treatments-xml/data/BD/07/87/BD0787EDD972FFC6D7F2C286FABA5C2C.xml

<document id="7B5EA7192221ACAAA373381B24ACC398" ID-CLB-Dataset="7487" ID-DOI="10.11646/zootaxa.4949.2.2" ID-GBIF-Dataset="085e7429-bd44-4056-8eb3-42466ec5f177" ID-ISSN="1175-5326" ID-Zenodo-Dep="4636318" ID-ZooBank="3E12070B-AC64-4B06-AE56-021D093CD7D6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616666884399" checkinUser="plazi" docAuthor="Joharchi, Omid, Döker, Ismail &amp; Khaustov, Vladimir A." docDate="2021" docId="BD0787EDD972FFC6D7F2C286FABA5C2C" docLanguage="en" docName="zootaxa.4949.2.2.pdf" docOrigin="Zootaxa 4949 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gaeolaelaps altaiensis Joharchi &amp; Döker &amp; Khaustov 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="243" masterDocId="413EFF95D973FFC5D765C739FFC45B5D" masterDocTitle="Two new species and a new record of Gaeolaelaps Evans &amp; Till (Acari: Laelapidae) from Altai Mountains, Russia" masterLastPageNumber="260" masterPageNumber="240" pageNumber="241" updateTime="1698934979109" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="58502945FD224E98ECA82ACCE88D6A11">Two new species and a new record of Gaeolaelaps Evans &amp; Till (Acari: Laelapidae) from Altai Mountains, Russia</mods:title>
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<mods:namePart id="6F8334830C12B339BEE4007D5C378F29">Joharchi, Omid</mods:namePart>
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<mods:affiliation id="C37C3B149BD57340AAAA4A2B0CF086A2">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia. &amp; J. omid 2000 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2741 - 4946</mods:affiliation>
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<mods:namePart id="13CB4C45DC6859EC3FBB9CD6E1CDAAE2">Döker, Ismail</mods:namePart>
<mods:affiliation id="CB640718BFE58C4E6886A84E74731073">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia. &amp; Cukurova University, Agricultural Faculty, Department of Plant Protection, Acarology Laboratory, 01330, Adana, Turkey https: // orcid. org / 0000 - 0002 - 1412 - 1554</mods:affiliation>
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<mods:namePart id="0B9855DD18E634E9BB1D4687A8DCDF7B">Khaustov, Vladimir A.</mods:namePart>
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<mods:affiliation id="CE978D2AA996E4036E6D6DA3A1C8ADB7">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia. &amp; https: // orcid. org / 0000 - 0003 - 2831 - 7213</mods:affiliation>
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<treatment id="BD0787EDD972FFC6D7F2C286FABA5C2C" ID-DOI="http://doi.org/10.5281/zenodo.4664799" ID-GBIF-Taxon="180876832" ID-Zenodo-Dep="4664799" LSID="urn:lsid:plazi:treatment:BD0787EDD972FFC6D7F2C286FABA5C2C" httpUri="http://treatment.plazi.org/id/BD0787EDD972FFC6D7F2C286FABA5C2C" lastPageId="3" lastPageNumber="243" pageId="1" pageNumber="241">
<subSubSection id="7DB46570D972FFC4D7F2C286FDBB5E87" box="[151,639,1471,1498]" pageId="1" pageNumber="241" type="nomenclature">
<paragraph id="351136FBD972FFC4D7F2C286FDBB5E87" blockId="1.[151,639,1471,1534]" box="[151,639,1471,1498]" pageId="1" pageNumber="241">
<heading id="6E598197D972FFC4D7F2C286FDBB5E87" bold="true" box="[151,639,1471,1498]" fontSize="11" level="1" pageId="1" pageNumber="241" reason="1">
<emphasis id="07DAEAE9D972FFC4D7F2C286FDBB5E87" bold="true" box="[151,639,1471,1498]" pageId="1" pageNumber="241">
<taxonomicName id="F2AE4D78D972FFC4D7F2C286FDDE5E87" authority="Joharchi" authorityName="Joharchi &amp; Döker &amp; Khaustov" authorityYear="2021" box="[151,538,1471,1498]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="241" phylum="Arthropoda" rank="species" species="altaiensis" status="sp. nov.">
<emphasis id="07DAEAE9D972FFC4D7F2C286FE615E87" bold="true" box="[151,421,1471,1498]" italics="true" pageId="1" pageNumber="241">Gaeolaelaps altaiensis</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="1CE95792D972FFC4D547C2F9FDBB5E87" box="[546,639,1472,1498]" pageId="1" pageNumber="241" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<paragraph id="351136FBD972FFC4D7F2C2DDFEF95EA3" blockId="1.[151,639,1471,1534]" box="[151,317,1508,1534]" pageId="1" pageNumber="241">
(
<figureCitation id="AD952A7ED972FFC4D7FAC2DDFEF05EA3" box="[159,308,1508,1534]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–10" captionStart-2="FIGURES 11–15" captionStart-3="FIGURES 16–19" captionStart-4="FIGURES 20–21" captionStartId-0="4.[151,264,1943,1967]" captionStartId-1="5.[151,264,1943,1967]" captionStartId-2="6.[151,264,1943,1967]" captionStartId-3="7.[151,264,1943,1967]" captionStartId-4="8.[151,264,1497,1521]" captionTargetBox-0="[178,1392,189,1903]" captionTargetBox-1="[194,1389,191,1907]" captionTargetBox-2="[187,1394,193,1910]" captionTargetBox-3="[187,1397,192,1902]" captionTargetBox-4="[174,1422,206,1456]" captionTargetId-0="figure-17@4.[171,1416,181,1919]" captionTargetId-1="figure-27@5.[179,1408,181,1920]" captionTargetId-2="figure-17@6.[179,1408,181,1920]" captionTargetId-3="figure-27@7.[167,1421,181,1919]" captionTargetId-4="figure-17@8.[151,1436,181,1473]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." captionText-1="FIGURES 7–10. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 7, idiosoma in dorsal view; 8, reticulation of dorsal shield, general view of some dorsal setae, absence of setae S1; 9, idiosoma in ventral view; 10, sternal &amp; genital shields." captionText-2="FIGURES 11–15. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 11, subcapitulum; 12, distal portion of palp, with a focus on apotele; 13, insemination structures; male. 14, idiosoma in ventral view; 15, chelicera." captionText-3="FIGURES 16–19. Gaeolaelaps altaiensis Joharchi sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." captionText-4="FIGURES 20–21. Gaeolaelaps altaiensis Joharchi sp. nov., male. 20, ventral idiosoma; 21, chelicera." figureDoi-0="http://doi.org/10.5281/zenodo.4636322" figureDoi-1="http://doi.org/10.5281/zenodo.4636326" figureDoi-2="http://doi.org/10.5281/zenodo.4636328" figureDoi-3="http://doi.org/10.5281/zenodo.4636330" figureDoi-4="http://doi.org/10.5281/zenodo.4636332" httpUri-0="https://zenodo.org/record/4636322/files/figure.png" httpUri-1="https://zenodo.org/record/4636326/files/figure.png" httpUri-2="https://zenodo.org/record/4636328/files/figure.png" httpUri-3="https://zenodo.org/record/4636330/files/figure.png" httpUri-4="https://zenodo.org/record/4636332/files/figure.png" pageId="1" pageNumber="241">Figures 1–21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7DB46570D972FFC4D7F2C115FC695DD3" pageId="1" pageNumber="241" type="materials_examined">
<paragraph id="351136FBD972FFC4D7F2C115FC695DD3" blockId="1.[151,1436,1580,2039]" pageId="1" pageNumber="241">
<emphasis id="07DAEAE9D972FFC4D7F2C115FE825D1B" bold="true" box="[151,326,1580,1606]" pageId="1" pageNumber="241">Type material:</emphasis>
<typeStatus id="EA158859D972FFC4D629C115FE735D1B" box="[332,439,1580,1606]" pageId="1" pageNumber="241" type="holotype">Holotype</typeStatus>
, female, Ulaganskiy Rayon, Chuysky Trakt, 
<collectingRegion id="F76AF819D972FFC4D4C2C115FB895D1B" box="[935,1101,1580,1606]" country="Russia" name="Altay" pageId="1" pageNumber="241">Altai Republic</collectingRegion>
, 
<collectingCountry id="4DB9766BD972FFC4D332C115FB605D1B" box="[1111,1188,1580,1606]" name="Russia" pageId="1" pageNumber="241">Russia</collectingCountry>
, 
<geoCoordinate id="509A503CD972FFC4D3CAC115FACB5D1A" box="[1199,1295,1580,1607]" degrees="50" direction="north" minutes="14" orientation="latitude" pageId="1" pageNumber="241" precision="925" value="50.233334">50°14’N</geoCoordinate>
<geoCoordinate id="509A503CD972FFC4D273C115FAB25D1B" box="[1302,1398,1580,1606]" degrees="87" direction="east" minutes="42" orientation="longitude" pageId="1" pageNumber="241" precision="925" value="87.7">87°42’E</geoCoordinate>
, 
<date id="4110103BD972FFC4D2E5C115FECD5D37" pageId="1" pageNumber="241" value="2020-07-30">30 July 2020</date>
, alt. 
<quantity id="F2569B1ED972FFC4D65AC169FE535D36" box="[319,407,1616,1643]" metricMagnitude="3" metricUnit="m" metricValue="1.533" pageId="1" pageNumber="241" unit="m" value="1533.0">1533 m</quantity>
, O. Joharchi coll., in the nest of 
<taxonomicName id="F2AE4D78D972FFC4D46CC168FC965D37" box="[777,850,1617,1642]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="241" phylum="Arthropoda" rank="genus">
<emphasis id="07DAEAE9D972FFC4D46CC168FC965D37" box="[777,850,1617,1642]" italics="true" pageId="1" pageNumber="241">Lasius</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="F2AE4D78D972FFC4D4ECC169FBEC5D37" box="[905,1064,1616,1642]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="241" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
: 
<taxonomicName id="F2AE4D78D972FFC4D356C169FB715D36" box="[1075,1205,1616,1643]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="241" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
) (in 
<collectionCode id="53BFAE3ED972FFC4D38FC169FA8A5D37" box="[1258,1358,1616,1642]" pageId="1" pageNumber="241">TSUMZ</collectionCode>
). 
<typeStatus id="EA158859D972FFC4D204C168FF115DD2" pageId="1" pageNumber="241" type="paratype">Paratypes</typeStatus>
, five females and two males same data as 
<typeStatus id="EA158859D972FFC4D5CBC14DFCD45DD3" box="[686,784,1652,1678]" pageId="1" pageNumber="241" type="holotype">holotype</typeStatus>
(in 
<collectionCode id="53BFAE3ED972FFC4D458C14DFC655DD3" box="[829,929,1652,1678]" pageId="1" pageNumber="241">TSUMZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="7DB46570D972FFC4D7A2C1A1FED85C8F" pageId="1" pageNumber="241" type="diagnosis">
<paragraph id="351136FBD972FFC4D7A2C1A1FED85C8F" blockId="1.[151,1436,1580,2039]" pageId="1" pageNumber="241">
<emphasis id="07DAEAE9D972FFC4D7A2C1A1FEFE5DEF" bold="true" box="[199,314,1688,1714]" pageId="1" pageNumber="241">Diagnosis</emphasis>
(adult female). Dorsal shield with distinct reticulation posterior to 
<emphasis id="07DAEAE9D972FFC4D349C1A1FB865DEF" box="[1068,1090,1688,1714]" italics="true" pageId="1" pageNumber="241">j6</emphasis>
, narrowing from level of setae 
<emphasis id="07DAEAE9D972FFC4D7F2C185FF745D8B" box="[151,176,1724,1750]" italics="true" pageId="1" pageNumber="241">r3</emphasis>
, progressively tapering until 
<emphasis id="07DAEAE9D972FFC4D69CC185FDD65D8B" box="[505,530,1724,1750]" italics="true" pageId="1" pageNumber="241">s6</emphasis>
, then more or less parallel-sided from 
<emphasis id="07DAEAE9D972FFC4D4A6C185FC185D8B" box="[963,988,1724,1750]" italics="true" pageId="1" pageNumber="241">r6</emphasis>
to level of 
<emphasis id="07DAEAE9D972FFC4D33EC185FBB35D8B" box="[1115,1143,1724,1750]" italics="true" pageId="1" pageNumber="241">S4</emphasis>
until rounded posteriorly; with 37 pairs of smooth setae (except 
<emphasis id="07DAEAE9D972FFC4D521C1D9FD9A5DA7" box="[580,606,1760,1786]" italics="true" pageId="1" pageNumber="241">J4</emphasis>
, 
<emphasis id="07DAEAE9D972FFC4D509C1D9FD435DA7" box="[620,647,1760,1786]" italics="true" pageId="1" pageNumber="241">J5</emphasis>
, 
<emphasis id="07DAEAE9D972FFC4D5F0C1D9FD775DA7" box="[661,691,1760,1786]" italics="true" pageId="1" pageNumber="241">Z5</emphasis>
, sparsely barbed), 
<emphasis id="07DAEAE9D972FFC4D4EDC1D9FC655DA7" box="[904,929,1760,1786]" italics="true" pageId="1" pageNumber="241">z3</emphasis>
and 
<emphasis id="07DAEAE9D972FFC4D4BCC1D9FC315DA7" box="[985,1013,1760,1786]" italics="true" pageId="1" pageNumber="241">S1</emphasis>
absent. Presternal area lightly sclerotised, punctate, with a few transverse curved lines, sternal shield with reticulate ornamentation laterally, smooth (or faintly reticulated) medially, anterior margin slightly concave, posterior margin more or less straight; ratio of shield length/width ≃ 0.86. Genital shield length/width ratio ≃ 2.5, faintly reticulated posterior to inverted-V ridge with 5–6 cells. Anal shield length/width ratio ≃ 1. Opisthosomal integument with eight pairs of smooth setae, peritremes short, reaching to mid-level of coxae II. Tarsus IV with two (
<emphasis id="07DAEAE9D972FFC4D431C0ADFCBA5CF3" box="[852,894,1940,1966]" italics="true" pageId="1" pageNumber="241">pd2</emphasis>
, 
<emphasis id="07DAEAE9D972FFC4D4EBC0ADFC7C5CF3" box="[910,952,1940,1966]" italics="true" pageId="1" pageNumber="241">pd3</emphasis>
) macrosetae. Fixed digit of chelicera with seven teeth.
</paragraph>
</subSubSection>
<subSubSection id="7DB46570D972FFC6D7A2C0E5FDE45F4C" lastPageId="3" lastPageNumber="243" pageId="1" pageNumber="241" type="description">
<paragraph id="351136FBD972FFC4D7A2C0E5FD3C5CAA" blockId="1.[151,1436,1580,2039]" box="[199,760,2012,2039]" pageId="1" pageNumber="241">
<emphasis id="07DAEAE9D972FFC4D7A2C0E5FE765CAB" bold="true" box="[199,434,2012,2038]" pageId="1" pageNumber="241">
<emphasis id="07DAEAE9D972FFC4D7A2C0E5FE895CAB" bold="true" box="[199,333,2012,2038]" italics="true" pageId="1" pageNumber="241">Description</emphasis>
. Female
</emphasis>
(n=6) (
<figureCitation id="AD952A7ED972FFC4D563C0E5FD595CAB" box="[518,669,2012,2038]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–10" captionStart-2="FIGURES 11–15" captionStartId-0="4.[151,264,1943,1967]" captionStartId-1="5.[151,264,1943,1967]" captionStartId-2="6.[151,264,1943,1967]" captionTargetBox-0="[178,1392,189,1903]" captionTargetBox-1="[194,1389,191,1907]" captionTargetBox-2="[187,1394,193,1910]" captionTargetId-0="figure-17@4.[171,1416,181,1919]" captionTargetId-1="figure-27@5.[179,1408,181,1920]" captionTargetId-2="figure-17@6.[179,1408,181,1920]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." captionText-1="FIGURES 7–10. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 7, idiosoma in dorsal view; 8, reticulation of dorsal shield, general view of some dorsal setae, absence of setae S1; 9, idiosoma in ventral view; 10, sternal &amp; genital shields." captionText-2="FIGURES 11–15. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 11, subcapitulum; 12, distal portion of palp, with a focus on apotele; 13, insemination structures; male. 14, idiosoma in ventral view; 15, chelicera." figureDoi-0="http://doi.org/10.5281/zenodo.4636322" figureDoi-1="http://doi.org/10.5281/zenodo.4636326" figureDoi-2="http://doi.org/10.5281/zenodo.4636328" httpUri-0="https://zenodo.org/record/4636322/files/figure.png" httpUri-1="https://zenodo.org/record/4636326/files/figure.png" httpUri-2="https://zenodo.org/record/4636328/files/figure.png" pageId="1" pageNumber="241">Figures 1–13</figureCitation>
, 
<figureCitation id="AD952A7ED972FFC4D5CCC0E5FD345CAA" box="[681,752,2012,2039]" captionStart="FIGURES 16–19" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[187,1397,192,1902]" captionTargetId="figure-27@7.[167,1421,181,1919]" captionTargetPageId="7" captionText="FIGURES 16–19. Gaeolaelaps altaiensis Joharchi sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi="http://doi.org/10.5281/zenodo.4636330" httpUri="https://zenodo.org/record/4636330/files/figure.png" pageId="1" pageNumber="241">16–19</figureCitation>
)
</paragraph>
<paragraph id="351136FBD971FFC7D7A2C7AEFEFB5AF0" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="242">
<emphasis id="07DAEAE9D971FFC7D7A2C7AEFE475BEC" box="[199,387,151,177]" italics="true" pageId="2" pageNumber="242">Dorsal idiosoma</emphasis>
(
<figureCitation id="AD952A7ED971FFC7D6F7C7AEFE1D5BEC" box="[402,473,151,177]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Figs 1</figureCitation>
, 
<figureCitation id="AD952A7ED971FFC7D682C7AEFDD15BEC" box="[487,533,151,177]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[194,1389,191,1907]" captionTargetId="figure-27@5.[179,1408,181,1920]" captionTargetPageId="5" captionText="FIGURES 7–10. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 7, idiosoma in dorsal view; 8, reticulation of dorsal shield, general view of some dorsal setae, absence of setae S1; 9, idiosoma in ventral view; 10, sternal &amp; genital shields." figureDoi="http://doi.org/10.5281/zenodo.4636326" httpUri="https://zenodo.org/record/4636326/files/figure.png" pageId="2" pageNumber="242">7–8</figureCitation>
). Dorsal shield 445–465 long, 228–250 wide, covering most of dorsal idiosoma; distinct reticulation posterior to 
<emphasis id="07DAEAE9D971FFC7D562C783FDD95B89" box="[519,541,186,212]" italics="true" pageId="2" pageNumber="242">j6</emphasis>
(
<figureCitation id="AD952A7ED971FFC7D54BC782FDB35B88" box="[558,631,187,213]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Figs 1</figureCitation>
, 
<figureCitation id="AD952A7ED971FFC7D5E2C782FD705B88" box="[647,692,187,213]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[194,1389,191,1907]" captionTargetId="figure-27@5.[179,1408,181,1920]" captionTargetPageId="5" captionText="FIGURES 7–10. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 7, idiosoma in dorsal view; 8, reticulation of dorsal shield, general view of some dorsal setae, absence of setae S1; 9, idiosoma in ventral view; 10, sternal &amp; genital shields." figureDoi="http://doi.org/10.5281/zenodo.4636326" httpUri="https://zenodo.org/record/4636326/files/figure.png" pageId="2" pageNumber="242">7–8</figureCitation>
). Shield narrowing from level of setae 
<emphasis id="07DAEAE9D971FFC7D312C782FB555B88" box="[1143,1169,187,213]" italics="true" pageId="2" pageNumber="242">r3</emphasis>
, progressively tapering until 
<emphasis id="07DAEAE9D971FFC7D7B1C7E7FF295BA5" box="[212,237,222,248]" italics="true" pageId="2" pageNumber="242">s6</emphasis>
, then subparallel from 
<emphasis id="07DAEAE9D971FFC7D69FC7E7FDD75BA5" box="[506,531,222,248]" italics="true" pageId="2" pageNumber="242">r6</emphasis>
to level of 
<emphasis id="07DAEAE9D971FFC7D5FEC7E6FD7C5BA4" box="[667,696,223,249]" italics="true" pageId="2" pageNumber="242">S4</emphasis>
until rounded posteriorly and with 37 pairs of setae; 21 pairs of podonotal setae, 
<emphasis id="07DAEAE9D971FFC7D630C63AFEAA5A40" box="[341,366,259,285]" italics="true" pageId="2" pageNumber="242">z3</emphasis>
absent, 16 pairs of opisthonotal setae, including two pairs of 
<emphasis id="07DAEAE9D971FFC7D343C63AFB865A41" box="[1062,1090,259,284]" italics="true" pageId="2" pageNumber="242">Zx</emphasis>
setae, 
<emphasis id="07DAEAE9D971FFC7D3F5C63AFB685A40" box="[1168,1196,259,285]" italics="true" pageId="2" pageNumber="242">S1</emphasis>
absent and 
<emphasis id="07DAEAE9D971FFC7D256C63BFA8F5A41" box="[1331,1355,258,284]" italics="true" pageId="2" pageNumber="242">r6</emphasis>
on lateral soft cuticle. All setae smooth (28–38) except 
<emphasis id="07DAEAE9D971FFC7D5B7C61EFD285A1C" box="[722,748,295,321]" italics="true" pageId="2" pageNumber="242">J4</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D599C61EFCD25A1C" box="[764,790,295,321]" italics="true" pageId="2" pageNumber="242">J5</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D443C61EFC805A1C" box="[806,836,295,321]" italics="true" pageId="2" pageNumber="242">Z5</emphasis>
, sparsely barbed, see 
<figureCitation id="AD952A7ED971FFC7D324C61EFB6D5A1C" box="[1089,1193,295,321]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">
Fig. 1, 
<emphasis id="07DAEAE9D971FFC7D3F1C61EFB6D5A1C" box="[1172,1193,295,321]" italics="true" pageId="2" pageNumber="242">j1</emphasis>
</figureCitation>
(22–24), 
<emphasis id="07DAEAE9D971FFC7D27EC61EFAF05A1C" box="[1307,1332,295,321]" italics="true" pageId="2" pageNumber="242">z1</emphasis>
(25–27), 
<emphasis id="07DAEAE9D971FFC7D7F2C672FF715A38" box="[151,181,331,357]" italics="true" pageId="2" pageNumber="242">Z5</emphasis>
(54–56) (
<figureCitation id="AD952A7ED971FFC7D640C672FEA85A38" box="[293,364,331,357]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Figs 1</figureCitation>
, 
<figureCitation id="AD952A7ED971FFC7D61EC672FE6C5A38" box="[379,424,331,357]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[194,1389,191,1907]" captionTargetId="figure-27@5.[179,1408,181,1920]" captionTargetPageId="5" captionText="FIGURES 7–10. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 7, idiosoma in dorsal view; 8, reticulation of dorsal shield, general view of some dorsal setae, absence of setae S1; 9, idiosoma in ventral view; 10, sternal &amp; genital shields." figureDoi="http://doi.org/10.5281/zenodo.4636326" httpUri="https://zenodo.org/record/4636326/files/figure.png" pageId="2" pageNumber="242">7–8</figureCitation>
). Shield with about 20 pairs of discernible pore-like structures, including 16 poroids (
<emphasis id="07DAEAE9D971FFC7D214C672FA515A38" box="[1393,1429,331,357]" italics="true" pageId="2" pageNumber="242">id1</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D7F2C656FF7F5AD4" box="[151,187,367,393]" italics="true" pageId="2" pageNumber="242">id2</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D7ACC657FEDA5AD5" box="[201,286,366,392]" italics="true" pageId="2" pageNumber="242">id4–id6</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D648C657FE5C5AD5" box="[301,408,366,392]" italics="true" pageId="2" pageNumber="242">idm-idm6</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D6C3C656FE035AD4" box="[422,455,367,393]" italics="true" pageId="2" pageNumber="242">is1</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D6B0C656FE335AD4" box="[469,503,367,393]" italics="true" pageId="2" pageNumber="242">idx</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D560C656FDF55AD4" box="[517,561,367,393]" italics="true" pageId="2" pageNumber="242">idl1</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D55AC656FDAE5AD4" box="[575,618,367,393]" italics="true" pageId="2" pageNumber="242">idl3</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D51DC656FD605AD4" box="[632,676,367,393]" italics="true" pageId="2" pageNumber="242">idl4</emphasis>
) and four gland openings (
<emphasis id="07DAEAE9D971FFC7D4B1C656FC3B5AD4" box="[980,1023,367,393]" italics="true" pageId="2" pageNumber="242">gd5</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D369C657FBF25AD5" box="[1036,1078,366,392]" italics="true" pageId="2" pageNumber="242">gd6</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D321C656FBAA5AD4" box="[1092,1134,367,393]" italics="true" pageId="2" pageNumber="242">gd8</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D310C656FB5B5AD4" box="[1141,1183,367,393]" italics="true" pageId="2" pageNumber="242">gd9</emphasis>
), others indistinct, see 
<figureCitation id="AD952A7ED971FFC7D7F2C6AAFEC45AF0" box="[151,256,403,429]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Figures 1</figureCitation>
, 
<figureCitation id="AD952A7ED971FFC7D66BC6AAFEFF5AF0" box="[270,315,403,429]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[194,1389,191,1907]" captionTargetId="figure-27@5.[179,1408,181,1920]" captionTargetPageId="5" captionText="FIGURES 7–10. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 7, idiosoma in dorsal view; 8, reticulation of dorsal shield, general view of some dorsal setae, absence of setae S1; 9, idiosoma in ventral view; 10, sternal &amp; genital shields." figureDoi="http://doi.org/10.5281/zenodo.4636326" httpUri="https://zenodo.org/record/4636326/files/figure.png" pageId="2" pageNumber="242">7–8</figureCitation>
.
</paragraph>
<paragraph id="351136FBD971FFC7D7A2C68EFBF45E6C" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="242">
<emphasis id="07DAEAE9D971FFC7D7A2C68EFE4D5A8C" box="[199,393,439,465]" italics="true" pageId="2" pageNumber="242">Ventral idiosoma</emphasis>
(
<figureCitation id="AD952A7ED971FFC7D6FEC68EFE235A8C" box="[411,487,439,465]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Figs 2</figureCitation>
, 
<figureCitation id="AD952A7ED971FFC7D692C68EFDF75A8C" box="[503,563,439,465]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[194,1389,191,1907]" captionTargetId="figure-27@5.[179,1408,181,1920]" captionTargetPageId="5" captionText="FIGURES 7–10. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 7, idiosoma in dorsal view; 8, reticulation of dorsal shield, general view of some dorsal setae, absence of setae S1; 9, idiosoma in ventral view; 10, sternal &amp; genital shields." figureDoi="http://doi.org/10.5281/zenodo.4636326" httpUri="https://zenodo.org/record/4636326/files/figure.png" pageId="2" pageNumber="242">9–10</figureCitation>
). Tritosternum with paired pilose laciniae (79–82), fused basally (2–4), columnar base 25–27 × 12–14 wide; presternal area lightly sclerotised, punctate, with a few transverse curved lines, sternal shield length 119–122, maximum width 137–141, narrowest between coxae II (85–92), with reticulate ornamentation laterally, central part smooth or faintly reticulated (
<figureCitation id="AD952A7ED971FFC7D40EC51AFC7B5960" box="[875,959,547,573]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[194,1389,191,1907]" captionTargetId="figure-27@5.[179,1408,181,1920]" captionTargetPageId="5" captionText="FIGURES 7–10. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 7, idiosoma in dorsal view; 8, reticulation of dorsal shield, general view of some dorsal setae, absence of setae S1; 9, idiosoma in ventral view; 10, sternal &amp; genital shields." figureDoi="http://doi.org/10.5281/zenodo.4636326" httpUri="https://zenodo.org/record/4636326/files/figure.png" pageId="2" pageNumber="242">Fig. 10</figureCitation>
), posterior margin almost straigh, anterior margin slightly concave; bearing three pairs of smooth setae (
<emphasis id="07DAEAE9D971FFC7D400C57EFC42593C" box="[869,902,583,609]" italics="true" pageId="2" pageNumber="242">st1</emphasis>
32–34, 
<emphasis id="07DAEAE9D971FFC7D482C57EFBC3593C" box="[999,1031,583,609]" italics="true" pageId="2" pageNumber="242">st2</emphasis>
32–34, 
<emphasis id="07DAEAE9D971FFC7D30CC57EFB4E593C" box="[1129,1162,583,609]" italics="true" pageId="2" pageNumber="242">st3</emphasis>
31–33), never reaching base of next setae, one pair of lyrifissures adjacent to setae 
<emphasis id="07DAEAE9D971FFC7D44BC552FC8B59D8" box="[814,847,619,645]" italics="true" pageId="2" pageNumber="242">st1</emphasis>
, and a pair of poroids between 
<emphasis id="07DAEAE9D971FFC7D3CAC552FB1459D8" box="[1199,1232,619,645]" italics="true" pageId="2" pageNumber="242">st2</emphasis>
and 
<emphasis id="07DAEAE9D971FFC7D263C552FAE359D8" box="[1286,1319,619,645]" italics="true" pageId="2" pageNumber="242">st3</emphasis>
; remnants of gland pores 
<emphasis id="07DAEAE9D971FFC7D620C5B6FEA959F4" box="[325,365,655,681]" italics="true" pageId="2" pageNumber="242">gv1</emphasis>
indistinct. Metasternal setae 
<emphasis id="07DAEAE9D971FFC7D5DBC5B6FD1B59F4" box="[702,735,655,681]" italics="true" pageId="2" pageNumber="242">st4</emphasis>
(26–28) and metasternal poroids located on soft integument; metasternal platelets absent. Endopodal plates II/III completely fused to sternal shield, endopodal plates III/IV elongate, narrow and curved, abutting posterioir part of endopodal plates II/III. Genital shield tongue-shaped, slightly expanded posteriorly past setae 
<emphasis id="07DAEAE9D971FFC7D564C5C2FDE55848" box="[513,545,763,789]" italics="true" pageId="2" pageNumber="242">st5</emphasis>
, length 166–180, maximum width 65–70, anterior margin of shield irregularly convex, overlapping posterior area of sternal shield, but not reaching base of setae 
<emphasis id="07DAEAE9D971FFC7D322C426FBAC5864" box="[1095,1128,799,825]" italics="true" pageId="2" pageNumber="242">st3</emphasis>
, posterior margin rounded, surface reticulated with irregular longitudinal lines in anterior region, posteriorly with 5–6 faint cells enclosed behind an inverted-V shaped ridge; bearing a pair of simple setae 
<emphasis id="07DAEAE9D971FFC7D40CC45EFC4E58DC" box="[873,906,871,897]" italics="true" pageId="2" pageNumber="242">st5</emphasis>
(24–26) (
<figureCitation id="AD952A7ED971FFC7D49EC45EFB8B58DC" box="[1019,1103,871,897]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[194,1389,191,1907]" captionTargetId="figure-27@5.[179,1408,181,1920]" captionTargetPageId="5" captionText="FIGURES 7–10. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 7, idiosoma in dorsal view; 8, reticulation of dorsal shield, general view of some dorsal setae, absence of setae S1; 9, idiosoma in ventral view; 10, sternal &amp; genital shields." figureDoi="http://doi.org/10.5281/zenodo.4636326" httpUri="https://zenodo.org/record/4636326/files/figure.png" pageId="2" pageNumber="242">Fig. 10</figureCitation>
). Shield flanked by two pairs of minute, elongate paragenital platelets; paragenital poroids 
<emphasis id="07DAEAE9D971FFC7D435C4B2FCB658F8" box="[848,882,907,933]" italics="true" pageId="2" pageNumber="242">iv5</emphasis>
located on soft cuticle lateral to shield near seta 
<emphasis id="07DAEAE9D971FFC7D7F2C496FF7C5894" box="[151,184,943,969]" italics="true" pageId="2" pageNumber="242">st5</emphasis>
. Anal shield subtriangular, rounded anteriorly, length 68–76, width 68–73, anterior half lineate-reticulate, paraanal setae (27–29) shorter than post-anal seta (35–37), cribrum consisting of a terminal tuft with 3–4 irregular rows of spicules and a pair of anterior arms extending slightly beyond post-anal seta (
<figureCitation id="AD952A7ED971FFC7D346C4CEFBA85F4C" box="[1059,1132,1015,1041]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Figs 2</figureCitation>
, 
<figureCitation id="AD952A7ED971FFC7D31CC4CEFB4D5F4C" box="[1145,1161,1015,1041]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[194,1389,191,1907]" captionTargetId="figure-27@5.[179,1408,181,1920]" captionTargetPageId="5" captionText="FIGURES 7–10. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 7, idiosoma in dorsal view; 8, reticulation of dorsal shield, general view of some dorsal setae, absence of setae S1; 9, idiosoma in ventral view; 10, sternal &amp; genital shields." figureDoi="http://doi.org/10.5281/zenodo.4636326" httpUri="https://zenodo.org/record/4636326/files/figure.png" pageId="2" pageNumber="242">9</figureCitation>
); anal gland pores (
<emphasis id="07DAEAE9D971FFC7D20FC4CEFA575F4C" box="[1386,1427,1015,1041]" italics="true" pageId="2" pageNumber="242">gv3</emphasis>
) on anterolateral margin of anal shield. Soft opisthogastric cuticle with pair of oval metapodal plates (18–20 long × 5–7 wide), an additional small round metapodal platelet more medially and eight pairs of smooth setae (
<emphasis id="07DAEAE9D971FFC7D25FC306FA525F04" box="[1338,1430,1087,1113]" italics="true" pageId="2" pageNumber="242">Jv1–Jv5</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D7F2C35AFF2F5F21" box="[151,235,1123,1149]" italics="true" pageId="2" pageNumber="242">Zv1–Zv</emphasis>
3) (23–26), 
<emphasis id="07DAEAE9D971FFC7D616C35AFE5E5F20" box="[371,410,1123,1149]" italics="true" pageId="2" pageNumber="242">Jv5</emphasis>
(36–38), and six pairs of poroids including 
<emphasis id="07DAEAE9D971FFC7D4F5C35AFC765F20" box="[912,946,1123,1149]" italics="true" pageId="2" pageNumber="242">iv5</emphasis>
and 
<emphasis id="07DAEAE9D971FFC7D48EC35AFBC95F21" box="[1003,1037,1123,1148]" italics="true" pageId="2" pageNumber="242">ivp</emphasis>
. Exopodal platelets between coxae II–III subtriangular, those behind coxae III–IV and parapodal platelets fused, curved, extending narrowly behind coxae IV, surrounding coxa IV, gland pores 
<emphasis id="07DAEAE9D971FFC7D51AC392FD6C5F98" box="[639,680,1195,1221]" italics="true" pageId="2" pageNumber="242">gv2</emphasis>
on parapodal platelets (
<figureCitation id="AD952A7ED971FFC7D4D7C392FC3E5F98" box="[946,1018,1195,1221]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Figs 2</figureCitation>
, 
<figureCitation id="AD952A7ED971FFC7D363C392FBD25F98" box="[1030,1046,1195,1221]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[194,1389,191,1907]" captionTargetId="figure-27@5.[179,1408,181,1920]" captionTargetPageId="5" captionText="FIGURES 7–10. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 7, idiosoma in dorsal view; 8, reticulation of dorsal shield, general view of some dorsal setae, absence of setae S1; 9, idiosoma in ventral view; 10, sternal &amp; genital shields." figureDoi="http://doi.org/10.5281/zenodo.4636326" httpUri="https://zenodo.org/record/4636326/files/figure.png" pageId="2" pageNumber="242">9</figureCitation>
). Peritreme extending anteriorly to mid-level of coxa II; peritrematal shield narrow, expanded anteriorly, fused to dorsal shield behind setae 
<emphasis id="07DAEAE9D971FFC7D25BC3F6FA935FB4" box="[1342,1367,1231,1257]" italics="true" pageId="2" pageNumber="242">z1</emphasis>
level, free from exopodal shields, bearing five pore-like structures, a lyrifissure 
<emphasis id="07DAEAE9D971FFC7D4ABC3CAFC205E51" box="[974,996,1267,1292]" italics="true" pageId="2" pageNumber="242">ip</emphasis>
and a gland pore 
<emphasis id="07DAEAE9D971FFC7D3CCC3CAFB015E51" box="[1193,1221,1267,1292]" italics="true" pageId="2" pageNumber="242">gp</emphasis>
at level near coxae II–III, two lyrifissures 
<emphasis id="07DAEAE9D971FFC7D6F3C22EFE685E6D" box="[406,428,1303,1328]" italics="true" pageId="2" pageNumber="242">ip</emphasis>
and a gland pore 
<emphasis id="07DAEAE9D971FFC7D511C22EFD545E6D" box="[628,656,1303,1328]" italics="true" pageId="2" pageNumber="242">gp</emphasis>
on post-stigmatic section (
<figureCitation id="AD952A7ED971FFC7D4DBC22EFBC25E6C" box="[958,1030,1303,1329]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Figs 2</figureCitation>
, 
<figureCitation id="AD952A7ED971FFC7D377C22EFBE65E6C" box="[1042,1058,1303,1329]" captionStart="FIGURES 7–10" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[194,1389,191,1907]" captionTargetId="figure-27@5.[179,1408,181,1920]" captionTargetPageId="5" captionText="FIGURES 7–10. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 7, idiosoma in dorsal view; 8, reticulation of dorsal shield, general view of some dorsal setae, absence of setae S1; 9, idiosoma in ventral view; 10, sternal &amp; genital shields." figureDoi="http://doi.org/10.5281/zenodo.4636326" httpUri="https://zenodo.org/record/4636326/files/figure.png" pageId="2" pageNumber="242">9</figureCitation>
).
</paragraph>
<paragraph id="351136FBD971FFC7D7A2C202FD475DBF" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="242">
<taxonomicName id="F2AE4D78D971FFC7D7A2C202FE925E08" authorityName="Krefft" authorityYear="1871" box="[199,342,1339,1365]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="242" phylum="Nematoda" rank="genus">
<emphasis id="07DAEAE9D971FFC7D7A2C202FE925E08" box="[199,342,1339,1365]" italics="true" pageId="2" pageNumber="242">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="AD952A7ED971FFC7D601C202FE0C5E08" box="[356,456,1339,1365]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Figs 3–6</figureCitation>
, 
<figureCitation id="AD952A7ED971FFC7D6B7C202FDDF5E08" box="[466,539,1339,1365]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[187,1394,193,1910]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 11–15. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 11, subcapitulum; 12, distal portion of palp, with a focus on apotele; 13, insemination structures; male. 14, idiosoma in ventral view; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636328" httpUri="https://zenodo.org/record/4636328/files/figure.png" pageId="2" pageNumber="242">11–12</figureCitation>
). Epistome subtriangular and irregularly denticulate (
<figureCitation id="AD952A7ED971FFC7D30CC202FB685E08" box="[1129,1196,1338,1365]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Fig. 5</figureCitation>
). Hypostomal groove with six transverse rows of denticles, each row with 7–12 small denticles, with smooth anterior and posterior transverse lines, groove very slightly wider anteriorly. Hypostome with four pairs of setae, internal posterior hypostomal setae 
<emphasis id="07DAEAE9D971FFC7D7B3C29EFF365E9C" box="[214,242,1447,1473]" italics="true" pageId="2" pageNumber="242">h3</emphasis>
(31–33), 
<emphasis id="07DAEAE9D971FFC7D63AC29EFEBE5E9C" box="[351,378,1447,1473]" italics="true" pageId="2" pageNumber="242">h1</emphasis>
(28–30), 
<emphasis id="07DAEAE9D971FFC7D682C29EFDC05E9C" box="[487,516,1447,1473]" italics="true" pageId="2" pageNumber="242">h2</emphasis>
(20–22), palpcoxal setae (
<emphasis id="07DAEAE9D971FFC7D44EC29EFC815E9D" box="[811,837,1447,1472]" italics="true" pageId="2" pageNumber="242">pc</emphasis>
) (23–25) (
<figureCitation id="AD952A7ED971FFC7D4D8C29EFBC05E9C" box="[957,1028,1447,1473]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Figs 3</figureCitation>
, 
<figureCitation id="AD952A7ED971FFC7D374C29EFBEA5E9C" box="[1041,1070,1447,1473]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[187,1394,193,1910]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 11–15. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 11, subcapitulum; 12, distal portion of palp, with a focus on apotele; 13, insemination structures; male. 14, idiosoma in ventral view; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636328" httpUri="https://zenodo.org/record/4636328/files/figure.png" pageId="2" pageNumber="242">11</figureCitation>
). Corniculi robust and horn-like, extending slightly beyond palptrochanter. Supralabral process with apically tri-tined anterior projection (
<figureCitation id="AD952A7ED971FFC7D222C2F2FA4B5EB8" box="[1351,1423,1483,1509]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Fig. 4</figureCitation>
). Internal malae with one pair of smooth median projections, flanked by lobes with fimbriate anterior margin; labrum with pilose surface (
<figureCitation id="AD952A7ED971FFC7D61CC12AFE055D70" box="[377,449,1555,1581]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Figs 3</figureCitation>
, 
<figureCitation id="AD952A7ED971FFC7D6ABC12AFE2F5D70" box="[462,491,1555,1581]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[187,1394,193,1910]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 11–15. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 11, subcapitulum; 12, distal portion of palp, with a focus on apotele; 13, insemination structures; male. 14, idiosoma in ventral view; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636328" httpUri="https://zenodo.org/record/4636328/files/figure.png" pageId="2" pageNumber="242">11</figureCitation>
). Chaetotaxy of palps (
<figureCitation id="AD952A7ED971FFC7D58AC12AFC805D70" box="[751,836,1555,1581]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[187,1394,193,1910]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 11–15. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 11, subcapitulum; 12, distal portion of palp, with a focus on apotele; 13, insemination structures; male. 14, idiosoma in ventral view; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636328" httpUri="https://zenodo.org/record/4636328/files/figure.png" pageId="2" pageNumber="242">Fig. 12</figureCitation>
): trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth and needle-like except 
<emphasis id="07DAEAE9D971FFC7D54AC10EFD975D0C" box="[559,595,1591,1617]" italics="true" pageId="2" pageNumber="242">al1</emphasis>
and 
<emphasis id="07DAEAE9D971FFC7D5E2C10EFD6F5D0C" box="[647,683,1591,1617]" italics="true" pageId="2" pageNumber="242">al2</emphasis>
on palpgenu thickened, paddle-like and 
<emphasis id="07DAEAE9D971FFC7D308C10EFB475D0C" box="[1133,1155,1591,1617]" italics="true" pageId="2" pageNumber="242">al</emphasis>
on palp palpfemur thickened, 
<emphasis id="07DAEAE9D971FFC7D7BEC162FF335D28" box="[219,247,1627,1653]" italics="true" pageId="2" pageNumber="242">d3</emphasis>
inserted on small tubercles, palp tarsal apotele two-tined (
<figureCitation id="AD952A7ED971FFC7D4EFC162FC1A5D28" box="[906,990,1627,1653]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[187,1394,193,1910]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 11–15. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 11, subcapitulum; 12, distal portion of palp, with a focus on apotele; 13, insemination structures; male. 14, idiosoma in ventral view; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636328" httpUri="https://zenodo.org/record/4636328/files/figure.png" pageId="2" pageNumber="242">Fig. 12</figureCitation>
). Fixed digit of chelicera with an offset distal tooth (gabelzahn), followed by six variously sized teeth, a setaceous pilus dentilis, dorsal cheliceral setae prostrate, arthrodial membrane with a rounded flap and normal filaments; cheliceral lyrifissures distinct; movable digit with two relatively large teeth (
<figureCitation id="AD952A7ED971FFC7D555C1FEFDB15DBF" box="[560,629,1735,1762]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1392,189,1903]" captionTargetId="figure-17@4.[171,1416,181,1919]" captionTargetPageId="4" captionText="FIGURES 1–6. Gaeolaelaps altaiensis Joharchi sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, supralabral process; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636322" httpUri="https://zenodo.org/record/4636322/files/figure.png" pageId="2" pageNumber="242">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="351136FBD971FFC6D7A2C1D2FD215A40" blockId="2.[151,1437,150,2013]" lastBlockId="3.[151,1437,151,1905]" lastPageId="3" lastPageNumber="243" pageId="2" pageNumber="242">
<emphasis id="07DAEAE9D971FFC7D7A2C1D2FF385C59" box="[199,252,1771,1796]" italics="true" pageId="2" pageNumber="242">Legs</emphasis>
(
<figureCitation id="AD952A7ED971FFC7D669C1D2FE4B5C5B" box="[268,399,1771,1798]" captionStart="FIGURES 16–19" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[187,1397,192,1902]" captionTargetId="figure-27@7.[167,1421,181,1919]" captionTargetPageId="7" captionText="FIGURES 16–19. Gaeolaelaps altaiensis Joharchi sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi="http://doi.org/10.5281/zenodo.4636330" httpUri="https://zenodo.org/record/4636330/files/figure.png" pageId="2" pageNumber="242">Figs 16–19</figureCitation>
). Legs II (293–297) and III (245–250) short, I (427–434) and IV (380–390) longer. Chaetotaxy normal for free-living 
<taxonomicName id="F2AE4D78D971FFC7D6A8C036FD885C74" box="[461,588,1807,1833]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="242" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
: Leg I (
<figureCitation id="AD952A7ED971FFC7D5C1C036FD3D5C77" box="[676,761,1807,1834]" captionStart="FIGURES 16–19" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[187,1397,192,1902]" captionTargetId="figure-27@7.[167,1421,181,1919]" captionTargetPageId="7" captionText="FIGURES 16–19. Gaeolaelaps altaiensis Joharchi sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi="http://doi.org/10.5281/zenodo.4636330" httpUri="https://zenodo.org/record/4636330/files/figure.png" pageId="2" pageNumber="242">Fig. 16</figureCitation>
): coxa 0–0/1, 0/1–0, trochanter 1–1/1, 0/2–1 (
<emphasis id="07DAEAE9D971FFC7D264C036FAD35C74" box="[1281,1303,1807,1833]" italics="true" pageId="2" pageNumber="242">al</emphasis>
thickened), femur 2–2/1, 3/3–2 (
<emphasis id="07DAEAE9D971FFC7D61AC00AFE6D5C10" box="[383,425,1843,1869]" italics="true" pageId="2" pageNumber="242">ad1</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D6D3C00AFE245C10" box="[438,480,1843,1869]" italics="true" pageId="2" pageNumber="242">pd2</emphasis>
, 
<emphasis id="07DAEAE9D971FFC7D68BC00AFDDC5C10" box="[494,536,1843,1869]" italics="true" pageId="2" pageNumber="242">pd3</emphasis>
slightly thickened), genu 2–3/2, 3/1–2, tibia 2–3/2, 3/1–2. Leg II (
<figureCitation id="AD952A7ED971FFC7D39AC00AFA965C10" box="[1279,1362,1843,1869]" captionStart="FIGURES 16–19" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[187,1397,192,1902]" captionTargetId="figure-27@7.[167,1421,181,1919]" captionTargetPageId="7" captionText="FIGURES 16–19. Gaeolaelaps altaiensis Joharchi sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi="http://doi.org/10.5281/zenodo.4636330" httpUri="https://zenodo.org/record/4636330/files/figure.png" pageId="2" pageNumber="242">Fig. 17</figureCitation>
): coxa 0–0/1, 0/1–0, trochanter 1–0/2, 0/1–1, femur 2–3/1, 2/2–1 (
<emphasis id="07DAEAE9D971FFC7D456C06EFC995C2C" box="[819,861,1879,1905]" italics="true" pageId="2" pageNumber="242">pd1</emphasis>
longer), genu 2–3/1, 2/1–2 (
<emphasis id="07DAEAE9D971FFC7D3C5C06EFB7E5C2D" box="[1184,1210,1879,1904]" italics="true" pageId="2" pageNumber="242">av</emphasis>
thickened, 
<emphasis id="07DAEAE9D971FFC7D25BC06EFAA35C2C" box="[1342,1383,1879,1905]" italics="true" pageId="2" pageNumber="242">pd2</emphasis>
longer), tibia 2–2/1, 2/1–2 (all ventral setae thickened). Leg III (
<figureCitation id="AD952A7ED971FFC7D42FC042FC5A5CC8" box="[842,926,1915,1941]" captionStart="FIGURES 16–19" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[187,1397,192,1902]" captionTargetId="figure-27@7.[167,1421,181,1919]" captionTargetPageId="7" captionText="FIGURES 16–19. Gaeolaelaps altaiensis Joharchi sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi="http://doi.org/10.5281/zenodo.4636330" httpUri="https://zenodo.org/record/4636330/files/figure.png" pageId="2" pageNumber="242">Fig. 18</figureCitation>
): coxa 0–0/1, 0/1–0, trochanter 1–1/1, 0/2–0, femur 1–2/1, 1/0–1 (all dorsal setae thickened), genu 2–2/1, 2/1–1, tibia: 2–1/1, 2/1–1 (all ventral setae thickened). Leg IV (
<figureCitation id="AD952A7ED971FFC7D79DC0FAFE885C80" box="[248,332,1987,2013]" captionStart="FIGURES 16–19" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[187,1397,192,1902]" captionTargetId="figure-27@7.[167,1421,181,1919]" captionTargetPageId="7" captionText="FIGURES 16–19. Gaeolaelaps altaiensis Joharchi sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi="http://doi.org/10.5281/zenodo.4636330" httpUri="https://zenodo.org/record/4636330/files/figure.png" pageId="2" pageNumber="242">Fig. 19</figureCitation>
): coxa 0–0/1, 0/0–0, trochanter 1–1/1, 0/2–0, femur 1–2/1, 1/0–1 (
<emphasis id="07DAEAE9D971FFC7D35EC0FAFBA25C80" box="[1083,1126,1987,2013]" italics="true" pageId="2" pageNumber="242">ad2</emphasis>
and 
<emphasis id="07DAEAE9D971FFC7D3F8C0FAFB7D5C80" box="[1181,1209,1987,2013]" italics="true" pageId="2" pageNumber="242">pd</emphasis>
thickened; 
<emphasis id="07DAEAE9D971FFC7D258C0FAFAA35C80" box="[1341,1383,1987,2013]" italics="true" pageId="2" pageNumber="242">ad1</emphasis>
longest), genu 2–2/1, 3/0–1, tibia 2–1/1, 3/1–2 (
<emphasis id="07DAEAE9D970FFC6D5EFC7AEFD605BED" box="[650,676,151,176]" italics="true" pageId="3" pageNumber="243">pv</emphasis>
thickened). Tarsi II-IV with 18 setae (3– 3/2, 3/2–3 + 
<emphasis id="07DAEAE9D970FFC6D267C7AEFAE75BED" box="[1282,1315,151,176]" italics="true" pageId="3" pageNumber="243">mv</emphasis>
, 
<emphasis id="07DAEAE9D970FFC6D255C7AEFA965BEC" box="[1328,1362,151,177]" italics="true" pageId="3" pageNumber="243">md</emphasis>
); with thickened some ventral and lateral setae, see 
<figureCitation id="AD952A7ED970FFC6D5EEC782FCEA5B88" box="[651,814,187,213]" captionStart="FIGURES 16–19" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[187,1397,192,1902]" captionTargetId="figure-27@7.[167,1421,181,1919]" captionTargetPageId="7" captionText="FIGURES 16–19. Gaeolaelaps altaiensis Joharchi sp. nov., female. 16, leg I (trochanter-tibia); 17, leg II; 18, leg III; 19, leg IV." figureDoi="http://doi.org/10.5281/zenodo.4636330" httpUri="https://zenodo.org/record/4636330/files/figure.png" pageId="3" pageNumber="243">Figures 17–19</figureCitation>
. Tarsus IV with two noticeably long setae (macrosetae), 
<emphasis id="07DAEAE9D970FFC6D7F2C7E6FF055BA4" box="[151,193,223,249]" italics="true" pageId="3" pageNumber="243">pd2</emphasis>
, 
<emphasis id="07DAEAE9D970FFC6D7B5C7E6FF3D5BA4" box="[208,249,223,249]" italics="true" pageId="3" pageNumber="243">pd3</emphasis>
(60–63) significantly longer than others, 
<emphasis id="07DAEAE9D970FFC6D5AAC7E6FD375BA4" box="[719,755,223,249]" italics="true" pageId="3" pageNumber="243">pl2</emphasis>
, 
<emphasis id="07DAEAE9D970FFC6D464C7E6FCE15BA4" box="[769,805,223,249]" italics="true" pageId="3" pageNumber="243">pl3</emphasis>
thick and blunt. All pretarsi with well-developed paired claws, rounded pulvilli and normal ambulacral stalk.
</paragraph>
<paragraph id="351136FBD970FFC6D7A2C61EFE0C5AD4" blockId="3.[151,1437,151,1905]" pageId="3" pageNumber="243">
<emphasis id="07DAEAE9D970FFC6D7A2C61EFE1F5A1C" bold="true" box="[199,475,295,321]" italics="true" pageId="3" pageNumber="243">Insemination structures</emphasis>
(
<figureCitation id="AD952A7ED970FFC6D68EC61EFDFA5A1C" box="[491,574,295,321]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[187,1394,193,1910]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 11–15. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 11, subcapitulum; 12, distal portion of palp, with a focus on apotele; 13, insemination structures; male. 14, idiosoma in ventral view; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636328" httpUri="https://zenodo.org/record/4636328/files/figure.png" pageId="3" pageNumber="243">Fig. 13</figureCitation>
). Laelapid-type sperm access system, tubulus long, wider at the solenostome level, mesad of coxa III, and entering sacculus. Proximal ends of the tubulus swollen at junction with ramus, leading to droplet-shaped sacculus.
</paragraph>
<paragraph id="351136FBD970FFC6D7A2C6AAFD385AF0" blockId="3.[151,1437,151,1905]" box="[199,764,402,429]" pageId="3" pageNumber="243">
<emphasis id="07DAEAE9D970FFC6D7A2C6AAFE895AF0" bold="true" box="[199,333,403,429]" italics="true" pageId="3" pageNumber="243">Description</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D63EC6AAFE295AF1" bold="true" box="[347,493,402,429]" pageId="3" pageNumber="243">Male (n= 2).</emphasis>
(
<figureCitation id="AD952A7ED970FFC6D69EC6AAFD645AF0" box="[507,672,402,429]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[187,1394,193,1910]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 11–15. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 11, subcapitulum; 12, distal portion of palp, with a focus on apotele; 13, insemination structures; male. 14, idiosoma in ventral view; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636328" httpUri="https://zenodo.org/record/4636328/files/figure.png" pageId="3" pageNumber="243">Figures 14–15</figureCitation>
, 
<figureCitation id="AD952A7ED970FFC6D5C9C6AAFD305AF0" box="[684,756,403,429]" captionStart="FIGURES 20–21" captionStartId="8.[151,264,1497,1521]" captionTargetBox="[174,1422,206,1456]" captionTargetId="figure-17@8.[151,1436,181,1473]" captionTargetPageId="8" captionText="FIGURES 20–21. Gaeolaelaps altaiensis Joharchi sp. nov., male. 20, ventral idiosoma; 21, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636332" httpUri="https://zenodo.org/record/4636332/files/figure.png" pageId="3" pageNumber="243">20–21</figureCitation>
)
</paragraph>
<paragraph id="351136FBD970FFC6D7A2C68EFAAF5A8C" blockId="3.[151,1437,151,1905]" box="[199,1387,438,465]" pageId="3" pageNumber="243">
<emphasis id="07DAEAE9D970FFC6D7A2C68EFE475A8C" box="[199,387,439,465]" italics="true" pageId="3" pageNumber="243">Dorsal idiosoma</emphasis>
. Dorsal shield 354–358 long, 208–213 wide; ornamentation and chaetotaxy as in female.
</paragraph>
<paragraph id="351136FBD970FFC6D7A2C6E2FA885864" blockId="3.[151,1437,151,1905]" pageId="3" pageNumber="243">
<emphasis id="07DAEAE9D970FFC6D7A2C6E2FE4E5AA8" box="[199,394,475,501]" italics="true" pageId="3" pageNumber="243">Ventral idiosoma</emphasis>
(
<figureCitation id="AD952A7ED970FFC6D6F9C6E2FE325AA8" box="[412,502,475,501]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[187,1394,193,1910]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 11–15. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 11, subcapitulum; 12, distal portion of palp, with a focus on apotele; 13, insemination structures; male. 14, idiosoma in ventral view; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636328" httpUri="https://zenodo.org/record/4636328/files/figure.png" pageId="3" pageNumber="243">Figs 14</figureCitation>
, 
<figureCitation id="AD952A7ED970FFC6D563C6E2FDE25AA8" box="[518,550,475,501]" captionStart="FIGURES 20–21" captionStartId="8.[151,264,1497,1521]" captionTargetBox="[174,1422,206,1456]" captionTargetId="figure-17@8.[151,1436,181,1473]" captionTargetPageId="8" captionText="FIGURES 20–21. Gaeolaelaps altaiensis Joharchi sp. nov., male. 20, ventral idiosoma; 21, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636332" httpUri="https://zenodo.org/record/4636332/files/figure.png" pageId="3" pageNumber="243">20</figureCitation>
). Sternal, genital, endopodal, ventral and anal shields fused into holoventral shield, 305–308 long from anterior to posterior margins of shield, (79–82) wide at level of 
<emphasis id="07DAEAE9D970FFC6D3F1C6C6FB705944" box="[1172,1204,511,537]" italics="true" pageId="3" pageNumber="243">st2</emphasis>
, (82–84) at 
<emphasis id="07DAEAE9D970FFC6D25BC6C6FA9B5944" box="[1342,1375,511,537]" italics="true" pageId="3" pageNumber="243">st3</emphasis>
level and (141–145) at broadest point (behind parapodals), anterolateral edges of shield fused to endopodals between coxae I–II; shield surface weakly reticulate, more distinct distinct behind 
<emphasis id="07DAEAE9D970FFC6D4BEC57EFC3F593C" box="[987,1019,583,609]" italics="true" pageId="3" pageNumber="243">st5</emphasis>
, with five pairs smooth sternal setae (
<emphasis id="07DAEAE9D970FFC6D7C4C552FF1A59D8" box="[161,222,619,645]" italics="true" pageId="3" pageNumber="243">st1–5</emphasis>
), and three pairs of smooth ventral setae (
<emphasis id="07DAEAE9D970FFC6D5DCC552FD2459D8" box="[697,736,619,645]" italics="true" pageId="3" pageNumber="243">Jv1</emphasis>
, 
<emphasis id="07DAEAE9D970FFC6D58BC552FCDC59D8" box="[750,792,619,645]" italics="true" pageId="3" pageNumber="243">Zv1</emphasis>
, 
<emphasis id="07DAEAE9D970FFC6D442C552FC9559D8" box="[807,849,619,645]" italics="true" pageId="3" pageNumber="243">Zv2</emphasis>
), plus three smooth circumanal setae, post-anal seta (27–29) longer and slightly thicker than para-anals (21–23), with five pairs of poroids and a pair of gland pores (
<emphasis id="07DAEAE9D970FFC6D7C4C58AFF0D5990" box="[161,201,691,717]" italics="true" pageId="3" pageNumber="243">gv3</emphasis>
) on lateral margins of shield at anterior level of anal opening, 
<emphasis id="07DAEAE9D970FFC6D4EDC58AFC6E5990" box="[904,938,691,717]" italics="true" pageId="3" pageNumber="243">iv1</emphasis>
slit-like; gland pore 
<emphasis id="07DAEAE9D970FFC6D3F0C58AFB795990" box="[1173,1213,691,717]" italics="true" pageId="3" pageNumber="243">gv2</emphasis>
behind coxa IV not discerned; cribrum with 3–4 irregular rows of spicules, restricted to region posterior to post-anal seta; metapodal platelets narrow and fused to shield, occasionally one side free. Soft opisthogastric cuticle with two pairs of simple setae, and four pairs of poroids. Peritremes and peritrematal shields similar to those in female (
<figureCitation id="AD952A7ED970FFC6D3D8C426FAD75864" box="[1213,1299,799,825]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[187,1394,193,1910]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 11–15. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 11, subcapitulum; 12, distal portion of palp, with a focus on apotele; 13, insemination structures; male. 14, idiosoma in ventral view; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636328" httpUri="https://zenodo.org/record/4636328/files/figure.png" pageId="3" pageNumber="243">Figs 14</figureCitation>
, 
<figureCitation id="AD952A7ED970FFC6D27AC426FAFB5864" box="[1311,1343,799,825]" captionStart="FIGURES 20–21" captionStartId="8.[151,264,1497,1521]" captionTargetBox="[174,1422,206,1456]" captionTargetId="figure-17@8.[151,1436,181,1473]" captionTargetPageId="8" captionText="FIGURES 20–21. Gaeolaelaps altaiensis Joharchi sp. nov., male. 20, ventral idiosoma; 21, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636332" httpUri="https://zenodo.org/record/4636332/files/figure.png" pageId="3" pageNumber="243">20</figureCitation>
).
</paragraph>
<paragraph id="351136FBD970FFC6D7A2C47AFECD58B0" blockId="3.[151,1437,151,1905]" pageId="3" pageNumber="243">
<taxonomicName id="F2AE4D78D970FFC6D7A2C47AFE925800" authorityName="Krefft" authorityYear="1871" box="[199,342,835,861]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="3" pageNumber="243" phylum="Nematoda" rank="genus">
<emphasis id="07DAEAE9D970FFC6D7A2C47AFE925800" box="[199,342,835,861]" italics="true" pageId="3" pageNumber="243">Gnathosoma</emphasis>
</taxonomicName>
. Epistome and subcapitulum similar to female. Fixed digit of chelicera with an offset distal tooth (gabelzahn), followed by 4–5 variously sized teeth and slender pilus dentilis. Movable digit of chelicera unidentate, spermadactyl stout, curved upward, with a large distal knob, and its free section more or less as long as movable digit (
<figureCitation id="AD952A7ED970FFC6D7BFC496FEF55894" box="[218,305,942,969]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[187,1394,193,1910]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 11–15. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 11, subcapitulum; 12, distal portion of palp, with a focus on apotele; 13, insemination structures; male. 14, idiosoma in ventral view; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636328" httpUri="https://zenodo.org/record/4636328/files/figure.png" pageId="3" pageNumber="243">Figs 15</figureCitation>
, 
<figureCitation id="AD952A7ED970FFC6D65BC496FE995894" box="[318,349,943,969]" captionStart="FIGURES 20–21" captionStartId="8.[151,264,1497,1521]" captionTargetBox="[174,1422,206,1456]" captionTargetId="figure-17@8.[151,1436,181,1473]" captionTargetPageId="8" captionText="FIGURES 20–21. Gaeolaelaps altaiensis Joharchi sp. nov., male. 20, ventral idiosoma; 21, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636332" httpUri="https://zenodo.org/record/4636332/files/figure.png" pageId="3" pageNumber="243">21</figureCitation>
); fringed hyaline arthrodial process at base of movable digit (
<figureCitation id="AD952A7ED970FFC6D376C496FBAE5894" box="[1043,1130,942,969]" captionStart="FIGURES 11–15" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[187,1394,193,1910]" captionTargetId="figure-17@6.[179,1408,181,1920]" captionTargetPageId="6" captionText="FIGURES 11–15. DIC micrographs of Gaeolaelaps altaiensis Joharchi sp. nov., female. 11, subcapitulum; 12, distal portion of palp, with a focus on apotele; 13, insemination structures; male. 14, idiosoma in ventral view; 15, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636328" httpUri="https://zenodo.org/record/4636328/files/figure.png" pageId="3" pageNumber="243">Figs 15</figureCitation>
, 
<figureCitation id="AD952A7ED970FFC6D313C496FB515894" box="[1142,1173,943,969]" captionStart="FIGURES 20–21" captionStartId="8.[151,264,1497,1521]" captionTargetBox="[174,1422,206,1456]" captionTargetId="figure-17@8.[151,1436,181,1473]" captionTargetPageId="8" captionText="FIGURES 20–21. Gaeolaelaps altaiensis Joharchi sp. nov., male. 20, ventral idiosoma; 21, chelicera." figureDoi="http://doi.org/10.5281/zenodo.4636332" httpUri="https://zenodo.org/record/4636332/files/figure.png" pageId="3" pageNumber="243">21</figureCitation>
). Palps similar to those of female.
</paragraph>
<paragraph id="351136FBD970FFC6D7A2C4CEFDE45F4C" blockId="3.[151,1437,151,1905]" box="[199,544,1015,1041]" pageId="3" pageNumber="243">
<emphasis id="07DAEAE9D970FFC6D7A2C4CEFF385F4D" box="[199,252,1015,1040]" italics="true" pageId="3" pageNumber="243">Legs</emphasis>
. Chaetotaxy as in female.
</paragraph>
</subSubSection>
<subSubSection id="7DB46570D970FFC6D7A2C322FABA5C2C" pageId="3" pageNumber="243" type="etymology">
<paragraph id="351136FBD970FFC6D7A2C322FE875F04" blockId="3.[151,1437,151,1905]" pageId="3" pageNumber="243">
<emphasis id="07DAEAE9D970FFC6D7A2C322FE8A5F68" bold="true" box="[199,334,1051,1077]" pageId="3" pageNumber="243">Etymology.</emphasis>
The specific name is derived from the name of the region (
<collectingRegion id="F76AF819D970FFC6D360C322FBFB5F68" box="[1029,1087,1051,1077]" country="Russia" name="Altay" pageId="3" pageNumber="243">Altai</collectingRegion>
Mountains) in which the 
<typeStatus id="EA158859D970FFC6D209C322FA585F68" box="[1388,1436,1051,1077]" pageId="3" pageNumber="243">type</typeStatus>
locality occurs.
</paragraph>
<paragraph id="351136FBD970FFC6D7A2C35AFABA5C2C" blockId="3.[151,1437,151,1905]" pageId="3" pageNumber="243">
<emphasis id="07DAEAE9D970FFC6D7A2C35AFED55F20" bold="true" box="[199,273,1123,1149]" pageId="3" pageNumber="243">Notes.</emphasis>
In most species of 
<taxonomicName id="F2AE4D78D970FFC6D68EC35AFDBD5F20" box="[491,633,1123,1149]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="genus">
<emphasis id="07DAEAE9D970FFC6D68EC35AFDBD5F20" box="[491,633,1123,1149]" italics="true" pageId="3" pageNumber="243">Gaeolaelaps</emphasis>
</taxonomicName>
the dorsal shield has 39 pairs of simple setae, including two pairs of 
<emphasis id="07DAEAE9D970FFC6D2E5C35AFA585F21" box="[1408,1436,1123,1148]" italics="true" pageId="3" pageNumber="243">Zx</emphasis>
setae, the peritremes usually reach anteriorly to the level of coxae I. Setae 
<emphasis id="07DAEAE9D970FFC6D483C3BEFC3B5FFC" box="[998,1023,1159,1185]" italics="true" pageId="3" pageNumber="243">z3</emphasis>
are occasionally absent, 
<emphasis id="07DAEAE9D970FFC6D279C3BEFAFC5FFC" box="[1308,1336,1159,1185]" italics="true" pageId="3" pageNumber="243">S1</emphasis>
is rarely absent as in 
<taxonomicName id="F2AE4D78D970FFC6D64CC392FE7C5F98" authorityName="Joharchi &amp; Döker &amp; Khaustov" authorityYear="2021" box="[297,440,1195,1221]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="07DAEAE9D970FFC6D64CC392FEF95F98" box="[297,317,1195,1221]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D628C392FE7C5F98" box="[333,440,1195,1221]" italics="true" pageId="3" pageNumber="243">altaiensis</emphasis>
</taxonomicName>
, and sometimes the peritremes are short, ending near the middle of coxae II (as in 
<taxonomicName id="F2AE4D78D970FFC6D2E4C392FEC65FB4" authorityName="Joharchi &amp; Döker &amp; Khaustov" authorityYear="2021" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="07DAEAE9D970FFC6D2E4C392FA515F98" box="[1409,1429,1195,1221]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D7F2C3F6FEC65FB4" box="[151,258,1231,1257]" italics="true" pageId="3" pageNumber="243">altaiensis</emphasis>
</taxonomicName>
) or even shorter, reaching to the anterior level of coxa III (e.g. 
<taxonomicName id="F2AE4D78D970FFC6D4B4C3F6FB3E5FB7" authority="Trach, 2016" authorityName="Trach" authorityYear="2016" box="[977,1274,1231,1258]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="heteroceri">
<emphasis id="07DAEAE9D970FFC6D4B4C3F6FC215FB4" box="[977,997,1231,1257]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D491C3F6FBA25FB4" box="[1012,1126,1231,1257]" italics="true" pageId="3" pageNumber="243">heteroceri</emphasis>
<bibRefCitation id="513F4B0AD970FFC6D30BC3F6FB3E5FB7" author="Trach, V. A." box="[1134,1274,1231,1258]" pageId="3" pageNumber="243" pagination="3 - 16" refId="ref11807" refString="Trach, V. A. (2016) Three new unusual beetle-associated species of the genus Gaeolaelaps (Acari, Mesostigmata, Laelapidae) from Ukraine. Vestnik Zoologii, 50, 3 - 16. https: // doi. org / 10.1515 / vzoo- 2016 - 0001" type="journal article" year="2016">Trach, 2016</bibRefCitation>
</taxonomicName>
). In the key to 
<taxonomicName id="F2AE4D78D970FFC6D7F2C3CAFEE15E50" box="[151,293,1267,1293]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="genus">
<emphasis id="07DAEAE9D970FFC6D7F2C3CAFEE15E50" box="[151,293,1267,1293]" italics="true" pageId="3" pageNumber="243">Gaeolaelaps</emphasis>
</taxonomicName>
species with short peritremes provided in 
<bibRefCitation id="513F4B0AD970FFC6D475C3CAFBE45E53" author="Vatankhah, F. &amp; Nemati, A. &amp; Esfandiari, M. &amp; Shishehbor, P." box="[784,1056,1267,1294]" pageId="3" pageNumber="243" pagination="566 - 574" refId="ref11857" refString="Vatankhah, F., Nemati, A., Esfandiari, M. &amp; Shishehbor, P. (2016) Description of a new species of Gaeolaelaps (Acari: Laelapidae) from Iran, with a key to world species of the genus with short peritremes. Zootaxa, 4121, 566 - 574. https: // doi. org / 10.11646 / zootaxa. 4121.5.6" type="journal article" year="2016">
Vatankhah 
<emphasis id="07DAEAE9D970FFC6D4EAC3CAFC075E50" box="[911,963,1267,1293]" italics="true" pageId="3" pageNumber="243">et al</emphasis>
. (2016)
</bibRefCitation>
, this species came to couplet 12, but the uncommon character state of 21 pairs of podonotal setae makes couplet choice problematic. According to 
<bibRefCitation id="513F4B0AD970FFC6D7F2C202FE5A5E0B" author="Vatankhah, F. &amp; Nemati, A. &amp; Esfandiari, M. &amp; Shishehbor, P." box="[151,414,1339,1366]" pageId="3" pageNumber="243" pagination="566 - 574" refId="ref11857" refString="Vatankhah, F., Nemati, A., Esfandiari, M. &amp; Shishehbor, P. (2016) Description of a new species of Gaeolaelaps (Acari: Laelapidae) from Iran, with a key to world species of the genus with short peritremes. Zootaxa, 4121, 566 - 574. https: // doi. org / 10.11646 / zootaxa. 4121.5.6" type="journal article" year="2016">
Vatankhah 
<emphasis id="07DAEAE9D970FFC6D671C202FE805E08" box="[276,324,1339,1365]" italics="true" pageId="3" pageNumber="243">et al</emphasis>
. (2016)
</bibRefCitation>
, the combination of short peritremes (reaching to middle of coxa II) and a dorsal shield with 37 pairs of setae occurs in only three described species of 
<taxonomicName id="F2AE4D78D970FFC6D45FC266FC0C5E24" box="[826,968,1375,1401]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="genus">
<emphasis id="07DAEAE9D970FFC6D45FC266FC0C5E24" box="[826,968,1375,1401]" italics="true" pageId="3" pageNumber="243">Gaeolaelaps</emphasis>
</taxonomicName>
: 
<taxonomicName id="F2AE4D78D970FFC6D4BCC266FADC5E24" authority="(Karg, 1994)" baseAuthorityName="Karg" baseAuthorityYear="1994" box="[985,1304,1375,1401]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="vertisimilis">
<emphasis id="07DAEAE9D970FFC6D4BCC266FC295E24" box="[985,1005,1375,1401]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D498C266FBBD5E24" box="[1021,1145,1375,1401]" italics="true" pageId="3" pageNumber="243">vertisimilis</emphasis>
(
<bibRefCitation id="513F4B0AD970FFC6D3EEC266FAD45E24" author="Karg, W." box="[1163,1296,1375,1401]" pageId="3" pageNumber="243" pagination="179 - 216" refId="ref10886" refString="Karg, W. (1994) Raubmilben der Cohors Gamasina Leach (Acarina, Parasitiformes) vom Galapagos - Archipel. Mitteilungen aus dem Zoologischen Museum in Berlin, 70, 179 - 216. https: // doi. org / 10.1002 / mmnz. 19940700202" type="journal article" year="1994">Karg, 1994</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="F2AE4D78D970FFC6D240C266FEEE5EC0" authority="(Karg, 1979)" baseAuthorityName="Karg" baseAuthorityYear="1979" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="verticis">
<emphasis id="07DAEAE9D970FFC6D240C266FAFD5E24" box="[1317,1337,1375,1401]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D22FC266FA585E25" box="[1354,1436,1375,1400]" italics="true" pageId="3" pageNumber="243">verticis</emphasis>
(
<bibRefCitation id="513F4B0AD970FFC6D7FAC2BAFEE65EC0" author="Karg, W." box="[159,290,1411,1437]" pageId="3" pageNumber="243" pagination="65 - 104" refId="ref10804" refString="Karg, W. (1979) Die Gattung Hypoaspis Canestrini 1884 (Acarina, Parasitiformes). Zoologische Jahrbucher Abteilung fur Systematik Okologie und Geographie der Tiere, 106, 65 - 104." type="journal article" year="1979">Karg, 1979</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="F2AE4D78D970FFC6D650C2BAFD425EC0" authority="(Ma, 1997)" baseAuthorityName="Ma" baseAuthorityYear="1997" box="[309,646,1411,1437]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="zhoumanshuae">
<emphasis id="07DAEAE9D970FFC6D650C2BAFE8D5EC0" box="[309,329,1411,1437]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D63DC2BAFE3A5EC0" box="[344,510,1411,1437]" italics="true" pageId="3" pageNumber="243">zhoumanshuae</emphasis>
(
<bibRefCitation id="513F4B0AD970FFC6D568C2BAFDBA5EC0" author="Ma, L." box="[525,638,1411,1437]" pageId="3" pageNumber="243" pagination="31 - 36" refId="ref11228" refString="Ma, L. (1997) Three new species of Gamasina in soll [soil] (Acari: Laelapidae, Parasitidae, Pachylaelaptidae). Acta Arachnologica Sinica, 6, 31 - 36." type="journal article" year="1997">Ma, 1997</bibRefCitation>
)
</taxonomicName>
. In both 
<taxonomicName id="F2AE4D78D970FFC6D58CC2BAFC995EC1" baseAuthorityName="Karg" baseAuthorityYear="1979" box="[745,861,1411,1437]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="verticis">
<emphasis id="07DAEAE9D970FFC6D58CC2BAFD395EC0" box="[745,765,1411,1437]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D46EC2BAFC995EC1" box="[779,861,1411,1436]" italics="true" pageId="3" pageNumber="243">verticis</emphasis>
</taxonomicName>
and 
<taxonomicName id="F2AE4D78D970FFC6D4F1C2BAFB985EC0" baseAuthorityName="Ma" baseAuthorityYear="1997" box="[916,1116,1411,1437]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="zhoumanshuae">
<emphasis id="07DAEAE9D970FFC6D4F1C2BAFC6C5EC0" box="[916,936,1411,1437]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D4D3C2BAFB985EC0" box="[950,1116,1411,1437]" italics="true" pageId="3" pageNumber="243">zhoumanshuae</emphasis>
</taxonomicName>
setae 
<emphasis id="07DAEAE9D970FFC6D3C6C2BAFB7F5EC0" box="[1187,1211,1411,1437]" italics="true" pageId="3" pageNumber="243">z3</emphasis>
is present and setae 
<emphasis id="07DAEAE9D970FFC6D7F2C29EFF085E9C" box="[151,204,1447,1473]" italics="true" pageId="3" pageNumber="243">s1–2</emphasis>
and 
<emphasis id="07DAEAE9D970FFC6D664C29EFED95E9D" box="[257,285,1447,1472]" italics="true" pageId="3" pageNumber="243">Zx</emphasis>
are absent, while in 
<taxonomicName id="F2AE4D78D970FFC6D567C29EFD4B5E9C" authorityName="Joharchi &amp; Döker &amp; Khaustov" authorityYear="2021" box="[514,655,1447,1473]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="07DAEAE9D970FFC6D567C29EFDD25E9C" box="[514,534,1447,1473]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D541C29EFD4B5E9C" box="[548,655,1447,1473]" italics="true" pageId="3" pageNumber="243">altaiensis</emphasis>
</taxonomicName>
setae 
<emphasis id="07DAEAE9D970FFC6D5B1C29EFD285E9C" box="[724,748,1447,1473]" italics="true" pageId="3" pageNumber="243">z3</emphasis>
and 
<emphasis id="07DAEAE9D970FFC6D447C29EFCFA5E9C" box="[802,830,1447,1473]" italics="true" pageId="3" pageNumber="243">S1</emphasis>
are absent. 
<taxonomicName id="F2AE4D78D970FFC6D4A7C29EFB055E9C" authorityName="Joharchi &amp; Döker &amp; Khaustov" authorityYear="2021" box="[962,1217,1447,1473]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="07DAEAE9D970FFC6D4A7C29EFB055E9C" box="[962,1217,1447,1473]" italics="true" pageId="3" pageNumber="243">Gaeolaelaps altaiensis</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="F2AE4D78D970FFC6D25BC29EFF245EB8" baseAuthorityName="Karg" baseAuthorityYear="1994" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="vertisimilis">
<emphasis id="07DAEAE9D970FFC6D25BC29EFA965E9C" box="[1342,1362,1447,1473]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D205C29EFF245EB8" italics="true" pageId="3" pageNumber="243">vertisimilis</emphasis>
</taxonomicName>
(couplet 
<quantity id="F2569B1ED970FFC6D62FC2F2FE475EB8" box="[330,387,1483,1509]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="3" pageNumber="243" unit="in" value="12.0">12 in</quantity>
the identification key to 
<taxonomicName id="F2AE4D78D970FFC6D5F9C2F2FCEE5EB8" box="[668,810,1483,1509]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="genus">
<emphasis id="07DAEAE9D970FFC6D5F9C2F2FCEE5EB8" box="[668,810,1483,1509]" italics="true" pageId="3" pageNumber="243">Gaeolaelaps</emphasis>
</taxonomicName>
species with short peritremes provided by 
<bibRefCitation id="513F4B0AD970FFC6D26EC2F2FF395D57" author="Vatankhah, F. &amp; Nemati, A. &amp; Esfandiari, M. &amp; Shishehbor, P." pageId="3" pageNumber="243" pagination="566 - 574" refId="ref11857" refString="Vatankhah, F., Nemati, A., Esfandiari, M. &amp; Shishehbor, P. (2016) Description of a new species of Gaeolaelaps (Acari: Laelapidae) from Iran, with a key to world species of the genus with short peritremes. Zootaxa, 4121, 566 - 574. https: // doi. org / 10.11646 / zootaxa. 4121.5.6" type="journal article" year="2016">
Vatankhah 
<emphasis id="07DAEAE9D970FFC6D2EDC2F2FF695D54" italics="true" pageId="3" pageNumber="243">et al</emphasis>
., 2016
</bibRefCitation>
), due to the dorsal shield having 37 pairs of setae (
<emphasis id="07DAEAE9D970FFC6D44EC2D6FC805D54" box="[811,836,1519,1545]" italics="true" pageId="3" pageNumber="243">z3</emphasis>
absent) and without 
<emphasis id="07DAEAE9D970FFC6D34EC2D6FBF85D55" box="[1067,1084,1519,1544]" italics="true" pageId="3" pageNumber="243">R</emphasis>
series setae on lateral cuticle of dorsal shield. 
<taxonomicName id="F2AE4D78D970FFC6D651C12AFDF05D70" authorityName="Joharchi &amp; Döker &amp; Khaustov" authorityYear="2021" box="[308,564,1555,1581]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="07DAEAE9D970FFC6D651C12AFDF05D70" box="[308,564,1555,1581]" italics="true" pageId="3" pageNumber="243">Gaeolaelaps altaiensis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="F2AE4D78D970FFC6D5ACC12AFC815D70" baseAuthorityName="Karg" baseAuthorityYear="1994" box="[713,837,1555,1581]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="vertisimilis">
<emphasis id="07DAEAE9D970FFC6D5ACC12AFC815D70" box="[713,837,1555,1581]" italics="true" pageId="3" pageNumber="243">vertisimilis</emphasis>
</taxonomicName>
by having 16 pairs of setae on opisthonotal region of dorsal shield (including two pairs 
<emphasis id="07DAEAE9D970FFC6D572C10EFDF75D0D" box="[535,563,1591,1616]" italics="true" pageId="3" pageNumber="243">Zx</emphasis>
setae, 
<emphasis id="07DAEAE9D970FFC6D5E4C10EFD595D0C" box="[641,669,1591,1617]" italics="true" pageId="3" pageNumber="243">S1</emphasis>
absent), while in 
<taxonomicName id="F2AE4D78D970FFC6D402C10EFBC25D0C" baseAuthorityName="Karg" baseAuthorityYear="1994" box="[871,1030,1591,1617]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="vertisimilis">
<emphasis id="07DAEAE9D970FFC6D402C10EFCBF5D0C" box="[871,891,1591,1617]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D4EFC10EFBC25D0C" box="[906,1030,1591,1617]" italics="true" pageId="3" pageNumber="243">vertisimilis</emphasis>
</taxonomicName>
opisthonotal region of dorsal shield has 18 pairs of setae (including two pairs 
<emphasis id="07DAEAE9D970FFC6D510C162FD555D29" box="[629,657,1627,1652]" italics="true" pageId="3" pageNumber="243">Zx</emphasis>
setae, 
<emphasis id="07DAEAE9D970FFC6D585C162FD385D28" box="[736,764,1627,1653]" italics="true" pageId="3" pageNumber="243">S1</emphasis>
and 
<emphasis id="07DAEAE9D970FFC6D450C162FC915D28" box="[821,853,1627,1653]" italics="true" pageId="3" pageNumber="243">R5</emphasis>
present on dorsal shield). 
<taxonomicName id="F2AE4D78D970FFC6D3E6C162FF1F5DC7" authority="(Karg, 1962)" baseAuthorityName="Karg" baseAuthorityYear="1962" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="nolli">
<emphasis id="07DAEAE9D970FFC6D3E6C162FA885D28" box="[1155,1356,1627,1653]" italics="true" pageId="3" pageNumber="243">Gaeolaelaps nolli</emphasis>
(
<bibRefCitation id="513F4B0AD970FFC6D239C162FF175DC7" author="Karg, W." pageId="3" pageNumber="243" pagination="23 - 119" refId="ref10755" refString="Karg, W. (1962) Zur Systematik und Postembryonalen Entwicklung der Gamasiden (Acarina, Parasitiformes) landwirtschaftlich genutzter Boden. Mitteilungen aus dem Zoologischen Museum in Berlin, 38, 23 - 119. https: // doi. org / 10.1002 / mmnz. 4830380103" type="journal article" year="1962">Karg, 1962</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="F2AE4D78D970FFC6D783C146FE265DC7" authority="(Costa, 1968)" baseAuthorityName="Costa" baseAuthorityYear="1968" box="[230,482,1663,1690]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="kargi">
<emphasis id="07DAEAE9D970FFC6D783C146FF3E5DC4" box="[230,250,1663,1689]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D66CC146FE875DC4" box="[265,323,1663,1689]" italics="true" pageId="3" pageNumber="243">kargi</emphasis>
(
<bibRefCitation id="513F4B0AD970FFC6D637C146FE1E5DC7" author="Costa, M." box="[338,474,1663,1690]" pageId="3" pageNumber="243" pagination="1 - 30" refId="ref9638" refString="Costa, M. (1968) Little known and new litter-inhabiting Laelapine mites (Acari: Mesostigmata) from Israel. Israel Journal of Zoology, 17, 1 - 30." type="journal article" year="1968">
<collectingRegion id="F76AF819D970FFC6D637C146FE525DC4" box="[338,406,1663,1689]" country="Tanzania" name="Pwani" pageId="3" pageNumber="243">Costa</collectingRegion>
, 1968
</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="F2AE4D78D970FFC6D57DC146FC595DC4" authority="(Willmann, 1949)" baseAuthorityName="Willmann" baseAuthorityYear="1949" box="[536,925,1663,1689]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="praesternalis">
<emphasis id="07DAEAE9D970FFC6D57DC146FDE85DC4" box="[536,556,1663,1689]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D55EC146FD0B5DC4" box="[571,719,1663,1689]" italics="true" pageId="3" pageNumber="243">praesternalis</emphasis>
(
<bibRefCitation id="513F4B0AD970FFC6D5BAC146FC515DC4" author="Willmann, C." box="[735,917,1663,1689]" pageId="3" pageNumber="243" pagination="106 - 142" refId="ref12069" refString="Willmann, C. (1949) Beitrage zur Kenntnis des Salzgebietes von Ciechocinek. 1. Milben aus den Salzwiesen und Salzmooren von Ciechocinek an der Weichsel. Voroffentlichungen aus dem Museum fur Natur-, Volker- und Handelskande in Bremen, Reihe A, 1, 106 - 142." type="journal article" year="1949">Willmann, 1949</bibRefCitation>
)
</taxonomicName>
may be similar to 
<taxonomicName id="F2AE4D78D970FFC6D30BC146FB3F5DC4" authorityName="Joharchi &amp; Döker &amp; Khaustov" authorityYear="2021" box="[1134,1275,1663,1689]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="07DAEAE9D970FFC6D30BC146FB465DC4" box="[1134,1154,1663,1689]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D3F5C146FB3F5DC4" box="[1168,1275,1663,1689]" italics="true" pageId="3" pageNumber="243">altaiensis</emphasis>
</taxonomicName>
in general appearance, but 
<taxonomicName id="F2AE4D78D970FFC6D650C19AFE565DE0" box="[309,402,1699,1725]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="kargi">
<emphasis id="07DAEAE9D970FFC6D650C19AFE8D5DE0" box="[309,329,1699,1725]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D63DC19AFE565DE0" box="[344,402,1699,1725]" italics="true" pageId="3" pageNumber="243">kargi</emphasis>
</taxonomicName>
and 
<taxonomicName id="F2AE4D78D970FFC6D6AFC19AFD465DE0" baseAuthorityName="Willmann" baseAuthorityYear="1949" box="[458,642,1699,1725]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="praesternalis">
<emphasis id="07DAEAE9D970FFC6D6AFC19AFE1A5DE0" box="[458,478,1699,1725]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D68BC19AFD465DE0" box="[494,642,1699,1725]" italics="true" pageId="3" pageNumber="243">praesternalis</emphasis>
</taxonomicName>
have a long peritremes nearly extending to coxa I and 
<taxonomicName id="F2AE4D78D970FFC6D397C19AFA8C5DE0" baseAuthorityName="Karg" baseAuthorityYear="1962" box="[1266,1352,1699,1725]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="nolli">
<emphasis id="07DAEAE9D970FFC6D397C19AFAC25DE0" box="[1266,1286,1699,1725]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D270C19AFA8C5DE0" box="[1301,1352,1699,1725]" italics="true" pageId="3" pageNumber="243">nolli</emphasis>
</taxonomicName>
having 39 pairs of setae on dorsal shield. Overall 
<taxonomicName id="F2AE4D78D970FFC6D50CC1FEFD325DBC" authorityName="Joharchi &amp; Döker &amp; Khaustov" authorityYear="2021" box="[617,758,1735,1761]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="07DAEAE9D970FFC6D50CC1FEFDB95DBC" box="[617,637,1735,1761]" italics="true" pageId="3" pageNumber="243">G</emphasis>
. 
<emphasis id="07DAEAE9D970FFC6D5EEC1FEFD325DBC" box="[651,758,1735,1761]" italics="true" pageId="3" pageNumber="243">altaiensis</emphasis>
</taxonomicName>
can be easily distinguished from all other members of 
<taxonomicName id="F2AE4D78D970FFC6D233C1FEFF2C5C58" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="243" phylum="Arthropoda" rank="genus">
<emphasis id="07DAEAE9D970FFC6D233C1FEFF2C5C58" italics="true" pageId="3" pageNumber="243">Gaeolaelaps</emphasis>
</taxonomicName>
with short peritremes (See 
<bibRefCitation id="513F4B0AD970FFC6D546C1D2FCE15C5B" author="Vatankhah, F. &amp; Nemati, A. &amp; Esfandiari, M. &amp; Shishehbor, P." box="[547,805,1771,1798]" pageId="3" pageNumber="243" pagination="566 - 574" refId="ref11857" refString="Vatankhah, F., Nemati, A., Esfandiari, M. &amp; Shishehbor, P. (2016) Description of a new species of Gaeolaelaps (Acari: Laelapidae) from Iran, with a key to world species of the genus with short peritremes. Zootaxa, 4121, 566 - 574. https: // doi. org / 10.11646 / zootaxa. 4121.5.6" type="journal article" year="2016">
Vatankhah 
<emphasis id="07DAEAE9D970FFC6D5C7C1D2FD115C58" box="[674,725,1771,1797]" italics="true" pageId="3" pageNumber="243">et al</emphasis>
., 2016
</bibRefCitation>
) by the following combination of characters: (1) dorsal shield with distinct reticulation posterior to 
<emphasis id="07DAEAE9D970FFC6D5E0C037FD5F5C75" box="[645,667,1806,1832]" italics="true" pageId="3" pageNumber="243">j6</emphasis>
, narrowing from level of setae 
<emphasis id="07DAEAE9D970FFC6D49AC036FBDC5C74" box="[1023,1048,1807,1833]" italics="true" pageId="3" pageNumber="243">r3</emphasis>
and with 37 pairs of setae (
<emphasis id="07DAEAE9D970FFC6D236C036FAA85C74" box="[1363,1388,1807,1833]" italics="true" pageId="3" pageNumber="243">z3</emphasis>
and 
<emphasis id="07DAEAE9D970FFC6D7F2C00AFF775C10" box="[151,179,1843,1869]" italics="true" pageId="3" pageNumber="243">S1</emphasis>
absent); (2) palptarsal claw two-tined; (3) peritremes short (ending near middle of coxae II); (4) tarsus and femur IV with two (
<emphasis id="07DAEAE9D970FFC6D64BC06EFE9C5C2C" box="[302,344,1879,1905]" italics="true" pageId="3" pageNumber="243">pd2</emphasis>
, 
<emphasis id="07DAEAE9D970FFC6D603C06EFE545C2C" box="[358,400,1879,1905]" italics="true" pageId="3" pageNumber="243">pd3</emphasis>
) and one (
<emphasis id="07DAEAE9D970FFC6D56CC06EFDF75C2C" box="[521,563,1879,1905]" italics="true" pageId="3" pageNumber="243">ad1</emphasis>
) elongated setae, respectively; (5) fixed digit of chelicera with seven teeth.
</paragraph>
</subSubSection>
</treatment>
</document>