166 lines
19 KiB
XML
166 lines
19 KiB
XML
<document id="8C809AF707C89ABE7CA822E9F8EF8F08" ID-ISSN="0024-4082" ID-ZooBank="54DD9ED6-0B56-4CA2-837D-A7E05307D81F" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632744255385" checkinUser="plazi" docAuthor="Arifin, Umilaela, Chan, Kin Onn, Smart, Utpal, Hertwig, Stefan T., Smith, Eric N., Iskandar, Djoko T. & Haas, Alexander" docDate="2021" docId="03F187ACCF68FFC8FEDAE543FC4BFEA4" docLanguage="en" docName="zlaa158.pdf" docOrigin="Zoological Journal of the Linnean Society 193" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Huia Yang 1991" docType="treatment" docVersion="3" lastPageNumber="687" masterDocId="FFC8FFD4CF66FFC6FFEFE74CFF92FFF2" masterDocTitle="Revisiting the phylogenetic predicament of the genus Huia (Amphibia: Ranidae) using molecular data and tadpole morphology" masterLastPageNumber="699" masterPageNumber="673" pageNumber="687" updateTime="1699415239833" updateUser="plazi">
|
||
<mods:mods id="D392474CF49C3AA2DF6D7B0E0386C0F0" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="B9879BE04E8B40548E4428C20BB2BA53">
|
||
<mods:title id="4B293614811035BC2B78BDF6A36E21FE">Revisiting the phylogenetic predicament of the genus Huia (Amphibia: Ranidae) using molecular data and tadpole morphology</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="2A84A002D10E182B776896F15BC938D1" type="personal">
|
||
<mods:role id="D5E735FCC0FDF055BECF90B115EDADBC">
|
||
<mods:roleTerm id="CCED0716DC369488F512234FF06AE11E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7618EE04E4B7432EC32D596BEF32F9FE">Arifin, Umilaela</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="B5ABB100B6EBD69A862782D319053700" type="personal">
|
||
<mods:role id="BCDF7077574F8EB9F2314DE6A9339406">
|
||
<mods:roleTerm id="B6A50DAC85FDD027D1CB286820E634DD">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9D747B98E479C3A8229365BE77730475">Chan, Kin Onn</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="B3F61DBC324A7433A3AAADDB65529F69" type="personal">
|
||
<mods:role id="D08471996537E4E6C3BBA540C9C1EFF3">
|
||
<mods:roleTerm id="E9C041106A319B9CBF2093E17730D880">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="18C284E82C49480DA8B30DD746FFE982">Smart, Utpal</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="93A9F01D173B82A00A3C4626312E663A" type="personal">
|
||
<mods:role id="F989D5D7AB5D1952C38CC10A48F3CB8E">
|
||
<mods:roleTerm id="A6F5BD1AE1130468D1BB907EEB76A332">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B0B37E900A06E7D7B21A23112211ACAB">Hertwig, Stefan T.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="904C3A0296C0F97626CEF21E35D84CFF" type="personal">
|
||
<mods:role id="848CCA5EA7783F833D1B09FFB64FC50C">
|
||
<mods:roleTerm id="261FA06597205E0F97DD7BEDC184E5E4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B6E61551C00695402ABE46793000D9A8">Smith, Eric N.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="40AF581891A6259CC51926EF63411EB2" type="personal">
|
||
<mods:role id="AE5C1129AF640AB60595130AD9AD021A">
|
||
<mods:roleTerm id="A8C0B9AA3864DFFA9200CFBC572FBDDC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="26AB6CA688AECE72B8E0370465F7DE2D">Iskandar, Djoko T.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="4DE8A9BABC5095BCF4C0440B2F03B8C4" type="personal">
|
||
<mods:role id="79391A1CFD321D8A2345E9FA2E749025">
|
||
<mods:roleTerm id="F585EA192CCF8068A47B0F5713CE7437">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4EC1B263CB002BAB00EBC4CA32A110B2">Haas, Alexander</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="D3593DC5138C83A52775F397444B6123">text</mods:typeOfResource>
|
||
<mods:relatedItem id="4B16B11D2C3B76A6CDF34D3FDFC4B1A3" type="host">
|
||
<mods:titleInfo id="BCC5BCB4E5858C5702213B03F8A66A79">
|
||
<mods:title id="841D874A00304A69825AE942EAE1E440">Zoological Journal of the Linnean Society</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="E7AFF47622683ABB08164FF214D2E94B">
|
||
<mods:date id="D4DAC131A7307A6B484346A4C5AB9861">2021</mods:date>
|
||
<mods:detail id="6605B2A83B2E7C6B0FE647D2525D1415" type="volume">
|
||
<mods:number id="5DAABCA6CA3295948D998478777DC938">193</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="CEE7964098FEFB5D8AD54E25CACB1E47" unit="page">
|
||
<mods:start id="C2A2AC93872F1994121EAB06E3560CCA">673</mods:start>
|
||
<mods:end id="C678C4A85D4C86D8C17E1CA30B707CF5">699</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="F90AE6E8BB516AABD2E23F4BC3D73B20">journal article</mods:classification>
|
||
<mods:identifier id="554E5FADB44ADB53B652ED631BD605C5" type="ISSN">0024-4082</mods:identifier>
|
||
<mods:identifier id="B506B9CF63D494EAF40FFF14E646822E" type="ZooBank">54DD9ED6-0B56-4CA2-837D-A7E05307D81F</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03F187ACCF68FFC8FEDAE543FC4BFEA4" ID-DOI="http://doi.org/10.5281/zenodo.5530639" ID-Zenodo-Dep="5530639" LSID="urn:lsid:plazi:treatment:03F187ACCF68FFC8FEDAE543FC4BFEA4" httpUri="http://treatment.plazi.org/id/03F187ACCF68FFC8FEDAE543FC4BFEA4" lastPageNumber="687" pageId="14" pageNumber="687">
|
||
<subSubSection id="C3426531CF68FFC8FEDAE543FDC6FDDA" box="[309,596,527,552]" pageId="14" pageNumber="687" type="nomenclature">
|
||
<paragraph id="8BE736BACF68FFC8FEDAE543FDC6FDDA" blockId="14.[309,596,527,552]" box="[309,596,527,552]" pageId="14" pageNumber="687">
|
||
<heading id="D0AF81D6CF68FFC8FEDAE543FDC6FDDA" box="[309,596,527,552]" centered="true" fontSize="9" level="2" pageId="14" pageNumber="687" reason="2">
|
||
GENUS
|
||
<taxonomicName id="4C584D39CF68FFC8FE63E55CFDC6FDDA" ID-CoL="4Z6D" authority="YANG, 1991" authorityName="Yang" authorityYear="1991" box="[396,596,527,552]" class="Amphibia" family="Ranidae" genus="Huia" kingdom="Animalia" order="Anura" pageId="14" pageNumber="687" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92CEAA8CF68FFC8FE63E55CFE5AFDD5" box="[396,456,528,551]" italics="true" pageId="14" pageNumber="687">HUIA</emphasis>
|
||
<bibRefCitation id="EFC94B4BCF68FFC8FE21E543FDC6FDDA" author="Yang DT" box="[462,596,527,552]" pageId="14" pageNumber="687" pagination="1 - 42" refId="ref15214" refString="Yang DT. 1991. Phylogenetic systematics of the Amolops group of ranid frogs of southeastern Asia and the Greater Sunda Islands. Fieldiana Zoology 63: 1 - 42." type="journal article" year="1991">YANG, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3426531CF68FFC8FE7EE577FC4BFEA4" pageId="14" pageNumber="687" type="description">
|
||
<paragraph id="8BE736BACF68FFC8FE7EE577FE6AFDA6" blockId="14.[401,504,571,596]" box="[401,504,571,596]" pageId="14" pageNumber="687">
|
||
(
|
||
<figureCitation id="13632A3FCF68FFC8FE75E577FE62FDA6" box="[410,496,571,596]" captionStart="Figure 6" captionStartId="15.[163,241,1574,1596]" captionTargetBox="[165,1441,197,1532]" captionTargetId="figure-26@15.[163,1443,195,1534]" captionTargetPageId="15" captionText="Figure 6. A, Huia cavitympanum male, Taman Nasional Bukit Baka Bukit Raya, Provinsi Kalimantan Barat, Indonesia. B, Wijayarana (cladeA); UA20140663, male, PadangAro, Taman Nasional Kerinci-Seblat, Provinsi Sumatera Barat, Indonesia. C, Wijayarana (clade B); UA20150202, male, Wiyono Waterfall, Provinsi Lampung, Indonesia. D, Wijayarana (clade C); UA20150464–65, male and female in amplexus, Batang Karangan, Provinsi Sumatera Barat, Indonesia. E, Wijayarana (clade E); UA20150032, male, Banyumas, Provinsi Jawa Tengah, Indonesia. F, Wijayarana (clade F); UA20150049; Palutungan, Provinsi Jawa Barat, Indonesia. Size not to scale. Photographs by U.A." pageId="14" pageNumber="687">FIG. 6A</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BE736BACF68FFC8FF7EE528FE89FD25" blockId="14.[145,762,612,727]" pageId="14" pageNumber="687">
|
||
<emphasis id="B92CEAA8CF68FFC8FF7EE528FEA0FD89" box="[145,306,612,635]" italics="true" pageId="14" pageNumber="687">Type species:</emphasis>
|
||
<taxonomicName id="4C584D39CF68FFC8FED5E52AFD60FD88" authority="Boulenger, 1893" authorityName="Boulenger" authorityYear="1893" box="[314,754,613,635]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="14" pageNumber="687" phylum="Chordata" rank="species" species="cavitympanum">
|
||
<emphasis id="B92CEAA8CF68FFC8FED5E52AFDBDFD89" box="[314,559,614,635]" italics="true" pageId="14" pageNumber="687">Rana cavitympanum</emphasis>
|
||
<materialsCitation id="3B303CE7CF68FFC8FDD8E529FD60FD88" box="[567,754,613,635]" collectionCode="BMNH" location="Proceedings" municipality="Zoological Society of London" pageId="14" pageNumber="687" specimenCode="BMNH 1893.5" specimenCount="1" typeStatus="holotype">Boulenger, 1893</materialsCitation>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B92CEAA8CF68FFC8FF7EE5C8FD28FD6B" box="[145,698,644,665]" italics="true" pageId="14" pageNumber="687">
|
||
<location id="8E876061CF68FFC8FF7EE5C8FE8EFD6B" LSID="urn:lsid:plazi:treatment:03F187ACCF68FFC8FEDAE543FC4BFEA4:8E876061CF68FFC8FF7EE5C8FE8EFD6B" box="[145,284,644,665]" municipality="Zoological Society of London" name="Proceedings" pageId="14" pageNumber="687">Proceedings</location>
|
||
of the
|
||
<collectingMunicipality id="6B83ACC0CF68FFC8FE83E5C8FD28FD6B" box="[364,698,644,665]" pageId="14" pageNumber="687">Zoological Society of London</collectingMunicipality>
|
||
</emphasis>
|
||
: 525.
|
||
<typeStatus id="54E38818CF68FFC8FF7EE5EEFF6FFD4A" box="[145,253,674,696]" pageId="14" pageNumber="687" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<specimenCode id="DBFE9EC1CF68FFC8FEE7E5EFFE20FD4A" box="[264,434,674,696]" collectionCode="BMNH" pageId="14" pageNumber="687">BMNH 1893.5</specimenCode>
|
||
.30.26 (BMNH 1947.2.4.16); adult male).
|
||
</paragraph>
|
||
<paragraph id="8BE736BACF68FFC8FF7EE454FDC2FCBF" blockId="14.[145,761,791,846]" pageId="14" pageNumber="687">
|
||
<materialsCitation id="3B303CE7CF68FFC8FF7EE454FECBFCBC" country="Malaysia" location="Kinabalu" pageId="14" pageNumber="687" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
|
||
<emphasis id="B92CEAA8CF68FFC8FF7EE454FEABFCDD" box="[145,313,791,815]" italics="true" pageId="14" pageNumber="687">
|
||
<typeStatus id="54E38818CF68FFC8FF7EE454FF5EFCDD" box="[145,204,792,815]" pageId="14" pageNumber="687">Type</typeStatus>
|
||
locality:
|
||
</emphasis>
|
||
<taxonomicName id="4C584D39CF68FFC8FEACE455FE55FCDD" authority="Baloo" authorityName="Baloo" box="[323,455,793,815]" class="Echinoidea" family="Schizasteridae" genus="Kina" kingdom="Animalia" order="Spatangoida" pageId="14" pageNumber="687" phylum="Echinodermata" rank="genus">Kina Baloo</taxonomicName>
|
||
, North Borneo (
|
||
<location id="8E876061CF68FFC8FD69E455FD67FCDD" LSID="urn:lsid:plazi:treatment:03F187ACCF68FFC8FEDAE543FC4BFEA4:8E876061CF68FFC8FD69E455FD67FCDD" box="[646,757,793,815]" country="Malaysia" name="Kinabalu" pageId="14" pageNumber="687" stateProvince="Sabah">Kinabalu</location>
|
||
,
|
||
<collectingRegion id="499CF858CF68FFC8FF7EE474FF48FCBC" box="[145,218,824,846]" country="Malaysia" name="Sabah" pageId="14" pageNumber="687">Sabah</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34F762ACF68FFC8FF0AE474FEC2FCBC" box="[229,336,824,846]" name="Malaysia" pageId="14" pageNumber="687">Malaysia</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
. Collector: A. Everett.
|
||
</paragraph>
|
||
<paragraph id="8BE736BACF68FFC8FF7EE4C2FD80FC11" blockId="14.[145,761,910,995]" pageId="14" pageNumber="687">
|
||
<emphasis id="B92CEAA8CF68FFC8FF7EE4C2FEC0FC54" box="[145,338,910,934]" italics="true" pageId="14" pageNumber="687">Common name:</emphasis>
|
||
Sabah
|
||
<taxonomicName id="4C584D39CF68FFC8FE46E4DCFE70FC54" authorityName="Yang" authorityYear="1991" box="[425,482,912,934]" class="Amphibia" family="Ranidae" genus="Huia" kingdom="Animalia" order="Anura" pageId="14" pageNumber="687" phylum="Chordata" rank="genus">Huia</taxonomicName>
|
||
frog (English) following
|
||
<bibRefCitation id="EFC94B4BCF68FFC8FF7EE4E2FE31FC36" author="Frank N & Ramus E" box="[145,419,942,964]" pageId="14" pageNumber="687" refId="ref13385" refString="Frank N, Ramus E. 1995. A complete guide to scientific and common names of reptiles and amphibians of the world. NG Publishing, Inc." type="book" year="1995">Frank & Ramus (1995)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFC94B4BCF68FFC8FE0FE4E3FDE1FC37" author="Frost DR" box="[480,627,943,965]" pageId="14" pageNumber="687" refId="ref13414" refString="Frost DR. 2019. Amphibian species of the world: an online reference. Version 6. 0. Available at: https: // amphibiansoftheworld. amnh. org" type="url" year="2019">Frost (2019)</bibRefCitation>
|
||
; Kongkang Jeram Sabah (Bahasa
|
||
<collectingCountry id="F34F762ACF68FFC8FE7CE481FD95FC11" box="[403,519,973,995]" name="Indonesia" pageId="14" pageNumber="687">Indonesia</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE736BACF68FFC8FF7EE36FFE6DFB26" blockId="14.[145,761,1059,1236]" pageId="14" pageNumber="687">
|
||
<emphasis id="B92CEAA8CF68FFC8FF7EE36FFD52FBC9" box="[145,704,1059,1083]" italics="true" pageId="14" pageNumber="687">
|
||
Phylogenetic definition and content:
|
||
<taxonomicName id="4C584D39CF68FFC8FD6CE36AFD52FBC9" authorityName="Yang" authorityYear="1991" box="[643,704,1062,1083]" class="Amphibia" family="Ranidae" genus="Huia" kingdom="Animalia" order="Anura" pageId="14" pageNumber="687" phylum="Chordata" rank="genus">Huia</taxonomicName>
|
||
</emphasis>
|
||
is a distinct and independently evolving genus, which shares a most recent common ancestor with the genus
|
||
<taxonomicName id="4C584D39CF68FFC8FF7EE3CEFEBFFB65" authorityName="Yang" authorityYear="1991" box="[145,301,1154,1175]" class="Amphibia" family="Ranidae" genus="Meristogenys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="687" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92CEAA8CF68FFC8FF7EE3CEFEBFFB65" box="[145,301,1154,1175]" italics="true" pageId="14" pageNumber="687">Meristogenys</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13632A3FCF68FFC8FEAFE3CDFE16FB64" box="[320,388,1153,1175]" captionStart="Figure 2" captionStartId="5.[163,245,1601,1623]" captionTargetBox="[166,1446,198,1558]" captionTargetId="figure-27@5.[163,1449,195,1561]" captionTargetPageId="5" captionText="Figure 2. Maximum likelihood tree (concatenated dataset) depicting phylogenetic relationship of frogs previously recognized as Huia (coloured branches, which also match with the island/locality of the samples; clades A–H) from Sumatra, Java, Borneo and the mainland to other closely related species within the family Ranidae. Black circles represent highly supported nodes [bootstrap support (BS)> 70] for unphased and phased dataset, whereas low support values are shown with a slash (/) for unphased dataset (BS)/phased dataset 1 (BS1)/phased dataset 2 (BS2). Tadpole samples are represented by ‘_Tad’. Topotypic samples are represented by numbers *1–*4 and the locality shown on the map. Pink triangles, orange circles, blue stars and purple squares on the map represent the sampling sites." pageId="14" pageNumber="687">Fig. 2</figureCitation>
|
||
). It is currently understood to be monotypic, containing
|
||
<taxonomicName id="4C584D39CF68FFC8FE54E3ECFD18FB47" authorityName="Boulenger" authorityYear="1893" box="[443,650,1184,1205]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="14" pageNumber="687" phylum="Chordata" rank="species" species="cavitympanum">
|
||
<emphasis id="B92CEAA8CF68FFC8FE54E3ECFD18FB47" box="[443,650,1184,1205]" italics="true" pageId="14" pageNumber="687">H. cavitympanum</emphasis>
|
||
</taxonomicName>
|
||
, which is endemic to the island of Borneo.
|
||
</paragraph>
|
||
<paragraph id="8BE736BACF68FFC8FF7EE259FC4BFEA4" blockId="14.[145,762,1301,1907]" lastBlockId="14.[809,1425,197,342]" pageId="14" pageNumber="687">
|
||
<emphasis id="B92CEAA8CF68FFC8FF7EE259FE85FADE" box="[145,279,1301,1324]" italics="true" pageId="14" pageNumber="687">Diagnosis:</emphasis>
|
||
(Tadpole characters presented here are based on observation by U.A., whereas morphological characters of adults are taken from published studies (e.g.
|
||
<bibRefCitation id="EFC94B4BCF68FFC8FF2BE23FFED0FA7A" author="Inger RF" box="[196,322,1394,1416]" pageId="14" pageNumber="687" pagination="1 - 402" refId="ref13706" refString="Inger RF. 1966. The systematics and zoogeography of the Amphibia of Borneo. Fieldiana Zoology 52: 1 - 402." type="journal article" year="1966">Inger, 1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC94B4BCF68FFC8FEA3E23FFE55FA7A" author="Yang DT" box="[332,455,1395,1416]" pageId="14" pageNumber="687" pagination="1 - 42" refId="ref15214" refString="Yang DT. 1991. Phylogenetic systematics of the Amolops group of ranid frogs of southeastern Asia and the Greater Sunda Islands. Fieldiana Zoology 63: 1 - 42." type="journal article" year="1991">Yang, 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC94B4BCF68FFC8FE3CE23EFD79FA7A" author="Manthey U & Denzer W" box="[467,747,1394,1416]" pageId="14" pageNumber="687" pagination="31 - 48" refId="ref14177" refString="Manthey U, Denzer W. 2014. Sudostasiatische Anuren im Fokus, Spezies der Gattung Huia (sensu lato) Yang, 1991 (Amphibia: Anura: Ranidae). Sauria 36: 31 - 48." type="journal article" year="2014">Manthey & Denzer, 2014</bibRefCitation>
|
||
). Larvae gastromyzophorous; upper beak Λ- shaped, lower beak V-shaped, both undivided; LTRF 10–12((3– 4)–(10–12))/5–6(0–2); postorbital glands present; an oblique cluster of glands immediately below the spiracular tube, running upward and backward present; prespiracular glands present in some individuals (
|
||
<bibRefCitation id="EFC94B4BCF68FFC8FECFE106FE0DF9AD" author="Inger RF" box="[288,415,1609,1631]" pageId="14" pageNumber="687" pagination="1 - 89" refId="ref13729" refString="Inger RF. 1985. Tadpoles of the forested regions of Borneo. Fieldiana Zoology 26: 1 - 89." type="journal article" year="1985">Inger, 1985</bibRefCitation>
|
||
); glands on the tail fin absent. Adults medium in size, body stocky; snout rounded; tympanum clear, deep in cavity; supra- and posttympanic folds present, dorsolateral folds present; ratio of tibia length to snout–vent length> 70%; outer metatarsal tubercle present or absent (absent according to
|
||
<bibRefCitation id="EFC94B4BCF68FFC8FEC5E04EFE3FF8E4" author="Yang DT" box="[298,429,1793,1815]" pageId="14" pageNumber="687" pagination="1 - 42" refId="ref15214" refString="Yang DT. 1991. Phylogenetic systematics of the Amolops group of ranid frogs of southeastern Asia and the Greater Sunda Islands. Fieldiana Zoology 63: 1 - 42." type="journal article" year="1991">Yang, 1991</bibRefCitation>
|
||
), inner metatarsal tubercle present, small; length of Finger I = Finger II; length of metacarpal of Finger I 54% of Finger II; width of crossbar terminal phalanx of Finger III 50% length; male with nuptial pad and paired lateral vocal sacs; humeral gland absent. In life, dorsum generally dark brown; flanks light brown with markings; head light brown with scattered dark brown markings; lips with dark markings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |