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<mods:title id="74E0FECEEB702DD4102FC6F3E6EF8420">Identification of Mediterranean marine gobies (Actinopterygii: Gobiidae) of the continental shelf from photographs of in situ individuals</mods:title>
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<heading id="B23881B13F7EFFE9FF76FF11FC3CFF50" bold="true" box="[151,988,151,178]" fontSize="11" level="2" pageId="75" pageNumber="76" reason="4">
<emphasis id="DBBBEACF3F7EFFE9FF76FF11FC3CFF50" bold="true" box="[151,988,151,178]" pageId="75" pageNumber="76">
<taxonomicName id="2ECF4D5E3F7EFFE9FF76FF11FD77FF50" ID-CoL="4LQXH" authority="(Risso, 1810)" authorityName="Benthic" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[151,663,151,178]" class="Actinopterygii" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="75" pageNumber="76" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="DBBBEACF3F7EFFE9FF76FF11FE10FF50" bold="true" box="[151,496,151,178]" italics="true" pageId="75" pageNumber="76">Pomatoschistus marmoratus</emphasis>
(
<bibRefCitation id="8D5E4B2C3F7EFFE9FE1EFF11FD6FFF50" author="Risso, A." box="[511,655,151,177]" pageId="75" pageNumber="76" refId="ref63775" refString="Risso, A. (1810) Ichtyologie de Nice, ou histoire naturelle des poissons du departement des Alpes Maritimes. F. Schoell, Paris, xxxvi + 388 pp. [reprint, 1966, Asher, Amsterdam] https: // doi. org / 10.5962 / bhl. title. 7052" type="book" year="1810">Risso, 1810</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="71F42A583F7EFFE9FD46FF11FD1CFF50" box="[679,764,151,177]" captionStart="FIGURE 65" captionStartId="75.[151,250,1970,1994]" captionTargetBox="[201,1386,556,1946]" captionTargetId="figure-140@75.[201,1386,556,1946]" captionTargetPageId="75" captionText="FIGURE 65. Live coloration in its natural habitat of Pomatoschistus marmoratus. A) Male and B) female, photos by R. Pillon, Rab Island, Croatia, northern Adriatic Sea. C) Female, photo by J. Renoult, Thau lagoon, Southern France." figureDoi="http://doi.org/10.5281/zenodo.6601763" httpUri="https://zenodo.org/record/6601763/files/figure.png" pageId="75" pageNumber="76">Fig. 65</figureCitation>
)—Marbled Goby
</emphasis>
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</paragraph>
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<paragraph id="E97036DD3F7EFFE9FF76FF5AFB7BFF15" blockId="75.[151,1179,220,244]" box="[151,1179,220,244]" pageId="75" pageNumber="76">
<treatmentCitationGroup id="C9DF11F33F7EFFE9FF76FF5AFB7BFF15" box="[151,1179,220,244]" pageId="75" pageNumber="76">
<treatmentCitation id="686E10CC3F7EFFE9FF76FF5AFDC6FF15" author="Risso, A." box="[151,550,220,244]" page="339" pageId="75" pageNumber="76" year="1810">
<taxonomicName id="2ECF4D5E3F7EFFE9FF76FF5AFDC6FF15" authority="Risso, 1810: 339" authorityName="Risso" authorityPageNumber="339" authorityYear="1810" box="[151,550,220,244]" class="Actinopterygii" family="Atherinidae" genus="Atherina" kingdom="Animalia" order="Atheriniformes" pageId="75" pageNumber="76" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="DBBBEACF3F7EFFE9FF76FF5AFE92FF12" box="[151,370,220,244]" italics="true" pageId="75" pageNumber="76">Atherina marmoratus</emphasis>
<bibRefCitation id="8D5E4B2C3F7EFFE9FE99FF5AFDC6FF15" author="Risso, A." box="[376,550,220,244]" pageId="75" pageNumber="76" refId="ref63775" refString="Risso, A. (1810) Ichtyologie de Nice, ou histoire naturelle des poissons du departement des Alpes Maritimes. F. Schoell, Paris, xxxvi + 388 pp. [reprint, 1966, Asher, Amsterdam] https: // doi. org / 10.5962 / bhl. title. 7052" type="book" year="1810">Risso, 1810: 339</bibRefCitation>
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;
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locality:
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,
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,
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.
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Size. Maximum size
<quantity id="2E379B383F7EFFE9FE65FEA4FE30FEDD" box="[388,464,290,317]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="75" pageNumber="76" unit="cm" value="6.5">6.5 cm</quantity>
total length (Miller 1986).
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Morphology. D V (VVII) + I,810;A I,810; P 1721 (Miller 1986). Moderately small goby with subcylindrical body, laterally compressed towards caudal fin, head slightly depressed and moderately large. Snout moderately large. Eyes dorsolateral, interorbital space moderately narrow. Caudal peduncle long and slender, clearly lower than body depth. Dorsal fins of similar height, the first dorsal fin may be slightly higher, with more or less rounded margin and no elongate spines. Caudal fin rounded. Scales present on body, sometimes visible on photographs, at least dorsally.
<emphasis id="DBBBEACF3F7EFFE9FED9FE7CFDD7FDF5" bold="true" box="[312,567,506,532]" pageId="75" pageNumber="76">Predorsal area naked</emphasis>
(Miller 1986).
</paragraph>
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<paragraph id="E97036DD3F7EFFE9FF76F834FB31F80F" blockId="75.[151,1436,1970,2031]" pageId="75" pageNumber="76">
<emphasis id="DBBBEACF3F7EFFE9FF76F834FEC0F82B" bold="true" box="[151,288,1970,1994]" pageId="75" pageNumber="76">FIGURE 65.</emphasis>
Live coloration in its natural habitat of
<taxonomicName id="2ECF4D5E3F7EFFE9FD50F834FC32F82B" authorityName="Benthic" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[689,978,1970,1994]" class="Actinopterygii" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="75" pageNumber="76" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="DBBBEACF3F7EFFE9FD50F834FC32F82B" box="[689,978,1970,1994]" italics="true" pageId="75" pageNumber="76">Pomatoschistus marmoratus</emphasis>
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. A) Male and B) female, photos by R. Pillon, Rab Island, Croatia, northern Adriatic Sea. C) Female, photo by J. Renoult, Thau lagoon, Southern France.
</paragraph>
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Live coloration. Background a pale sandy color, with a more or less marbled pattern of dark and white markings. On pale fine sand, light colors prevail, with almost no recognizable patterns (
<figureCitation id="71F42A583F79FFEEFBCDFF3DFAE6FF34" box="[1068,1286,186,213]" captionStart="FIGURE 65" captionStartId="75.[151,250,1970,1994]" captionTargetBox="[201,1386,556,1946]" captionTargetId="figure-140@75.[201,1386,556,1946]" captionTargetPageId="75" captionText="FIGURE 65. Live coloration in its natural habitat of Pomatoschistus marmoratus. A) Male and B) female, photos by R. Pillon, Rab Island, Croatia, northern Adriatic Sea. C) Female, photo by J. Renoult, Thau lagoon, Southern France." figureDoi="http://doi.org/10.5281/zenodo.6601763" httpUri="https://zenodo.org/record/6601763/files/figure.png" pageId="76" pageNumber="77">Figs. 65a and 65b</figureCitation>
). On coarser sediments, more contrasting patterns are usually displayed:
<emphasis id="DBBBEACF3F79FFEEFCD6FF58FB91FF18" bold="true" box="[823,1137,222,249]" pageId="76" pageNumber="77">6 pale saddles on the back</emphasis>
, 1 below first dorsal fin, 2 below second dorsal fin, 2 on caudal peduncle and one, narrower and sometimes faint, on nape above pectoral-fin base; variable number of faint rounded dark spots along midline, the last longitudinal and more obvious at caudalfin base (
<figureCitation id="71F42A583F79FFEEFF1EFECDFE80FE84" box="[255,352,330,357]" captionStart="FIGURE 65" captionStartId="75.[151,250,1970,1994]" captionTargetBox="[201,1386,556,1946]" captionTargetId="figure-140@75.[201,1386,556,1946]" captionTargetPageId="75" captionText="FIGURE 65. Live coloration in its natural habitat of Pomatoschistus marmoratus. A) Male and B) female, photos by R. Pillon, Rab Island, Croatia, northern Adriatic Sea. C) Female, photo by J. Renoult, Thau lagoon, Southern France." figureDoi="http://doi.org/10.5281/zenodo.6601763" httpUri="https://zenodo.org/record/6601763/files/figure.png" pageId="76" pageNumber="77">Fig. 65c</figureCitation>
). A preorbital dark bar, especially obvious in females, extends from each eye, through upper lip, to connect with that from the opposite side, forming a
<emphasis id="DBBBEACF3F79FFEEFD10FEE8FBFDFE68" bold="true" box="[753,1053,366,393]" pageId="76" pageNumber="77">distinctive dark chin spot</emphasis>
(
<figureCitation id="71F42A583F79FFEEFBCDFEE9FB6EFE68" box="[1068,1166,366,393]" captionStart="FIGURE 65" captionStartId="75.[151,250,1970,1994]" captionTargetBox="[201,1386,556,1946]" captionTargetId="figure-140@75.[201,1386,556,1946]" captionTargetPageId="75" captionText="FIGURE 65. Live coloration in its natural habitat of Pomatoschistus marmoratus. A) Male and B) female, photos by R. Pillon, Rab Island, Croatia, northern Adriatic Sea. C) Female, photo by J. Renoult, Thau lagoon, Southern France." figureDoi="http://doi.org/10.5281/zenodo.6601763" httpUri="https://zenodo.org/record/6601763/files/figure.png" pageId="76" pageNumber="77">Fig. 65b</figureCitation>
). Often a whitish to pale blue dot on nape, just behind eyes, generally more or less circled with an interrupted dark line, often like between brackets (such a marking may also appear in
<taxonomicName id="2ECF4D5E3F79FFEEFD6DFE31FCE0FE31" authorityName="Benthic" baseAuthorityName="Kroyer" baseAuthorityYear="1838" box="[652,768,439,464]" class="Actinopterygii" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="microps">
<emphasis id="DBBBEACF3F79FFEEFD6DFE31FCE0FE31" box="[652,768,439,464]" italics="true" pageId="76" pageNumber="77">P. microps</emphasis>
</taxonomicName>
or
<taxonomicName id="2ECF4D5E3F79FFEEFCC5FE31FB61FE30" authorityName="Benthic" baseAuthorityName="Valenciennes" baseAuthorityYear="1837" box="[804,1153,439,465]" class="Actinopterygii" family="Gobiidae" genus="Deltentosteus" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="quadrimaculatus">
<emphasis id="DBBBEACF3F79FFEEFCC5FE31FB61FE30" box="[804,1153,439,465]" italics="true" pageId="76" pageNumber="77">Deltentosteus quadrimaculatus</emphasis>
</taxonomicName>
).
<emphasis id="DBBBEACF3F79FFEEFB76FE31FD8CFE14" bold="true" pageId="76" pageNumber="77">In males, cheeks often densely sprinkled with small black dots</emphasis>
; first dorsal fin may be large, with brownish and pale stripes, and a blueblack spot on its rear upper corner (
<figureCitation id="71F42A583F79FFEEFDC4FE79FD66FDF8" box="[549,646,510,537]" captionStart="FIGURE 65" captionStartId="75.[151,250,1970,1994]" captionTargetBox="[201,1386,556,1946]" captionTargetId="figure-140@75.[201,1386,556,1946]" captionTargetPageId="75" captionText="FIGURE 65. Live coloration in its natural habitat of Pomatoschistus marmoratus. A) Male and B) female, photos by R. Pillon, Rab Island, Croatia, northern Adriatic Sea. C) Female, photo by J. Renoult, Thau lagoon, Southern France." figureDoi="http://doi.org/10.5281/zenodo.6601763" httpUri="https://zenodo.org/record/6601763/files/figure.png" pageId="76" pageNumber="77">Fig. 65a</figureCitation>
).
</paragraph>
<paragraph id="E97036DD3F79FFEEFF26FDA5FC5BFDDC" blockId="76.[151,1437,151,826]" box="[199,955,547,573]" pageId="76" pageNumber="77">
Similar species.
<taxonomicName id="2ECF4D5E3F79FFEEFE9EFDA5FD48FDDC" authorityName="Miller" authorityYear="1973" box="[383,680,547,573]" class="Actinopterygii" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="adriaticus">
<emphasis id="DBBBEACF3F79FFEEFE9EFDA5FD48FDDC" box="[383,680,547,573]" italics="true" pageId="76" pageNumber="77">Pomatoschistus adriaticus</emphasis>
</taxonomicName>
,
<taxonomicName id="2ECF4D5E3F79FFEEFD57FDA5FCCEFDDD" authorityName="Benthic" baseAuthorityName="Kroyer" baseAuthorityYear="1838" box="[694,814,547,573]" class="Actinopterygii" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="microps">
<emphasis id="DBBBEACF3F79FFEEFD57FDA5FD27FDDD" box="[694,711,547,572]" italics="true" pageId="76" pageNumber="77">P</emphasis>
.
<emphasis id="DBBBEACF3F79FFEEFD34FDA5FCCEFDDD" box="[725,814,547,572]" italics="true" pageId="76" pageNumber="77">microps</emphasis>
</taxonomicName>
,
<taxonomicName id="2ECF4D5E3F79FFEEFCDDFDA5FC54FDDD" authorityName="Benthic" baseAuthorityName="Pallas" baseAuthorityYear="1770" box="[828,948,547,573]" class="Actinopterygii" family="Gobiidae" genus="Pomatoschistus" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="minutus">
<emphasis id="DBBBEACF3F79FFEEFCDDFDA5FCADFDDD" box="[828,845,547,572]" italics="true" pageId="76" pageNumber="77">P</emphasis>
.
<emphasis id="DBBBEACF3F79FFEEFCBAFDA5FC54FDDD" box="[859,948,547,572]" italics="true" pageId="76" pageNumber="77">minutus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="E97036DD3F79FFEEFF26FDC1FAFFFD67" blockId="76.[151,1437,151,826]" pageId="76" pageNumber="77">
Habitat. Infralittoral species, often common on shallow soft bottoms (mostly fine to coarse sand) from
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depth (
<bibRefCitation id="8D5E4B2C3F79FFEEFEE0FDEDFE17FD64" author="Apostolou, A. &amp; Ivanova, P. &amp; Velkov, B. &amp; Vassilev, M. &amp; Dobrev, D. &amp; Dobrovolov, I." box="[257,503,619,645]" pageId="76" pageNumber="77" pagination="289 - 294" refId="ref51172" refString="Apostolou, A., Ivanova, P., Velkov, B., Vassilev, M., Dobrev, D. &amp; Dobrovolov, I. (2011) Pomatoschistus marmoratus (Risso 1810), is it really a &quot; New &quot; species for Bulgarian Ichthyofauna? Acta Zoologica Bulgarica, 63 (3), 289 - 294." type="journal article" year="2011">
Apostolou
<emphasis id="DBBBEACF3F79FFEEFE9FFDEDFE4FFD64" box="[382,431,619,645]" italics="true" pageId="76" pageNumber="77">et al</emphasis>
. 2011
</bibRefCitation>
;
<bibRefCitation id="8D5E4B2C3F79FFEEFDE2FDEDFD27FD64" author="Pillon, R. &amp; Pavon, D. &amp; Berenger, L." box="[515,711,618,645]" pageId="76" pageNumber="77" refId="ref62997" refString="Pillon, R., Pavon, D. &amp; Berenger, L. (2015) Signalisation de / Record of Pomatoschistus marmoratus, 16 November 2015 (obs. 17 June 2013, photo # 1147). Fish Watch Forum, Louisy, P. &amp; Francour, P. (Ed.). Available from: https: // www. fish-watch. org (accessed 28 September 2021)" type="url" year="2015">
Pillon
<emphasis id="DBBBEACF3F79FFEEFDACFDEDFD9EFD64" box="[589,638,619,645]" italics="true" pageId="76" pageNumber="77">et al</emphasis>
. 2015
</bibRefCitation>
;
<bibRefCitation id="8D5E4B2C3F79FFEEFD32FDEDFC8AFD64" author="Patzner, R. A." box="[723,874,619,645]" pageId="76" pageNumber="77" refId="ref62855" refString="Patzner, R. A. (2021) Gobiidae. Available from: http: // www. patzner. sbg. ac. at / Gobiidae / GobiidaeSpecies. htm (accessed 1 October 2021)" type="url" year="2021">Patzner 2021</bibRefCitation>
), may occur to
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(Miller 1986).
</paragraph>
<paragraph id="E97036DD3F79FFEEFF26FD09FE1AFCDB" blockId="76.[151,1437,151,826]" pageId="76" pageNumber="77">
Geographic distribution. Black Sea, Mediterranean and adjacentAtlantic Ocean.Widespread in the Mediterranean from
<collectingCountry id="91D8764D3F79FFEEFF35FD35FED9FD2C" box="[212,313,691,717]" name="Gibraltar" pageId="76" pageNumber="77">Gibraltar</collectingCountry>
to Levant, also common in the Black Sea (Miller 1986;
<bibRefCitation id="8D5E4B2C3F79FFEEFC44FD35FB3CFD2C" author="Boltachev, A. R. &amp; Karpova, E. P." box="[933,1244,691,717]" pageId="76" pageNumber="77" refId="ref52296" refString="Boltachev, A. R. &amp; Karpova, E. P. (2017) Marine fishes of the Crimean Peninsula, second edition. Business-Inform, Simpheropol, 376 pp." type="book" year="2017">Boltachev &amp; Karpova 2017</bibRefCitation>
). Along the south Mediterranean coast between the Alboran Sea and
<collectingRegion id="2B0BF83F3F79FFEEFD3AFD51FCF1FD10" box="[731,785,727,753]" country="Egypt" name="As Suways" pageId="76" pageNumber="77">Suez</collectingRegion>
Canal in
<collectingCountry id="91D8764D3F79FFEEFC62FD51FC26FD10" box="[899,966,727,753]" name="Egypt" pageId="76" pageNumber="77">Egypt</collectingCountry>
(Miller 1986) the species was confirmed from northern and southern
<collectingCountry id="91D8764D3F79FFEEFE3CFD7DFDD0FCF4" box="[477,560,763,789]" name="Tunisia" pageId="76" pageNumber="77">Tunisia</collectingCountry>
(
<bibRefCitation id="8D5E4B2C3F79FFEEFDA3FD7DFCFBFCF4" author="Mejri, R. &amp; Kovacic, M. &amp; Ben Hassine, O. K." box="[578,795,763,789]" pageId="76" pageNumber="77" pagination="171 - 172" refId="ref60911" refString="Mejri, R., Kovacic, M. &amp; Ben Hassine, O. K. (2009 a) First record of Pomatoschistus marmoratus (Gobiidae) on the Tunisian coast. Cybium, 33, 171 - 172. https: // doi. org / 10.26028 / cybium / 2009 - 331 - 012" type="journal article" year="2009">
Mejri
<emphasis id="DBBBEACF3F79FFEEFD6DFD7DFD21FCF4" box="[652,705,763,789]" italics="true" pageId="76" pageNumber="77">et al</emphasis>
. 2009a
</bibRefCitation>
). In the Atlantic Ocean, known from
<collectingCountry id="91D8764D3F79FFEEFB31FD7DFAD5FCF4" box="[1232,1333,763,789]" name="Gibraltar" pageId="76" pageNumber="77">Gibraltar</collectingCountry>
to south Portuguese coast (Miller 1986).
</paragraph>
</subSubSection>
</treatment>
</document>