpomeres, IV elongated and fusiform (
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<figureCitation id="7BE02A30FFE3FF98F30AFEF4FAE0FECF" box="[1332,1401,263,283]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 25–28. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 27–28, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 25–28 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195497" httpUri="https://zenodo.org/record/13195497/files/figure.png" pageId="3" pageNumber="211">Fig. 27</figureCitation>
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). Labium, mentum with apex concave; post-mentum with elongated dense setae laterally; palp inserted into a lateral cavity in pre-mentum, with 3 palpomeres, III elongate (
|
||
<figureCitation id="7BE02A30FFE3FF98F2BBFEA8FB5FFEBA" box="[1157,1222,347,366]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 29–32. Mouthparts. 29–32, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 29–32 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195499" httpUri="https://zenodo.org/record/13195499/files/figure.png" pageId="3" pageNumber="211">Fig. 31</figureCitation>
|
||
).
|
||
<emphasis id="D1AFEAA7FFE3FF98F2E6FEA9FAA0FEBA" box="[1240,1337,346,366]" italics="true" pageId="3" pageNumber="211">Pronotum</emphasis>
|
||
: Convex. Length
|
||
<quantity id="24239B50FFE3FF98F57FFE84FC14FE5E" box="[833,909,375,394]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="211" unit="mm" value="6.3">6.3 mm</quantity>
|
||
, anterior width
|
||
<quantity id="24239B50FFE3FF98F205FE84FB1EFE5E" box="[1083,1159,375,394]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="3" pageNumber="211" unit="mm" value="7.6">7.6 mm</quantity>
|
||
, mean width
|
||
<quantity id="24239B50FFE3FF98F322FE84FAECFE5E" box="[1308,1397,375,394]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="211" unit="mm" value="11.5">11.5 mm</quantity>
|
||
, posterior width
|
||
<quantity id="24239B50FFE3FF98F5B8FE60FC79FE72" box="[902,992,403,422]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="211" unit="mm" value="11.5">11.5 mm</quantity>
|
||
. Punctures dense and moderately wide. Anterior margin concave, with anterior angles acute. Posterior margin convex.Lateral margin rounded. Membranous border produced weakly anterior to the anterior marginal bead.
|
||
<emphasis id="D1AFEAA7FFE3FF98F2E6FE15FAA1FE2E" box="[1240,1336,486,506]" italics="true" pageId="3" pageNumber="211">Scutellum</emphasis>
|
||
: Width
|
||
<quantity id="24239B50FFE3FF98F3B1FE14FA42FE2E" box="[1423,1499,487,506]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="211" unit="mm" value="0.3">0.3 mm</quantity>
|
||
, total length 1.0 mm. Punctures moderate and moderately large. Posterior margin rounded. Lacking setae.
|
||
<emphasis id="D1AFEAA7FFE3FF98F2D3FDEEFAADFDE5" box="[1261,1332,541,561]" italics="true" pageId="3" pageNumber="211">Elytron</emphasis>
|
||
: Humeral width
|
||
<quantity id="24239B50FFE3FF98F57FFDC9FC03FD9A" box="[833,922,570,590]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="3" pageNumber="211" unit="mm" value="12.9">12.9 mm</quantity>
|
||
, total length
|
||
<quantity id="24239B50FFE3FF98F21FFDC9FBE2FD9A" box="[1057,1147,570,590]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="3" pageNumber="211" unit="mm" value="17.2">17.2 mm</quantity>
|
||
. Surface smooth (punctures sparse to moderate, and small to moderate, slightly organized longitudinally), lacking setae. Membranous border present at the anterior half. Two dark spots present, one in humerus and other in lateral margin. Apex with punctures moderate and moderately wide. Elytra with dark brown suture and truncate apex (
|
||
<figureCitation id="7BE02A30FFE3FF98F369FD35FA05FD0D" box="[1367,1436,710,729]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 33–40. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 33–36, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 37–40, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 33–36 =0.500 mm, Figs. 37–40 =1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13195501" httpUri="https://zenodo.org/record/13195501/files/figure.png" pageId="3" pageNumber="211">Fig. 35</figureCitation>
|
||
).
|
||
<emphasis id="D1AFEAA7FFE3FF98F38CFD36FCC5FD21" italics="true" pageId="3" pageNumber="211">Venter</emphasis>
|
||
: Punctures dense and moderately wide. Moderately covered by brown setae. Prosternum elongated in the shape of a tubercle but not extending beyond procoxae. Mesoventrite with anterior process between mesocoxae. Mesoventrite medial area with sparse setae.
|
||
<emphasis id="D1AFEAA7FFE3FF98F541FCA3FC31FCB0" box="[895,936,848,868]" italics="true" pageId="3" pageNumber="211">Legs</emphasis>
|
||
: Punctures dense. Protibia with dark outer margin and 3 teeth, apical tooth slightly recurved and larger than the other two; inner apical spur present; dorsal longitudinal ridge present and with a row of 8 brown setae (
|
||
<figureCitation id="7BE02A30FFE3FF98F255FC56FB34FC6C" box="[1131,1197,933,952]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 41–46. Protibia and pygidium in dorsal view. 41–44, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 45–46, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 41–42 = 1.0mm, Figs. 45–76 =2.7 mm." figureDoi="http://doi.org/10.5281/zenodo.13195503" httpUri="https://zenodo.org/record/13195503/files/figure.png" pageId="3" pageNumber="211">Fig. 43</figureCitation>
|
||
); inner protarsal claw slightly recurved and larger than outer claw (
|
||
<figureCitation id="7BE02A30FFE3FF98F280FC32FA98FC00" box="[1214,1281,961,980]" captionStart="Figs" captionStartId="10.[85,120,1787,1801]" captionTargetBox="[197,1362,153,1753]" captionTargetId="figure-13@10.[197,1362,150,1756]" captionTargetPageId="10" captionText="Figs. 47–52. Anterior tarsal claws of male and left metatibia of female in lateral view.47–50, anterior tarsal claws of male;47,Pelidnota beckeri sp.nov.; 48,P.unicolor (Drury); 49, P. nordestina sp. nov.; 50, P. pernambucana sp. nov.; 51 and 52, metatibia of female; 51, P. beckeri sp. nov., a spine-like seta; 52, P. unicolor (Drury), a spine (cuticular projection). Scale bars: Figs.47–50 = 0.500 mm, Figs.51 and 52 = 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195505" httpUri="https://zenodo.org/record/13195505/files/figure.png" pageId="3" pageNumber="211">Fig. 49</figureCitation>
|
||
). Metacoxa with posterior projection beyond base of metatrochanter; anteroexternal angle rounded and slightly sharp (
|
||
<figureCitation id="7BE02A30FFE3FF98F293FC0AFB6AFBD8" box="[1197,1267,1017,1036]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 33–40. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 33–36, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 37–40, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 33–36 =0.500 mm, Figs. 37–40 =1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13195501" httpUri="https://zenodo.org/record/13195501/files/figure.png" pageId="3" pageNumber="211">Fig. 39</figureCitation>
|
||
). Metafemur flattened medially. Mesotibia with setae moderate on inner surface, spine-like setae sparse on outer surface; apical margin dark, inner margin with two larger spurs interleaved with two smaller spine-like setae, outer margin with 10 smaller spine-like setae. Metatibia with apical margin dark, inner margin with two larger spurs interleaved with two smaller spine-like setae, outer margin with one small spine-like setae; external longitudinal carinae with six spinae-like setae and inner longitudinal carinae with six setae present (
|
||
<figureCitation id="7BE02A30FFE3FF98F3AFFB2BFA4BFB3F" box="[1425,1490,1240,1259]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 53–56. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.53–56= 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195507" httpUri="https://zenodo.org/record/13195507/files/figure.png" pageId="3" pageNumber="211">Fig.55</figureCitation>
|
||
). Tarsomeres with small ventral tooth, most evident on meso- and metatarsomeres V.Tarsomeres 2–4, with two inner spine-like setae on all legs. Tarsal claws simple. Outer claws of meso- and metatarsus larger than inner claws.
|
||
<emphasis id="D1AFEAA7FFE3FF98F25EFAB5FB25FA8E" box="[1120,1212,1350,1370]" italics="true" pageId="3" pageNumber="211">Abdomen</emphasis>
|
||
: Abdominal ventrite convex and punctures moderate to dense, and with sparse setae. Ventrite VI ending before posterior margin of pygidium.
|
||
<emphasis id="D1AFEAA7FFE3FF98F31AFA8DFA19FA46" box="[1316,1408,1406,1426]" italics="true" pageId="3" pageNumber="211">Pygidium</emphasis>
|
||
: (
|
||
<figureCitation id="7BE02A30FFE3FF98F3ABFA8CFA41FA46" box="[1429,1496,1407,1426]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 41–46. Protibia and pygidium in dorsal view. 41–44, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 45–46, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 41–42 = 1.0mm, Figs. 45–76 =2.7 mm." figureDoi="http://doi.org/10.5281/zenodo.13195503" httpUri="https://zenodo.org/record/13195503/files/figure.png" pageId="3" pageNumber="211">Fig. 45</figureCitation>
|
||
) slightly convex in lateral view. Surface densely striate and with moderately dense setae. Setae whitish, moderately dense. A gradual concavity present on posterior margin.
|
||
<emphasis id="D1AFEAA7FFE3FF98F2F5FA21FAA8FA32" box="[1227,1329,1490,1510]" italics="true" pageId="3" pageNumber="211">Parameres</emphasis>
|
||
: (
|
||
<figureCitation id="7BE02A30FFE3FF98F378FA20FA41FA32" box="[1350,1496,1491,1510]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 57–68. Male genitalia and paramers margin drawing. 57–64, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 65–68, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 57–64 =2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13195511" httpUri="https://zenodo.org/record/13195511/files/figure.png" pageId="3" pageNumber="211">Figs. 61, 62, 67</figureCitation>
|
||
) symmetrical, fused; distal margin V-shaped, gradually acute laterally; proximal margin gradually semi-quadrate.
|
||
</paragraph>
|
||
<paragraph id="E36436B5FFE3FF98F55EF9D6FA46F9A5" blockId="3.[833,1504,152,1984]" pageId="3" pageNumber="211">
|
||
<emphasis id="D1AFEAA7FFE3FF98F55EF9D6FC72F9ED" box="[864,1003,1573,1593]" italics="true" pageId="3" pageNumber="211">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="53B33CE8FFE3FF98F5C4F9D4FA46F9A5" collectingDate="2015-03-25" collectionCode="MZFS" collectorName="E. M. Menezes" country="Brazil" elevation="510" location="Cachoeira Veu da Noiva" municipality="Bonito" pageId="3" pageNumber="211" specimenCode="MZFS#55726" specimenCount="1" stateProvince="Pernambuco" typeStatus="holotype">
|
||
<typeStatus id="3C608817FFE3FF98F5C4F9D4FBCEF9EE" box="[1018,1111,1575,1594]" pageId="3" pageNumber="211" type="holotype">Holotype</typeStatus>
|
||
male,
|
||
<collectingCountry id="9BCC7625FFE3FF98F29DF9D4FB74F9EE" box="[1187,1261,1575,1594]" name="Brazil" pageId="3" pageNumber="211">BRAZIL</collectingCountry>
|
||
,
|
||
<emphasis id="D1AFEAA7FFE3FF98F2C4F9D6FAEAF9ED" box="[1274,1395,1573,1593]" italics="true" pageId="3" pageNumber="211">
|
||
<collectingRegion id="211FF857FFE3FF98F2C4F9D6FAEAF9ED" box="[1274,1395,1573,1593]" country="Brazil" name="Pernambuco" pageId="3" pageNumber="211">Pernambuco</collectingRegion>
|
||
</emphasis>
|
||
: (“
|
||
<collectingMunicipality id="0300ACCFFFE3FF98F3A8F9D4FA45F9EE" box="[1430,1500,1575,1594]" pageId="3" pageNumber="211">Bonito</collectingMunicipality>
|
||
,
|
||
<location id="E604606EFFE3FF98F57FF9B1FBDBF981" LSID="urn:lsid:plazi:treatment:6B7287A3FFE3FF9FF6ACFB70FD10F8E7:E604606EFFE3FF98F57FF9B1FBDBF981" box="[833,1090,1602,1622]" country="Brazil" municipality="Bonito" name="Cachoeira Veu da Noiva" pageId="3" pageNumber="211" stateProvince="Pernambuco">Cachoeira Véu da Noiva</location>
|
||
”, “
|
||
<quantity id="24239B50FFE3FF98F25CF9B1FB38F982" box="[1122,1185,1602,1622]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="211" unit="m" value="510.0">
|
||
<elevation id="68F6D186FFE3FF98F25CF9B1FB38F982" box="[1122,1185,1602,1622]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="211" unit="m" value="510.0">510 m</elevation>
|
||
</quantity>
|
||
”) (8
|
||
<emphasis id="D1AFEAA7FFE3FF98F2ECF9CDFB43F998" bold="true" box="[1234,1242,1598,1612]" pageId="3" pageNumber="211">
|
||
<superScript id="14AE9BFDFFE3FF98F2ECF9CDFB43F998" attach="none" box="[1234,1242,1598,1612]" fontSize="6" pageId="3" pageNumber="211">◦</superScript>
|
||
</emphasis>
|
||
32
|
||
<emphasis id="D1AFEAA7FFE3FF98F2CBF9CDFB60F998" bold="true" box="[1269,1273,1598,1612]" pageId="3" pageNumber="211">
|
||
<superScript id="14AE9BFDFFE3FF98F2CBF9CDFB60F998" attach="none" box="[1269,1273,1598,1612]" fontSize="6" pageId="3" pageNumber="211">l</superScript>
|
||
</emphasis>
|
||
32.4
|
||
<emphasis id="D1AFEAA7FFE3FF98F318F9CDFAB6F998" bold="true" box="[1318,1327,1598,1612]" pageId="3" pageNumber="211">
|
||
<superScript id="14AE9BFDFFE3FF98F318F9CDFAB6F998" attach="right" box="[1318,1327,1598,1612]" fontSize="6" pageId="3" pageNumber="211">ll</superScript>
|
||
</emphasis>
|
||
S/35
|
||
<emphasis id="D1AFEAA7FFE3FF98F363F9CDFAFCF998" bold="true" box="[1373,1381,1598,1612]" pageId="3" pageNumber="211">
|
||
<superScript id="14AE9BFDFFE3FF98F363F9CDFAFCF998" attach="none" box="[1373,1381,1598,1612]" fontSize="6" pageId="3" pageNumber="211">◦</superScript>
|
||
</emphasis>
|
||
42
|
||
<emphasis id="D1AFEAA7FFE3FF98F3BEF9CDFA1DF998" bold="true" box="[1408,1412,1598,1612]" pageId="3" pageNumber="211">
|
||
<superScript id="14AE9BFDFFE3FF98F3BEF9CDFA1DF998" attach="none" box="[1408,1412,1598,1612]" fontSize="6" pageId="3" pageNumber="211">l</superScript>
|
||
</emphasis>
|
||
53.9
|
||
<emphasis id="D1AFEAA7FFE3FF98F38FF9CDFA23F998" bold="true" box="[1457,1466,1598,1612]" pageId="3" pageNumber="211">
|
||
<superScript id="14AE9BFDFFE3FF98F38FF9CDFA23F998" attach="right" box="[1457,1466,1598,1612]" fontSize="6" pageId="3" pageNumber="211">ll</superScript>
|
||
</emphasis>
|
||
W),
|
||
<date id="97651075FFE3FF98F57FF9ADFC37F9A5" box="[833,942,1630,1649]" pageId="3" pageNumber="211" value="2015-03-25">
|
||
<collectingDate id="8721E99DFFE3FF98F57FF9ADFC37F9A5" box="[833,942,1630,1649]" pageId="3" pageNumber="211" value="2015-03-25">25.III.2015</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="4E2E5363FFE3FF98F588F9ACFBD8F9A6" box="[950,1089,1631,1650]" pageId="3" pageNumber="211">E.M. Menezes</collectorName>
|
||
col. (
|
||
<specimenCode id="B37D9ECEFFE3FF98F24AF9ADFB66F9A5" box="[1140,1279,1630,1649]" collectionCode="MZFS" pageId="3" pageNumber="211">MZFS#55726</specimenCode>
|
||
) (
|
||
<collectionCode id="85CAAE70FFE3FF98F32FF9ADFAD4F9A5" box="[1297,1357,1630,1649]" pageId="3" pageNumber="211">MZFS</collectionCode>
|
||
) (
|
||
<figureCitation id="7BE02A30FFE3FF98F35EF9ADFA4BF9A5" box="[1376,1490,1630,1649]" captionStart="Figs" captionStartId="13.[114,149,1725,1739]" captionTargetBox="[179,720,149,1695]" captionTargetId="figure-13@13.[179,720,149,1695]" captionTargetPageId="13" captionText="Figs.69–77. Labels attachedtothe hotypes.69–71,Pelidnota beckerisp.nov.;72–74, P. nordestina sp. nov.; 75–77, P.pernambucana sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13195513" httpUri="https://zenodo.org/record/13195513/files/figure.png" pageId="3" pageNumber="211">Figs. 72–74</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="E36436B5FFE3FF98F55EF98AFA82F959" blockId="3.[833,1504,152,1984]" box="[864,1307,1657,1678]" pageId="3" pageNumber="211">
|
||
<emphasis id="D1AFEAA7FFE3FF98F55EF98AFC45F959" box="[864,988,1657,1677]" italics="true" pageId="3" pageNumber="211">Distribution.</emphasis>
|
||
Known only from
|
||
<typeStatus id="3C608817FFE3FF98F2A9F988FB5DF95A" box="[1175,1220,1659,1678]" pageId="3" pageNumber="211">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="E36436B5FFE3FF98F55EF966FBD8F911" blockId="3.[833,1504,152,1984]" pageId="3" pageNumber="211">
|
||
<emphasis id="D1AFEAA7FFE3FF98F55EF966FC5EF97D" box="[864,967,1685,1705]" italics="true" pageId="3" pageNumber="211">Etymology</emphasis>
|
||
. The specific epithet
|
||
<taxonomicName id="24DB4D36FFE3FF98F293F966FA8CF97D" authorityName="Ferreira & Almeida & Bravo" authorityYear="2017" box="[1197,1301,1685,1705]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="211" phylum="Arthropoda" rank="species" species="nordestina">
|
||
<emphasis id="D1AFEAA7FFE3FF98F293F966FA8CF97D" box="[1197,1301,1685,1705]" italics="true" pageId="3" pageNumber="211">nordestina</emphasis>
|
||
</taxonomicName>
|
||
refers to its origin from northeastern
|
||
<collectingCountry id="9BCC7625FFE3FF98F23FF941FBA4F911" box="[1025,1085,1714,1733]" name="Brazil" pageId="3" pageNumber="211">Brazil</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E36436B5FFE3FF9FF55EF93EFECAF928" blockId="3.[833,1504,152,1984]" lastBlockId="4.[85,756,1434,1844]" lastPageId="4" lastPageNumber="212" pageId="3" pageNumber="211">
|
||
<emphasis id="D1AFEAA7FFE3FF98F55EF93EFB11F935" box="[864,1160,1741,1761]" italics="true" pageId="3" pageNumber="211">
|
||
Remarks.
|
||
<taxonomicName id="24DB4D36FFE3FF98F581F93EFB11F935" authorityName="Ferreira & Almeida & Bravo" authorityYear="2017" box="[959,1160,1741,1761]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="211" phylum="Arthropoda" rank="species" species="nordestina">Pelidnota nordestina</taxonomicName>
|
||
</emphasis>
|
||
is distinguished as follows (
|
||
<taxonomicName id="24DB4D36FFE3FF98F399F93EFC22F929" authorityName="Ferreira & Almeida & Bravo" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="211" phylum="Arthropoda" rank="species" species="beckeri">
|
||
<emphasis id="D1AFEAA7FFE3FF98F399F93EFC22F929" italics="true" pageId="3" pageNumber="211">Pelidnota beckeri</emphasis>
|
||
</taxonomicName>
|
||
characters in brackets): golden-brown with metallic green reflections, especially on the pronotum and ventral surface, and a black spot is present on both sides of pronotum, with two black spots on the elytron (
|
||
<figureCitation id="7BE02A30FFE3FF98F272F8CEFB20F885" box="[1100,1209,1853,1873]" captionStart="Figs" captionStartId="4.[85,120,1355,1369]" captionTargetBox="[250,1305,180,1309]" captionTargetId="figure-13@4.[249,1309,149,1324]" captionTargetPageId="4" captionText="Figs. 11–16. Pelidnota. 11–13, Pelidnota nordestina sp. nov., male holotype (dorsal, ventral, lateral); 14–16, P. pernambucana sp. nov., male holotype (dorsal, ventral, lateral). Scale bars: Figs. 11–13 =5.5 mm, Figs.14–16= 5.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195493" httpUri="https://zenodo.org/record/13195493/files/figure.png" pageId="3" pageNumber="211">Figs.11, 13</figureCitation>
|
||
) (pronotum and ventral body surface dark brown, with no spots on pronotum but three black spots on elytron (
|
||
<figureCitation id="7BE02A30FFE3FF98F5CDF886FBD3F85C" box="[1011,1098,1909,1928]" captionStart="Figs" captionStartId="2.[85,120,1920,1934]" captionTargetBox="[253,1300,156,1885]" captionTargetId="figure-13@2.[252,1307,149,1889]" captionTargetPageId="2" captionText="Figs. 1–10. Pelidnota. 1–8, Pelidnota beckeri sp. nov.; 1–3, male holotype (dorsal, ventral, lateral); 4–6, female paratype (dorsal, ventral, lateral); 7–8, clypeus in dorsal view (male, female); 9–10, P.unicolor (Drury, 1778) (dorsal, lateral). Scale bars: Figs. 1–6 =9.0 mm, Figs.7–8 =2.5 mm, Figs. 9–10= 8.2mm." figureDoi="http://doi.org/10.5281/zenodo.13195491" httpUri="https://zenodo.org/record/13195491/files/figure.png" pageId="3" pageNumber="211">Figs. 1, 3</figureCitation>
|
||
)); labrum triangular (
|
||
<figureCitation id="7BE02A30FFE3FF98F317F886FAF5F85C" box="[1321,1388,1909,1928]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 17–24. Mouthparts. 17–20, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 21–24, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.17–24 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195495" httpUri="https://zenodo.org/record/13195495/files/figure.png" pageId="3" pageNumber="211">Fig. 19</figureCitation>
|
||
) (transversal (
|
||
<figureCitation id="7BE02A30FFE3FF98F552F862FC36F870" box="[876,943,1937,1956]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 17–24. Mouthparts. 17–20, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 21–24, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.17–24 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195495" httpUri="https://zenodo.org/record/13195495/files/figure.png" pageId="3" pageNumber="211">Fig. 17</figureCitation>
|
||
)); inner teeth of mandible more acuminate and larger (
|
||
<figureCitation id="7BE02A30FFE3FF98F577F85EFC15F814" box="[841,908,1965,1984]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 17–24. Mouthparts. 17–20, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 21–24, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.17–24 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195495" httpUri="https://zenodo.org/record/13195495/files/figure.png" pageId="3" pageNumber="211">Fig. 23</figureCitation>
|
||
) (small (
|
||
<figureCitation id="7BE02A30FFE3FF98F5D9F85EFBB3F814" box="[999,1066,1965,1984]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 17–24. Mouthparts. 17–20, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 21–24, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.17–24 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195495" httpUri="https://zenodo.org/record/13195495/files/figure.png" pageId="3" pageNumber="211">Fig. 21</figureCitation>
|
||
)); first and second teeth of maxillary galea sawed, and IV palpomere more elongated (
|
||
<figureCitation id="7BE02A30FFE4FF9FF424FA69FDC6FA79" box="[538,607,1434,1453]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 25–28. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 27–28, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 25–28 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195497" httpUri="https://zenodo.org/record/13195497/files/figure.png" pageId="4" pageNumber="212">Fig. 27</figureCitation>
|
||
) (only second teeth sawed, and IV palpomere less longer (
|
||
<figureCitation id="7BE02A30FFE4FF9FF419FA45FDF4FA1D" box="[551,621,1462,1481]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 25–28. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 27–28, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 25–28 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195497" httpUri="https://zenodo.org/record/13195497/files/figure.png" pageId="4" pageNumber="212">Fig. 25</figureCitation>
|
||
)); excavated area on the surface of pre-mentum larger, and lateral margin of post-mentum less rounded and with elongated dense setae (
|
||
<figureCitation id="7BE02A30FFE4FF9FF492FA1EFD75F9D5" box="[684,748,1517,1537]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 29–32. Mouthparts. 29–32, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 29–32 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195499" httpUri="https://zenodo.org/record/13195499/files/figure.png" pageId="4" pageNumber="212">Fig.31</figureCitation>
|
||
) (excavated area small, lateral margin more rounded, and with moderate setae (
|
||
<figureCitation id="7BE02A30FFE4FF9FF6E8F9D6FE82F9EC" box="[214,283,1573,1592]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 29–32. Mouthparts. 29–32, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 29–32 = 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195499" httpUri="https://zenodo.org/record/13195499/files/figure.png" pageId="4" pageNumber="212">Fig. 29</figureCitation>
|
||
)); anteroexternal angle of metacoxa rounded and slightly sharp (
|
||
<figureCitation id="7BE02A30FFE4FF9FF723F9B2FEFBF980" box="[285,354,1601,1620]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 33–40. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 33–36, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 37–40, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 33–36 =0.500 mm, Figs. 37–40 =1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13195501" httpUri="https://zenodo.org/record/13195501/files/figure.png" pageId="4" pageNumber="212">Fig. 39</figureCitation>
|
||
) (acute (
|
||
<figureCitation id="7BE02A30FFE4FF9FF781F9B2FD9AF980" box="[447,515,1601,1620]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 33–40. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 33–36, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 37–40, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 33–36 =0.500 mm, Figs. 37–40 =1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13195501" httpUri="https://zenodo.org/record/13195501/files/figure.png" pageId="4" pageNumber="212">Fig. 37</figureCitation>
|
||
)); protarsal tubercle of male located more on the claw apex (
|
||
<figureCitation id="7BE02A30FFE4FF9FF7DCF9AEFDBEF9A4" box="[482,551,1629,1648]" captionStart="Figs" captionStartId="10.[85,120,1787,1801]" captionTargetBox="[197,1362,153,1753]" captionTargetId="figure-13@10.[197,1362,150,1756]" captionTargetPageId="10" captionText="Figs. 47–52. Anterior tarsal claws of male and left metatibia of female in lateral view.47–50, anterior tarsal claws of male;47,Pelidnota beckeri sp.nov.; 48,P.unicolor (Drury); 49, P. nordestina sp. nov.; 50, P. pernambucana sp. nov.; 51 and 52, metatibia of female; 51, P. beckeri sp. nov., a spine-like seta; 52, P. unicolor (Drury), a spine (cuticular projection). Scale bars: Figs.47–50 = 0.500 mm, Figs.51 and 52 = 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195505" httpUri="https://zenodo.org/record/13195505/files/figure.png" pageId="4" pageNumber="212">Fig. 49</figureCitation>
|
||
) (located not so on the claw apex (
|
||
<figureCitation id="7BE02A30FFE4FF9FF6D1F98AFEABF958" box="[239,306,1657,1676]" captionStart="Figs" captionStartId="10.[85,120,1787,1801]" captionTargetBox="[197,1362,153,1753]" captionTargetId="figure-13@10.[197,1362,150,1756]" captionTargetPageId="10" captionText="Figs. 47–52. Anterior tarsal claws of male and left metatibia of female in lateral view.47–50, anterior tarsal claws of male;47,Pelidnota beckeri sp.nov.; 48,P.unicolor (Drury); 49, P. nordestina sp. nov.; 50, P. pernambucana sp. nov.; 51 and 52, metatibia of female; 51, P. beckeri sp. nov., a spine-like seta; 52, P. unicolor (Drury), a spine (cuticular projection). Scale bars: Figs.47–50 = 0.500 mm, Figs.51 and 52 = 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195505" httpUri="https://zenodo.org/record/13195505/files/figure.png" pageId="4" pageNumber="212">Fig. 47</figureCitation>
|
||
)); external surface of metatibia less punctuated, and inner longitudinal carinae with six setae (
|
||
<figureCitation id="7BE02A30FFE4FF9FF45BF966FD31F97C" box="[613,680,1685,1704]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 53–56. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.53–56= 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195507" httpUri="https://zenodo.org/record/13195507/files/figure.png" pageId="4" pageNumber="212">Fig. 55</figureCitation>
|
||
) (more punctuated and with more than six setae (
|
||
<figureCitation id="7BE02A30FFE4FF9FF434F942FDD7F910" box="[522,590,1713,1732]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 53–56. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.53–56= 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.13195507" httpUri="https://zenodo.org/record/13195507/files/figure.png" pageId="4" pageNumber="212">Fig. 53</figureCitation>
|
||
)); shapes of the parameres distal margin, any less acute laterally (
|
||
<figureCitation id="7BE02A30FFE4FF9FF47CF93EFD36F934" box="[578,687,1741,1760]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 57–68. Male genitalia and paramers margin drawing. 57–64, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 65–68, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 57–64 =2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13195511" httpUri="https://zenodo.org/record/13195511/files/figure.png" pageId="4" pageNumber="212">Figs. 61, 67</figureCitation>
|
||
) (apex more acute (
|
||
<figureCitation id="7BE02A30FFE4FF9FF6EBF91BFEDFF92F" box="[213,326,1768,1787]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 57–68. Male genitalia and paramers margin drawing. 57–64, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 65–68, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 57–64 =2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13195511" httpUri="https://zenodo.org/record/13195511/files/figure.png" pageId="4" pageNumber="212">Figs. 57, 65</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B7A4663DFFE4FF9FF66BFAB8FD94FAA4" ID-DOI="http://doi.org/10.5281/zenodo.13195493" ID-Zenodo-Dep="13195493" httpUri="https://zenodo.org/record/13195493/files/figure.png" pageId="4" pageNumber="212" startId="4.[85,120,1355,1369]" targetBox="[250,1305,180,1309]" targetPageId="4" targetType="figure">
|
||
<paragraph id="E36436B5FFE4FF9FF66BFAB8FD94FAA4" blockId="4.[85,1476,1354,1392]" pageId="4" pageNumber="212">
|
||
<emphasis id="D1AFEAA7FFE4FF9FF66BFAB8FF21FA8D" bold="true" box="[85,184,1355,1369]" pageId="4" pageNumber="212">Figs. 11–16.</emphasis>
|
||
<taxonomicName id="24DB4D36FFE4FF9FF6FFFAB9FE93FA8D" authorityName="MacLeay" authorityYear="1819" box="[193,266,1354,1369]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="212" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D1AFEAA7FFE4FF9FF6FFFAB9FE93FA8D" box="[193,266,1354,1369]" italics="true" pageId="4" pageNumber="212">Pelidnota</emphasis>
|
||
</taxonomicName>
|
||
. 11–13,
|
||
<taxonomicName id="24DB4D36FFE4FF9FF770FAB9FE77FA8D" authorityName="Ferreira & Almeida & Bravo" authorityYear="2017" box="[334,494,1354,1369]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="212" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
|
||
<emphasis id="D1AFEAA7FFE4FF9FF770FAB9FE77FA8D" box="[334,494,1354,1369]" italics="true" pageId="4" pageNumber="212">Pelidnota nordestina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CA9C57DCFFE4FF9FF7CDFAB8FDAAFA8D" box="[499,563,1355,1369]" pageId="4" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male holotype (dorsal, ventral, lateral); 14–16,
|
||
<taxonomicName id="24DB4D36FFE4FF9FF588FAB9FBA4FA8D" authorityName="Ferreira & Almeida & Bravo" authorityYear="2017" box="[950,1085,1354,1369]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="212" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
|
||
<emphasis id="D1AFEAA7FFE4FF9FF588FAB9FBA4FA8D" box="[950,1085,1354,1369]" italics="true" pageId="4" pageNumber="212">P. pernambucana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CA9C57DCFFE4FF9FF27FFAB8FB18FA8D" box="[1089,1153,1355,1369]" pageId="4" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male holotype (dorsal, ventral, lateral). Scale bars: Figs. 11–13 = 5.5 mm, Figs. 14–16 = 5.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E36436B5FFE4FF9FF64BF8F0FD10F8E7" blockId="4.[85,756,1434,1844]" pageId="4" pageNumber="212">
|
||
<emphasis id="D1AFEAA7FFE4FF9FF64BF8F0FF7BF8C3" box="[117,226,1795,1815]" italics="true" pageId="4" pageNumber="212">Comments.</emphasis>
|
||
The specimen was collected in a tropical forest in an enclave in the Caatinga ecoregion (
|
||
<bibRefCitation id="874A4B44FFE4FF9FF789F8D3FDE2F8E7" author="Instituto LIFE" box="[439,635,1824,1844]" pageId="4" pageNumber="212" pagination="35 - 52" refId="ref10592" refString="Instituto LIFE, 2015. Bahia, in: Ecorregioes do Brasil - Prioridades Terrestres e Marinhas. Serie cadernos tecnicos, Volume 3, Brasil., pp. 35 - 52." type="book chapter" year="2015">Instituto LIFE, 2015</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |