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<document id="E7B22A64F6DA3C71F9EB15B989033E57" ID="10.11646/zootaxa.4169.1.5" ID-DOI="10.11646/zootaxa.4169.1.5" ID-GBIF-Dataset="e35dbd92-b156-4ae3-9f90-9996c91ee626" ID-ISSN="1175-5326" ID-Zenodo-Dep="265168" ID-ZooBank="62419F0E-128A-4E60-932C-DD746BA57906" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474268893273" checkinUser="plazi" docAuthor="Thoni, R. J., Gurung, D. B. &amp; Mayden, R. L." docDate="2016" docId="03F187D8FFBDFF83FF1BFA1FFB84F84C" docLanguage="en" docName="zootaxa.4169.1.5.pdf" docOrigin="Zootaxa 4169 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Garra birostris Nebeshwar &amp; Vishwanath 2014" docType="treatment" docVersion="7" lastPageNumber="122" masterDocId="FFC8FFA0FFBAFF84FF8CFFB8FFA3FF9D" masterDocTitle="A review of the genus Garra Hamilton 1822 of Bhutan, including the descriptions of two new species and three additional records (Cypriniformes: Cyprinidae)" masterLastPageNumber="132" masterPageNumber="115" pageNumber="122" updateTime="1698386207380" updateUser="plazi">
<mods:mods id="B1D853BA9186C51A3DB9B80A16602358" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5DC90DDB9660A4BC7FDCBBC32F0216BC">A review of the genus Garra Hamilton 1822 of Bhutan, including the descriptions of two new species and three additional records (Cypriniformes: Cyprinidae)</mods:title>
</mods:titleInfo>
<mods:name id="A7159070D98FBBD1BA0EB6B8B10C29C3" type="personal">
<mods:role id="DD1E7773785CCB08F62A5C18CC0763F0">
<mods:roleTerm id="63E35FC4D70BB09A9BD7C397FB05F948">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="28EE24F5897279CB5272E6A571747E53">Thoni, R. J.</mods:namePart>
</mods:name>
<mods:name id="77715CD472A5E19C64A400187D6488F2" type="personal">
<mods:role id="F11900EDE2AFD7D669243C0B7CC4499E">
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<mods:namePart id="60CCF1DB393576E2A6604FCCE5C0676E">Gurung, D. B.</mods:namePart>
</mods:name>
<mods:name id="8DC5ED303F750D2E7BCE5928C15B3C7B" type="personal">
<mods:role id="501FE98D387348872D5EA5F523C7423A">
<mods:roleTerm id="40EF3DEF88CF0EF79855F608E84F2618">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4FBEC707168C1369DF28C7883C90F6A3">Mayden, R. L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2E3F255646A219F324EA3A41411E063C">text</mods:typeOfResource>
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<mods:title id="4220E632F760E060A0152B9ABFA2AE6F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A5F5DF7AC95D57EDC3F12E17ED9E47F6">
<mods:date id="8B5FF008668ACD1D8D94635DEA1E1767">2016</mods:date>
<mods:detail id="57BA0522FDDD3F86C84911405F61FB5C" type="volume">
<mods:number id="EFCA05481DAE83A7135909447F6E9C92">4169</mods:number>
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<mods:detail id="A99F5D6861B9ED324BD0E40E5D7FBAF3" type="issue">
<mods:number id="68E8E134ADD97BCFFAFAE424B3106F16">1</mods:number>
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<mods:classification id="910FC40F06A26C83B4A58F96FAD8FCB3">journal article</mods:classification>
<mods:identifier id="5134724A2097EA800B26D24F2F09ED48" type="DOI">10.11646/zootaxa.4169.1.5</mods:identifier>
<mods:identifier id="51A1A48324402F7A99C0375A35CD0A9D" type="GBIF-Dataset">e35dbd92-b156-4ae3-9f90-9996c91ee626</mods:identifier>
<mods:identifier id="29385FE8A89C0C6B81A6F03E9ED2EC4F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="99A73632502AD691FDE417560FE253CF" type="Zenodo-Dep">265168</mods:identifier>
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<treatment id="03F187D8FFBDFF83FF1BFA1FFB84F84C" ID-DOI="http://doi.org/10.5281/zenodo.6087319" ID-GBIF-Taxon="124551992" ID-Zenodo-Dep="6087319" LSID="urn:lsid:plazi:treatment:03F187D8FFBDFF83FF1BFA1FFB84F84C" httpUri="http://treatment.plazi.org/id/03F187D8FFBDFF83FF1BFA1FFB84F84C" lastPageNumber="122" pageId="7" pageNumber="122">
<subSubSection id="C3426545FFBDFF83FF1BFA1FFF48FA7E" pageId="7" pageNumber="122" type="nomenclature">
<paragraph id="8BE736CEFFBDFF83FF1BFA1FFD40FA5F" blockId="7.[151,739,1447,1507]" box="[151,739,1447,1474]" pageId="7" pageNumber="122">
<heading id="D0AF81A2FFBDFF83FF1BFA1FFD40FA5F" bold="true" box="[151,739,1447,1474]" fontSize="11" level="1" pageId="7" pageNumber="122" reason="1">
<taxonomicName id="4C584D4DFFBDFF83FF1BFA1FFD40FA5F" ID-CoL="3FBH4" authority="Nebeshwar &amp; Vishwanath, 2014" authorityName="Nebeshwar &amp; Vishwanath" authorityYear="2014" box="[151,739,1447,1474]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="birostris">
<emphasis id="B92CEADCFFBDFF83FF1BFA1FFD40FA5F" bold="true" box="[151,739,1447,1474]" pageId="7" pageNumber="122">
<emphasis id="B92CEADCFFBDFF83FF1BFA1FFEE9FA5C" bold="true" box="[151,330,1447,1473]" italics="true" pageId="7" pageNumber="122">Garra birostris</emphasis>
Nebeshwar &amp; Vishwanath, 2014
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE736CEFFBDFF83FF1BFA72FF48FA7E" blockId="7.[151,739,1447,1507]" box="[151,235,1482,1507]" pageId="7" pageNumber="122">
(
<figureCitation id="13632A4BFFBDFF83FF13FA72FF40FA7E" box="[159,227,1482,1507]" captionStart="FIGURE 6" captionStartId="8.[151,250,1323,1345]" captionTargetBox="[161,1436,195,1287]" captionTargetId="figure@8.[151,1436,195,1301]" captionTargetPageId="8" captionText="FIGURE 6. Garra birostris. CNR 13037, 80.7 mm SL; Bhutan: Samdrup Jongkhar Dzongkhag: Tikrikhola, D. B. Gurung &amp; R. J. Thoni, October 20, 2013." httpUri="https://zenodo.org/record/265174/files/figure.png" pageId="7" pageNumber="122">Fig. 6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3426545FFBDFF83FF1BF9ABFD6EF9DA" pageId="7" pageNumber="122" type="reference_group">
<paragraph id="8BE736CEFFBDFF83FF1BF9ABFD6EF9DA" blockId="7.[151,1436,1554,1607]" pageId="7" pageNumber="122">
<materialsCitation id="3B303C93FFBDFF83FF1BF9ABFD6EF9DA" ID-GBIF-Occurrence="1313779502" collectionCode="MUMF" country="India" location="Dikrong River" municipality="Papum Pare district" pageId="7" pageNumber="122" specimenCode="MUMF 4302" specimenCount="2" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<taxonomicName id="4C584D4DFFBDFF83FF1BF9ABFD16F9B5" ID-CoL="3FBH4" authority="Nebeshwar &amp; Vishwanath 2014: 104" authorityName="Nebeshwar &amp; Vishwanath" authorityPageNumber="104" authorityYear="2014" box="[151,693,1554,1576]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="birostris">
<emphasis id="B92CEADCFFBDFF83FF1BF9ABFE91F9B5" box="[151,306,1554,1576]" italics="true" pageId="7" pageNumber="122">Garra birostris</emphasis>
Nebeshwar &amp; Vishwanath 2014: 104
</taxonomicName>
, fig. 5.
<typeStatus id="54E3886CFFBDFF83FC89F9ABFC94F9B5" box="[773,823,1555,1576]" pageId="7" pageNumber="122">Type</typeStatus>
locality:
<location id="8E876015FFBDFF83FC15F9AAFB8FF9B5" LSID="urn:lsid:plazi:treatment:03F187D8FFBDFF83FF1BFA1FFB84F84C:8E876015FFBDFF83FC15F9AAFB8FF9B5" box="[921,1068,1554,1576]" country="India" municipality="Papum Pare district" name="Dikrong River" pageId="7" pageNumber="122" stateProvince="Arunachal Pradesh">Dikrong River</location>
and Doimukh,
<collectingMunicipality id="6B83ACB4FFBDFF83FB41F9ABFA3BF9B5" box="[1229,1432,1554,1576]" pageId="7" pageNumber="122">Papum Pare district</collectingMunicipality>
,
<collectingRegion id="499CF82CFFBDFF83FF4BF989FE2AF9DA" box="[199,393,1585,1607]" country="India" name="Arunachal Pradesh" pageId="7" pageNumber="122">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="F34F765EFFBDFF83FE1FF989FE69F9DA" box="[403,458,1585,1607]" name="India" pageId="7" pageNumber="122">India</collectingCountry>
.
<typeStatus id="54E3886CFFBDFF83FE58F989FD95F9DA" box="[468,566,1585,1607]" pageId="7" pageNumber="122" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBFE9EB5FFBDFF83FDCCF98AFD6AF9DA" box="[576,713,1585,1607]" collectionCode="MUMF" country="India" httpUri="http://grbio.org/cool/x8gv-47he" name="Department of Life Sciences" pageId="7" pageNumber="122">
<collectionCode id="ED49AE0BFFBDFF83FDCCF98AFD2EF9DA" box="[576,653,1586,1607]" country="India" httpUri="http://grbio.org/cool/x8gv-47he" name="Department of Life Sciences" pageId="7" pageNumber="122">MUMF</collectionCode>
4302
</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3426545FFBDFF83FF1BF9CCFDA1F92D" pageId="7" pageNumber="122" type="materials_examined">
<paragraph id="8BE736CEFFBDFF83FF1BF9CCFDA1F92D" blockId="7.[151,1437,1652,2001]" pageId="7" pageNumber="122">
<materialsCitation id="3B303C93FFBDFF83FF1BF9CCFDA1F92D" ID-GBIF-Occurrence="1313779503" collectingDate="2013-10-20" collectionCode="CNR" collectorName="Tikrikhola, D. B. &amp; R. J. Thoni" country="Bhutan" location="Samdrup Jongkhar" pageId="7" pageNumber="122" specimenCount="1" stateProvince="Samdrup Jongkhar">
<emphasis id="B92CEADCFFBDFF83FF1BF9CCFE22F910" bold="true" box="[151,385,1652,1677]" pageId="7" pageNumber="122">Material examined.</emphasis>
<specimenCount id="9D5EFD47FFBDFF83FE04F9CCFDADF910" box="[392,526,1652,1677]" pageId="7" pageNumber="122" type="generic">1 Specimen</specimenCount>
,
<quantity id="4CA09B2BFFBDFF83FD96F9CCFDDCF911" box="[538,639,1652,1676]" metricMagnitude="-2" metricUnit="m" metricValue="8.87" pageId="7" pageNumber="122" unit="mm" value="88.7">88.7 mm</quantity>
SL,
<collectionCode id="ED49AE0BFFBDFF83FD39F9CCFD51F911" box="[693,754,1652,1676]" country="Ukraine" httpUri="http://grbio.org/cool/5zp2-7vs1" name="Crimean Natural Reserve" pageId="7" pageNumber="122">CNR</collectionCode>
13037;
<collectingCountry id="F34F765EFFBDFF83FCC8F9CCFC3AF910" box="[836,921,1652,1677]" name="Bhutan" pageId="7" pageNumber="122">Bhutan</collectingCountry>
:
<collectingRegion id="499CF82CFFBDFF83FC28F9CCFBD8F910" box="[932,1147,1652,1677]" country="Bhutan" name="Samdrup Jongkhar" pageId="7" pageNumber="122">Samdrup Jongkhar</collectingRegion>
:
<collectorName id="26AD5318FFBDFF83FB0AF9CCFA9CF911" box="[1158,1343,1652,1677]" pageId="7" pageNumber="122">Tikrikhola, D.B.</collectorName>
Gurung &amp;
<collectorName id="26AD5318FFBDFF83FF38F920FE8FF92D" box="[180,300,1687,1712]" pageId="7" pageNumber="122">R.J. Thoni</collectorName>
,
<collectingDate id="EFA2E9E6FFBDFF83FEBBF92FFE5DF92D" box="[311,510,1687,1712]" pageId="7" pageNumber="122" value="2013-10-20">
<date id="FFE6100EFFBDFF83FEBBF92FFE5DF92D" box="[311,510,1687,1712]" pageId="7" pageNumber="122" value="2013-10-20">October 20, 2013</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3426545FFBDFF83FF4BF904FB84F84C" pageId="7" pageNumber="122" type="discussion">
<paragraph id="8BE736CEFFBDFF83FF4BF904FB84F84C" blockId="7.[151,1437,1652,2001]" pageId="7" pageNumber="122">
<emphasis id="B92CEADCFFBDFF83FF4BF904FE98F948" bold="true" box="[199,315,1724,1749]" pageId="7" pageNumber="122">Remarks.</emphasis>
This species was encountered in a stream in southeastern
<collectingCountry id="F34F765EFFBDFF83FC4BF904FBB8F948" box="[967,1051,1724,1749]" name="Bhutan" pageId="7" pageNumber="122">Bhutan</collectingCountry>
. It was identified by its prominent bi-lobed proboscis, tubercle pattern on the rostral lobe and depressed rostral surface, caudal-fin pigmentation, and overall body shape. There is a slight difference in snout shape when viewed ventrally compared to Nebeshwar &amp; Vishwanath’s specimen, however, based on only one specimen, we cannot definitively separate it from
<emphasis id="B92CEADCFFBDFF83FA95F890FA3FF8DD" box="[1305,1436,1831,1856]" italics="true" pageId="7" pageNumber="122">
<taxonomicName id="4C584D4DFFBDFF83FA95F890FA3BF8DD" box="[1305,1432,1831,1856]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="birostris">G. birostris</taxonomicName>
.
</emphasis>
It was collected in close proximity to localities in
<collectingRegion id="499CF82CFFBDFF83FD49F8F4FC39F8F8" box="[709,922,1868,1893]" country="India" name="Arunachal Pradesh" pageId="7" pageNumber="122">Arunachal Pradesh</collectingRegion>
where Nebeshwar &amp; Vishwanath collected
<emphasis id="B92CEADCFFBDFF83FA08F8F5FF5FF815" italics="true" pageId="7" pageNumber="122">
<taxonomicName id="4C584D4DFFBDFF83FA08F8F5FF5BF815" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="122" phylum="Chordata" rank="species" species="birostris">G. birostris</taxonomicName>
.
</emphasis>
It is possible that this species, and others described by Nebeshwar &amp; Vishwanath (2014), have already been described by early ichthyologists who’s descriptions remain vague, and without
<typeStatus id="54E3886CFFBDFF83FBD9F82CFB65F830" box="[1109,1222,1940,1965]" pageId="7" pageNumber="122" type="holotype">holotypes</typeStatus>
. We therefore gave priority to those species names available with ample and precise diagnostic traits.
</paragraph>
</subSubSection>
</treatment>
</document>
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