395 lines
55 KiB
XML
395 lines
55 KiB
XML
<document id="7CE56330C6DB473128EC76AB0B3538F0" ID-DOI="10.5281/zenodo.185553" ID-GBIF-Dataset="3ec6241f-1e53-4eda-abef-b9f59fa98c18" ID-ISSN="1175-5326" ID-Zenodo-Dep="185553" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459994708828" checkinUser="plazi" docAuthor="Kok, Philippe J. R." docDate="2009" docId="235D9D0CFFF0FFC1FF73F959FE6EFE36" docLanguage="en" docName="zt01992p067.pdf" docOrigin="Zootaxa 1992" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pantepuisaurus rodriguesi" docType="treatment" docVersion="9" lastPageNumber="66" masterDocId="DF64E574FFF4FFCCFFE4FFE4FFCAFFAC" masterDocTitle="Lizard in the clouds: a new highland genus and species of Gymnophthalmidae (Reptilia: Squamata) from Maringma tepui, western Guyana" masterLastPageNumber="67" masterPageNumber="53" pageNumber="57" updateTime="1698232753826" updateUser="plazi">
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<mods:titleInfo id="BF981BC5E0A877A40436DA64D5D68AA9">
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<mods:title id="628E3688763FD12EA4EF68A87251571D">Lizard in the clouds: a new highland genus and species of Gymnophthalmidae (Reptilia: Squamata) from Maringma tepui, western Guyana</mods:title>
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<mods:name id="C598789B96D5B291998AE5A5D603ECE3" type="personal">
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<mods:roleTerm id="1A6F359A6D71A8030BE788AFA5FD4C1F">Author</mods:roleTerm>
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<mods:namePart id="E8900E90F81FD714C1FE3C0C15C57118">Kok, Philippe J. R.</mods:namePart>
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<mods:typeOfResource id="79AAFC8952A3955B20D7B774EDAFB20F">text</mods:typeOfResource>
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<mods:title id="FFFFAD7141EEA0AB2E01A12EAFB8278E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D5365B55DBB86716588726201E54B41A">
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<mods:date id="C81EA4905CCD10DF214E7E8DB76030C3">2009</mods:date>
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<mods:number id="786F4CC20E51346F201E2C398C4A2708">1992</mods:number>
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<mods:classification id="175D7F27A4233B64D080CD5B37507F41">journal article</mods:classification>
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<mods:identifier id="C1590F9E9F61B5985F3A1476890B8E04" type="DOI">10.5281/zenodo.185553</mods:identifier>
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<mods:identifier id="277C4DE55472856FE74D443D3C8AF398" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="6F4E22681418FE72671691FAAA9CD4AB" type="Zenodo-Dep">185553</mods:identifier>
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<treatment id="235D9D0CFFF0FFC1FF73F959FE6EFE36" ID-DOI="http://doi.org/10.5281/zenodo.5673891" ID-GBIF-Taxon="119380225" ID-Zenodo-Dep="5673891" LSID="urn:lsid:plazi:treatment:235D9D0CFFF0FFC1FF73F959FE6EFE36" httpUri="http://treatment.plazi.org/id/235D9D0CFFF0FFC1FF73F959FE6EFE36" lastPageId="13" lastPageNumber="66" pageId="4" pageNumber="57">
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<subSubSection id="E3EE7F91FFF0FFC8FF73F959FECEF956" pageId="4" pageNumber="57" type="nomenclature">
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<paragraph id="AB4B2C1AFFF0FFC8FF73F959FD5EF97B" blockId="4.[151,660,1725,1786]" box="[151,660,1725,1751]" pageId="4" pageNumber="57">
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<heading id="F0039B76FFF0FFC8FF73F959FD5EF97B" bold="true" box="[151,660,1725,1751]" fontSize="11" level="1" pageId="4" pageNumber="57" reason="1">
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<emphasis id="9980F008FFF0FFC8FF73F959FD5EF97B" bold="true" box="[151,660,1725,1751]" pageId="4" pageNumber="57">
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<taxonomicName id="6CF45799FFF0FFC8FF73F959FE13F97B" box="[151,473,1725,1751]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="rodriguesi">
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<emphasis id="9980F008FFF0FFC8FF73F959FE13F97B" bold="true" box="[151,473,1725,1751]" italics="true" pageId="4" pageNumber="57">Pantepuisaurus rodriguesi</emphasis>
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</taxonomicName>
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species novum
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="AB4B2C1AFFF0FFC8FF73F904FECEF956" blockId="4.[151,660,1725,1786]" box="[151,260,1760,1786]" pageId="4" pageNumber="57">
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<figureCitation id="33CF309FFFF0FFC8FF73F904FECEF956" box="[151,260,1760,1786]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStart-4="FIGURE 9" captionStartId-0="8.[151,255,1941,1965]" captionStartId-1="9.[151,255,1976,2000]" captionStartId-2="10.[151,257,1440,1464]" captionStartId-3="11.[151,258,1083,1107]" captionStartId-4="11.[151,257,1914,1938]" captionTargetBox-0="[151,1436,194,1915]" captionTargetBox-1="[247,1361,199,1925]" captionTargetBox-2="[151,1435,194,1415]" captionTargetBox-3="[151,1434,194,1059]" captionTargetBox-4="[151,1435,1195,1889]" captionTargetId-0="figure@8.[151,1436,194,1917]" captionTargetId-1="figure@9.[175,1419,148,1977]" captionTargetId-2="figure@10.[151,1436,194,1416]" captionTargetId-3="figure@11.[151,1436,194,1059]" captionTargetId-4="figure@11.[151,1436,1195,1890]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="11" captionText-0="FIGURE 5. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. A: Dorsolateral view. B: Detail of head and neck region." captionText-1="FIGURE 6. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm." captionText-2="FIGURE 7. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. A: Ventral view. B: Detail of the gular region." captionText-3="FIGURE 8. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. Detail of perianal region." captionText-4="FIGURE 9. Everted hemipenis of the holotype of Pantepuisaurus rodriguesi gen. nov. sp. nov., (IRSNB 2650, left organ). A: Asulcate view. B: Sulcate view. C. Detail of mineralized spicules showing series of similar spicules interrupted by one enlarged central spicule. Scale bar is 1 mm. Photographs by Pedro Nunes." httpUri-0="https://zenodo.org/record/185558/files/figure.png" httpUri-1="https://zenodo.org/record/185559/files/figure.png" httpUri-2="https://zenodo.org/record/185560/files/figure.png" httpUri-3="https://zenodo.org/record/185561/files/figure.png" httpUri-4="https://zenodo.org/record/185562/files/figure.png" pageId="4" pageNumber="57">Figs. 5–9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="E3EE7F91FFF0FFC8FF73F8C9FCF5F839" pageId="4" pageNumber="57" type="materials_examined">
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<paragraph id="AB4B2C1AFFF0FFC8FF73F8C9FCF5F839" blockId="4.[151,1436,1837,1941]" pageId="4" pageNumber="57">
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<emphasis id="9980F008FFF0FFC8FF73F8C9FED8F8EB" bold="true" box="[151,274,1837,1863]" pageId="4" pageNumber="57">
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<typeStatus id="744F92B8FFF0FFC8FF73F8C9FEC4F8EB" box="[151,270,1837,1863]" pageId="4" pageNumber="57" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="CDE5B4DFFFF0FFC8FEFFF8C9FEB9F8EB" box="[283,371,1837,1863]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="4" pageNumber="57">IRSNB</collectionCode>
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2650 (field number PK 2044), an adult male collected by Paul Benjamin, Philippe J. R. Kok and Claudius Perry,
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<date id="DF4A0ADAFFF0FFC8FE5FF8B0FD57F8C2" box="[443,669,1876,1902]" pageId="4" pageNumber="57" value="2007-11-28">28 November 2007</date>
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at 14h55, summit plateau of Mount Maringma, Cuyuni-Mazaruni District,
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<collectingCountry id="D3E36C8AFFF0FFC8FF19F89FFE92F839" box="[253,344,1915,1941]" name="Guyana" pageId="4" pageNumber="57">Guyana</collectingCountry>
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(
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<geoCoordinate id="CEC04ADDFFF0FFC8FE8DF89FFDC0F839" box="[361,522,1915,1941]" direction="north" orientation="latitude" pageId="4" pageNumber="57" precision="15" value="5.215833">05° 12’ 57”N</geoCoordinate>
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,
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<geoCoordinate id="CEC04ADDFFF0FFC8FDF2F89FFD00F839" box="[534,714,1915,1941]" direction="west" orientation="longitude" pageId="4" pageNumber="57" precision="15" value="-60.585278">060° 35’ 07”W</geoCoordinate>
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,
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<quantity id="6C0C81FFFFF0FFC8FD32F89FFCF8F839" box="[726,818,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="2.08" pageId="4" pageNumber="57" unit="m" value="2080.0">2080 m</quantity>
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).
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</paragraph>
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</subSubSection>
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<caption id="FF8B7C92FFF1FFC9FF73FBD1FBF0FBE1" box="[151,1082,1077,1101]" httpUri="https://zenodo.org/record/185555/files/figure.png" pageId="5" pageNumber="58" targetBox="[151,1435,194,1053]" targetPageId="5">
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<paragraph id="AB4B2C1AFFF1FFC9FF73FBD1FBF0FBE1" blockId="5.[151,1082,1077,1101]" box="[151,1082,1077,1101]" pageId="5" pageNumber="58">
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<emphasis id="9980F008FFF1FFC9FF73FBD1FED8FBE1" bold="true" box="[151,274,1077,1101]" pageId="5" pageNumber="58">FIGURE 2</emphasis>
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. Maringma tepui, view of the mountain from the east (20 November 2007).
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</paragraph>
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</caption>
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<subSubSection id="E3EE7F91FFF1FFC9FF21FB9FFB5EFB4F" pageId="5" pageNumber="58" type="etymology">
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<paragraph id="AB4B2C1AFFF1FFC9FF21FB9FFB5EFB4F" blockId="5.[151,1437,1147,2024]" pageId="5" pageNumber="58">
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<emphasis id="9980F008FFF1FFC9FF21FB9FFE99FB39" bold="true" box="[197,339,1147,1173]" pageId="5" pageNumber="58">Etymology.</emphasis>
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The specific epithet is a noun in the genitive case, honoring Brazilian herpetologist Miguel Trefaut Rodrigues (Universidade de São Paulo, São Paulo,
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<collectingCountry id="D3E36C8AFFF1FFC9FCAAFB46FC5DFB10" box="[846,919,1186,1212]" name="Brazil" pageId="5" pageNumber="58">Brazil</collectingCountry>
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) for his huge contribution to the knowledge of the family
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<taxonomicName id="6CF45799FFF1FFC9FED1FB2DFDD4FB4F" box="[309,542,1225,1251]" class="Reptilia" family="Gymnophthalmidae" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="58" phylum="Chordata" rank="family">Gymnophthalmidae</taxonomicName>
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using both morphological and molecular approaches.
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</paragraph>
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</subSubSection>
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<subSubSection id="E3EE7F91FFF1FFC9FF21FB0BFD62FAD1" pageId="5" pageNumber="58" type="diagnosis">
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<paragraph id="AB4B2C1AFFF1FFC9FF21FB0BFD62FAD1" blockId="5.[151,1437,1147,2024]" pageId="5" pageNumber="58">
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<emphasis id="9980F008FFF1FFC9FF21FB0BFE8EFAA5" bold="true" box="[197,324,1263,1289]" pageId="5" pageNumber="58">Diagnosis.</emphasis>
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In addition to the generic characteristics, the new species is also characterized by the following features: four supraoculars; 15 smooth temporal scales; scales around midbody 32; dorsal scales sharply keeled, in 33 transverse rows; ventral scales in 20 transverse rows; trunk length 2.6x length of forelimb; dorsum and flanks completely black, iris red.
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</paragraph>
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</subSubSection>
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<subSubSection id="E3EE7F91FFF1FFC1FF21FA6DFD3AFF74" lastPageId="13" lastPageNumber="66" pageId="5" pageNumber="58" type="description">
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<paragraph id="AB4B2C1AFFF1FFC9FF21FA6DFBD3F920" blockId="5.[151,1437,1147,2024]" pageId="5" pageNumber="58">
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<emphasis id="9980F008FFF1FFC9FF21FA6DFDE1FA0F" bold="true" box="[197,555,1417,1443]" pageId="5" pageNumber="58">
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Description of the
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<typeStatus id="744F92B8FFF1FFC9FE54FA6DFDEFFA0F" box="[432,549,1417,1443]" pageId="5" pageNumber="58" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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An adult male
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<quantity id="6C0C81FFFFF1FFC9FD01FA6EFC85FA08" box="[741,847,1418,1444]" metricMagnitude="-2" metricUnit="m" metricValue="5.83" pageId="5" pageNumber="58" unit="mm" value="58.3">58.3 mm</quantity>
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SVL. Head length 19 % SVL, 1.38x head width, head width 1.45x head height. Head slightly wider than neck, temporal region not distinctly swollen, neck approximately as wide as anterior body. Snout blunt. Body oval in transverse section, slightly depressed, slightly wider than high. Tail complete,
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<quantity id="6C0C81FFFFF1FFC9FD9CFA1AFD2FF9B4" box="[632,741,1534,1560]" metricMagnitude="-2" metricUnit="m" metricValue="9.33" pageId="5" pageNumber="58" unit="mm" value="93.3">93.3 mm</quantity>
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, 1.60x SVL, circular in cross-section, tapering toward the tip. Limbs well-developed, forelimbs 19% SVL, hindlimbs 29% SVL. Limbs pentadactyl, all fingers and toes with terminal claw. Limbs short, not overlapping when adpressed along the body. Distance between forelimbs and hindlimbs (trunk length) =
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<quantity id="6C0C81FFFFF1FFC9FDE2F996FDBAF920" box="[518,624,1650,1676]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="5" pageNumber="58" unit="mm" value="29.2">29.2 mm</quantity>
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, 50 % SVL, 2.6x forelimbs (
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<figureCitation id="33CF309FFFF1FFC9FC26F996FBC0F920" box="[962,1034,1650,1676]" captionStart="FIGURE 5" captionStartId="8.[151,255,1941,1965]" captionTargetBox="[151,1436,194,1915]" captionTargetId="figure@8.[151,1436,194,1917]" captionTargetPageId="8" captionText="FIGURE 5. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. A: Dorsolateral view. B: Detail of head and neck region." httpUri="https://zenodo.org/record/185558/files/figure.png" pageId="5" pageNumber="58">Fig. 5</figureCitation>
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).
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</paragraph>
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<paragraph id="AB4B2C1AFFF1FFC9FF21F97DFBF1F8AC" blockId="5.[151,1437,1147,2024]" pageId="5" pageNumber="58">Tongue lanceolate, anterodorsally covered by oblique (chevron-shaped), anteriorly converging plicae, posterodorsally covered by large scalelike papillae; tip bifid, smooth; one pair of infralingual plicae after fork. Anterior teeth conical, most of posterior ones bicuspid, a few of them tricuspid.</paragraph>
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<paragraph id="AB4B2C1AFFF1FFCAFF21F8E8FBA9FD98" blockId="5.[151,1437,1147,2024]" lastBlockId="6.[151,1437,152,564]" lastPageId="6" lastPageNumber="59" pageId="5" pageNumber="58">
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Rostral roughly hexagonal, twice as wide as high, visible from above, in broad contact with frontonasal. Frontonasal single, heptagonal, laterally in contact with nasal and loreal, not in contact with first supraocular. Prefrontals roughly pentagonal, as broad as long, with a moderately long medial suture, laterally in broad contact with first supraocular and the loreal, touching the anterior point of the second supraocular on the right, separated from it by first supraocular-frontal contact on the left. Frontal hexagonal, 1.4x as long as wide, anteriorly as wide as posteriorly, laterally markedly concave; frontal laterally in contact over the entire length of second supraocular, touching the first supraocular on the left, separated from it by prefrontal-second supraocular contact on the right, in contact with third supraocular. Frontoparietals pentagonal, 1.3x as long as wide with moderately long medial suture; each frontoparietal in contact with frontal, distinctly separated from second supraocular by frontal-third supraocular contact, in broad contact with third supraocular, parietal and interparietal, and narrow contact with the fourth supraocular. Interparietal heptagonal, posteriorly projecting, 1.7x as long as wide, with parallel sides, distinctly longer than parietals. Interparietal and parietals form a jagged, irregular, posterior margin. A row of five smooth occipitals of various shapes, shorter and wider than dorsal neck scales, the central smallest. Four supraoculars, first smallest, about 3/4 of the fourth; second and third larger, subequal. Five superciliaries, first much larger than the others, with a short incomplete suture running dorsad from ciliaries on the right side and a similar very short suture on the left side; second and third smallest, subequal, fourth distinctly larger, about twice the size of the fifth (
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<figureCitation id="33CF309FFFF2FFCAFBE9FDFEFB9FFD98" box="[1037,1109,538,564]" captionStart="FIGURE 6" captionStartId="9.[151,255,1976,2000]" captionTargetBox="[247,1361,199,1925]" captionTargetId="figure@9.[175,1419,148,1977]" captionTargetPageId="9" captionText="FIGURE 6. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm." httpUri="https://zenodo.org/record/185559/files/figure.png" pageId="6" pageNumber="59">Fig. 6</figureCitation>
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).
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</paragraph>
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<caption id="FF8B7C92FFF2FFCAFF73FA1FFCCBF931" httpUri="https://zenodo.org/record/185556/files/figure.png" pageId="6" pageNumber="59" targetBox="[194,1398,609,1513]" targetPageId="6">
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<paragraph id="AB4B2C1AFFF2FFCAFF73FA1FFCCBF931" blockId="6.[151,1436,1531,1693]" pageId="6" pageNumber="59">
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<emphasis id="9980F008FFF2FFCAFF73FA1FFED8F9BF" bold="true" box="[151,274,1531,1555]" pageId="6" pageNumber="59">FIGURE 3</emphasis>
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. Vegetation on the summit plateau of Maringma tepui. A: Habitat of
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<taxonomicName id="6CF45799FFF2FFCAFC14FA18FAC1F9BF" box="[1008,1291,1532,1555]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="59" phylum="Chordata" rank="species" species="rodriguesi" status="gen. nov.">
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<emphasis id="9980F008FFF2FFCAFC14FA18FAC1F9BF" box="[1008,1291,1532,1555]" italics="true" pageId="6" pageNumber="59">Pantepuisaurus rodriguesi</emphasis>
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</taxonomicName>
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||
<emphasis id="9980F008FFF2FFCAFAF6FA1FFF0CF99A" bold="true" pageId="6" pageNumber="59">
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||
<taxonomicNameLabel id="82B34D73FFF2FFCAFAF6FA1FFABCF9BF" box="[1298,1398,1531,1555]" pageId="6" pageNumber="59" rank="genus">gen. nov.</taxonomicNameLabel>
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sp. nov.
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||
</emphasis>
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, the typical tepui meadow, consisting of very dense low vegetation with plants mainly in the families
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<taxonomicName id="6CF45799FFF2FFCAFAE2F9FAFA52F999" box="[1286,1432,1566,1589]" class="Magnoliopsida" family="Bonnetiaceae" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="family">Bonnetiaceae</taxonomicName>
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,
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<taxonomicName id="6CF45799FFF2FFCAFF73F9A5FEE7F9F4" box="[151,301,1601,1624]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="family">Bromeliaceae</taxonomicName>
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,
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<taxonomicName id="6CF45799FFF2FFCAFED3F9A5FE67F9F4" box="[311,429,1601,1624]" class="Magnoliopsida" family="Clusiaceae" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="family">Clusiaceae</taxonomicName>
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,
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<taxonomicName id="6CF45799FFF2FFCAFE5DF9A5FD8BF9F4" box="[441,577,1601,1624]" class="Liliopsida" family="Orchidaceae" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="family">Orchidaceae</taxonomicName>
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,
|
||
<taxonomicName id="6CF45799FFF2FFCAFDA8F9A5FD1EF9F4" box="[588,724,1601,1624]" class="Liliopsida" family="Rapateaceae" kingdom="Plantae" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="family">Rapateaceae</taxonomicName>
|
||
,
|
||
<taxonomicName id="6CF45799FFF2FFCAFD3BF9A5FCB5F9F4" box="[735,895,1601,1624]" class="Magnoliopsida" family="Sarraceniaceae" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="family">Sarraceniaceae</taxonomicName>
|
||
and
|
||
<taxonomicName id="6CF45799FFF2FFCAFC50F9A5FBFBF9F4" box="[948,1073,1601,1624]" class="Liliopsida" family="Xyridaceae" kingdom="Plantae" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="family">Xyridaceae</taxonomicName>
|
||
; view from the summit plateau of the mountain, looking to the south (25 November 2007). B: Dense vegetation and
|
||
<taxonomicName id="6CF45799FFF2FFCAFBE9F980FA4AF9D7" box="[1037,1408,1636,1659]" class="Magnoliopsida" family="Bonnetiaceae" genus="Bonnetia" kingdom="Animalia" order="Malpighiales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="species" species="roraimae">
|
||
<emphasis id="9980F008FFF2FFCAFBE9F980FB1DF9D7" box="[1037,1239,1636,1659]" italics="true" pageId="6" pageNumber="59">Bonnetia roraimae</emphasis>
|
||
(Bonnetiaceae)
|
||
</taxonomicName>
|
||
in the mist on the very top of the tepui (26 November 2007).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="AB4B2C1AFFF2FFCBFF22F92FFC22FF74" blockId="6.[151,1437,1739,2036]" lastBlockId="7.[151,1436,152,216]" lastPageId="7" lastPageNumber="60" pageId="6" pageNumber="59">
|
||
Nasal entire, with two short divisions running ventrad from naris to supralabial on the left side, these two short divisions are incomplete, running from the first supralabial, on the right side; naris just below centre, directed laterally. Loreal quadrangular, slightly anteriorly inclined, in contact with nasal, frenocular, one preocular, first superciliary, first supraocular, prefrontal and frontonasal, separated from supralabials by frenocular-nasal contact. Two small preoculars between first superciliary and first subocular. Frenocular roughly quadrangular, longer than wide, about half-size the loreal, in contact with first supralabial. Four suboculars. Two postoculars, the lower smallest. Seven supralabials on both sides, fifth highest, fourth below middle of the eye; a very short, inconspicuous division on the first supralabial on the right side (running ventrad from nasal); suture between second and third supralabials in contact with first subocular, suture between first and second supralabials in contact with frenocular (
|
||
<figureCitation id="33CF309FFFF3FFCBFC75FF5AFC13FF74" box="[913,985,190,216]" captionStart="FIGURE 6" captionStartId="9.[151,255,1976,2000]" captionTargetBox="[247,1361,199,1925]" captionTargetId="figure@9.[175,1419,148,1977]" captionTargetPageId="9" captionText="FIGURE 6. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm." httpUri="https://zenodo.org/record/185559/files/figure.png" pageId="7" pageNumber="60">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="FF8B7C92FFF3FFCBFF73FC88FBC7FC47" httpUri="https://zenodo.org/record/185557/files/figure.png" pageId="7" pageNumber="60" targetBox="[181,1397,273,852]" targetPageId="7">
|
||
<paragraph id="AB4B2C1AFFF3FFCBFF73FC88FBC7FC47" blockId="7.[151,1436,876,1003]" pageId="7" pageNumber="60">
|
||
<emphasis id="9980F008FFF3FFCBFF73FC88FED9FC28" bold="true" box="[151,275,876,900]" pageId="7" pageNumber="60">FIGURE 4</emphasis>
|
||
. Schematic illustration of the ventral scale condition in
|
||
<taxonomicName id="6CF45799FFF3FFCBFC91FC88FBD4FC2F" box="[885,1054,876,899]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="60" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="9980F008FFF3FFCBFC91FC88FBD4FC2F" box="[885,1054,876,899]" italics="true" pageId="7" pageNumber="60">Pantepuisaurus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="82B34D73FFF3FFCBFBC1FC88FB41FC28" box="[1061,1163,876,900]" pageId="7" pageNumber="60" rank="genus">
|
||
<emphasis id="9980F008FFF3FFCBFBC1FC88FB41FC28" bold="true" box="[1061,1163,876,900]" pageId="7" pageNumber="60">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="6CF45799FFF3FFCBFB25FC88FAB9FC2F" box="[1217,1395,876,899]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="60" phylum="Chordata" rank="genus">
|
||
<emphasis id="9980F008FFF3FFCBFB25FC88FAB9FC2F" box="[1217,1395,876,899]" italics="true" pageId="7" pageNumber="60">Kaieteurosaurus</emphasis>
|
||
</taxonomicName>
|
||
. A: Four posterior ventral scales of
|
||
<taxonomicName id="6CF45799FFF3FFCBFE0EFC6BFCCCFC0A" box="[490,774,911,934]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="rodriguesi" status="gen. nov.">
|
||
<emphasis id="9980F008FFF3FFCBFE0EFC6BFCCCFC0A" box="[490,774,911,934]" italics="true" pageId="7" pageNumber="60">Pantepuisaurus rodriguesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9980F008FFF3FFCBFCE9FC6AFC07FC0A" bold="true" box="[781,973,910,934]" pageId="7" pageNumber="60">
|
||
<taxonomicNameLabel id="82B34D73FFF3FFCBFCE9FC6AFCBBFC0A" box="[781,881,910,934]" pageId="7" pageNumber="60" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
sp. nov.
|
||
</emphasis>
|
||
showing the broad keel (in white). B: Four posterior ventral scales of
|
||
<taxonomicName id="6CF45799FFF3FFCBFE51FC56FD7BFC65" box="[437,689,946,969]" class="Reptilia" family="Gymnophthalmidae" genus="Kaieteurosaurus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="hindsi">
|
||
<emphasis id="9980F008FFF3FFCBFE51FC56FD7BFC65" box="[437,689,946,969]" italics="true" pageId="7" pageNumber="60">Kaieteurosaurus hindsi</emphasis>
|
||
</taxonomicName>
|
||
showing the broad keel (in white) that significantly extends laterally giving an aspect of smooth unkeeled lanceolate ventrals. Drawings are not at scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="AB4B2C1AFFF3FFCBFF22FBFEFE7DFB04" blockId="7.[151,1437,1050,2004]" pageId="7" pageNumber="60">
|
||
Upper eyelid with three ciliaries on both sides, second very large, apparently resulting from the fusion of several smaller scales. Ocular recess with median row of three scales separating median ciliary from superciliaries. Lower eyelid with a semi-transparent, slightly pigmented, palpebral disc of two long vertical palpebral scales (
|
||
<figureCitation id="33CF309FFFF3FFCBFE85FB6AFE63FB04" box="[353,425,1166,1192]" captionStart="FIGURE 6" captionStartId="9.[151,255,1976,2000]" captionTargetBox="[247,1361,199,1925]" captionTargetId="figure@9.[175,1419,148,1977]" captionTargetPageId="9" captionText="FIGURE 6. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm." httpUri="https://zenodo.org/record/185559/files/figure.png" pageId="7" pageNumber="60">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="AB4B2C1AFFF3FFCBFF21FB50FDA3FAEE" blockId="7.[151,1437,1050,2004]" pageId="7" pageNumber="60">
|
||
Temporal region with 15 irregularly shaped, juxtaposed, smooth scales on both sides. Temporals arranged in four oblique rows. Ear opening moderately large, vertically semi-circular with a straight posterior margin, bordered by a few small, juxtaposed, smooth scales. Auditory meatus moderately deep, tympanum mostly transparent, slightly pigmented (
|
||
<figureCitation id="33CF309FFFF3FFCBFDF7FACCFD90FAEE" box="[531,602,1320,1346]" captionStart="FIGURE 6" captionStartId="9.[151,255,1976,2000]" captionTargetBox="[247,1361,199,1925]" captionTargetId="figure@9.[175,1419,148,1977]" captionTargetPageId="9" captionText="FIGURE 6. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm." httpUri="https://zenodo.org/record/185559/files/figure.png" pageId="7" pageNumber="60">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="AB4B2C1AFFF3FFCBFF21FAABFB9EF940" blockId="7.[151,1437,1050,2004]" pageId="7" pageNumber="60">
|
||
Mental trapezoidal, its posterior edge almost straight, slightly concave. Postmental large, heptagonal, laterally in contact with first and second infralabials. Three pairs of large genials, first two pairs in contact with each other medially. First pair in lateral contact with second and third infralabials; second pair in lateral contact with third infralabial only, separated from fourth and fifth infralabials by a large sublabial scale, medial suture of second pair slightly longer than that of first pair; third pair in lateral contact with fifth infralabial, separated medially by two small scales. A pair of enlarged postgenials separated medially by scales of varying sizes and widely separated from infralabials. Seven infralabials on the right side, six on the left side, fourth scale highly reduced on both sides. One row of imbricate, smooth pregulars with rounded posterior margins. Five rows of gulars, four of them with two transversely enlarged scales. Collar row with seven scales, central one distinctly larger, others decreasing in size laterally, forming a slightly scalloped margin. Gular fold distinct, concealing one row of very small scales (
|
||
<figureCitation id="33CF309FFFF3FFCBFC26F936FBDFF940" box="[962,1045,1746,1772]" captionStart="FIGURE 6" captionStartId="9.[151,255,1976,2000]" captionTargetBox="[247,1361,199,1925]" captionTargetId="figure@9.[175,1419,148,1977]" captionTargetPageId="9" captionText="FIGURE 6. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm." httpUri="https://zenodo.org/record/185559/files/figure.png" pageId="7" pageNumber="60">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="33CF309FFFF3FFCBFBC5F936FBFBF940" box="[1057,1073,1746,1772]" captionStart="FIGURE 7" captionStartId="10.[151,257,1440,1464]" captionTargetBox="[151,1435,194,1415]" captionTargetId="figure@10.[151,1436,194,1416]" captionTargetPageId="10" captionText="FIGURE 7. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. A: Ventral view. B: Detail of the gular region." httpUri="https://zenodo.org/record/185560/files/figure.png" pageId="7" pageNumber="60">7</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
<paragraph id="AB4B2C1AFFF3FFCBFF21F91CFBA5F878" blockId="7.[151,1437,1050,2004]" pageId="7" pageNumber="60">Head scales with numerous small pits, especially concentrated on rostral, frontonasal, supraoculars, and lateral head scales, also present on ventral head scales, and body and limbs scales. Scales on nape posterior to occipitals longer than wide, with a broad, flat keel, mucronate, in transverse rows, grading posteriorly into dorsal body scales. Sides of neck with medium-sized, roughly subequal, smooth, juxtaposed scales in about eight oblique rows, not distinctly decreasing in size posteriorly but interrupted by a row of smaller scales 2/3 the way before arm insertion, which weakly form annuli with nape and gular scales.</paragraph>
|
||
<caption id="FF8B7C92FFFCFFC4FF73F871FD8CF863" httpUri="https://zenodo.org/record/185558/files/figure.png" pageId="8" pageNumber="61" targetBox="[151,1436,194,1915]" targetPageId="8">
|
||
<paragraph id="AB4B2C1AFFFCFFC4FF73F871FD8CF863" blockId="8.[151,1436,1941,1999]" pageId="8" pageNumber="61">
|
||
<emphasis id="9980F008FFFCFFC4FF73F871FED8F801" bold="true" box="[151,274,1941,1965]" pageId="8" pageNumber="61">FIGURE 5</emphasis>
|
||
.
|
||
<taxonomicName id="6CF45799FFFCFFC4FEFBF871FDF1F800" box="[287,571,1941,1964]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="rodriguesi" status="gen. nov.">
|
||
<emphasis id="9980F008FFFCFFC4FEFBF871FDF1F800" box="[287,571,1941,1964]" italics="true" pageId="8" pageNumber="61">Pantepuisaurus rodriguesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9980F008FFFCFFC4FDA5F871FCCAF801" bold="true" box="[577,768,1941,1965]" pageId="8" pageNumber="61">
|
||
<taxonomicNameLabel id="82B34D73FFFCFFC4FDA5F871FD6FF801" box="[577,677,1941,1965]" pageId="8" pageNumber="61" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
sp. nov.
|
||
</emphasis>
|
||
, male holotype (IRSNB 2650), 58.3 mm SVL. A: Dorsolateral view. B: Detail of head and neck region.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="FF8B7C92FFFDFFC5FF73F85CFDBFF85E" httpUri="https://zenodo.org/record/185559/files/figure.png" pageId="9" pageNumber="62" targetBox="[247,1361,199,1925]" targetPageId="9">
|
||
<paragraph id="AB4B2C1AFFFDFFC5FF73F85CFDBFF85E" blockId="9.[151,1436,1976,2034]" pageId="9" pageNumber="62">
|
||
<emphasis id="9980F008FFFDFFC5FF73F85CFED8F87C" bold="true" box="[151,274,1976,2000]" pageId="9" pageNumber="62">FIGURE 6</emphasis>
|
||
.
|
||
<taxonomicName id="6CF45799FFFDFFC5FEFBF85DFDF1F87C" box="[287,571,1977,2000]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="62" phylum="Chordata" rank="species" species="rodriguesi" status="gen. nov.">
|
||
<emphasis id="9980F008FFFDFFC5FEFBF85DFDF1F87C" box="[287,571,1977,2000]" italics="true" pageId="9" pageNumber="62">Pantepuisaurus rodriguesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9980F008FFFDFFC5FDA5F85CFCCAF87C" bold="true" box="[577,768,1976,2000]" pageId="9" pageNumber="62">
|
||
<taxonomicNameLabel id="82B34D73FFFDFFC5FDA5F85CFD6FF87C" box="[577,677,1976,2000]" pageId="9" pageNumber="62" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
sp. nov.
|
||
</emphasis>
|
||
, male holotype (IRSNB 2650), 58.3 mm SVL. Dorsal, ventral, and lateral views of head. Scale bar is 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="FF8B7C92FFFEFFC6FF73FA44FDC1FA76" httpUri="https://zenodo.org/record/185560/files/figure.png" pageId="10" pageNumber="63" targetBox="[151,1435,194,1415]" targetPageId="10">
|
||
<paragraph id="AB4B2C1AFFFEFFC6FF73FA44FDC1FA76" blockId="10.[151,1437,1440,1498]" pageId="10" pageNumber="63">
|
||
<emphasis id="9980F008FFFEFFC6FF73FA44FEDCFA14" bold="true" box="[151,278,1440,1464]" pageId="10" pageNumber="63">FIGURE 7</emphasis>
|
||
.
|
||
<taxonomicName id="6CF45799FFFEFFC6FEC1FA44FD81FA1B" box="[293,587,1440,1463]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="63" phylum="Chordata" rank="species" species="rodriguesi" status="gen. nov.">
|
||
<emphasis id="9980F008FFFEFFC6FEC1FA44FD81FA1B" box="[293,587,1440,1463]" italics="true" pageId="10" pageNumber="63">Pantepuisaurus rodriguesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9980F008FFFEFFC6FDB6FA44FCD0FA14" bold="true" box="[594,794,1440,1464]" pageId="10" pageNumber="63">
|
||
<taxonomicNameLabel id="82B34D73FFFEFFC6FDB6FA44FD70FA14" box="[594,698,1440,1464]" pageId="10" pageNumber="63" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
sp. nov.
|
||
</emphasis>
|
||
, male holotype (IRSNB 2650), 58.3 mm SVL. A: Ventral view. B: Detail of the gular region.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="AB4B2C1AFFFEFFC6FF21F9ECFD5EF860" blockId="10.[151,1437,1544,1996]" pageId="10" pageNumber="63">
|
||
Dorsals imbricate, pentagonal, some appearing hexagonal because of imbrication, elongate, sharply keeled, mucronate, except very few isolated scales. Dorsal scales in transverse rows only, 33 rows between the interparietal and the posterior edge of hindlimbs (including row of occipitals). Middorsal scales not distinctly narrower than adjacent dorsals. A zone of small juxtaposed scales before arm insertion. Flank scales similar to, but slightly narrower than, dorsal scales, most of flank scales are not mucronate but have a straight or slightly rounded posterior margin; no abrupt demarcation between dorsals and laterals. Lateral scales becoming slightly smaller just before leg insertion. No row of ventrolateral scales (see
|
||
<bibRefCitation id="CF6551EBFFFEFFC6FB5CF914FF1EF89D" author="Myers" pageId="10" pageNumber="67" refString="Myers, C. W. & Donnelly, M. A. (2008) The summit herpetofauna of Auyantepui, Venezuela: report from the Robert G. Goelet American Museum - Terramar expedition. Bulletin of the American Museum of Natural History, 308, 1 - 147." type="journal article" year="2008">Myers & Donnelly 2008</bibRefCitation>
|
||
) can be detected; some very small, longitudinally oval or rounded scales intercalated in the interstitial skin between the transverse rows of lateral; these small scales are more conspicuous along the first anterior third of flank and before leg insertion. All transverse rows of dorsals correspond to one row of laterals, except at midbody on the right side where one transverse row of dorsals corresponds to two rows of laterals, and before leg insertion and after arm insertion.
|
||
</paragraph>
|
||
<caption id="FF8B7C92FFFFFFC7FF73FBDFFEF5FBD9" httpUri="https://zenodo.org/record/185561/files/figure.png" pageId="11" pageNumber="64" targetBox="[151,1434,194,1059]" targetPageId="11">
|
||
<paragraph id="AB4B2C1AFFFFFFC7FF73FBDFFEF5FBD9" blockId="11.[151,1436,1083,1141]" pageId="11" pageNumber="64">
|
||
<emphasis id="9980F008FFFFFFC7FF73FBDFFEDDFBFF" bold="true" box="[151,279,1083,1107]" pageId="11" pageNumber="64">FIGURE 8</emphasis>
|
||
.
|
||
<taxonomicName id="6CF45799FFFFFFC7FEC2FBD8FD9AFBFF" box="[294,592,1084,1107]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="64" phylum="Chordata" rank="species" species="rodriguesi" status="gen. nov.">
|
||
<emphasis id="9980F008FFFFFFC7FEC2FBD8FD9AFBFF" box="[294,592,1084,1107]" italics="true" pageId="11" pageNumber="64">Pantepuisaurus rodriguesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9980F008FFFFFFC7FDBCFBDFFCE9FBFF" bold="true" box="[600,803,1083,1107]" pageId="11" pageNumber="64">
|
||
<taxonomicNameLabel id="82B34D73FFFFFFC7FDBCFBDFFD0BFBFF" box="[600,705,1083,1107]" pageId="11" pageNumber="64" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
sp. nov.
|
||
</emphasis>
|
||
, male holotype (IRSNB 2650), 58.3 mm SVL. Detail of perianal region.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="FF8B7C92FFFFFFC7FF73F89EFBBFF87B" httpUri="https://zenodo.org/record/185562/files/figure.png" pageId="11" pageNumber="64" targetBox="[151,1435,1195,1889]" targetPageId="11">
|
||
<paragraph id="AB4B2C1AFFFFFFC7FF73F89EFBBFF87B" blockId="11.[151,1437,1914,2007]" pageId="11" pageNumber="64">
|
||
<emphasis id="9980F008FFFFFFC7FF73F89EFEDCF83E" bold="true" box="[151,278,1914,1938]" pageId="11" pageNumber="64">FIGURE 9</emphasis>
|
||
. Everted hemipenis of the holotype of
|
||
<taxonomicName id="6CF45799FFFFFFC7FD2EF89FFC3BF83E" box="[714,1009,1915,1938]" class="Reptilia" family="Gymnophthalmidae" genus="Pantepuisaurus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="64" phylum="Chordata" rank="species" species="rodriguesi" status="gen. nov.">
|
||
<emphasis id="9980F008FFFFFFC7FD2EF89FFC3BF83E" box="[714,1009,1915,1938]" italics="true" pageId="11" pageNumber="64">Pantepuisaurus rodriguesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9980F008FFFFFFC7FC1CF89EFB0BF83E" bold="true" box="[1016,1217,1914,1938]" pageId="11" pageNumber="64">
|
||
<taxonomicNameLabel id="82B34D73FFFFFFC7FC1CF89EFBABF83E" box="[1016,1121,1914,1938]" pageId="11" pageNumber="64" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
sp. nov.
|
||
</emphasis>
|
||
, (IRSNB 2650, left organ). A: Asulcate view. B: Sulcate view. C. Detail of mineralized spicules showing series of similar spicules interrupted by one enlarged central spicule. Scale bar is 1 mm. Photographs by Pedro Nunes.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="AB4B2C1AFFF8FFC0FF22FF7CFBFEFEDF" blockId="12.[151,1437,152,1995]" pageId="12" pageNumber="65">
|
||
Ventrals imbricate, broadly keeled, longer than wide, mucronate except very few isolated scales, in transverse rows only (
|
||
<figureCitation id="33CF309FFFF8FFC0FE73FF5AFE16FF74" box="[407,476,190,216]" captionStart="FIGURE 7" captionStartId="10.[151,257,1440,1464]" captionTargetBox="[151,1435,194,1415]" captionTargetId="figure@10.[151,1436,194,1416]" captionTargetPageId="10" captionText="FIGURE 7. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. A: Ventral view. B: Detail of the gular region." httpUri="https://zenodo.org/record/185560/files/figure.png" pageId="12" pageNumber="65">Fig. 7</figureCitation>
|
||
A). Most transverse rows of ventrals correspond to a transverse row of laterals and of dorsals (exceptions occur at midbody). Twenty transverse rows between collar and preanal plate, 32 scales around midbody. Preanal plate consisting of six scales, one anterior, five posterior scales, most lateral posterior scales narrow, distinctly smaller than adjacent scales (
|
||
<figureCitation id="33CF309FFFF8FFC0FC9CFED6FC74FEE0" box="[888,958,306,332]" captionStart="FIGURE 8" captionStartId="11.[151,258,1083,1107]" captionTargetBox="[151,1434,194,1059]" captionTargetId="figure@11.[151,1436,194,1059]" captionTargetPageId="11" captionText="FIGURE 8. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. Detail of perianal region." httpUri="https://zenodo.org/record/185561/files/figure.png" pageId="12" pageNumber="65">Fig. 8</figureCitation>
|
||
). Four preanal pores. Four femoral pores on the right side, three on the left side, each femoral pore between three scales.
|
||
</paragraph>
|
||
<paragraph id="AB4B2C1AFFF8FFC0FF21FE64FBC2FE4B" blockId="12.[151,1437,152,1995]" pageId="12" pageNumber="65">Scales on tail keeled, not mucronate (except proximally), slightly imbricate, roughly rectangular, as long as dorsal scales, in transverse rows only, in uninterrupted annuli. Pigmented subcaudal scales keeled, slightly imbricate, slightly mucronate, roughly rectangular, in transverse rows only.</paragraph>
|
||
<paragraph id="AB4B2C1AFFF8FFC0FF21FE10FBCCFCC6" blockId="12.[151,1437,152,1995]" pageId="12" pageNumber="65">Dorsal surfaces of upper and lower arms with smooth, variably polygonal scales, ventral surface of upper arms with small, juxtaposed scales, ventral surface of lower arms with scales similar to those on dorsal surface, except at the base of hands where scales are juxtaposed and distinctly smaller. Thighs with a row of enlarged, smooth, roughly hexagonal scales on anterior surface, bordered ventrally by two rows of smaller polygonal scales diminishing in size ventrad (scales of first row smooth, scales of second row broadly keeled); bordered dorsally by 2–3 rows of keeled scales diminishing in size dorsad, grading into small, smooth (but with many very small asperities and pits that give a rugose aspect), slightly granular, juxtaposed scales, which cover posterodorsal and posterior surfaces of thighs; these scales increase in size ventrally. Shanks ventrally and posteriorly with large, smooth, polygonal, imbricate scales. Scales on upper and anterior surfaces of shanks slightly smaller, imbricate, rugose (see above), and strongly keeled.</paragraph>
|
||
<paragraph id="AB4B2C1AFFF8FFC0FF21FC92FE5DFBFE" blockId="12.[151,1437,152,1995]" pageId="12" pageNumber="65">Dorsal surface of hands and digits with imbricate, smooth scales, dorsal surface of feet with imbricate, broadly keeled scales, except a few anterodorsally; palm and soles covered with small, smooth, slightly protuberant, juxtaposed, polygonal scales. Two enlarged, subequal, thenar scales on inner margin of palm below pollex, each with produced inner edge, similar scales occur at the base of Toes I and III–IV; two slightly enlarged hypothenar scales on outer margin of palm. Subdigital lamellae divided basally and single on distal halves on most digits.</paragraph>
|
||
<paragraph id="AB4B2C1AFFF8FFC0FF21FBBAFC8BFB6A" blockId="12.[151,1437,152,1995]" pageId="12" pageNumber="65">
|
||
Subdigital lamellae as follows (Roman numbers indicate digits, Arabic numbers indicate paired or single lamellae on left/right; lower ungual sheath scale is omitted from the counts): hands
|
||
<date id="DF4A0ADAFFF8FFC0FB99FB61FB6AFB33" box="[1149,1184,1157,1183]" pageId="12" pageNumber="65">
|
||
<emphasis id="9980F008FFF8FFC0FB99FB61FB43FB33" bold="true" box="[1149,1161,1157,1183]" pageId="12" pageNumber="65">I</emphasis>
|
||
4
|
||
</date>
|
||
/4,
|
||
<date id="DF4A0ADAFFF8FFC0FB22FB61FB3EFB33" box="[1222,1268,1157,1183]" pageId="12" pageNumber="65">
|
||
<emphasis id="9980F008FFF8FFC0FB22FB61FB17FB33" bold="true" box="[1222,1245,1157,1183]" pageId="12" pageNumber="65">II</emphasis>
|
||
7
|
||
</date>
|
||
/7,
|
||
<date id="DF4A0ADAFFF8FFC0FAFFFB61FA9FFB33" box="[1307,1365,1157,1183]" pageId="12" pageNumber="65">
|
||
<emphasis id="9980F008FFF8FFC0FAFFFB61FAF4FB33" bold="true" box="[1307,1342,1157,1183]" pageId="12" pageNumber="65">III</emphasis>
|
||
9
|
||
</date>
|
||
/9,
|
||
<date id="DF4A0ADAFFF8FFC0FA9FFB61FF10FB6A" pageId="12" pageNumber="65" value="1911-04-10">
|
||
<emphasis id="9980F008FFF8FFC0FA9FFB61FA56FB33" bold="true" box="[1403,1436,1157,1183]" pageId="12" pageNumber="65">IV</emphasis>
|
||
10/11
|
||
</date>
|
||
,
|
||
<emphasis id="9980F008FFF8FFC0FF03FB4FFF36FB69" bold="true" box="[231,252,1195,1221]" pageId="12" pageNumber="65">V</emphasis>
|
||
6/7; feet
|
||
<date id="DF4A0ADAFFF8FFC0FE88FB4FFE47FB6A" box="[364,397,1195,1222]" pageId="12" pageNumber="65">
|
||
<emphasis id="9980F008FFF8FFC0FE88FB4FFEBDFB69" bold="true" box="[364,375,1195,1221]" pageId="12" pageNumber="65">I</emphasis>
|
||
5
|
||
</date>
|
||
/5,
|
||
<date id="DF4A0ADAFFF8FFC0FE57FB4FFE15FB6A" box="[435,479,1195,1222]" pageId="12" pageNumber="65">
|
||
<emphasis id="9980F008FFF8FFC0FE57FB4FFE00FB69" bold="true" box="[435,458,1195,1221]" pageId="12" pageNumber="65">II</emphasis>
|
||
8
|
||
</date>
|
||
/8,
|
||
<date id="DF4A0ADAFFF8FFC0FDE1FB4FFDB8FB6A" box="[517,626,1195,1222]" pageId="12" pageNumber="65" value="1910-03-11">
|
||
<emphasis id="9980F008FFF8FFC0FDE1FB4FFDEDFB69" bold="true" box="[517,551,1195,1221]" pageId="12" pageNumber="65">III</emphasis>
|
||
11/10
|
||
</date>
|
||
,
|
||
<date id="DF4A0ADAFFF8FFC0FD9AFB4FFD26FB6A" box="[638,748,1195,1222]" pageId="12" pageNumber="65" value="1916-04-16">
|
||
<emphasis id="9980F008FFF8FFC0FD9AFB4FFD54FB69" bold="true" box="[638,670,1195,1221]" pageId="12" pageNumber="65">IV</emphasis>
|
||
16/16
|
||
</date>
|
||
,
|
||
<emphasis id="9980F008FFF8FFC0FD1CFB4FFCC7FB69" bold="true" box="[760,781,1195,1221]" pageId="12" pageNumber="65">V</emphasis>
|
||
9/8.
|
||
</paragraph>
|
||
<paragraph id="AB4B2C1AFFF8FFC0FF21FB36FC85FA79" blockId="12.[151,1437,152,1995]" pageId="12" pageNumber="65">
|
||
<emphasis id="9980F008FFF8FFC0FF21FB36FDC9FB40" bold="true" box="[197,515,1234,1260]" pageId="12" pageNumber="65">
|
||
Color of
|
||
<typeStatus id="744F92B8FFF8FFC0FED3FB36FE61FB40" box="[311,427,1234,1260]" pageId="12" pageNumber="65" type="holotype">Holotype</typeStatus>
|
||
in life.
|
||
</emphasis>
|
||
Dorsal and lateral surfaces completely black; closer examination shows that black scales on body and neck are peppered with brown or reddish brown. Upper surface of head black. Tympanum light pink. Arms, legs and tail black. Iris red (
|
||
<figureCitation id="33CF309FFFF8FFC0FCA4FAC4FC42FA96" box="[832,904,1312,1338]" captionStart="FIGURE 5" captionStartId="8.[151,255,1941,1965]" captionTargetBox="[151,1436,194,1915]" captionTargetId="figure@8.[151,1436,194,1917]" captionTargetPageId="8" captionText="FIGURE 5. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. A: Dorsolateral view. B: Detail of head and neck region." httpUri="https://zenodo.org/record/185558/files/figure.png" pageId="12" pageNumber="65">Fig. 5</figureCitation>
|
||
). Tongue dark gray in its anterior two-thirds, whitish with gray flecks on its posterior third. Palms and soles black. Underside of head and throat black with irregular light gray markings; venter orangish brown, ventral scales peppered with black; underside of upper legs black; underside of lower legs black with irregular orangish brown markings; underside of tail black, except proximally where some scales are white (
|
||
<figureCitation id="33CF309FFFF8FFC0FD34FA5FFC88FA79" box="[720,834,1467,1493]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="10.[151,257,1440,1464]" captionStartId-1="11.[151,258,1083,1107]" captionTargetBox-0="[151,1435,194,1415]" captionTargetBox-1="[151,1434,194,1059]" captionTargetId-0="figure@10.[151,1436,194,1416]" captionTargetId-1="figure@11.[151,1436,194,1059]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 7. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. A: Ventral view. B: Detail of the gular region." captionText-1="FIGURE 8. Pantepuisaurus rodriguesi gen. nov. sp. nov., male holotype (IRSNB 2650), 58.3 mm SVL. Detail of perianal region." httpUri-0="https://zenodo.org/record/185560/files/figure.png" httpUri-1="https://zenodo.org/record/185561/files/figure.png" pageId="12" pageNumber="65">Figs. 7–8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="AB4B2C1AFFF8FFC0FF21FA04FC43F9E4" blockId="12.[151,1437,152,1995]" pageId="12" pageNumber="65">
|
||
<emphasis id="9980F008FFF8FFC0FF21FA04FDBCFA56" bold="true" box="[197,630,1504,1530]" pageId="12" pageNumber="65">
|
||
Color of
|
||
<typeStatus id="744F92B8FFF8FFC0FED2FA04FE63FA56" box="[310,425,1504,1530]" pageId="12" pageNumber="65" type="holotype">Holotype</typeStatus>
|
||
in preservative.
|
||
</emphasis>
|
||
After four months in 70 % ethanol, the
|
||
<typeStatus id="744F92B8FFF8FFC0FBBCFA05FB0AFA57" box="[1112,1216,1505,1531]" pageId="12" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
became very dark brown, the light gray markings on the underside of head and chest are white; venter is whitish peppered with dark gray; tympanum is transparent, no other noticeable change.
|
||
</paragraph>
|
||
<paragraph id="AB4B2C1AFFF8FFC1FF21F9B1FD3AFF74" blockId="12.[151,1437,152,1995]" lastBlockId="13.[151,1437,152,410]" lastPageId="13" lastPageNumber="66" pageId="12" pageNumber="65">
|
||
<emphasis id="9980F008FFF8FFC0FF21F9B1FE9FF9C3" bold="true" box="[197,341,1621,1647]" pageId="12" pageNumber="65">Hemipenis.</emphasis>
|
||
The hemipenes of the
|
||
<typeStatus id="744F92B8FFF8FFC0FD8FF9B1FD1EF9C3" box="[619,724,1621,1647]" pageId="12" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
were everted in the field. The left organ was subsequently removed, sent to and prepared by Pedro Nunes (Universidade de São Paulo, São Paulo,
|
||
<collectingCountry id="D3E36C8AFFF8FFC0FB2EF998FAD3F93A" box="[1226,1305,1660,1686]" name="Brazil" pageId="12" pageNumber="65">Brazil</collectingCountry>
|
||
), currently working on a PhD thesis on the hemipenial morphology in the family
|
||
<taxonomicName id="6CF45799FFF8FFC0FC20F947FB7BF911" box="[964,1201,1699,1725]" class="Reptilia" family="Gymnophthalmidae" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="family">Gymnophthalmidae</taxonomicName>
|
||
. The hemipenis was inflated with blue petroleum jelly and the spicules were stained with Alizarin Red (
|
||
<figureCitation id="33CF309FFFF8FFC0FB87F92DFB60F94F" box="[1123,1194,1737,1763]" captionStart="FIGURE 9" captionStartId="11.[151,257,1914,1938]" captionTargetBox="[151,1435,1195,1889]" captionTargetId="figure@11.[151,1436,1195,1890]" captionTargetPageId="11" captionText="FIGURE 9. Everted hemipenis of the holotype of Pantepuisaurus rodriguesi gen. nov. sp. nov., (IRSNB 2650, left organ). A: Asulcate view. B: Sulcate view. C. Detail of mineralized spicules showing series of similar spicules interrupted by one enlarged central spicule. Scale bar is 1 mm. Photographs by Pedro Nunes." httpUri="https://zenodo.org/record/185562/files/figure.png" pageId="12" pageNumber="65">Fig. 9</figureCitation>
|
||
). The left hemipenis is
|
||
<quantity id="6C0C81FFFFF8FFC0FF55F914FEC1F8A6" box="[177,267,1776,1802]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="12" pageNumber="65" unit="mm" value="6.9">6.9 mm</quantity>
|
||
long, with a greatest width of
|
||
<quantity id="6C0C81FFFFF8FFC0FD8AF914FD01F8A6" box="[622,715,1776,1802]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="12" pageNumber="65" unit="mm" value="4.5">4.5 mm</quantity>
|
||
. The organ is weakly bilobed, with each of the short divisions terminating in 2–3 protuberances. Sulcus spermaticus medial, broad, poorly defined. Asulcate surface with a conspicuous series of 15–18 curved transverse plicae (tissue ridges) encircling the side of the organ and terminating laterally on the sulcate surface, most plicae being laterally shortly interrupted by a narrow nude space. A wide nude space between the
|
||
<emphasis id="9980F008FFF8FFC0FDB8F86FFDB5F808" box="[604,639,1931,1956]" italics="true" pageId="12" pageNumber="65">ca.</emphasis>
|
||
8–9 distal plicae on the asulcate surface. All plicae bear mineralized spicules; plicae located on the proximal part of the hemipenis bear a single spicule, the number of spicules is increasing from the proximal part towards the tip resulting in plicae bearing a series of several uniform spicules interrupted by a larger central one (
|
||
<figureCitation id="33CF309FFFF9FFC1FD7EFF5AFD2BFF74" box="[666,737,190,216]" captionStart="FIGURE 9" captionStartId="11.[151,257,1914,1938]" captionTargetBox="[151,1435,1195,1889]" captionTargetId="figure@11.[151,1436,1195,1890]" captionTargetPageId="11" captionText="FIGURE 9. Everted hemipenis of the holotype of Pantepuisaurus rodriguesi gen. nov. sp. nov., (IRSNB 2650, left organ). A: Asulcate view. B: Sulcate view. C. Detail of mineralized spicules showing series of similar spicules interrupted by one enlarged central spicule. Scale bar is 1 mm. Photographs by Pedro Nunes." httpUri="https://zenodo.org/record/185562/files/figure.png" pageId="13" pageNumber="66">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E3EE7F91FFF9FFC1FF21FF01FE6EFE36" pageId="13" pageNumber="66" type="distribution">
|
||
<paragraph id="AB4B2C1AFFF9FFC1FF21FF01FE6EFE36" blockId="13.[151,1437,152,410]" pageId="13" pageNumber="66">
|
||
<emphasis id="9980F008FFF9FFC1FF21FF01FDC1FF53" bold="true" box="[197,523,229,255]" pageId="13" pageNumber="66">Distribution and ecology.</emphasis>
|
||
The species is currently known only from the
|
||
<typeStatus id="744F92B8FFF9FFC1FBAFFF01FBB5FF53" box="[1099,1151,229,255]" pageId="13" pageNumber="66">type</typeStatus>
|
||
locality, the summit of Maringma tepui in
|
||
<collectingCountry id="D3E36C8AFFF9FFC1FE66FEE8FE2AFE8A" box="[386,480,268,294]" name="Guyana" pageId="13" pageNumber="66">Guyana</collectingCountry>
|
||
(
|
||
<figureCitation id="33CF309FFFF9FFC1FE16FEE8FDF0FE8A" box="[498,570,268,294]" captionStart="FIGURE 1" captionStartId="3.[151,256,1233,1257]" captionTargetBox="[151,1435,197,1208]" captionTargetId="figure@3.[151,1436,197,1208]" captionTargetPageId="3" captionText="FIGURE 1. Area map of Mount Maringma showing the type locality of Pantepuisaurus rodriguesi gen. nov. sp. nov. (red dot). Coordinates and boundary with Brazil are approximate and based on the available map (note that there is a substantial difference in the country boundary between this map and that shown in Google Earth). Maps elaborated after “ Roraima Sheet 40 SE ” published by the Survey Department of Guyana, 1972, and a radar image of South America by NASA / JPL / NIMA available at http: // photojournal. jpl. nasa. gov / catalog / PIA 03388." httpUri="https://zenodo.org/record/185554/files/figure.png" pageId="13" pageNumber="66">Fig. 1</figureCitation>
|
||
). The only specimen available was collected in the afternoon (14h55), during a sunny spell, in a sunny spot, on the ground, among dense vegetation (
|
||
<figureCitation id="33CF309FFFF9FFC1FBC8FED6FBB9FEE0" box="[1068,1139,306,332]" captionStart="FIGURE 3" captionStartId="6.[151,255,1531,1555]" captionTargetBox="[194,1398,609,1513]" captionTargetId="figure@6.[194,1400,609,1513]" captionTargetPageId="6" captionText="FIGURE 3. Vegetation on the summit plateau of Maringma tepui. A: Habitat of Pantepuisaurus rodriguesi gen. nov. sp. nov., the typical tepui meadow, consisting of very dense low vegetation with plants mainly in the families Bonnetiaceae, Bromeliaceae, Clusiaceae, Orchidaceae, Rapateaceae, Sarraceniaceae and Xyridaceae; view from the summit plateau of the mountain, looking to the south (25 November 2007). B: Dense vegetation and Bonnetia roraimae (Bonnetiaceae) in the mist on the very top of the tepui (26 November 2007)." httpUri="https://zenodo.org/record/185556/files/figure.png" pageId="13" pageNumber="66">Fig. 3</figureCitation>
|
||
). No other specimen was observed. The only other lizard species recorded from the summit of Mount Maringma is
|
||
<taxonomicName id="6CF45799FFF9FFC1FAE0FEBDFE6AFE36" authority="Kok, 2008" authorityName="Kok" authorityYear="2008" class="Reptilia" family="Gymnophthalmidae" genus="Arthrosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="66" phylum="Chordata" rank="species" species="hoogmoedi">
|
||
<emphasis id="9980F008FFF9FFC1FAE0FEBDFED0FE35" italics="true" pageId="13" pageNumber="66">Arthrosaura hoogmoedi</emphasis>
|
||
<bibRefCitation id="CF6551EBFFF9FFC1FEC4FE64FE6AFE36" author="Kok" box="[288,416,384,410]" pageId="13" pageNumber="66" refString="Kok, P. J. R. (2008) A new highland species of Arthrosaura Boulenger, 1885 (Squamata: Gymnophthalmidae) from Maringma tepui on the border of Guyana and Brazil. Zootaxa, 1909, 1 - 15." type="journal article" year="2008">Kok, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |