164 lines
18 KiB
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164 lines
18 KiB
XML
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<mods:title id="199F9E181AA50A8AF2A566D61A1D3959">Taxonomic review of weevils of the tribe Celeuthetini, mainly from Sundaland, Sulawesi and the Lesser Sunda Islands (Coleoptera, Curculionidae, Entiminae)</mods:title>
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<mods:namePart id="9B1A084E3530656D81DA94326B58455E">Riedel, Alexander</mods:namePart>
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<mods:date id="51A12110BBE08754316D3792F71B7F9C">2024</mods:date>
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<paragraph id="B8080432FFE2252FFF6FBB33FE7FFCFA" blockId="42.[151,525,788,815]" box="[151,525,788,815]" pageId="42" pageNumber="43">
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<emphasis id="8AC3D820FFE2252FFF6FBB33FE7FFCFA" bold="true" box="[151,525,788,815]" pageId="42" pageNumber="43">
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<taxonomicName id="7FB77FB1FFE2252FFF6FBB33FDC6FCFB" authority="Riedel" authorityName="Riedel" authorityYear="2024" box="[151,436,788,815]" class="Insecta" family="Curculionidae" genus="Nusasyntrophus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="8AC3D820FFE2252FFF6FBB33FD2EFCFA" bold="true" box="[151,348,788,815]" italics="true" pageId="42" pageNumber="43">Nusasyntrophus</emphasis>
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Riedel
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</taxonomicName>
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<taxonomicNameLabel id="91F0655BFFE2252FFE43BB32FE7FFCFA" box="[443,525,789,815]" pageId="42" pageNumber="43" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="B8080432FFE2252FFF6FBB7EF830FCA4" blockId="42.[151,1090,857,881]" box="[151,1090,857,881]" pageId="42" pageNumber="43">
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http://zoobank.org/
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<uuid id="CC113EE7FFE2252FFDACBB7EF830FCA4" box="[596,1090,857,881]" pageId="42" pageNumber="43">40D08E0A-8547-4AF7-92E5-2AE63E7AD3DF</uuid>
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</paragraph>
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<subSubSection id="F0AD57B9FFE2252FFF6FBBBBFFDFFC61" box="[151,941,923,948]" pageId="42" pageNumber="43" type="type_taxon">
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<paragraph id="B8080432FFE2252FFF6FBBBBFFDFFC61" blockId="42.[151,941,923,948]" box="[151,941,923,948]" pageId="42" pageNumber="43">
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<emphasis id="8AC3D820FFE2252FFF6FBBBBFD54FC66" bold="true" box="[151,294,923,948]" pageId="42" pageNumber="43">
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<typeStatus id="670CBA90FFE2252FFF6FBBBBFCBEFC61" box="[151,204,924,948]" pageId="42" pageNumber="43">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="7FB77FB1FFE2252FFED4BBBBFECAFC61" authority="Heller, 1900" authorityName="Heller" authorityYear="1900" box="[300,696,924,948]" class="Insecta" family="Curculionidae" genus="Heteroglymma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="echinatum">
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<emphasis id="8AC3D820FFE2252FFED4BBBBFE40FC61" box="[300,562,924,948]" italics="true" pageId="42" pageNumber="43">Heteroglymma echinatum</emphasis>
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<bibRefCitation id="DC2679C3FFE2252FFDC0BBBBFECAFC61" author="Heller, K. M." box="[568,696,924,948]" pageId="42" pageNumber="43" pagination="1 - 46" refId="ref41001" refString="Heller, K. M. (1900) Neue Kafer von Celebes. IV. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden, 9 (5), 1 - 46, 1 pl." type="journal article" year="1900">Heller, 1900</bibRefCitation>
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</taxonomicName>
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, by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="F0AD57B9FFE2252FFF6FBBC6F89FFAB1" pageId="42" pageNumber="43" type="description">
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<paragraph id="B8080432FFE2252FFF6FBBC6F89FFAB1" blockId="42.[151,1437,993,1632]" pageId="42" pageNumber="43">
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<emphasis id="8AC3D820FFE2252FFF6FBBC6FD53FC2E" bold="true" box="[151,289,993,1019]" pageId="42" pageNumber="43">Description</emphasis>
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. Integument covered with appressed, microgranulate, polygoniform scales; elytra with rows of long erect setiform scales, their length exceeding width of elytral interval. Head with angulate, transverse sulcus separating rostral base; vertex with radial furrows partly obscured by scales. Eyes protruding, of conical shape; separated from apex of prothorax by ca. length of eye. Rostrum. Dorsum medially with weak costa; apical declivity distinct, with sparse setae, with sunk-in scales; dorsal profile subangulate. Antenna. Scape curved, squamose, with suberect setae. Funicle with article 2 slightly longer (1.2 X) than article 1. Pronotum slightly longer than wide (1.04 X); coarsely punctate. Metanepisternum narrow, subglabrous, anteriorly with impression. Elytra subovate; with 12 striae, some abbreviated. Legs. Protibia ventrally simple, not denticulate. Venter. Procoxae subcontiguous, positioned slightly closer to apex than base of prothorax. Mesoventral process trapezoid, 1.25 X as long as posterior width. Abdomen. Female ventrites 3–5 exposed, laterally slightly shorter than ventrites 1–2. Genitalia. Penis (
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<figureCitation id="208C18B7FFE2252FFB4BBD01F953FA95" box="[1203,1313,1318,1344]" captionStart="FIGURE 24" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[260,1336,188,1913]" captionTargetId="figure-32@44.[252,1336,181,1920]" captionTargetPageId="44" captionText="FIGURE 24. Nusasyntrophus echinatus (Heller), a–c, Heteroglymma echinatum Heller, lectotype: (a) Habitus (b) Penis (c) Labels; d–f, Trigonops bipusulosa Heller, lectotype: (a) Habitus (b) Terminalia (c) Labels." figureDoi="http://doi.org/10.5281/zenodo.11612902" httpUri="https://zenodo.org/record/11612902/files/figure.png" pageId="42" pageNumber="43">Figs. 24b</figureCitation>
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,
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<figureCitation id="208C18B7FFE2252FFAD6BD01F928FA94" box="[1326,1370,1318,1345]" captionStart="FIGURE 25" captionStartId="45.[151,250,1961,1985]" captionTargetBox="[161,1427,355,1937]" captionTargetId="figure-86@45.[151,1436,345,1937]" captionTargetPageId="45" captionText="FIGURE 25. Nusasyntrophus virescens (Heller). Heteroglymma echinatus virescens Heller, lectotype: (a) Habitus (b) Penis (c) Labels." figureDoi="http://doi.org/10.5281/zenodo.11612910" httpUri="https://zenodo.org/record/11612910/files/figure.png" pageId="42" pageNumber="43">25b</figureCitation>
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) with apodeme firmly attached to body. Transfer apparatus spiniform, curved, shorter than body of penis.
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</paragraph>
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</subSubSection>
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<subSubSection id="F0AD57B9FFE2252FFF3FBD49FFE9F9B5" pageId="42" pageNumber="43" type="etymology">
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<paragraph id="B8080432FFE2252FFF3FBD49FD0FFA79" blockId="42.[151,1437,993,1632]" pageId="42" pageNumber="43">
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<emphasis id="8AC3D820FFE2252FFF3FBD49FD3CFA5D" bold="true" box="[199,334,1390,1416]" pageId="42" pageNumber="43">Etymology.</emphasis>
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The name is a combination of the Malay noun
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<emphasis id="8AC3D820FFE2252FFC9DBD48FFE8FA5D" box="[869,922,1391,1416]" italics="true" pageId="42" pageNumber="43">nusa</emphasis>
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(island) and the genus name
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<taxonomicName id="7FB77FB1FFE2252FFB1EBD49F911FA5D" box="[1254,1379,1390,1416]" class="Insecta" family="Curculionidae" genus="Syntrophus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
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<emphasis id="8AC3D820FFE2252FFB1EBD49F911FA5D" box="[1254,1379,1390,1416]" italics="true" pageId="42" pageNumber="43">Syntrophus</emphasis>
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</taxonomicName>
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. The gender is masculine.
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</paragraph>
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<paragraph id="B8080432FFE2252FFF3FBD91FFE9F9B5" blockId="42.[151,1437,993,1632]" pageId="42" pageNumber="43">
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<emphasis id="8AC3D820FFE2252FFF3FBD91FD78FA05" bold="true" box="[199,266,1462,1488]" pageId="42" pageNumber="43">Notes</emphasis>
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. The morphology of the male genital and the consistent placement in molecular phylogenies (
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<bibRefCitation id="DC2679C3FFE2252FFAD7BD91FD62FA20" author="Toussaint, E. F. A. & Tanzler, R. & Rahmadi, C. & Balke, M. & Riedel, A." pageId="42" pageNumber="43" pagination="632 - 644" refId="ref41812" refString="Toussaint, E. F. A., Tanzler, R., Rahmadi, C., Balke, M. & Riedel, A. (2015) Biogeography of Australasian flightless weevils (Curculionidae, Celeuthetini) suggests permeability of Lydekker's and Wallace's Lines. Zoologica Scripta, 44 (6), 632 - 644. https: // doi. org / 10.1111 / zsc. 12127" type="journal article" year="2015">
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Toussaint
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<emphasis id="8AC3D820FFE2252FFF6FBDFCFCBBFA21" box="[151,201,1498,1524]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
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. 2015
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</bibRefCitation>
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and additional unpublished ones) indicate that this new genus is closely related to
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<taxonomicName id="7FB77FB1FFE2252FFB46BDFDFCA7F9CD" authority="Marshall. Since" authorityName="Marshall. Since" authorityYear="1956" class="Insecta" family="Curculionidae" genus="Javaulius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
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<emphasis id="8AC3D820FFE2252FFB46BDFDF95BFA21" box="[1214,1321,1498,1524]" italics="true" pageId="42" pageNumber="43">Javaulius</emphasis>
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Marshall. Since
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</taxonomicName>
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the species of the latter genus are found in leaf litter, their habitus and general morphology differs greatly from
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<taxonomicName id="7FB77FB1FFE2252FFF6FBE05FD38F9E9" authority="Riedel, 2024" authorityName="Riedel" authorityYear="2024" box="[151,330,1570,1596]" class="Insecta" family="Curculionidae" genus="Nusasyntrophus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="8AC3D820FFE2252FFF6FBE05FD38F9E9" box="[151,330,1570,1596]" italics="true" pageId="42" pageNumber="43">Nusasyntrophus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="91F0655BFFE2252FFEABBE05FDEDF9E9" box="[339,415,1570,1596]" pageId="42" pageNumber="43" rank="genus">gen. n.</taxonomicNameLabel>
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living on foliage. The inclusion in
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<taxonomicName id="7FB77FB1FFE2252FFCCABE05F86FF9E9" authority="Faust" authorityName="Faust. A" authorityYear="1897" box="[818,1053,1570,1596]" class="Insecta" family="Curculionidae" genus="Heteroglymma" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
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<emphasis id="8AC3D820FFE2252FFCCABE05FFA5F9E9" box="[818,983,1570,1596]" italics="true" pageId="42" pageNumber="43">Heteroglymma</emphasis>
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Faust
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</taxonomicName>
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is impossible by the absence of a protibial denticle. Therefore, it is necessary to establish a new genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="F0AD57B9FFE2252FFF6FBE95F9EFF8F6" pageId="42" pageNumber="43" type="key">
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<paragraph id="B8080432FFE2252FFF6FBE95FECEF919" blockId="42.[151,700,1713,1740]" box="[151,700,1713,1740]" pageId="42" pageNumber="43">
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<emphasis id="8AC3D820FFE2252FFF6FBE95FECEF919" bold="true" box="[151,700,1713,1740]" pageId="42" pageNumber="43">
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Key to the species of
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<taxonomicName id="7FB77FB1FFE2252FFE65BE96FE10F919" authority="Riedel, 2024" authorityName="Riedel" authorityYear="2024" box="[413,610,1713,1740]" class="Insecta" family="Curculionidae" genus="Nusasyntrophus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="8AC3D820FFE2252FFE65BE96FE10F919" bold="true" box="[413,610,1713,1740]" italics="true" pageId="42" pageNumber="43">Nusasyntrophus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="91F0655BFFE2252FFD91BE95FECEF919" box="[617,700,1714,1740]" pageId="42" pageNumber="43" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<keyStep id="03431C97FFE2252FFF6FBED7F9EFF8F6" pageId="42" pageNumber="43">
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<key id="CC265303FFE2252FFF6FBED7F9EFF8F6" pageId="42" pageNumber="43">
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<paragraph id="B8080432FFE2252FFF6FBED7F9EFF8D3" blockId="42.[151,1437,1776,1827]" box="[151,1437,1776,1798]" pageId="42" pageNumber="43">
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<keyLead id="0346A707FFE2252FFF6FBED7F9EFF8D3" box="[151,1437,1776,1798]" pageId="42" pageNumber="43">
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1 Vestiture greyish (
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<figureCitation id="208C18B7FFE2252FFE69BED7FD89F8D3" box="[401,507,1776,1798]" captionStart="FIGURE 24" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[260,1336,188,1913]" captionTargetId="figure-32@44.[252,1336,181,1920]" captionTargetPageId="44" captionText="FIGURE 24. Nusasyntrophus echinatus (Heller), a–c, Heteroglymma echinatum Heller, lectotype: (a) Habitus (b) Penis (c) Labels; d–f, Trigonops bipusulosa Heller, lectotype: (a) Habitus (b) Terminalia (c) Labels." figureDoi="http://doi.org/10.5281/zenodo.11612902" httpUri="https://zenodo.org/record/11612902/files/figure.png" pageId="42" pageNumber="43">Fig. 24a, d</figureCitation>
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). Lombok Isl.......................................
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<taxonomicName id="7FB77FB1FFE2252FFBB6BED7F9EFF8D3" authority="(Heller)" authorityName="Riedel" authorityYear="2024" baseAuthorityName="Heller" baseAuthorityYear="2024" box="[1102,1437,1776,1798]" class="Insecta" family="Curculionidae" genus="Nusasyntrophus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="echinatus">
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||
<emphasis id="8AC3D820FFE2252FFBB6BED7F938F8D0" box="[1102,1354,1776,1797]" italics="true" pageId="42" pageNumber="43">Nusasyntrophus echinatus</emphasis>
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(Heller)
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="B8080432FFE2252FFF6FBF2AF9EFF8F6" blockId="42.[151,1437,1776,1827]" box="[151,1437,1805,1827]" pageId="42" pageNumber="43">
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<keyLead id="0346A707FFE2252FFF6FBF2AF9EFF8F6" box="[151,1437,1805,1827]" pageId="42" pageNumber="43">
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1´Vestiture greenish (
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||
<figureCitation id="208C18B7FFE2252FFE64BF2AFD9DF8F6" box="[412,495,1805,1827]" captionStart="FIGURE 25" captionStartId="45.[151,250,1961,1985]" captionTargetBox="[161,1427,355,1937]" captionTargetId="figure-86@45.[151,1436,345,1937]" captionTargetPageId="45" captionText="FIGURE 25. Nusasyntrophus virescens (Heller). Heteroglymma echinatus virescens Heller, lectotype: (a) Habitus (b) Penis (c) Labels." figureDoi="http://doi.org/10.5281/zenodo.11612910" httpUri="https://zenodo.org/record/11612910/files/figure.png" pageId="42" pageNumber="43">Fig. 25a</figureCitation>
|
||
).
|
||
<collectingRegion id="7A73CAD0FFE2252FFE07BF2AFE55F8F6" box="[511,551,1805,1827]" country="Indonesia" name="Bali" pageId="42" pageNumber="43">Bali</collectingRegion>
|
||
Isl............................................
|
||
<taxonomicName id="7FB77FB1FFE2252FFBAABF2AF9EFF8F6" authority="(Heller)" baseAuthorityName="Heller" box="[1106,1437,1805,1827]" class="Insecta" family="Curculionidae" genus="Nusasyntrophus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="virescens">
|
||
<emphasis id="8AC3D820FFE2252FFBAABF2AF938F8F6" box="[1106,1354,1805,1827]" italics="true" pageId="42" pageNumber="43">Nusasyntrophus virescens</emphasis>
|
||
(Heller)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |