366 lines
45 KiB
XML
366 lines
45 KiB
XML
<document id="C8ED40656EB1632E1850DF1FDF996FC4" ID-DOI="10.5281/zenodo.185553" ID-GBIF-Dataset="3ec6241f-1e53-4eda-abef-b9f59fa98c18" ID-ISSN="1175-5326" ID-Zenodo-Dep="185553" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459994708828" checkinUser="plazi" docAuthor="Kok, Philippe J. R." docDate="2009" docId="235D9D0CFFF5FFC8FF73FA3DFCBEFE36" docLanguage="en" docName="zt01992p067.pdf" docOrigin="Zootaxa 1992" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pantepuisaurus" docType="treatment" docVersion="9" lastPageNumber="56" masterDocId="DF64E574FFF4FFCCFFE4FFE4FFCAFFAC" masterDocTitle="Lizard in the clouds: a new highland genus and species of Gymnophthalmidae (Reptilia: Squamata) from Maringma tepui, western Guyana" masterLastPageNumber="67" masterPageNumber="53" pageNumber="54" updateTime="1698232753826" updateUser="plazi">
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<mods:titleInfo id="B277B9DE246CA435E107A97B0D6B2794">
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<mods:title id="813A6FD63E8CF7ACD6E931E912CF8BF8">Lizard in the clouds: a new highland genus and species of Gymnophthalmidae (Reptilia: Squamata) from Maringma tepui, western Guyana</mods:title>
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<mods:name id="0E44CA041FBEC43C36AD9516635705B7" type="personal">
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<mods:namePart id="DA834EEE3C07E6C9A6F85AE74FA6F725">Kok, Philippe J. R.</mods:namePart>
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<mods:title id="75BA56F1823AD56B9FBEC5A6BBEE3929">Zootaxa</mods:title>
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<mods:part id="73B042FEE58E9284E6DB298F9685DE4F">
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<mods:date id="532832D2807481744AAE437504323E5C">2009</mods:date>
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<mods:number id="F3E783A77D459DFABD3F65785E0DE2E5">1992</mods:number>
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<mods:classification id="15C023ECE47690451F79BD2D4FC4AA39">journal article</mods:classification>
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<mods:identifier id="66AD8725AB8704928513F8ADFFF19323" type="DOI">10.5281/zenodo.185553</mods:identifier>
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<mods:identifier id="F491645594D245DAD3B11B29FE6FA5BA" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B0425B4911704A5F6EA0A7C11E00AF0B" type="Zenodo-Dep">185553</mods:identifier>
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<treatment id="235D9D0CFFF5FFC8FF73FA3DFCBEFE36" ID-DOI="http://doi.org/10.5281/zenodo.5673885" ID-GBIF-Taxon="119380223" ID-Zenodo-Dep="5673885" LSID="urn:lsid:plazi:treatment:235D9D0CFFF5FFC8FF73FA3DFCBEFE36" httpUri="http://treatment.plazi.org/id/235D9D0CFFF5FFC8FF73FA3DFCBEFE36" lastPageId="4" lastPageNumber="56" pageId="1" pageNumber="54">
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<subSubSection id="E3EE7F91FFF5FFCDFF73FA3DFDCBFA5E" box="[151,513,1496,1523]" pageId="1" pageNumber="54" type="nomenclature">
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<paragraph id="AB4B2C1AFFF5FFCDFF73FA3DFDCBFA5E" blockId="1.[151,513,1496,1523]" box="[151,513,1496,1523]" pageId="1" pageNumber="54">
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<heading id="F0039B76FFF5FFCDFF73FA3DFDCBFA5E" bold="true" box="[151,513,1496,1523]" fontSize="11" level="1" pageId="1" pageNumber="54" reason="1">
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<emphasis id="9980F008FFF5FFCDFF73FA3DFDCBFA5E" bold="true" box="[151,513,1496,1523]" pageId="1" pageNumber="54">
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<taxonomicName id="6CF45799FFF5FFCDFF73FA3DFE9CFA5F" box="[151,342,1497,1523]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="54" phylum="Chordata" rank="genus">
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<emphasis id="9980F008FFF5FFCDFF73FA3DFE9CFA5F" bold="true" box="[151,342,1497,1523]" italics="true" pageId="1" pageNumber="54">Pantepuisaurus</emphasis>
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</taxonomicName>
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genus novum
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E3EE7F91FFF5FFCDFF73F9C6FCFEF990" box="[151,820,1570,1596]" pageId="1" pageNumber="54" type="materials_examined">
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<paragraph id="AB4B2C1AFFF5FFCDFF73F9C6FCFEF990" blockId="1.[151,1436,1570,2022]" box="[151,820,1570,1596]" pageId="1" pageNumber="54">
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||
<emphasis id="9980F008FFF5FFCDFF73F9C6FEF3F990" bold="true" box="[151,313,1570,1596]" pageId="1" pageNumber="54">
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<typeStatus id="744F92B8FFF5FFCDFF73F9C6FF1EF990" box="[151,212,1570,1596]" pageId="1" pageNumber="54">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="6CF45799FFF5FFCDFEA4F9C7FDB3F990" box="[320,633,1571,1596]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="rodriguesi">
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<emphasis id="9980F008FFF5FFCDFEA4F9C7FDB3F990" box="[320,633,1571,1596]" italics="true" pageId="1" pageNumber="54">Pantepuisaurus rodriguesi</emphasis>
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</taxonomicName>
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<emphasis id="9980F008FFF5FFCDFD64F9C6FCFEF990" bold="true" box="[640,820,1570,1596]" pageId="1" pageNumber="54">species novum</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="E3EE7F91FFF5FFCDFF21F9ACFC0BF91C" pageId="1" pageNumber="54" type="etymology">
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<paragraph id="AB4B2C1AFFF5FFCDFF21F9ACFAD7F926" blockId="1.[151,1436,1570,2022]" pageId="1" pageNumber="54">
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<emphasis id="9980F008FFF5FFCDFF21F9ACFE9FF9CE" bold="true" box="[197,341,1608,1634]" pageId="1" pageNumber="54">Etymology.</emphasis>
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A noun in apposition, derived from “Pantepui” referring to the phytogeographic province where the
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<typeStatus id="744F92B8FFF5FFCDFEF5F994FE89F926" box="[273,323,1648,1674]" pageId="1" pageNumber="54">type</typeStatus>
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species was discovered, and the Greek
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<emphasis id="9980F008FFF5FFCDFCF0F994FCABF925" box="[788,865,1648,1673]" italics="true" pageId="1" pageNumber="54">sauros</emphasis>
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meaning “lizard”. Gender masculine.
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</paragraph>
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<paragraph id="AB4B2C1AFFF5FFCDFF21F972FC0BF91C" blockId="1.[151,1436,1570,2022]" box="[197,961,1686,1712]" pageId="1" pageNumber="54">
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<emphasis id="9980F008FFF5FFCDFF21F972FEFBF91C" bold="true" box="[197,305,1686,1712]" pageId="1" pageNumber="54">Content.</emphasis>
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<taxonomicName id="6CF45799FFF5FFCDFEDCF973FDBBF91C" box="[312,625,1687,1712]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="rodriguesi">
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<emphasis id="9980F008FFF5FFCDFEDCF973FDBBF91C" box="[312,625,1687,1712]" italics="true" pageId="1" pageNumber="54">Pantepuisaurus rodriguesi</emphasis>
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</taxonomicName>
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<emphasis id="9980F008FFF5FFCDFD9CF972FCE1F91C" bold="true" box="[632,811,1686,1712]" pageId="1" pageNumber="54">species novum</emphasis>
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; monotypic.
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</paragraph>
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</subSubSection>
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<subSubSection id="E3EE7F91FFF5FFCEFF21F959FCDAFF74" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54" type="diagnosis">
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<paragraph id="AB4B2C1AFFF5FFCEFF21F959FCDAFF74" blockId="1.[151,1436,1570,2022]" lastBlockId="2.[151,1438,152,2034]" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54">
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<emphasis id="9980F008FFF5FFCDFF21F959FE8EF97B" bold="true" box="[197,324,1725,1751]" pageId="1" pageNumber="54">Diagnosis.</emphasis>
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An elongate gymnophthalmid (known SVL in male
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<quantity id="6C0C81FFFFF5FFCDFC55F959FBD6F97B" box="[945,1052,1725,1751]" metricMagnitude="-2" metricUnit="m" metricValue="5.83" pageId="1" pageNumber="54" unit="mm" value="58.3">58.3 mm</quantity>
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) with a long tail (1.6x SVL) and the unique following combination of characters: (1) distinctive ear opening and moveable eyelids, (2) limbs pentadactyl with all digits clawed, (3) nasal scales separated by a single frontonasal, (4) prefrontals present, (5) frontoparietals, parietals and interparietal present, (6) parietals and interparietal longer than wide, (7) interparietal and parietals forming a jagged, irregular posterior margin, (8) occipitals present, (9) three pairs of genials, second pair in contact with only one infralabial, (10) enlarged median pairs of gulars, (11) dorsal scales hexagonal, keeled, in transverse rows only, (12) ventral scales imbricate, hexagonal, mucronate, broadly keeled, in transverse rows only, (13) tongue anterodorsally covered by oblique, anteriorly converging plicae, posterodorsally covered by large scalelike papillae (14) hemipenis weakly bilobed with series of curved transverse plicae bearing mineralized spicules.
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</paragraph>
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</subSubSection>
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<subSubSection id="E3EE7F91FFF6FFCEFF21FF01FD18F911" pageId="2" pageNumber="55" type="reference_group">
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<paragraph id="AB4B2C1AFFF6FFCEFF21FF01FD18F911" blockId="2.[151,1438,152,2034]" pageId="2" pageNumber="55">
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<taxonomicName id="6CF45799FFF6FFCEFF21FF01FEB7FF52" box="[197,381,229,254]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
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<emphasis id="9980F008FFF6FFCEFF21FF01FEB7FF52" box="[197,381,229,254]" italics="true" pageId="2" pageNumber="55">Pantepuisaurus</emphasis>
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</taxonomicName>
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differs from members of Alopoglossinae in having the tongue anterodorsally covered by oblique plicae and posterodorsally covered by large scalelike papillae (entirely covered by oblique plicae in Alopoglossinae), the interparietal and the parietals forming a jagged, irregular line (straight posterior margin), and in having ventral scales in transverse rows only (in transverse and longitudinal rows in Alopoglossinae); from members of Gymnophthalminae in having hands pentadactyl with the first finger clawed (first finger absent, reduced and/or clawless in Gymnophthalminae); from Rhachisaurinae in having external ear openings (absent in Rhachisaurinae). Within Cercosaurinae (
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<emphasis id="9980F008FFF6FFCEFD19FE29FCF5FE4A" box="[765,831,461,486]" italics="true" pageId="2" pageNumber="55">sensu</emphasis>
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<bibRefCitation id="CF6551EBFFF6FFCEFCACFE29FB86FE4B" author="Pellegrino" box="[840,1100,461,487]" pageId="2" pageNumber="67" refString="Pellegrino, K. C. M., Rodrigues, M. T., Yonenaga-Yassuda, Y. & Sites Jr., J. W. (2001) A molecular perspective on the evolution of microteiids lizards (Squamata, Gymnophthalmidae), and a new classification for the family. Biological Journal of the Linnean Society, 74, 315 - 338." type="journal article" year="2001">
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Pellegrino
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<emphasis id="9980F008FFF6FFCEFC2FFE29FBCDFE4A" box="[971,1031,461,486]" italics="true" pageId="2" pageNumber="55">et al.</emphasis>
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2001
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</bibRefCitation>
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and
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<bibRefCitation id="CF6551EBFFF6FFCEFB63FE29FA45FE4B" author="Rodrigues" box="[1159,1423,461,487]" pageId="2" pageNumber="67" refString="Rodrigues, M. T., Freire, E. M. X., Pellegrino, K. C. M & Sites Jr., J. W. (2005) Phylogenetic relationships of a new genus and species of microteiid lizard from the Atlantic forest of north-eastern Brazil (Squamata, Gymnophthalmidae). Zoological Journal of the Linnean Society, 144, 543 - 557." type="journal article" year="2005">
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Rodrigues
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<emphasis id="9980F008FFF6FFCEFAEEFE29FA8DFE4A" box="[1290,1351,461,486]" italics="true" pageId="2" pageNumber="55">et al.</emphasis>
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2005
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</bibRefCitation>
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),
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<taxonomicName id="6CF45799FFF6FFCEFF73FE10FE9EFDA1" box="[151,340,500,525]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
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<emphasis id="9980F008FFF6FFCEFF73FE10FE9EFDA1" box="[151,340,500,525]" italics="true" pageId="2" pageNumber="55">Pantepuisaurus</emphasis>
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</taxonomicName>
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differs from all genera except
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<taxonomicName id="6CF45799FFF6FFCEFD2FFE10FC82FDA1" box="[715,840,500,525]" class="Reptilia" family="Gymnophthalmidae" genus="Ecpleopus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
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<emphasis id="9980F008FFF6FFCEFD2FFE10FC82FDA1" box="[715,840,500,525]" italics="true" pageId="2" pageNumber="55">Ecpleopus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="6CF45799FFF6FFCEFC60FE10FC35FDA1" ID-CoL="8HHKH" box="[900,1023,500,525]" class="Insecta" family="Curculionidae" genus="Leposoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="9980F008FFF6FFCEFC60FE10FC35FDA1" box="[900,1023,500,525]" italics="true" pageId="2" pageNumber="55">Leposoma</emphasis>
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</taxonomicName>
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of the
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<emphasis id="9980F008FFF6FFCEFBB2FE10FB19FDA1" box="[1110,1235,500,525]" italics="true" pageId="2" pageNumber="55">scincoides</emphasis>
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group in having hexagonal, mucronate ventral scales arranged in transverse rows only.
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<taxonomicName id="6CF45799FFF6FFCEFC2AFDFEFB4CFD9F" box="[974,1158,538,563]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
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<emphasis id="9980F008FFF6FFCEFC2AFDFEFB4CFD9F" box="[974,1158,538,563]" italics="true" pageId="2" pageNumber="55">Pantepuisaurus</emphasis>
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</taxonomicName>
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is readily distinguished from
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<taxonomicName id="6CF45799FFF6FFCEFF3CFDA5FE98FDF6" box="[216,338,577,602]" class="Reptilia" family="Gymnophthalmidae" genus="Ecpleopus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
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<emphasis id="9980F008FFF6FFCEFF3CFDA5FE98FDF6" box="[216,338,577,602]" italics="true" pageId="2" pageNumber="55">Ecpleopus</emphasis>
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</taxonomicName>
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(characters of
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<taxonomicName id="6CF45799FFF6FFCEFDE0FDA5FDB4FDF6" box="[516,638,577,602]" class="Reptilia" family="Gymnophthalmidae" genus="Ecpleopus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
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||
<emphasis id="9980F008FFF6FFCEFDE0FDA5FDB4FDF6" box="[516,638,577,602]" italics="true" pageId="2" pageNumber="55">Ecpleopus</emphasis>
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</taxonomicName>
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in parentheses) notably by the presence of a broad keel on ventrals (absent), quadrangular gular scales (mucronate), complete superciliary series (incomplete), and the presence of femoral pores in males (absent).
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<taxonomicName id="6CF45799FFF6FFCEFDADFD6BFCCCFD04" box="[585,774,655,680]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
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<emphasis id="9980F008FFF6FFCEFDADFD6BFCCCFD04" box="[585,774,655,680]" italics="true" pageId="2" pageNumber="55">Pantepuisaurus</emphasis>
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</taxonomicName>
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||
is quickly separated from
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<taxonomicName id="6CF45799FFF6FFCEFBA8FD6BFB02FD04" ID-CoL="8HHKH" box="[1100,1224,655,680]" class="Insecta" family="Curculionidae" genus="Leposoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="9980F008FFF6FFCEFBA8FD6BFB02FD04" box="[1100,1224,655,680]" italics="true" pageId="2" pageNumber="55">Leposoma</emphasis>
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</taxonomicName>
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of the
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<emphasis id="9980F008FFF6FFCEFAFAFD6BFA56FD04" box="[1310,1436,655,680]" italics="true" pageId="2" pageNumber="55">scincoides</emphasis>
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group (characters of
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<taxonomicName id="6CF45799FFF6FFCEFE7EFD51FDD2FD62" ID-CoL="8HHKH" box="[410,536,693,718]" class="Insecta" family="Curculionidae" genus="Leposoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="genus">
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||
<emphasis id="9980F008FFF6FFCEFE7EFD51FDD2FD62" box="[410,536,693,718]" italics="true" pageId="2" pageNumber="55">Leposoma</emphasis>
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||
</taxonomicName>
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in parentheses) in having quadrangular gular scales (mucronate), by the absence of distinct striations on head scales (present), the interparietal and the parietals forming a jagged, irregular line (round posterior margin), and the presence of occipitals (absent). Two additional gymnophthalmid genera,
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<taxonomicName id="6CF45799FFF6FFCEFE2EFCCDFDA4FCEE" box="[458,622,809,834]" class="Reptilia" family="Gymnophthalmidae" genus="Adercosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
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<emphasis id="9980F008FFF6FFCEFE2EFCCDFDA4FCEE" box="[458,622,809,834]" italics="true" pageId="2" pageNumber="55">Adercosaurus</emphasis>
|
||
</taxonomicName>
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||
and
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||
<taxonomicName id="6CF45799FFF6FFCEFD4EFCCDFCBAFCEE" box="[682,880,809,834]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
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||
<emphasis id="9980F008FFF6FFCEFD4EFCCDFCBAFCEE" box="[682,880,809,834]" italics="true" pageId="2" pageNumber="55">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
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, have not been assigned to any subfamily yet, but might be related each other and with
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<taxonomicName id="6CF45799FFF6FFCEFD9AFCB4FCFDFCC5" box="[638,823,848,873]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
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||
<emphasis id="9980F008FFF6FFCEFD9AFCB4FCFDFCC5" box="[638,823,848,873]" italics="true" pageId="2" pageNumber="55">Pantepuisaurus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="6CF45799FFF6FFCEFCA2FCB4FC35FCC5" box="[838,1023,848,873]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFCA2FCB4FC35FCC5" box="[838,1023,848,873]" italics="true" pageId="2" pageNumber="55">Pantepuisaurus</emphasis>
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</taxonomicName>
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is immediately distinguished from
|
||
<taxonomicName id="6CF45799FFF6FFCEFF73FC93FE89FC3C" box="[151,323,887,912]" class="Reptilia" family="Gymnophthalmidae" genus="Adercosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFF73FC93FE89FC3C" box="[151,323,887,912]" italics="true" pageId="2" pageNumber="55">Adercosaurus</emphasis>
|
||
</taxonomicName>
|
||
(characters of
|
||
<taxonomicName id="6CF45799FFF6FFCEFDE4FC93FD66FC3C" box="[512,684,887,912]" class="Reptilia" family="Gymnophthalmidae" genus="Adercosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFDE4FC93FD66FC3C" box="[512,684,887,912]" italics="true" pageId="2" pageNumber="55">Adercosaurus</emphasis>
|
||
</taxonomicName>
|
||
in parentheses) in having imbricate, hexagonal, mucronate, broadly keeled ventral scales in transverse rows only (smooth, in transverse and longitudinal rows), in having a highly reduced fourth infralabial (not reduced) and the second pair of genials partly separated from the infralabials and in contact with one infralabial only (not separated from the infralabials, in contact with two infralabials), in lacking oblique plicae at the rear of the tongue, which is posteriorly covered by large scalelike papillae (rear of tongue covered by oblique plicae), and in having mineralized spicules on hemipenes’ plicae (absent).
|
||
<taxonomicName id="6CF45799FFF6FFCEFEE3FBBBFE08FBD4" box="[263,450,1119,1144]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFEE3FBBBFE08FBD4" box="[263,450,1119,1144]" italics="true" pageId="2" pageNumber="55">Pantepuisaurus</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="6CF45799FFF6FFCEFD47FBBBFCA2FBD4" box="[675,872,1119,1144]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFD47FBBBFCA2FBD4" box="[675,872,1119,1144]" italics="true" pageId="2" pageNumber="55">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
in having hexagonal, mucronate ventral scales arranged in transverse rows only, but it is readily distinguished from it (characters of
|
||
<taxonomicName id="6CF45799FFF6FFCEFB57FB61FAB6FB32" box="[1203,1404,1157,1182]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFB57FB61FAB6FB32" box="[1203,1404,1157,1182]" italics="true" pageId="2" pageNumber="55">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
in parentheses) by the presence of prefrontal scales (absent), three pairs of genials (two), in having a highly reduced fourth infralabial (not reduced) and the second pair of genials partly separated from the infralabials and in contact with one infralabial only (not separated from the infralabials, in contact with two infralabials), the common suture of the interparietal and parietals forming a jagged, irregular line (more or less straight posterior margin), an entire nasal (divided), and the aspect of the ventral scales that are much less lanceolate and mucronate than in
|
||
<taxonomicName id="6CF45799FFF6FFCEFE48FA89FDB8FA2A" box="[428,626,1389,1414]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFE48FA89FDB8FA2A" box="[428,626,1389,1414]" italics="true" pageId="2" pageNumber="55">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
. It should be emphasized here that
|
||
<taxonomicName id="6CF45799FFF6FFCEFBC5FA89FB2DFA2A" box="[1057,1255,1389,1414]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFBC5FA89FB2DFA2A" box="[1057,1255,1389,1414]" italics="true" pageId="2" pageNumber="55">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
also has broad “keels” on ventral scales (
|
||
<emphasis id="9980F008FFF6FFCEFE3FFA70FDCBFA01" box="[475,513,1428,1453]" italics="true" pageId="2" pageNumber="55">i.e.</emphasis>
|
||
the central part of the scale is broadly elevated), a character that was only noticed while closely comparing the two genera. This character is not conspicuous in all
|
||
<taxonomicName id="6CF45799FFF6FFCEFB30FA5FFA56FA78" box="[1236,1436,1467,1492]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFB30FA5FFA56FA78" box="[1236,1436,1467,1492]" italics="true" pageId="2" pageNumber="55">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
specimens and is mostly noticeable on anteriormost and posteriormost ventral scales. In
|
||
<taxonomicName id="6CF45799FFF6FFCEFB49FA05FABAFA56" box="[1197,1392,1505,1530]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFB49FA05FABAFA56" box="[1197,1392,1505,1530]" italics="true" pageId="2" pageNumber="55">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
the significant lateral extension of the flat “keel” gives a lanceolate, smooth, “unkeeled” aspect to the ventral scales as described by
|
||
<bibRefCitation id="CF6551EBFFF6FFCEFE48F9CAFDF6F9E4" author="Kok" box="[428,572,1582,1608]" pageId="2" pageNumber="66" refString="Kok, P. J. R. (2005) A new genus and species of gymnophthalmid lizard (Squamata: Gymnophthalmidae) from Kaieteur National Park, Guyana. Bulletin de L'Institut Royal des Sciences Naturelles de Belgique, Biologie, 75, 35 - 45." type="journal article" year="2005">Kok (2005)</bibRefCitation>
|
||
, while the keel is more prominent and conspicuous in
|
||
<taxonomicName id="6CF45799FFF6FFCEFB33F9CBFA5EF9E4" box="[1239,1428,1583,1608]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFB33F9CBFA5EF9E4" box="[1239,1428,1583,1608]" italics="true" pageId="2" pageNumber="55">Pantepuisaurus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<figureCitation id="33CF309FFFF6FFCEFF73F9B1FF34F9C3" box="[151,254,1621,1647]" captionStart="FIGURE 4" captionStartId="7.[151,255,876,900]" captionTargetBox="[181,1397,273,852]" captionTargetId="figure@7.[151,1436,237,880]" captionTargetPageId="7" captionText="FIGURE 4. Schematic illustration of the ventral scale condition in Pantepuisaurus gen. nov. and Kaieteurosaurus. A: Four posterior ventral scales of Pantepuisaurus rodriguesi gen. nov. sp. nov. showing the broad keel (in white). B: Four posterior ventral scales of Kaieteurosaurus hindsi showing the broad keel (in white) that significantly extends laterally giving an aspect of smooth unkeeled lanceolate ventrals. Drawings are not at scale." httpUri="https://zenodo.org/record/185557/files/figure.png" pageId="2" pageNumber="55">Figure 4</figureCitation>
|
||
schematically illustrates the difference in the aspect of the broad keel on ventral scales between
|
||
<taxonomicName id="6CF45799FFF6FFCEFF73F998FE94F939" box="[151,350,1660,1685]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFF73F998FE94F939" box="[151,350,1660,1685]" italics="true" pageId="2" pageNumber="55">Pantepuisaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6CF45799FFF6FFCEFE79F998FDA5F939" box="[413,623,1660,1685]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFE79F998FDA5F939" box="[413,623,1660,1685]" italics="true" pageId="2" pageNumber="55">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
. A summary of main diagnostic characters for
|
||
<taxonomicName id="6CF45799FFF6FFCEFB2AF998FA5FF939" box="[1230,1429,1660,1685]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFB2AF998FA5FF939" box="[1230,1429,1660,1685]" italics="true" pageId="2" pageNumber="55">Pantepuisaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6CF45799FFF6FFCEFF73F947FEF3F910" box="[151,313,1699,1724]" class="Reptilia" family="Gymnophthalmidae" genus="Adercosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFF73F947FEF3F910" box="[151,313,1699,1724]" italics="true" pageId="2" pageNumber="55">Adercosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6CF45799FFF6FFCEFE95F947FDFEF910" box="[369,564,1699,1724]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF6FFCEFE95F947FDFEF910" box="[369,564,1699,1724]" italics="true" pageId="2" pageNumber="55">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
is in
|
||
<tableCitation id="E67619A1FFF6FFCEFD90F947FD06F911" box="[628,716,1699,1725]" captionStart="TABLE 1" captionStartId="4.[151,243,457,481]" captionTargetBox="[159,1426,617,1611]" captionTargetPageId="4" captionText="TABLE 1. Summary of main diagnostic characters for Pantepuisaurus gen. nov. and two possibly related genera (Adercosaurus and Kaieteurosaurus). The diagnostic characters are those currently used to separate genera in the tribe Ecpleopini [sensu Pellegrino et al. (2001); Ecpleopinae sensu Castoe et al. (2004)]. Based on Myers and Donnelly (2001), Kok (2005) and the present paper." httpUri="http://table.plazi.org/id/FF8B7C92FFF0FFC8FF73FE2DFD9DFDE4" pageId="2" pageNumber="55" tableUuid="FF8B7C92FFF0FFC8FF73FE2DFD9DFDE4">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E3EE7F91FFF6FFC8FF21F92DFCBEFE36" lastPageId="4" lastPageNumber="57" pageId="2" pageNumber="55" type="distribution">
|
||
<paragraph id="AB4B2C1AFFF6FFCEFF21F92DFE92F8A6" blockId="2.[151,1438,152,2034]" pageId="2" pageNumber="55">
|
||
<emphasis id="9980F008FFF6FFCEFF21F92DFEAFF94F" bold="true" box="[197,357,1737,1763]" pageId="2" pageNumber="55">Distribution.</emphasis>
|
||
The genus is currently known only from the
|
||
<typeStatus id="744F92B8FFF6FFCEFC9CF92DFC60F94F" box="[888,938,1737,1763]" pageId="2" pageNumber="55">type</typeStatus>
|
||
locality, the summit of Maringma tepui in
|
||
<collectingCountry id="D3E36C8AFFF6FFCEFF73F914FF38F8A6" box="[151,242,1776,1802]" name="Guyana" pageId="2" pageNumber="55">Guyana</collectingCountry>
|
||
(
|
||
<figureCitation id="33CF309FFFF6FFCEFEE6F914FE8DF8A6" box="[258,327,1776,1802]" captionStart="FIGURE 1" captionStartId="3.[151,256,1233,1257]" captionTargetBox="[151,1435,197,1208]" captionTargetId="figure@3.[151,1436,197,1208]" captionTargetPageId="3" captionText="FIGURE 1. Area map of Mount Maringma showing the type locality of Pantepuisaurus rodriguesi gen. nov. sp. nov. (red dot). Coordinates and boundary with Brazil are approximate and based on the available map (note that there is a substantial difference in the country boundary between this map and that shown in Google Earth). Maps elaborated after “ Roraima Sheet 40 SE ” published by the Survey Department of Guyana, 1972, and a radar image of South America by NASA / JPL / NIMA available at http: // photojournal. jpl. nasa. gov / catalog / PIA 03388." httpUri="https://zenodo.org/record/185554/files/figure.png" pageId="2" pageNumber="55">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="AB4B2C1AFFF6FFCEFF21F8F2FE97F85E" blockId="2.[151,1438,152,2034]" pageId="2" pageNumber="55">
|
||
<emphasis id="9980F008FFF6FFCEFF21F8F2FE15F89C" bold="true" box="[197,479,1814,1840]" pageId="2" pageNumber="55">Taxonomic comments.</emphasis>
|
||
No less than eight new gymnophthalmid genera have been described (
|
||
<bibRefCitation id="CF6551EBFFF6FFCEFACBF8F2FE87F8FB" author="Myers" pageId="2" pageNumber="67" refString="Myers, C. W. & Donnelly, M. A. (2001) Herpetofauna of the Yutaje-Corocoro Massif, Venezuela: Second report from the Robert G. Goelet American Museum-Terramar Expedition to the northwestern tepuis. Bulletin of the American Museum of Natural History, 261, 1 - 85." type="journal article" year="2001">Myers & Donnelly 2001</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="CF6551EBFFF6FFCEFEBEF8D9FD91F8FB" author="Doan" box="[346,603,1853,1879]" pageId="2" pageNumber="66" refString="Doan, T. M. & Castoe, T. A. (2005) Phylogenetic taxonomy of the Cercosaurini (Squamata: Gymnophthalmidae), with new genera for species of Neusticurus and Proctoporus. Zoological Journal of the Linnean Society, 143, 405 - 416." type="journal article" year="2005">Doan & Castoe 2005</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="CF6551EBFFF6FFCEFD8CF8D9FD28F8FB" author="Kok" box="[616,738,1853,1879]" pageId="2" pageNumber="66" refString="Kok, P. J. R. (2005) A new genus and species of gymnophthalmid lizard (Squamata: Gymnophthalmidae) from Kaieteur National Park, Guyana. Bulletin de L'Institut Royal des Sciences Naturelles de Belgique, Biologie, 75, 35 - 45." type="journal article" year="2005">Kok 2005</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="CF6551EBFFF6FFCEFD0BF8D9FC3CF8FB" author="Rodrigues" box="[751,1014,1853,1879]" pageId="2" pageNumber="67" refString="Rodrigues, M. T., Freire, E. M. X., Pellegrino, K. C. M & Sites Jr., J. W. (2005) Phylogenetic relationships of a new genus and species of microteiid lizard from the Atlantic forest of north-eastern Brazil (Squamata, Gymnophthalmidae). Zoological Journal of the Linnean Society, 144, 543 - 557." type="journal article" year="2005">
|
||
Rodrigues
|
||
<emphasis id="9980F008FFF6FFCEFC96F8D9FC65F8FA" box="[882,943,1853,1878]" italics="true" pageId="2" pageNumber="55">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="CF6551EBFFF6FFCEFBE7F8D9FB88F8FB" author="Rodrigues" box="[1027,1090,1853,1879]" pageId="2" pageNumber="67" refString="Rodrigues, M. T., Pellegrino, K. C. M, Dixo, M., Kruth Verdade, V., Pavan, D., Argolo, A. J. S. & Sites Jr., J. W. (2007) A new genus of microteiid lizard from the Atlantic forests of State of Bahia, Brazil, with a new generic name for Colobosaura mentalis, and a discussion of relationships among the Heterodactylini (Squamata, Gymnophthalmidae). American Museum Novitates, 3565, 1 - 27." type="journal article" year="2007">2007</bibRefCitation>
|
||
, Rodrigues & Maranhão dos Santos 2008) since the molecular-based phylogenetic hypothesis of
|
||
<bibRefCitation id="CF6551EBFFF6FFCEFBF9F880FA92F8D2" box="[1053,1368,1892,1918]" pageId="2" pageNumber="67" refString="Pellegrino, K. C. M., Rodrigues, M. T., Yonenaga-Yassuda, Y. & Sites Jr., J. W. (2001) A molecular perspective on the evolution of microteiids lizards (Squamata, Gymnophthalmidae), and a new classification for the family. Biological Journal of the Linnean Society, 74, 315 - 338." type="journal article">
|
||
Pellegrino
|
||
<emphasis id="9980F008FFF6FFCEFB4AF880FB39F8D1" box="[1198,1267,1892,1917]" italics="true" pageId="2" pageNumber="55">et al.</emphasis>
|
||
(2001)
|
||
</bibRefCitation>
|
||
, who demonstrated the monophyly of
|
||
<taxonomicName id="6CF45799FFF6FFCEFDF0F86EFCCBF808" box="[532,769,1930,1956]" class="Reptilia" family="Gymnophthalmidae" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="family">Gymnophthalmidae</taxonomicName>
|
||
. Some higher-level taxonomic changes are still in debate (see
|
||
<bibRefCitation id="CF6551EBFFF6FFCEFF29F855FE6FF867" author="Castoe" box="[205,421,1969,1995]" pageId="2" pageNumber="66" refString="Castoe, T. A., Doan, T. M. & Parkinson, C. L. (2004) Data partitions and complex models in Bayesian analysis: The phylogeny of gymnophthalmid lizards. Systematic Biology, 53, 448 - 469." type="journal article" year="2004">
|
||
Castoe
|
||
<emphasis id="9980F008FFF6FFCEFEC1F855FEAAF866" box="[293,352,1969,1994]" italics="true" pageId="2" pageNumber="55">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="CF6551EBFFF6FFCEFE55F855FD65F867" author="Rodrigues" box="[433,687,1969,1995]" pageId="2" pageNumber="67" refString="Rodrigues, M. T., Pellegrino, K. C. M, Dixo, M., Kruth Verdade, V., Pavan, D., Argolo, A. J. S. & Sites Jr., J. W. (2007) A new genus of microteiid lizard from the Atlantic forests of State of Bahia, Brazil, with a new generic name for Colobosaura mentalis, and a discussion of relationships among the Heterodactylini (Squamata, Gymnophthalmidae). American Museum Novitates, 3565, 1 - 27." type="journal article" year="2007">
|
||
Rodrigues
|
||
<emphasis id="9980F008FFF6FFCEFDD5F855FDA6F866" box="[561,620,1969,1994]" italics="true" pageId="2" pageNumber="55">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
) and refinement of higher-level taxonomy of gymnophthalmids is still necessary.
|
||
</paragraph>
|
||
<caption id="FF8B7C92FFF7FFCFFF73FB35FBCDFADF" httpUri="https://zenodo.org/record/185554/files/figure.png" pageId="3" pageNumber="56" targetBox="[151,1435,197,1208]" targetPageId="3">
|
||
<paragraph id="AB4B2C1AFFF7FFCFFF73FB35FBCDFADF" blockId="3.[151,1436,1233,1395]" pageId="3" pageNumber="56">
|
||
<emphasis id="9980F008FFF7FFCFFF73FB35FEDEFB45" bold="true" box="[151,276,1233,1257]" pageId="3" pageNumber="56">FIGURE 1</emphasis>
|
||
. Area map of Mount Maringma showing the type locality of
|
||
<taxonomicName id="6CF45799FFF7FFCFFC54FB36FB1AFB45" box="[944,1232,1234,1257]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="rodriguesi" status="gen. nov.">
|
||
<emphasis id="9980F008FFF7FFCFFC54FB36FB1AFB45" box="[944,1232,1234,1257]" italics="true" pageId="3" pageNumber="56">Pantepuisaurus rodriguesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9980F008FFF7FFCFFB3CFB35FA56FB45" bold="true" box="[1240,1436,1233,1257]" pageId="3" pageNumber="56">
|
||
<taxonomicNameLabel id="82B34D73FFF7FFCFFB3CFB35FAF4FB45" box="[1240,1342,1233,1257]" pageId="3" pageNumber="56" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
sp. nov.
|
||
</emphasis>
|
||
(red dot). Coordinates and boundary with Brazil are approximate and based on the available map (note that there is a substantial difference in the country boundary between this map and that shown in Google Earth). Maps elaborated after “Roraima Sheet 40 SE” published by the Survey Department of Guyana, 1972, and a radar image of South America by NASA/JPL/NIMA available at http://photojournal.jpl.nasa.gov/catalog/PIA03388.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="AB4B2C1AFFF7FFCFFF21FA46FB62F95D" blockId="3.[151,1437,1442,2009]" pageId="3" pageNumber="56">
|
||
Morphologically, the new genus shows some similarities with
|
||
<taxonomicName id="6CF45799FFF7FFCFFC7BFA46FB8BFA17" box="[927,1089,1442,1467]" class="Reptilia" family="Gymnophthalmidae" genus="Adercosaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF7FFCFFC7BFA46FB8BFA17" box="[927,1089,1442,1467]" italics="true" pageId="3" pageNumber="56">Adercosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6CF45799FFF7FFCFFB9EFA46FAF6FA17" box="[1146,1340,1442,1467]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF7FFCFFB9EFA46FAF6FA17" box="[1146,1340,1442,1467]" italics="true" pageId="3" pageNumber="56">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
. Giving the low number of specimens available, any speculation on the relationships between
|
||
<taxonomicName id="6CF45799FFF7FFCFFB72FA2DFAF3FA4E" box="[1174,1337,1481,1506]" class="Reptilia" family="Gymnophthalmidae" genus="Adercosaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF7FFCFFB72FA2DFAF3FA4E" box="[1174,1337,1481,1506]" italics="true" pageId="3" pageNumber="56">Adercosaurus</emphasis>
|
||
</taxonomicName>
|
||
(known only from the
|
||
<typeStatus id="744F92B8FFF7FFCFFEA6FA0BFE65F9A5" box="[322,431,1519,1545]" pageId="3" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
),
|
||
<taxonomicName id="6CF45799FFF7FFCFFE20FA0BFD43F9A4" box="[452,649,1519,1544]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF7FFCFFE20FA0BFD43F9A4" box="[452,649,1519,1544]" italics="true" pageId="3" pageNumber="56">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
(known only from three specimens), and
|
||
<taxonomicName id="6CF45799FFF7FFCFFB9BFA0BFAF0F9A4" box="[1151,1338,1519,1544]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF7FFCFFB9BFA0BFAF0F9A4" box="[1151,1338,1519,1544]" italics="true" pageId="3" pageNumber="56">Pantepuisaurus</emphasis>
|
||
</taxonomicName>
|
||
(known only from the
|
||
<typeStatus id="744F92B8FFF7FFCFFEAFF9F2FE70F99C" box="[331,442,1558,1584]" pageId="3" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
) seems premature. More specimens of these three genera are clearly needed to elucidate relationships between them. However, preliminary molecular analysis and comparisons do indicate affinities between
|
||
<taxonomicName id="6CF45799FFF7FFCFFE97F987FDE7F9D0" box="[371,557,1635,1660]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF7FFCFFE97F987FDE7F9D0" box="[371,557,1635,1660]" italics="true" pageId="3" pageNumber="56">Pantepuisaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6CF45799FFF7FFCFFD8DF987FCE4F9D0" box="[617,814,1635,1660]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF7FFCFFD8DF987FCE4F9D0" box="[617,814,1635,1660]" italics="true" pageId="3" pageNumber="56">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
(unfortunately, molecular data on
|
||
<taxonomicName id="6CF45799FFF7FFCFFB28F987FABAF9D0" box="[1228,1392,1635,1660]" class="Reptilia" family="Gymnophthalmidae" genus="Adercosaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF7FFCFFB28F987FABAF9D0" box="[1228,1392,1635,1660]" italics="true" pageId="3" pageNumber="56">Adercosaurus</emphasis>
|
||
</taxonomicName>
|
||
are not available), but reveal significant genetic divergences between these two genera that give further support to the erection of a new genus. Molecular analysis also suggests assignment of
|
||
<taxonomicName id="6CF45799FFF7FFCFFB73F955FAAFF966" box="[1175,1381,1713,1738]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF7FFCFFB73F955FAAFF966" box="[1175,1381,1713,1738]" italics="true" pageId="3" pageNumber="56">Pantepuisaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6CF45799FFF7FFCFFF73F933FE93F95C" box="[151,345,1751,1776]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF7FFCFFF73F933FE93F95C" box="[151,345,1751,1776]" italics="true" pageId="3" pageNumber="56">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
to the tribe Ecpleopini (Cercosaurinae) (M. T. Rodrigues, pers. comm.).
|
||
</paragraph>
|
||
<paragraph id="AB4B2C1AFFF7FFC8FF21F91AFCBEFE36" blockId="3.[151,1437,1442,2009]" lastBlockId="4.[151,1437,152,410]" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56">
|
||
A more “conservative” attitude would have been to accommodate the new species in an already known monotypic genus sharing some morphological similarities (
|
||
<emphasis id="9980F008FFF7FFCFFCBEF8C0FBE3F891" box="[858,1065,1828,1853]" italics="true" pageId="3" pageNumber="56">
|
||
i.e.
|
||
<taxonomicName id="6CF45799FFF7FFCFFC62F8C0FBE3F891" box="[902,1065,1828,1853]" class="Reptilia" family="Gymnophthalmidae" genus="Adercosaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">Adercosaurus</taxonomicName>
|
||
</emphasis>
|
||
or
|
||
<taxonomicName id="6CF45799FFF7FFCFFBB7F8C0FADDF891" box="[1107,1303,1828,1853]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF7FFCFFBB7F8C0FADDF891" box="[1107,1303,1828,1853]" italics="true" pageId="3" pageNumber="56">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
). However this would have given rise to two important problems: (1) assignment of the new species to either of these two genera would have been arbitrary and poorly supported because the new species differs from both of them in a combination of important characteristics currently used to diagnose gymnophthalmid genera (
|
||
<emphasis id="9980F008FFF7FFCFFB39F87DFACDF81E" box="[1245,1287,1945,1970]" italics="true" pageId="3" pageNumber="56">e.g.</emphasis>
|
||
condition of the posterior margin of the interparietal and parietals, condition of ventral scales, number of genials, presence of prefrontals, hemipenial morphology, tongue morphology) (2) because the new species differs from these two genera in morphological characteristics used to separate other genera (especially in the tribe Ecpleopini), placing the new species in one of them would have rendered the taxonomic status of these and other genera dubious by suggesting that these characters are variable intragenerically (thus making taxonomy of the group more complicated instead of simplifying it). One could suggest that
|
||
<taxonomicName id="6CF45799FFF0FFC8FC1EFED6FB00FEE7" box="[1018,1226,306,331]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF0FFC8FC1EFED6FB00FEE7" box="[1018,1226,306,331]" italics="true" pageId="4" pageNumber="57">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
is a synonym of
|
||
<taxonomicName id="6CF45799FFF0FFC8FF73FEBDFEF6FEDE" box="[151,316,345,370]" class="Reptilia" family="Gymnophthalmidae" genus="Adercosaurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF0FFC8FF73FEBDFEF6FEDE" box="[151,316,345,370]" italics="true" pageId="4" pageNumber="57">Adercosaurus</emphasis>
|
||
</taxonomicName>
|
||
, however this is not supported by any evidence, these two genera being differentiated on the basis of many important diagnostic characters (see
|
||
<bibRefCitation id="CF6551EBFFF0FFC8FD0AFE64FCADFE36" author="Kok" box="[750,871,384,410]" pageId="4" pageNumber="66" refString="Kok, P. J. R. (2005) A new genus and species of gymnophthalmid lizard (Squamata: Gymnophthalmidae) from Kaieteur National Park, Guyana. Bulletin de L'Institut Royal des Sciences Naturelles de Belgique, Biologie, 75, 35 - 45." type="journal article" year="2005">Kok 2005</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |