780 lines
89 KiB
XML
780 lines
89 KiB
XML
<document id="6A24ADE3E7B9FC70DF4F521922BEDDF8" ID-DOI="10.3897/zookeys.1207.117225" ID-ZooBank="94F18819-5AF5-4100-AB35-AA3C3976EE80" ID-publisher-id="117225" URI-arpha="255F6420-5705-5F50-9167-509E6C76FA1C" URI-zoobank="http://zoobank.org/94F18819-5AF5-4100-AB35-AA3C3976EE80" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1721676390257" checkinUser="pensoft" docAuthor="Angus, Robert B., Maté, Jason F., Angus, Elizabeth M. & Král, David" docDate="2024" docId="236DE0689AE65F0B9E4185F645D1F480" docLanguage="en" docName="ZooKeys 1207: 205-299" docOrigin="ZooKeys 1207" docSource="https://zookeys.pensoft.net/article/117225/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Aphodius (Liothoxax) muscorum Adam 1994" docType="treatment" docVersion="2" lastPageNumber="299" masterDocId="255F642057055F509167509E6C76FA1C" masterDocTitle="Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" masterLastPageNumber="299" masterPageNumber="205" pageNumber="205" updateTime="1721687169892" updateUser="admin">
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<mods:title id="1408BB93B846D5F8E5DE48205ED3878D">Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)</mods:title>
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<mods:namePart id="0A2650261EC707885AD8645B34FFB0BB">Angus, Robert B.</mods:namePart>
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<mods:affiliation id="EDEA57D4376DEAC00712CA11B6EA987F">Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="B37D25FAD85D17AC3ABC2533B3BD81BF">Maté, Jason F.</mods:namePart>
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<mods:affiliation id="BECF7154FC4B4BA5C66A1EE26B88F26D">c / Henares 16, Velilla de San Antonio, Madrid, 28891, Spain</mods:affiliation>
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<mods:namePart id="C67DA49B7AE98F3A4ADB3A9CEDC33500">Angus, Elizabeth M.</mods:namePart>
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<mods:affiliation id="E072C07551FDE61AB98C30C78F781948">Biomedical Imaging Unit, Level B South Block, Mail point 12, General Hospital, Southampton SO 16 6 YD, UK</mods:affiliation>
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<mods:namePart id="BF13B8270822A67DBD647A8AD9B29DAB">Král, David</mods:namePart>
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<mods:affiliation id="AFD0A21CFCAF99DBA815DFA69F9B0996">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czech Republic</mods:affiliation>
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<mods:date id="401D4D87234FB3393DE447097AEB2D40">2024</mods:date>
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<taxonomicName id="D7DA216CAA733FDA4A592E7A22A972EC" authority="Adam, 1994" authorityName="Adam" authorityYear="1994" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="muscorum" subGenus="Liothoxax">
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<emphasis id="2B083601C0E00A144A754064F92EA6A6" italics="true">Aphodius</emphasis>
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(
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<emphasis id="3872F192A6821A5313FC855B9DCB95C9" italics="true">Liothoxax</emphasis>
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)
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<emphasis id="D115AD1CDC14742DA36A7142080C202F" italics="true">muscorum</emphasis>
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<bibRefCitation id="CA00830964FCCB13C5363961C1195BFB" author="Ádám L" journalOrPublisher="Folia Entomologica Hungarica" pagination="5-17" refId="B2" refString="Adam L (1994) A check-list of the Hungarian Scarabaeoidea with the description of ten new taxa. Folia Entomologica Hungarica 55: 5 - 17." title="A check-list of the Hungarian Scarabaeoidea with the description of ten new taxa." volume="55" year="1994">Ádám, 1994</bibRefCitation>
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<figureCitation id="03CDD9F6DC2F634DBC8529CAFAF0039D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">Figs 4 b</figureCitation>
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,
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<figureCitation id="3EBD78B25F28EA1FAD412E5BECAE7D2E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 f</figureCitation>
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,
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<figureCitation id="67C503EF00ABAB3998F4ED9BB41D0D82" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. a – i epipharynxes a – c A. niger a SV, Tullgarn b, c GB, Hampshire, New Forest d A. niger?, CZ, Hradec Králové e A. muscorum, HU, Hortobagyi f, g A. bellumgerens sp. nov., IT Piano Battaglia, paratypes h, i A. wilsonae paratypes, SP h El Vellón i Manzanares el Real. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure9" httpUri="https://binary.pensoft.net/fig/1097468">9 e</figureCitation>
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,
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<figureCitation id="CD69FE283EAB6DB8139332256BAFBC98" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Aphodius niger group, pronotal bases 1 a A. niger, Sweden, Södermanland, Tullgarn b A. cf. niger, CZ, České Budějovice c A. muscorum, HU, Hortobágyi d A. felix sp. nov., paratype." figureDoi="10.3897/zookeys.1207.117225.figure15" httpUri="https://binary.pensoft.net/fig/1097476">15 c</figureCitation>
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,
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<figureCitation id="E537DBD62B9B0625E4F7F75D29FDA693" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 n-q</figureCitation>
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<figureCitation id="36075E6D3FD35745DC27C0DE2197184A" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 j</figureCitation>
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,
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<figureCitation id="4F603B6C40FFBB77D17338486ED52ABE" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 e, g, i, j</figureCitation>
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,
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<figureCitation id="5F602FBFF342F56ED0B3FD24E9DB85D7" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">23 u</figureCitation>
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,
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<figureCitation id="5E7060204DB5BEB3450FB12BAEF7A048" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Aedeagal encasements (9 th abdominal sternite) a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. felix sp. nov., IT, Campo Felice d A. alberti sp. nov., TU, Ovit Dağı e A. bellumgerens sp. nov., IT, Sicily, Piano Battaglia f A. bameuli sp. nov., FR, Corsica, Haute-Corse g A. wilsonae, SP, Cantabria, Corconte h A. kraatzi, RU, Chita Reg. i A. plagiatus, UK, Norfolk, Hunstanton j A. plagiatus, CN, Beijing k A. rodrigoi sp. nov., SP, Madrid, Aranjuez l A. chellala sp. nov., AG, Chellala, paratype m A. rutilipennis sp. nov., CY, Konakli, paratype n A. discoides sp. nov., paratype. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure25" httpUri="https://binary.pensoft.net/fig/1097496">25 b</figureCitation>
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,
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<figureCitation id="F5E8CDE43987A3D887D04CBD8498EF4A" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure26" httpUri="https://binary.pensoft.net/fig/1097499">26 u</figureCitation>
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,
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<figureCitation id="3112097A1FD98913AAAEAB0AE618451B" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Mitotic chromosomes, arranged as karyotypes a, b A. niger, ♂, SV, Tullgarn a plain b the same nucleus C-banded (1 replicate of autosome 5 missing) c A. niger ♀, SV, Tullgarn, C-banded d, e A. niger, ♂♂, GB, Hampshire, New Forest, C-banded e with 3 B-chromosomes f, g A. muscorum, ♂, HU, Hortobágyi f plain g the same nucleus C-banded h A. muscorum, ♂, HU, Hortobágyi, C-banded i, j A. muscorum, ♀, HU, Kisújszállás i plain j C-banded k, l A. muscorum, ♀, HU, Kisújszállás k plain l the same nucleus C-banded. Scale bar: 5 μm" figureDoi="10.3897/zookeys.1207.117225.figure31" httpUri="https://binary.pensoft.net/fig/1097515">31 f – l</figureCitation>
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,
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<figureCitation id="66302996C42729053EA6334B28C9B789" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Meiotic chromosomes, metaphase I from testis a – d A. niger, GB, Hampshire, New Forest, White Moor a, c plain b, d the same nuclei C-banded e – g A. muscorum, HU, Hortobágyi e plain f, g the same nucleus f plain g C-banded h A. bameuli paratype, FR, Corsica, by Lac de Melo, plain i, j A. krelli IT, Sardinia, paratype i plain j the same nucleus C-banded k, l A. wilsonae, SP, Areños k plain l the same nucleus C-banded. The sex bivalent is indicated by an arrow. Scale bar: 5 μm." figureDoi="10.3897/zookeys.1207.117225.figure34" httpUri="https://binary.pensoft.net/fig/1097526">34 e – g</figureCitation>
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</paragraph>
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<treatmentCitation id="29F9895050F2209B4DEE31D73B6D2BCF" author="Ádám L" page="6" year="1994">
|
||
<taxonomicName id="9573B54CF796C13B572894006BE57B47" authority="Adam, 1994: 6" authorityName="Adam" authorityPageNumber="6" authorityYear="1994" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="muscorum" subGenus="Liothorax">
|
||
<emphasis id="A057C8D406E194C5469CDD4DB81E0333" italics="true">Aphodius</emphasis>
|
||
(
|
||
<emphasis id="6043F12F65F115969400602244BB6650" italics="true">Liothorax</emphasis>
|
||
)
|
||
<emphasis id="9D7A1731FCD58A49DE56A26F59644562" italics="true">muscorum</emphasis>
|
||
<comment id="5089A39B547512D2DE98C68374F99689">
|
||
<bibRefCitation id="D7654B3595AF65E2D063948D42DEE919" author="Ádám L" journalOrPublisher="Folia Entomologica Hungarica" pagination="5-17" refId="B2" refString="Adam L (1994) A check-list of the Hungarian Scarabaeoidea with the description of ten new taxa. Folia Entomologica Hungarica 55: 5 - 17." title="A check-list of the Hungarian Scarabaeoidea with the description of ten new taxa." volume="55" year="1994">Ádám, 1994: 6</bibRefCitation>
|
||
.
|
||
</comment>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EBTAM" type="type material">
|
||
<paragraph id="48B500BFF275213C4BD6737093017C4D">
|
||
<heading id="418DADFD8FA83A79C48D46326057D1A4" reason="title">Type material examined.</heading>
|
||
</paragraph>
|
||
<paragraph id="E4A95B971AE10457FBA24986C7930F9C">
|
||
The
|
||
<typeStatus id="5BEAF44D39EEAEBBD544950785C28044" type="holotype">holotype</typeStatus>
|
||
(Fig.
|
||
<figureCitation id="70F3253012B0DF83319715661064F1C4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">4 b</figureCitation>
|
||
) has five labels: 1: Hung.
|
||
<collectingRegion id="B4F08F208B9BB22E88B588780D5919DB" country="Hungary" name="Borsod-Abauj-Zemplen">Borsod-Abaúj-Zemplén</collectingRegion>
|
||
m., Aggtelek Vörös-tó,
|
||
<specimenCount id="45CB915881F7426E29E0E02D059BB12D" count="350" type="generic">
|
||
<quantity id="D3D68F18DAFB4DB79506C7C4464EAFDD" metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">350 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
2: Typhetum angustifoliae-latifoliae. 3: iszapból (= from mud)
|
||
<date id="DB07E5E9030831969A5C43FFAD897899" value="1989-05-19">1989. v. 19.</date>
|
||
, leg Ádám
|
||
<emphasis id="BCE17CD81540915981607BC16B37BBD0" italics="true">L.</emphasis>
|
||
4:
|
||
<typeStatus id="2999104A0D391A12FE1EDD6B3A8E5E02" type="paratype">Paratypus</typeStatus>
|
||
<taxonomicName id="D497880B474BBB204612FC04364E4265" authority="Adam, 1994" authorityName="Adam" authorityYear="1994" class="Insecta" family="Scarabaeidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="muscorum">
|
||
<emphasis id="50B90039E02E83A92C0A699103397725" italics="true">Liothorax muscorum</emphasis>
|
||
<bibRefCitation id="6F6A62AF5D685D9D3970A13A9EC0102A" author="Ádám L" journalOrPublisher="Folia Entomologica Hungarica" pagination="5-17" refId="B2" refString="Adam L (1994) A check-list of the Hungarian Scarabaeoidea with the description of ten new taxa. Folia Entomologica Hungarica 55: 5 - 17." title="A check-list of the Hungarian Scarabaeoidea with the description of ten new taxa." volume="55" year="1994">Ádám, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
(A red label, incorrect because Adam mentioned only a single
|
||
<typeStatus id="7604EB9E408C7A1DA4A39D134825A32E" type="holotype">holotype</typeStatus>
|
||
with these data). 5:
|
||
<typeStatus id="0E026E061592B7133BBD66F883F4C893" type="holotype">HOLOTYPE</typeStatus>
|
||
<taxonomicName id="DA2DB145A51134166B7F2BD7E57C295E" authority="Adam. R. B." authorityName="Adam. R. B." class="Insecta" family="Scarabaeidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="muscorum" subGenus="Liothorax">
|
||
<emphasis id="4C7F73A7D916794BFFD795A9E912376E" italics="true">A.</emphasis>
|
||
(
|
||
<emphasis id="6616D99BEE9E566B4A732784B0D6914E" italics="true">Liothorax</emphasis>
|
||
)
|
||
<emphasis id="585EDA9CF0D2C03A0EE576FC40EEA3FC" italics="true">muscorum</emphasis>
|
||
Adam. R. B.
|
||
</taxonomicName>
|
||
<collectingRegion id="CEDD371990757E746756C50AD88E5839" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||
det. 2017. It is a female, so only somatic characters are available for assessment (
|
||
<collectionCode id="C53F939A0423A8928AEE6FD5E2230F42" country="Hungary" httpUri="http://grbio.org/cool/9j47-dvjw" name="Museum of Natural Sciences" type="">MNSB</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EKUAM" type="material">
|
||
<paragraph id="B5B1222314FAB661B0AE514D9212330A">
|
||
<heading id="50D2FB4CB886C29AB2367ACC3C050BFF" reason="title">Additional material examined.</heading>
|
||
</paragraph>
|
||
<paragraph id="03702C9410D67046647ADFE8A900B3A1">
|
||
HU: Kisújszállás,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETUAM" specific-use="{"type":"Point","coordinates":[20.700000,47.212000]}">
|
||
<geoCoordinate id="0D667E964A074B27930F8688BB2AC41F" degrees="47.212" direction="north" orientation="latitude" precision="55" value="47.212">47.212 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="8D015201AB237EBAE261467B9F83D89C" degrees="20.700" direction="east" orientation="longitude" precision="55" value="20.7">20.700 ° E</geoCoordinate>
|
||
</named-content>
|
||
,
|
||
<date id="74B8F085B875D60385A8B287F5EEE943" value="2012-04">iv. 2012</date>
|
||
, R. B.
|
||
<collectingRegion id="A82B459D0625FC2F81A77446DFC94413" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||
,
|
||
<specimenCount id="014B6A6CDCDF9E8B44911D868D63A3FA" count="2" type="female">2 ♀♀</specimenCount>
|
||
(
|
||
<named-content id="99A78C03658CE38A23B62CDDD3090AD1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||
<collectionCode id="8445D32AA8C5423CE7DA1C2AFD3D92FE" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||
</named-content>
|
||
). Hortobagy National Park, Kis-Kecskés area,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6UAM" specific-use="{"type":"Point","coordinates":[21.060670,47.677210]}">
|
||
<geoCoordinate id="ACD2CD197E13ACCC747128724C03D26B" degrees="47.67721" direction="north" orientation="latitude" precision="1" value="47.67721">47.67721 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="4A5F395189EC80C529463582572DAE9C" degrees="21.06067" direction="east" orientation="longitude" precision="1" value="21.06067">21.06067 ° E</geoCoordinate>
|
||
</named-content>
|
||
. Washed from mud at edge of a lake, and swarming in mud at edge of lake,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHVAM" specific-use="{"type":"Point","coordinates":[21.074510,47.677720]}">
|
||
<geoCoordinate id="00917C2D4181DF2AE947A74CB22A852B" degrees="47.67772" direction="north" orientation="latitude" precision="1" value="47.67772">47.67772 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F886CEDD85D829B2AB9217306DE70817" degrees="21.07451" direction="east" orientation="longitude" precision="1" value="21.07451">21.07451 ° E</geoCoordinate>
|
||
</named-content>
|
||
.
|
||
<date id="5160E0EBE3855678BCACFAD1B4485D3D" value="2015-04-24">24. iv. 2015</date>
|
||
,
|
||
<specimenCount id="50AE188DF4CA1122669B457B350A6509" count="16" type="male">16 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="3C1EE3566C68948EF4111816E86BA14A" count="18" type="female">18 ♀♀</specimenCount>
|
||
. R. B.
|
||
<collectingRegion id="03498A12D478E7D3A33434634F529C89" country="United Kingdom" name="Angus">Angus</collectingRegion>
|
||
(
|
||
<named-content id="6A95105C871D7C70B9CD7DB27A4AEA09" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||
<collectionCode id="A8A476F981F5D42A681BA73ABAFFFF33" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||
</named-content>
|
||
,
|
||
<collectionCode id="A10EDAF2945D2C563EA7F53EA79293FC">JFM</collectionCode>
|
||
). Aggteleki NP, Josvafö, Szelce-völgy
|
||
<specimenCount id="D62B526151EC91B6C68F38CF8BECEBCC" count="1" type="female">1 ♀</specimenCount>
|
||
, 1 unsexed (
|
||
<collectionCode id="118B71CE870DE0CFBD233C1C9E08101C" country="Hungary" httpUri="http://grbio.org/cool/9j47-dvjw" name="Museum of Natural Sciences" type="">MNSB</collectionCode>
|
||
). Aggteleki NP, Josvafö, Menes-völgy. 1 unsexed (
|
||
<collectionCode id="6D74F0342B955C0FDE2DA1F34078115E" country="Hungary" httpUri="http://grbio.org/cool/9j47-dvjw" name="Museum of Natural Sciences" type="">MNSB</collectionCode>
|
||
). Bukk NP
|
||
<specimenCount id="EFFF592C663DCFD8D422D545497A2BDD" count="1" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="B1D3B0E692A4D12644C396194BC37312" count="2" type="female">2 ♀♀</specimenCount>
|
||
, 3 unsexed (
|
||
<collectionCode id="87E1A02F888B1C4D39CEFF7016DA47C8" country="Hungary" httpUri="http://grbio.org/cool/9j47-dvjw" name="Museum of Natural Sciences" type="">MNSB</collectionCode>
|
||
). Hortobágyi N. P.,
|
||
<date id="EE2934C9E88F7C2D28A0174539EA56A3" value="1980-06-03">3. vi. 1980</date>
|
||
, leg. U. Göllner, 2 unsexed (
|
||
<abbrev id="ABBRID0EQVAM" xlink_title="Axel Bellmann Collection">
|
||
<collectionCode id="555FFAD6169691F2AB79591D91D40FA9">ABC</collectionCode>
|
||
</abbrev>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EUVAM" type="Differential diagnosis">
|
||
<paragraph id="D0BBBDAA53D1745A4921AB147DF88F35">
|
||
<heading id="94AEA159657C427733653E9057602435" reason="title">Differential diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="FF6C05D9E70AF017FCE8E2C892BD854A">
|
||
Closely resembling
|
||
<taxonomicName id="E3BBFB6D23EC691625C16941ECA5C67D" authorityName="Illiger" authorityYear="1798" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="3F1CDF1F09A576EC1990E486D1BE5F03" italics="true">A. niger</emphasis>
|
||
</taxonomicName>
|
||
but clearly distinguished as a separate species by the short, almost dot-like, Y chromosome (Fig.
|
||
<figureCitation id="C42597040814A27CDB43ED8B5B0D8960" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Mitotic chromosomes, arranged as karyotypes a, b A. niger, ♂, SV, Tullgarn a plain b the same nucleus C-banded (1 replicate of autosome 5 missing) c A. niger ♀, SV, Tullgarn, C-banded d, e A. niger, ♂♂, GB, Hampshire, New Forest, C-banded e with 3 B-chromosomes f, g A. muscorum, ♂, HU, Hortobágyi f plain g the same nucleus C-banded h A. muscorum, ♂, HU, Hortobágyi, C-banded i, j A. muscorum, ♀, HU, Kisújszállás i plain j C-banded k, l A. muscorum, ♀, HU, Kisújszállás k plain l the same nucleus C-banded. Scale bar: 5 μm" figureDoi="10.3897/zookeys.1207.117225.figure31" httpUri="https://binary.pensoft.net/fig/1097515">31 f – h</figureCitation>
|
||
). Elytral interstices often with fine but distinct reticulation, giving the surface a leaden-grey silky sheen. Metaventrite of females variously depressed medially, depressed area often partly reticulate (Fig.
|
||
<figureCitation id="0324797DA14E91E83CD9DBDBF47AE465" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 e, g – j</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ENWAM" type="redescription">
|
||
<paragraph id="D54E6A3B605223F5E8337FD77592EDC3">
|
||
<heading id="DFBCED6F754DDC4E1DBCE95DC4E69EFC" reason="title">Redescription.</heading>
|
||
</paragraph>
|
||
<paragraph id="5A537DE4AC3CC899E75A9309B3A59477">
|
||
General appearance (Fig.
|
||
<figureCitation id="8E180F6E7D7ECE28C216EB392DBD5117" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure4" httpUri="https://binary.pensoft.net/fig/1097462">4 b</figureCitation>
|
||
). Length:
|
||
<quantity id="27CB6EAA8734A94F118C06C1DE39E3A9" metricMagnitude="-3" metricUnit="m" metricValue="4.7" metricValueMax="5.2" metricValueMin="4.2" unit="mm" value="4.7" valueMax="5.2" valueMin="4.2">4.2–5.2 mm</quantity>
|
||
(
|
||
<specimenCount id="F9D92CFAE071D0FD0646F5BE69D47ABE" count="1" type="male">♂</specimenCount>
|
||
),
|
||
<quantity id="5CEA57A4EB59E0CF4D88E8757E4C37DE" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.2" metricValueMin="4.3" unit="mm" value="4.75" valueMax="5.2" valueMin="4.3">4.3–5.2 mm</quantity>
|
||
(
|
||
<specimenCount id="9B1B87E78769F88705EDD66B9084EE72" count="1" type="female">♀</specimenCount>
|
||
). Width:
|
||
<quantity id="2C338C877DC38FB1AC0D2C1D8595D540" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.35" metricValueMin="1.85" unit="mm" value="2.1" valueMax="2.35" valueMin="1.85">1.85–2.35 mm</quantity>
|
||
(
|
||
<specimenCount id="19B8429ACF47F4E22E88F709F04E12E2" count="1" type="male">♂</specimenCount>
|
||
),
|
||
<quantity id="EB96E1C64F5D5AA90D2B61350A029173" metricMagnitude="-3" metricUnit="m" metricValue="2.0999999999999996" metricValueMax="2.3" metricValueMin="1.9" unit="mm" value="2.0999999999999996" valueMax="2.3" valueMin="1.9">1.9–2.3 mm</quantity>
|
||
(
|
||
<specimenCount id="42494B1F34CAB9B3CA3F6B5B9AAD95B1" count="1" type="female">♀</specimenCount>
|
||
). Black, head, pronotum and scutellum glossy (pronotum sometimes less so), elytra with interstices distinctly reticulate, giving a silky lead-grey sheen. Head: glossy black, rather finely punctate, the punctures separated by 2–3 × their diameter. Some slightly larger punctures are present. Frontoclypeal suture very fine, with a glossy impunctate area either side of it. Median 2 / 3 almost straight, its mid-point and lateral ends slightly raised. Suture angled forward outside the lateral elevations, reaching the margin of the head at the front of the genae. Genae (preocular lobes) protrude laterally at an angle of ca 45 ˚ in front of eyes, then curve inwards to meet the frontoclypeal suture ca 1.5 × eye-diameter in front of eyes. Clypeal margin obtusely angled outward from this point, then curved inwards to the median excision. Median area of clypeus bulging upwards. Genae laterally with a narrow raised margin, this continued across the clypeus, becoming weaker medially. The punctation of the frons is sometimes stronger, with a clear distinction between larger punctures and fine dot-like ones. Antennae and maxillary palpi black to blackish brown. Epipharynx (Fig.
|
||
<figureCitation id="6940F910B07D6EDB58B98C57B7A7B2DC" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. a – i epipharynxes a – c A. niger a SV, Tullgarn b, c GB, Hampshire, New Forest d A. niger?, CZ, Hradec Králové e A. muscorum, HU, Hortobagyi f, g A. bellumgerens sp. nov., IT Piano Battaglia, paratypes h, i A. wilsonae paratypes, SP h El Vellón i Manzanares el Real. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure9" httpUri="https://binary.pensoft.net/fig/1097468">9 e</figureCitation>
|
||
) similar to that of
|
||
<taxonomicName id="0DF2831F88D986039ECDBFD76D2EE55C" authorityName="Illiger" authorityYear="1798" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="D83344EEBF24699872F39BD671272685" italics="true">A. niger</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="6204C8AA1D6DBDA17877DE5809831EFD" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. a – i epipharynxes a – c A. niger a SV, Tullgarn b, c GB, Hampshire, New Forest d A. niger?, CZ, Hradec Králové e A. muscorum, HU, Hortobagyi f, g A. bellumgerens sp. nov., IT Piano Battaglia, paratypes h, i A. wilsonae paratypes, SP h El Vellón i Manzanares el Real. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure9" httpUri="https://binary.pensoft.net/fig/1097468">9 a – c</figureCitation>
|
||
). Head and pronotum (Fig.
|
||
<figureCitation id="EF40A5CAB544DAC9BDD14840839EC416" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure6" httpUri="https://binary.pensoft.net/fig/1097465">6 f</figureCitation>
|
||
). Pronotum: hemi-cylindrical, glossy black and with double punctation. The intensity of the punctation is variable, with the larger punctures in some specimens confined to the lateral and posterior 1 / 4 s of its surface, while in others it extends over the whole surface and in some cases both the large and small punctures are heavily impressed, giving a roughened appearance to the surface. There is often an ill-defined narrow median band more or less free of punctures, extending from behind the middle of the disc almost to the hind margin. Lateral margins with a distinct raised bead, this either black or dark brown and extending round the hind corners often giving a continuous narrow raised border over the entire basal margin. It is necessary to tilt the specimen to trace the full course of this border, which is sometimes absent either side of the median 1 / 3, or from the middle 1 / 3 of the margin (Fig.
|
||
<figureCitation id="38446757F792358DA97A3714340A3B3D" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Aphodius niger group, pronotal bases 1 a A. niger, Sweden, Södermanland, Tullgarn b A. cf. niger, CZ, České Budějovice c A. muscorum, HU, Hortobágyi d A. felix sp. nov., paratype." figureDoi="10.3897/zookeys.1207.117225.figure15" httpUri="https://binary.pensoft.net/fig/1097476">15 c</figureCitation>
|
||
). Lateral margins visible from above over the anterior 1 / 2, behind this varying from being completely visible to totally obscured by the bulging pronotal surface. Scutellum: elongate, sides parallel in basal 1 / 2, then angled inwards to bluntly rounded apex. Surface glossy black, with sparse fine punctures. Elytra: black but with lead-grey silky sheen resulting from fine but distinct reticulation on the interstices. The strength of the reticulation varies from very weak in the
|
||
<typeStatus id="B3EF0B4A7614703E095AD1C9F0807DF4" type="holotype">holotype</typeStatus>
|
||
(Fig.
|
||
<figureCitation id="975AC917092147C615A782BE6E3FBA55" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 n</figureCitation>
|
||
) to rather stronger in some Hortobágyi material (Fig.
|
||
<figureCitation id="441BE0D1B10B46F3F816C71017D4DC34" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 o</figureCitation>
|
||
) and stronger still in the
|
||
<specimenCount id="11480F730903DCE17C7E0CD7D3BA766D" count="2" type="female">two females</specimenCount>
|
||
from Kisújszállás (Fig.
|
||
<figureCitation id="CD448ECB3DF6E1E5BFE7BF11773F28CE" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1207.117225.figure17" httpUri="https://binary.pensoft.net/fig/1097480">17 p, q</figureCitation>
|
||
). This reticulation may be obscured if the specimen is at all greasy. Interstices flat, ca 6 × wider than the striae, with very fine punctures, these ca the same size as the meshes of the reticulation. Striae narrow, vertical-sided, their bottoms glossy. Strial punctures in single rows, small, separated by ca 3 × their diameter, and excising the strial margins.
|
||
</paragraph>
|
||
<paragraph id="85BDF7D01B959D649DD19DBB3CD437C6">
|
||
Metaventrite: median diamond-shaped plate in male finely and sparsely punctate, concave, depressed to median longitudinal groove (Fig.
|
||
<figureCitation id="59646AC41BE28770613D150A90928BB8" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">
|
||
<quantity id="2363ED07E5137B9E850AABF32D0803AA" metricMagnitude="-2" metricUnit="kg" metricValue="2.1" unit="g" value="21.0">21 g</quantity>
|
||
</figureCitation>
|
||
); in female variable, sometimes median longitudinal groove flattened, but with the median area depressed anteriorly and sometimes with some reticulate sculpture (Fig.
|
||
<figureCitation id="24E45F54BD8012A0EBB43E71C14304D4" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1207.117225.figure21" httpUri="https://binary.pensoft.net/fig/1097488">21 i, j</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8D857D0B44BD024F99BA4B9421BF6A5B">
|
||
Legs dark brown, often blackish, basal segment of mesotarsi normally longer than longer mesotibial spur (Fig.
|
||
<figureCitation id="2AB1BC8514B8F565BDA799BA5FFC0A72" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1207.117225.figure18" httpUri="https://binary.pensoft.net/fig/1097482">18 j</figureCitation>
|
||
) but sometimes equal in length or slightly shorter.
|
||
</paragraph>
|
||
<paragraph id="C3D9475AC62EFF15058477426EDF2BC2">
|
||
Aedeagus: Fig.
|
||
<figureCitation id="F340E9D0345B00FA5A2B7DC76514E629" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." figureDoi="10.3897/zookeys.1207.117225.figure23" httpUri="https://binary.pensoft.net/fig/1097492">23 u</figureCitation>
|
||
. Endophallus with heavy recurved teeth. Length ca
|
||
<quantity id="7BC760609EB7B6A2CD7AD1F96D3429D0" metricMagnitude="-3" metricUnit="m" metricValue="1.06" unit="mm" value="1.06">1.06 mm</quantity>
|
||
, paramere length ca
|
||
<quantity id="CCD346428BACD21614ABC2B997626F4F" metricMagnitude="-4" metricUnit="m" metricValue="5.4" unit="mm" value="0.54">0.54 mm</quantity>
|
||
, basal piece length ca
|
||
<quantity id="1D17B8FE1C131C25B53425DBE747437E" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" unit="mm" value="0.69">0.69 mm</quantity>
|
||
. Length of tooth-field ca
|
||
<quantity id="032CEFC0891139BF26B0E8A893FAFD14" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" unit="mm" value="0.57">0.57 mm</quantity>
|
||
, of longest teeth on endophallus 47–58 μm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EUYAM" type="remarks">
|
||
<paragraph id="F642AC7AA22EE124518D5F2148707B70">
|
||
<heading id="CBF8E00F780303A3CFB0805CD1071A36" reason="title">Remarks.</heading>
|
||
</paragraph>
|
||
<paragraph id="C8535B508EAC7EB17332F6DEA4203EA8">
|
||
As already mentioned, Ádám described
|
||
<taxonomicName id="3844D70DED7D877F13AA8F651334E5E9" class="Insecta" family="Scarabaeidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="muscorum">
|
||
<emphasis id="1606FE1BB383ABCFC32BB0D546919697" italics="true">A. muscorum</emphasis>
|
||
</taxonomicName>
|
||
as a new species, intending it to be the valid name for the species then known as
|
||
<taxonomicName id="CEE679A31826D57BC0A016A39F0CEE5F" authority="(Panzer, 1797)" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger" subGenus="Liothorax">
|
||
<emphasis id="0BC713228998BE477FED69BAB03358E9" italics="true">Aphodius</emphasis>
|
||
(
|
||
<emphasis id="3841B4A2A2B400794F2104A670761DBD" italics="true">Liothorax</emphasis>
|
||
)
|
||
<emphasis id="7628277766DC3DD2FE1DBFDEB1796282" italics="true">niger</emphasis>
|
||
(Panzer, 1797)
|
||
</taxonomicName>
|
||
. However, chromosomal research by RBA has shown that Hungarian material is in fact different from true
|
||
<taxonomicName id="46AA40FD3F9D6F652B46D63AD2C8B339" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="4AAFFCFA5EAE27C30392EB59C9C53478" italics="true">A. niger</emphasis>
|
||
</taxonomicName>
|
||
, so Ádám’s
|
||
<taxonomicName id="D6C1938B032FC6939579CF60E47D6DBE" class="Insecta" family="Scarabaeidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="muscorum">
|
||
<emphasis id="04EA1A82C67BD48A55E6A0FA83D872C0" italics="true">Liothorax muscorum</emphasis>
|
||
</taxonomicName>
|
||
must be regarded as a valid species. Ádám gives a description which, from its reference to variation in the strength of the elytral sculpture, is based on a fairly wide selection of Hungarian material. However, he lists only
|
||
<specimenCount id="C0DA94E6EC183A637B3BAFAF51E1F650" count="1" type="generic">one type</specimenCount>
|
||
specimen, the
|
||
<typeStatus id="8B397BD9818DBF134584432E8B0330E3" type="holotype">holotype</typeStatus>
|
||
from the lake Vörös-tó in the Aggteleki national park in NE
|
||
<collectingCountry id="83D6899E8C0F7AF1FBBA2C7BC456B77A" name="Hungary">Hungary</collectingCountry>
|
||
, adding that he took the unsexed specimen from mud in a drying out
|
||
<taxonomicName id="0B1797FD6CF635227469A942E0A1C6DA" class="Liliopsida" family="Typhaceae" genus="Typha" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="46074A5332A9584D7B264CDD7CC5732B" italics="true">Typha</emphasis>
|
||
</taxonomicName>
|
||
bed. Vörös-tó is now a fairly deep lake with limited development of
|
||
<taxonomicName id="E04E85D192C0CA05D7E667EDB3F77920" class="Liliopsida" family="Poaceae" genus="Phragmites" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="688D935B1DE1E7A8674A60AAE1E8E290" italics="true">Phragmites</emphasis>
|
||
</taxonomicName>
|
||
round its edge. It was dredged out when the nearby entrance to the Baradla Barlang (cave) was developed (Sandor Rozsa, Chief Warden of the National Park, pers. comm. to RBA,
|
||
<date id="037E61CB8F563F205367A09E6B808538" value="2012-04">April 2012</date>
|
||
). The nearby Aggteleki-tó has extensive development of mud and
|
||
<taxonomicName id="96E183A3FC0488E0F890EE45F38FDEED" class="Liliopsida" family="Typhaceae" genus="Typha" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8D4CA0B92776C35621155A36989841B1" italics="true">Typha</emphasis>
|
||
</taxonomicName>
|
||
, but though RBA has taken
|
||
<taxonomicName id="A43052E321AC3DBB1C93B4561B423A15" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="varians" subGenus="Nialus">
|
||
<emphasis id="C887451E9C81F8006AB669214631AA31" italics="true">A.</emphasis>
|
||
(
|
||
<emphasis id="70FF90191182FA255C310354F94C423A" italics="true">Nialus</emphasis>
|
||
)
|
||
<emphasis id="E0D8CCAB5D97B301880A2F3329A414A4" italics="true">varians</emphasis>
|
||
</taxonomicName>
|
||
in both lakes he failed to find the
|
||
<taxonomicName id="301D7061B6D87AC30E352A069C3D284E" class="Insecta" family="Scarabaeidae" genus="Liothorax" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E17F9FF0EF924818A4B18AE8AE4C0893" italics="true">Liothorax</emphasis>
|
||
</taxonomicName>
|
||
. It is therefore necessary to rely on the
|
||
<typeStatus id="52B6FE905679DAA58DE21AB2D735D2F9" type="holotype">holotype</typeStatus>
|
||
and a limited amount of material from the neighbourhood, in the MNSB, for comparison with extensive material from the Hortobágyi National Park, from which chromosomes and DNA are available.
|
||
</paragraph>
|
||
<paragraph id="1878D4678D47A9326DE26B840F3C9F12">
|
||
<taxonomicName id="DE5244397E4C061C21811A80EC6F9549" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="muscorum" status="stat. nov.">
|
||
<emphasis id="4360C89C8B1F7C191B1EA2F1136FA2E3" italics="true">Aphodius muscorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C06B07C94C8829FE1664FF387B4A2C3B">stat. nov.</taxonomicNameLabel>
|
||
is at present reliably identified only from
|
||
<collectingCountry id="81A0CFEE10ABC0F39B778178690C6FC1" name="Hungary">Hungary</collectingCountry>
|
||
. It appears to be a steppe species which may have a wider distribution eastwards but this needs to be verified with
|
||
<taxonomicName id="8F1E38801CEA7383273958FEBE32479B" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="E39AA61A683E763C84DC613A8AD31380" italics="true">A. niger</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BBFF7A4FB3A572FD957E9358AC79E666" sensu="lato">s. l.</taxonomicNameLabel>
|
||
material from adjacent areas further east (Fig.
|
||
<figureCitation id="1B3F199BE53B2F73DF99600733B0AF0C" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29 a</figureCitation>
|
||
). RBA took the species abundantly in lakeside mud in the Hortobágy National Park in
|
||
<date id="60BD0511B3F0798D4841F60853F5F365" value="2015-04">April 2015</date>
|
||
(Fig.
|
||
<figureCitation id="23EA44CC4C63899DCDCA08A7EBD6F886" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Habitat pictures a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. niger, SW, Tullgarn d A. krelli sp. nov., IT, Sardinia, Badde Salighes e A. bameuli sp. nov., FR, Corsica, Haute-Corse, Lac de Melo f A. wilsonae, SP, Cantabria, Corconte g A. bellumgerens sp. nov., IT, Sicily, Nebrodi." figureDoi="10.3897/zookeys.1207.117225.figure27" httpUri="https://binary.pensoft.net/fig/1097502">27 b</figureCitation>
|
||
), and
|
||
<specimenCount id="4CB6955C2A16E50E177A4D5F43ECA5C7" count="2" type="generic">2 specimens</specimenCount>
|
||
in the mud of a drying out ditch near Kisújszállás in
|
||
<date id="AA0FC17E9D427BF477553B62407664ED" value="2012-04">April 2012</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4A2B6CC145DE0C27FBC346076A981FB2">
|
||
<title id="24613FD9987581C0324F83967D2F32AB">
|
||
<taxonomicName id="B691F59E72F7D95EBBD91837FC9BC3AE" authorityName="Hellwig" authorityYear="1798" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B689AB7B2C4A5E65C58BE38F65C6CCA9" italics="true">Aphodius</emphasis>
|
||
</taxonomicName>
|
||
" cf
|
||
<taxonomicName id="8974E1CE6B1BFD2BDAE6BE91CCD2DC6D" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="61E505DCC4868FDABC2E68D575753C4F" italics="true">A. niger</emphasis>
|
||
</taxonomicName>
|
||
": species resembling
|
||
<taxonomicName id="20B77C4437F5D2559349C6B20AAD81FD" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="1C1CB245A88D01701BD2933E7A9EFC31" italics="true">A. niger</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E807972F1B2E849267910D19D8EDD99B" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="muscorum">
|
||
<emphasis id="EDA3F8D41F464679031F1E72A04EF592" italics="true">A. muscorum</emphasis>
|
||
</taxonomicName>
|
||
, but of uncertain identity
|
||
</title>
|
||
</paragraph>
|
||
<paragraph id="3EE661FD778215264652785235941151">
|
||
<emphasis id="B358DA55042073F61660491C66AFFF5D" bold="true">Material examined.</emphasis>
|
||
RU:
|
||
<collectingRegion id="960D9077FA862BE4F8D09E1CB81524C5" country="Russia" name="Volgograd">Volgograd</collectingRegion>
|
||
,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EH5AM" specific-use="{"type":"Point","coordinates":[44.529000,48.739000]}">
|
||
<geoCoordinate id="DBCB2923709BD9BE43ADCF461404BDF6" degrees="48.739" direction="north" orientation="latitude" precision="55" value="48.739">48.739 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F2572F34914C7F5CEA70F69E9C1F3ABE" degrees="44.529" direction="east" orientation="longitude" precision="55" value="44.529">44.529 ° E</geoCoordinate>
|
||
</named-content>
|
||
.
|
||
<specimenCount id="236A1086D889702A439DE4D995F6C6D7" count="1" type="male">1 ♂</specimenCount>
|
||
(
|
||
<named-content id="BBD5B205160CD01614633E246740F02E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||
<collectionCode id="9D68E222708267862E7E4A88C56CF490" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||
</named-content>
|
||
).
|
||
<collectingRegion id="6509751023CD51081C461087A6D61D54" country="Russia" name="Tyumen'">Tyumen</collectingRegion>
|
||
.
|
||
<geoCoordinate id="6730B1D2D49CD7D0B4F293A3C7A4285C" degrees="57.215" direction="north" orientation="latitude" precision="55" value="57.215">57.215 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="927F4F13ACB8256AADDD0B4633C67A9E" degrees="65.574" direction="east" orientation="longitude" precision="55" value="65.574">65.574 ° E</geoCoordinate>
|
||
.
|
||
<specimenCount id="90D37C0447701B9B5BD7BE4D813B0CF7" count="1" type="male">1 ♂</specimenCount>
|
||
. (
|
||
<named-content id="BFC6938E49CB84E763F25610BD36CEB4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||
<collectionCode id="71469CB3D4D64B37F1BF4655CB956CDA" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||
</named-content>
|
||
). Tuyanso, Chernomorsk oblast’.
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EX5AM" specific-use="{"type":"Point","coordinates":[30.517000,44.310000]}">
|
||
<geoCoordinate id="CC9127DC4DAB8AE61A4070956E180D10" degrees="44.310" direction="north" orientation="latitude" precision="55" value="44.31">44.310 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="65D4EFBF1F04CFE8FB26B0B078B14165" degrees="30.517" direction="east" orientation="longitude" precision="55" value="30.517">30.517 ° E</geoCoordinate>
|
||
</named-content>
|
||
. 1915.
|
||
<specimenCount id="A1FDC6549F2B24FC80C50DCB58B4DA97" count="1" type="male">1 ♂</specimenCount>
|
||
(
|
||
<named-content id="BA01D96C93B0EE332C5F30DD29ACF69F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||
<collectionCode id="63CD443AB37788BF86FB02BFACE77643" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||
</named-content>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="B59E59AAA214A2BCB5F3EADDE6710A67">
|
||
KZ: 10 KM SE of Akmolinsk (Astana)
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EF6AM" specific-use="{"type":"Point","coordinates":[71.720000,51.085000]}">
|
||
<geoCoordinate id="460E50547D9D143394AEA22DE4ABB4D0" degrees="51.085" direction="north" orientation="latitude" precision="55" value="51.085">51.085 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3F5666CCDBFD8D4DA4A76BAF6E470B75" degrees="71.720" direction="east" orientation="longitude" precision="55" value="71.72">71.720 ° E</geoCoordinate>
|
||
</named-content>
|
||
. Under detritus by the water.
|
||
<date id="7AC7511070639A761D0336758715BCB5" value="1957-05-05">5. v. 1957</date>
|
||
. E.
|
||
<emphasis id="A74DB1B4366145443B7C66A4AC044D11" italics="true">L.</emphasis>
|
||
Gurjeva.
|
||
<specimenCount id="F7E37418786E37D274C1A479D6B12649" count="1" type="female">1 ♀</specimenCount>
|
||
(
|
||
<named-content id="36E3B2C2BF2CA3C5147DA21147A3CFEC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||
<collectionCode id="2EBA8AB1009B3D54125C0B54E4A93391" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||
</named-content>
|
||
). Akmola obl. Kokshipan. By R. Tirs, Marsh on floodplain with
|
||
<taxonomicName id="ECE151D111145E0002AD886D8EC98798" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" kingdom="Plantae" order="Ranunculales" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B23BA746D7F88DBD391B9C8B699AD32A" italics="true">Ranunculus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3544C954239555304ED9D3F19B033DCB" authorityName="Fol" authorityYear="1872" class="Appendicularia" family="Fritillariidae" genus="Fritillaria" kingdom="Animalia" order="Copelata" phylum="Chordata" rank="genus">
|
||
<emphasis id="95258D45C8E056720D83330B3E58F379" italics="true">Fritillaria</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<date id="774919319B7AE5A6D26B314C5EB3274D" value="1957-05-13">13. v. 1957</date>
|
||
. E.
|
||
<emphasis id="7ED28C470A370554FF9CC536106598E1" italics="true">L.</emphasis>
|
||
Gurjeva.
|
||
<specimenCount id="A9DF7B34DC1050BFCA0CD77438C385D5" count="1" type="male">1 ♂</specimenCount>
|
||
(
|
||
<named-content id="81C5497DB55FE28A59C93C524D6AA5D0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||
<collectionCode id="2F525B2FE05615F0503CE157D65299E0" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||
</named-content>
|
||
). Akmola oblast’, dolina Basaga-ozik.,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPABM" specific-use="{"type":"Point","coordinates":[65.991000,49.840000]}">
|
||
<geoCoordinate id="143C0F544FE3216F818660170E3DC17D" degrees="49.840" direction="north" orientation="latitude" precision="55" value="49.84">49.840 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="FF7867CF9536F4CE6369D44C32BC5975" degrees="65.991" direction="east" orientation="longitude" precision="55" value="65.991">65.991 ° E</geoCoordinate>
|
||
</named-content>
|
||
. In a Solonchak (saltmarsh).
|
||
<emphasis id="B453D53E8C5840266F8DE8F5C462EB2D" italics="true">L.</emphasis>
|
||
Arnoldi, 27,
|
||
<date id="1EF11B69CE7B563B7F5C464F079214B8" value="1957-05">v. 1957</date>
|
||
. (
|
||
<named-content id="1FB02E8FA4E25243A7ADA04474D853EA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/russian-academy-sciences-zoological-institute-zoological-museum" xlink_title="Zoological Institute, Russian Academy of Sciences St Petersburg">
|
||
<collectionCode id="4C44626943866D55011CA774D0336DBC" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||
</named-content>
|
||
); Uralsk
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBBBM" specific-use="{"type":"Point","coordinates":[51.271000,51.214000]}">
|
||
<geoCoordinate id="9969BB3BF64ECD6B69972F7EEB33B877" degrees="51.214" direction="north" orientation="latitude" precision="55" value="51.214">51.214 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="0588D74CA7D372B42A62246301718B0C" degrees="51.271" direction="east" orientation="longitude" precision="55" value="51.271">51.271 ° E</geoCoordinate>
|
||
</named-content>
|
||
.
|
||
<specimenCount id="B4790F31CE6A62206305BED769E0548B" count="1" type="male">1 ♂</specimenCount>
|
||
(
|
||
<named-content id="97E4DDD71A5EBF89B1AF7D12D8E5E374" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-0" xlink_title="National Museum Prague">
|
||
<collectionCode id="7651F32614ABAEDF35ABE8A8ED5330A4" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||
</named-content>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E59174FEDF5E914755EAEDCC732D1FF0">
|
||
CZ:
|
||
<collectingRegion id="AF633CB2DCB1DAB57FD43961CA5234E0" country="Czech Republic" name="Kralovehradecky">Hradec Králové</collectingRegion>
|
||
, Na Plachtě.
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPBBM" specific-use="{"type":"Point","coordinates":[15.859000,50.186000]}">
|
||
<geoCoordinate id="40270BE1DF5E81B67FB2A4C36DCD66FD" degrees="50.186" direction="north" orientation="latitude" precision="55" value="50.186">50.186 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="596901D4C8F4B6F89D76ED94F2872D56" degrees="15.859" direction="east" orientation="longitude" precision="55" value="15.859">15.859 ° E</geoCoordinate>
|
||
</named-content>
|
||
.
|
||
<specimenCount id="C900D3DFD31CCDAC7363DC8217DFFC4A" count="1" type="male">1 ♂</specimenCount>
|
||
, 1 unsexed (
|
||
<named-content id="652EE3ABA7283057107243960E0181A2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||
<collectionCode id="C6ED428602799F5E7049A7BD0EF94276" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||
</named-content>
|
||
). České Budějovice, Hosín aerodrome. 2 unsexed (
|
||
<named-content id="A9BE6915136E7873D969DB169D95EEF2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||
<collectionCode id="41F6F88C5D01D30B29168C3254E57484" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||
</named-content>
|
||
). Břeclav Pohansko
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6BBM" specific-use="{"type":"Point","coordinates":[18.918000,48.730000]}">
|
||
<geoCoordinate id="953080C6179947E926EC485A9155BED5" degrees="48.73" direction="north" orientation="latitude" precision="555" value="48.73">48.73 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1F924F7102EB31A4DEA7596FB2377ED9" degrees="18.918" direction="east" orientation="longitude" precision="55" value="18.918">18.918 ° E</geoCoordinate>
|
||
</named-content>
|
||
. Martinů Ivo leg.
|
||
<date id="1348DA92A9502A585B4DDD44BF1BBB9F" value="2008-04-20">20. iv. 2008</date>
|
||
.
|
||
<specimenCount id="D3E59F74A1D34634B1D5E1AEE50A7F85" count="1" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="5B5470D4F989DBA3822064B775B98412" count="4" type="female">4 ♀♀</specimenCount>
|
||
, 4 unsexed (
|
||
<named-content id="98C24A53FF9EFA2805E755978A241C54" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||
<collectionCode id="427DDC42675E9F04982BC22E83E85CC5" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||
</named-content>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="809CDD28C0846B91A5F59D960F6E2B55">
|
||
SK: Bol.
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENCBM" specific-use="{"type":"Point","coordinates":[21.949000,48.494000]}">
|
||
<geoCoordinate id="052020C0481EB7742FA029DFA4F71FBF" degrees="48.494" direction="north" orientation="latitude" precision="55" value="48.494">48.494 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="518109A0C6BE7EDE69754FA2E2D03175" degrees="21.949" direction="east" orientation="longitude" precision="55" value="21.949">21.949 ° E</geoCoordinate>
|
||
</named-content>
|
||
. 2 unsexed (
|
||
<named-content id="E5612EB86FCE24762D4EB108AC96A246" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
|
||
<collectionCode id="A02FC5908CBEA0F1C646A6E4EE5304C2" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
|
||
</named-content>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="3ABB660372E412D6E12EC8ABAE7F609B">
|
||
AU: Feldkirch Moosbrugger Vbg.
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2CBM" specific-use="{"type":"Point","coordinates":[9.581000,47.258000]}">
|
||
<geoCoordinate id="C0267B5C8C4C4007D621E6026BB50F2A" degrees="47.258" direction="north" orientation="latitude" precision="55" value="47.258">47.258 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E2AD16006A25D6761D918E0A8178EDC0" degrees="9.581" direction="east" orientation="longitude" precision="55" value="9.581">9.581 ° E</geoCoordinate>
|
||
</named-content>
|
||
.
|
||
<specimenCount id="F778AB5C62759BBB5F67D1C5B4E46605" count="1" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="CAAB46EBBF0FD498969F17A9FCF2D0BE" count="1" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="2EC116EECB86FB506342C21A650FDE8B">JHMC</collectionCode>
|
||
). Purgstall a. E.
|
||
<collectingCountry id="BBEE5F3F5E5849C4CEE6DEE8D0C1D40F" name="Austria">Austria</collectingCountry>
|
||
inf.
|
||
<named-content id="95FDEDF18BDCDE730F645F63A5794DF3" content-type="dwc:verbatimCoordinates">
|
||
<geoCoordinate id="9CC4CB51CA45D5EE5794E733ABA24B3E" degrees="48.063" direction="north" orientation="latitude" precision="55" value="48.063">48.063 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2E8126F178D5653987603A30BA26AEF9" degrees="15.132" direction="east" orientation="longitude" precision="55" value="15.132">15.132 ° E</geoCoordinate>
|
||
</named-content>
|
||
&
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELDBM" specific-use="{"type":"Point","coordinates":[15.133000,48.064000]}">
|
||
<geoCoordinate id="D53E17D39F7F866576A9DBE1C39177EF" degrees="48.064" direction="north" orientation="latitude" precision="55" value="48.064">48.064 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2C3ACBA051A39332402EA1DEE6BE1467" degrees="15.133" direction="east" orientation="longitude" precision="55" value="15.133">15.133 ° E</geoCoordinate>
|
||
</named-content>
|
||
. 2, unsexed. Leg. Ressi (
|
||
<abbrev id="ABBRID0EQDBM" xlink_title="Jason Maté Collection">
|
||
<collectionCode id="65004831A6B14EF64A31A8589D2229B6">JFMC</collectionCode>
|
||
</abbrev>
|
||
). Umgebung
|
||
<collectingRegion id="D518870C56F10F091ACC7DADFD858649" country="Austria" name="Wien">Wien</collectingRegion>
|
||
Mauerbach,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXDBM" specific-use="{"type":"Point","coordinates":[16.165000,48.246000]}">
|
||
<geoCoordinate id="B9271DE199432BD61CF87EAFB0F5500C" degrees="48.246" direction="north" orientation="latitude" precision="55" value="48.246">48.246 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CC0AB31A779AFA57FE00BC2190743BD1" degrees="16.165" direction="east" orientation="longitude" precision="55" value="16.165">16.165 ° E</geoCoordinate>
|
||
</named-content>
|
||
&
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6DBM" specific-use="{"type":"Point","coordinates":[16.354000,48.264000]}">
|
||
<geoCoordinate id="294F924D67B349A7AC455DBE7321DEDC" degrees="48.264" direction="north" orientation="latitude" precision="55" value="48.264">48.264 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="48B58BC613C6C3EBE021AC5ECDBD5AC6" degrees="16.354" direction="east" orientation="longitude" precision="55" value="16.354">16.354 ° E</geoCoordinate>
|
||
</named-content>
|
||
. 2, unsexed (
|
||
<abbrev id="ABBRID0EEEBM" xlink_title="Jason Maté Collection">
|
||
<collectionCode id="AA77CFF8DD1879811CE7CD0C97CA6B7E">JFMC</collectionCode>
|
||
</abbrev>
|
||
).
|
||
<collectingRegion id="63D412D643070224A18AE256333E8027" country="Austria" name="Wien">Wien</collectingRegion>
|
||
Umgebung, 4
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELEBM" specific-use="{"type":"Point","coordinates":[16.392000,8.209000]}">
|
||
<geoCoordinate id="1FFE4E2255D82FFCD52F0FA36CA2D6A8" degrees="8.209" direction="north" orientation="latitude" precision="55" value="8.209">8.209 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E6167DF7CF512DDA5AD4152B279FF96E" degrees="16.392" direction="east" orientation="longitude" precision="55" value="16.392">16.392 ° E</geoCoordinate>
|
||
</named-content>
|
||
. 1, unsexed (
|
||
<abbrev id="ABBRID0EQEBM" xlink_title="Jason Maté Collection">
|
||
<collectionCode id="077174205B0E907C63D2E32C1DC4F026">JFMC</collectionCode>
|
||
</abbrev>
|
||
). Neusiedler See.
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXEBM" specific-use="{"type":"Point","coordinates":[16.280067,47.712000]}">
|
||
<geoCoordinate id="CB080456550CDDC7465D207111FB2AE9" degrees="47.712" direction="north" orientation="latitude" precision="55" value="47.712">47.712 ° N</geoCoordinate>
|
||
, 16.16. 804 ° E
|
||
</named-content>
|
||
1, unsexed (
|
||
<abbrev id="ABBRID0E2EBM" xlink_title="Jason Maté Collection">
|
||
<collectionCode id="ADE04C68F80AB30B49C80C7CAFB6DF3A">JFMC</collectionCode>
|
||
</abbrev>
|
||
). Ullig Hart bei Graz,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECFBM" specific-use="{"type":"Point","coordinates":[15.440000,47.082000]}">
|
||
<geoCoordinate id="F3DA1B175D646FBD67B37CF431AE2DC2" degrees="47.082" direction="north" orientation="latitude" precision="55" value="47.082">47.082 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="8D7E8423B4616ED60B2E91F9EAB9AF6D" degrees="15.440" direction="east" orientation="longitude" precision="55" value="15.44">15.440 ° E</geoCoordinate>
|
||
</named-content>
|
||
. 1, unsexed (
|
||
<abbrev id="ABBRID0EHFBM" xlink_title="Jason Maté Collection">
|
||
<collectionCode id="01FCE286C09BA4A32D79D5CA6B03B8E5">JFMC</collectionCode>
|
||
</abbrev>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BF675CBC3571FEC67D17DF130F4EA594">
|
||
GE: Masselund, Emsland,
|
||
<date id="92C44F81C50159B022BA228146599ED6" value="1969-05-08">8. 5. 1969</date>
|
||
, Kerstens 1 unsexed; Hasslünne, Emsland, 8.5. 59, Kerstens, 1 unsexed (
|
||
<abbrev id="ABBRID0ENFBM" xlink_title="Axel Bellmann Collection">
|
||
<collectionCode id="C9AE8701FB3BAF1F121AAC220E5E4EE0">ABC</collectionCode>
|
||
</abbrev>
|
||
); Garz Dr Minarz, “ 594
|
||
<taxonomicName id="2B176C951E708468A5B891609B03F834" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="87E2E510CAFEF52F2ADF96EA8137DB19" italics="true">niger</emphasis>
|
||
</taxonomicName>
|
||
”, 1 unsexed; c. Epplsh Steind. d.,
|
||
<specimenCount id="9C6B2CC9C8C78AF509F5A7EA44A8DC53" count="1" type="male">1 ♂</specimenCount>
|
||
, 6 unsexed (
|
||
<named-content id="CF47A523867E1D2892F0CCCDC83887B4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum, Wien">
|
||
<collectionCode id="346BD412F36143900BFB305B812C11E1" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
|
||
</named-content>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FD78FC5C676470A30AEE9C8B11B03F03">
|
||
PL: Hinter
|
||
<collectingRegion id="F9E500EC0AC57DC345623B245BC4849E" country="Poland" name="Zachodniopomorskie">Pommern</collectingRegion>
|
||
Kołobrzeg (Zieleniewo), 1 unsexed (
|
||
<named-content id="BAFBD43D7D42827EDDB9B3E664F0903B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-ciencias-naturales" xlink_title="Museo Nacional de Ciencias Naturales">
|
||
<collectionCode id="FCFF348C8D907F301E75C7BF1805B935" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" type="Museum">MNCN</collectionCode>
|
||
</named-content>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="21111F4950962E326CC049A0CF62E6DD">
|
||
RU:
|
||
<collectingRegion id="FC33976BC3B4DAAE70E97E10E9AF57AF" country="Russia" name="Yaroslavl'">Yaroslavl</collectingRegion>
|
||
oblast’, Berditsino,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJGBM" specific-use="{"type":"Point","coordinates":[40.110800,57.454000]}">
|
||
<geoCoordinate id="CA6A629865AB0A020F4FC4EA64609FF8" degrees="57.454" direction="north" orientation="latitude" precision="55" value="57.454">57.454 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F064E5B955A8F67FAA9A629F35A09919" degrees="40.1108" direction="east" orientation="longitude" precision="5" value="40.1108">40.1108 ° E</geoCoordinate>
|
||
</named-content>
|
||
. leg. Yakovlev.
|
||
<specimenCount id="5C1371769A0632AA38841AC04CCEC223" count="1" type="male">1 ♂</specimenCount>
|
||
(
|
||
<named-content id="1D36C7DBFD86581EC6EFA102C41F5759" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum, Wien">
|
||
<collectionCode id="86E962E0381560AC98EA203F2337D2AE" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
|
||
</named-content>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F0BBAA4695A26C8438F21F07E70115CD">
|
||
<emphasis id="99A3E5F78E4132CA33598127A231D1FB" bold="true">Remarks.</emphasis>
|
||
Material from
|
||
<collectingCountry id="4FE27A41309734414064605CFD05B362" name="France">France</collectingCountry>
|
||
,
|
||
<collectingCountry id="629F7231993F6DCC047B2918D29403A8" name="Italy">Italy</collectingCountry>
|
||
,
|
||
<collectingCountry id="F0767EB79C7368696575E18B3B3331B4" name="Bosnia and Herzegovina">Bosnia</collectingCountry>
|
||
, and
|
||
<collectingCountry id="2A408FF8AC3E15FC0DB69F6C73038961" name="Turkey">Turkey</collectingCountry>
|
||
is listed by
|
||
<bibRefCitation id="E7C02749E80F7F32A5853BA6A26ED497" author="Maté JF & Angus RB" firstAuthor="Mate" journalOrPublisher="Koleopterologische Rundschau" pagination="325-336" refId="B33" refString="Maté JF, Angus RB (2005) Description of a new species of Aphodius Illiger from the Iberian Peninsula and comments regarding the biogeography and ecology of the subgenus Liothorax Motschulsky. Koleopterologische Rundschau 75: 325–336." title="Description of a new species of Aphodius Illiger from the Iberian Peninsula and comments regarding the biogeography and ecology of the subgenus Liothorax Motschulsky." volume="75" year="2005">Maté and Angus (2005)</bibRefCitation>
|
||
. It is not listed here but is mapped (Fig.
|
||
<figureCitation id="291F5BF6B72DA21A149F10912E2D3C62" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." figureDoi="10.3897/zookeys.1207.117225.figure29" httpUri="https://binary.pensoft.net/fig/1097509">29</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="31E9819765D73E1A705D6A2C6719172C">
|
||
Material from coastal or near-costal localities from
|
||
<collectingCountry id="D56D83162824E7657B3CE13A7CAA8D35" name="Poland">Poland</collectingCountry>
|
||
,
|
||
<collectingCountry id="AC1F4C8752352113A65D2860EA346760" name="Germany">Germany</collectingCountry>
|
||
, and
|
||
<collectingCountry id="F096ACAE01CF703365DA4943D472C857" name="France">France</collectingCountry>
|
||
is likely to be
|
||
<taxonomicName id="6E84726F4A1308F2017716D6C8751AF3" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="F60BA31469556AACC8B93567F9C512F7" italics="true">A. niger</emphasis>
|
||
</taxonomicName>
|
||
. The Yaroslavl specimen is taken as
|
||
<taxonomicName id="CAAEE46585964D2F71933DDF2583F583" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="9C7EE1BB7039289DB99ED4E5BF278639" italics="true">A. niger</emphasis>
|
||
</taxonomicName>
|
||
because locality seems to be ecologically similar to the English and Swedish ones. This might also be true of the Tyumen male.
|
||
</paragraph>
|
||
<paragraph id="2C1B62A773115544FC1978237C3AD961">
|
||
The specimens from Břeclav Pohansko and Bol are likely to be
|
||
<taxonomicName id="7E1959314AD516F754CFF3E57FCAFF4D" authorityName="Angus & Maté & Angus & Král" authorityYear="2024" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="muscorum">
|
||
<emphasis id="7A8A16050BA86414B68CF3F45FC8F530" italics="true">A. muscorum</emphasis>
|
||
</taxonomicName>
|
||
because these lowland localities are on the edge of the Hungarian plain, and the same is true of all the eastern Austrian material.
|
||
</paragraph>
|
||
<paragraph id="2545627210D0FFD13343D47FFF4D044D">
|
||
The outstanding problem with Central European material is the identity of the “
|
||
<taxonomicName id="D891C2E835C16307C008544A22DF330C" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="BFDB9C6205F0015902DB28DDCD8F783A" italics="true">A. niger</emphasis>
|
||
</taxonomicName>
|
||
” from the German Mittelgeberge and the northern and central parts of
|
||
<collectingCountry id="AB8231D8AFC658033608159049AD530E" name="Czech Republic">Czechia</collectingCountry>
|
||
, as well as the French Massif Central. The map given by
|
||
<bibRefCitation id="AFE145737E94F03F7A7F4B6037385F64" author="Rößner E" firstAuthor="Rossner" refId="B45" refString="Rößner E (2012) Die Hirschkäfer und Blatthornkäfer Ostdeutschlands (Coleoptera: Scarabaeoidea). Herausgegeben und zu beziehen vom Verein der Freunde & Förderer des Naturkundemuseums Erfurt e. V. Große Arche 14. 99084 Erfurt." year="2012">Rößner (2012)</bibRefCitation>
|
||
shows German records, mainly pre- 1950, concentrated either in coastal regions or somewhat further inland (the Mittelgeberge) with few records connecting the two areas.
|
||
<bibRefCitation id="B17AC6F8C6046A5F9E57548CEF5F9944" author="Rößner E & Frenzel E & Hillert O & Zeigler W" etAl="et al." firstAuthor="Rossner" journalOrPublisher="Entomologische Nachrichten und Berichte" pagination="191-196" refId="B46" refString="Rößner E, Frenzel E, Hillert O, Zeigler W (2016) Neue Nachweise von Liothorax niger (Illiger, 1798) (Coleoptera, Scarabaeidae, Aphodiinae) in Deutschland. Entomologische Nachrichten und Berichte 60: 191–196. https://www.academia.edu/49478986/%C3%9Fner_Frenzel_Hillert_Ziegler_2017?auto=download" title="Neue Nachweise von Liothorax niger (Illiger, 1798) (Coleoptera, Scarabaeidae, Aphodiinae) in Deutschland." url="https://www.academia.edu/49478986/%C3%9Fner_Frenzel_Hillert_Ziegler_2017?auto=download" volume="60" year="2016">Rößner et al. (2016)</bibRefCitation>
|
||
give updated records, with some west German localities in the Mittlegeberge post 1950.
|
||
<bibRefCitation id="8B207D34C2451D63D6619D5A5220897B" author="Costessèque R" firstAuthor="Costesseque" refId="B10" refString="Costessèque R (2005) Les Aphodius de France Une clé de détermination. Magellanes, 79 pp." year="2005">Costessèque (2005)</bibRefCitation>
|
||
records it from all the Départements of the Massif Central, but RBA failed to find it in the southern Auvergne (Cantal, Haute-Loire) in
|
||
<date id="D05B494ACC442B2671898E18BBF58DE6" value="2017-04">April 2017</date>
|
||
. While all the material may be
|
||
<taxonomicName id="DF62BF994C30EF17E70ADD35C5B2D348" baseAuthorityName="Panzer" baseAuthorityYear="1797" class="Insecta" family="Scarabaeidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="D0E59B41BF435B828CB6834AB000F6B8" italics="true">A. niger</emphasis>
|
||
</taxonomicName>
|
||
, there remains the possibility that a further, as yet unknown species may be present. We have, unfortunately, not been able to obtain living material from this inland area.
|
||
</paragraph>
|
||
<paragraph id="5E8747460AA8E38273DD86EAC08C5C10">
|
||
The material from Astana and the Akmola oblast and Uralsk may include a new species characterised by a reticulate metaventrite in the males, but a male from a saline depression (Solonchak) by the river Basaga-ozik lacks any trace of reticulation. The
|
||
<collectingRegion id="2348D5AF9BB682C8E916682EF9FF223F" country="Russia" name="Volgograd">Volgograd</collectingRegion>
|
||
male also lacks any trace of reticulation on the central portion of the metaventrite. The males from the Chernomorsk oblast’ and
|
||
<collectingRegion id="6E97E09BF894ABB89897245193F16326" country="Russia" name="Volgograd">Volgograd</collectingRegion>
|
||
also lack any reticulation on the metaventrites and their identities are unclear.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |