271 lines
35 KiB
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271 lines
35 KiB
XML
<document id="CE059950A452E61541D881D15221AE30" ID-DOI="10.11646/zootaxa.5523.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13934113" ID-ZooBank="D350E64F-BA2B-4115-B28E-9739502553DB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728986145680" checkinUser="plazi" docAuthor="Villalobos-Hiriart, José Luis, Moreno-Juárez, Eric G. & Álvarez, Fernando" docDate="2024" docId="953E8795FFA8FFC6C8A19BF1FE865FCA" docLanguage="en" docName="zootaxa.5523.2.2.pdf" docOrigin="Zootaxa 5523 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5523.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudothelphusinae Ortmann 1893" docType="treatment" docVersion="2" lastPageNumber="177" masterDocId="6907FFEDFFADFFC0C8369E65FE595D4E" masterDocTitle="Xoconochcothelphusa, a new genus for Ehecatusa chiapensis (Rodríguez & Smalley, 1972), with notes on Spirothelphusa Pretzmann, 1965 (Crustacea: Decapoda: Pseudothelphusidae)" masterLastPageNumber="191" masterPageNumber="171" pageNumber="176" updateTime="1728986889405" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D9D84BFDE1F0894994803669CF9D3BA3">Xoconochcothelphusa, a new genus for Ehecatusa chiapensis (Rodríguez & Smalley, 1972), with notes on Spirothelphusa Pretzmann, 1965 (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<mods:namePart id="F89AAC3C39ABBA4E503E97AD0D95BB90">Villalobos-Hiriart, José Luis</mods:namePart>
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<mods:affiliation id="247EAFF8309B12D19A6D7A826A7CABC6">Colección Nacional de Crustáceos, Instituto de Biología, Universidad Nacional Autónoma de México, 04510 Ciudad de México, Mexico.</mods:affiliation>
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<mods:namePart id="8D366ACB0F017E2115A247B2ED196F7F">Moreno-Juárez, Eric G.</mods:namePart>
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<mods:affiliation id="336635FF1396C30068579871C3D18A6A">Posgrado en Ciencias Biológicas, Universidad Nacional Autónoma de México, 04510 Ciudad de México, Mexico.</mods:affiliation>
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<mods:namePart id="1D0CAE6E3FD4FE746033E1A50C1335A8">Álvarez, Fernando</mods:namePart>
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<mods:affiliation id="606678D909E9E927FF49A0814D660395">Colección Nacional de Crustáceos, Instituto de Biología, Universidad Nacional Autónoma de México, 04510 Ciudad de México, Mexico.</mods:affiliation>
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<mods:title id="38613E14FCCC064AFD3024271009D3D6">Zootaxa</mods:title>
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<mods:part id="BC8FB341006131747D57DA63F2961A96">
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<mods:date id="184A11FF7C7953325A110B6C44163404">2024</mods:date>
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<mods:number id="A632F9BB67A5EE1A33AAD8CFD4AB7864">2024-10-15</mods:number>
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<mods:number id="8412F1FD61F7023F5EE68A02420AAF54">5523</mods:number>
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<mods:identifier id="FD00DBC33F7BEF385F600D0CD69061E1" type="DOI">10.11646/zootaxa.5523.2.2</mods:identifier>
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<treatment id="953E8795FFA8FFC6C8A19BF1FE865FCA" ID-DOI="http://doi.org/10.5281/zenodo.13934162" ID-Zenodo-Dep="13934162" LSID="urn:lsid:plazi:treatment:953E8795FFA8FFC6C8A19BF1FE865FCA" httpUri="http://treatment.plazi.org/id/953E8795FFA8FFC6C8A19BF1FE865FCA" lastPageId="6" lastPageNumber="177" pageId="5" pageNumber="176">
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<subSubSection id="558D6508FFA8FFC5C8A19BF1FC8158E0" box="[151,728,1428,1454]" pageId="5" pageNumber="176" type="nomenclature">
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<paragraph id="1D283683FFA8FFC5C8A19BF1FC8158E0" blockId="5.[151,728,1428,1454]" box="[151,728,1428,1454]" pageId="5" pageNumber="176">
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<heading id="466081EFFFA8FFC5C8A19BF1FC8158E0" bold="true" box="[151,728,1428,1454]" fontSize="11" level="1" pageId="5" pageNumber="176" reason="1">
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<emphasis id="2FE3EA91FFA8FFC5C8A19BF1FC8158E0" bold="true" box="[151,728,1428,1454]" pageId="5" pageNumber="176">
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Subfamily
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<taxonomicName id="DA974D00FFA8FFC5C9299BF1FC8158E0" ID-CoL="847JP" authority="Ortmann 1893" authorityName="Ortmann" authorityYear="1893" box="[287,728,1428,1454]" class="Malacostraca" family="Pseudothelphusidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="subFamily" subFamily="Pseudothelphusinae">
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Pseudothelphusinae
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<bibRefCitation id="79064B72FFA8FFC5CA169BF1FC8158E0" author="Ortmann, A." box="[544,728,1428,1454]" pageId="5" pageNumber="176" pagination="411 - 495" refId="ref15356" refString="Ortmann, A. (1893) Die Decapoden-Krebse des Strasbourg Museums, mit besonderer Berucksichtingung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen. VII. Theil. Abtheilung: Brachyura (Brachyura genuina Boas) II. Unterabtheilung: Cancroidea, 2. Section: Cancrinea, 1. Gruppe: Cyclometopa. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 7, 411 - 495. https: // doi. org / 10.5962 / bhl. part. 24064" type="journal article" year="1893">Ortmann 1893</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="558D6508FFA8FFC5C8A19BB9FDC05AA0" pageId="5" pageNumber="176" type="diagnosis">
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<paragraph id="1D283683FFA8FFC5C8A19BB9FDC05AA0" blockId="5.[151,1437,1500,2031]" pageId="5" pageNumber="176">
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<emphasis id="2FE3EA91FFA8FFC5C8A19BB9FF5358B8" bold="true" box="[151,266,1500,1526]" pageId="5" pageNumber="176">Diagnosis</emphasis>
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(modified from
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<bibRefCitation id="79064B72FFA8FFC5C9F29BB8FCF958B8" author="Alvarez, F. & Ojeda, J. C. & Souza-Carvalho, E. & Villalobos, J. L. & Magalhaes, C. & Wehrtmann, I. S. & Mantelatto, F. L." box="[452,672,1500,1528]" pageId="5" pageNumber="176" pagination="973 - 1001" refId="ref13779" refString="Alvarez, F., Ojeda, J. C., Souza-Carvalho, E., Villalobos, J. L., Magalhaes, C., Wehrtmann, I. S. & Mantelatto, F. L. (2020) Revision of the higher taxonomy of Neotropical freshwater crabs of the family Pseudothelphusidae, based on multi-gene and morphological analyses. Zoological Journal of the Linnean Society, 93 (3), 973 - 1001. https: // doi. org / 10.1093 / zoolinnean / zlaa 162" type="journal article" year="2020">
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Álvarez
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<emphasis id="2FE3EA91FFA8FFC5CA129BB8FC0C58B8" box="[548,597,1500,1526]" italics="true" pageId="5" pageNumber="176">et al</emphasis>
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. 2020
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</bibRefCitation>
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). Anterolateral margin often finely serrate or granulated to unarmed, lateral armature indistinct. Exopod of third maxilliped always more than half of ischium length, ratio exopod/ischium length ranging between 0.50 and 0.84. Branchial efferent channel without spine or tooth next to aperture; orifice of efferent channel open, gap between jugal and lateral angles. Gonopod slender to robust, with obvious torsion towards median axis of body. Marginal plate and caudal surface fused to end apically in caudo-marginal projection, directed cephalically with one to three lobes. Lateral lobes along main axis absent, except in
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<taxonomicName id="DA974D00FFA8FFC5CC1398F5FAC35BE4" authorityName="Ng & Low" authorityYear="2010" box="[1061,1178,1680,1706]" class="Malacostraca" family="Pseudothelphusidae" genus="Ehecatusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
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<emphasis id="2FE3EA91FFA8FFC5CC1398F5FAC35BE4" box="[1061,1178,1680,1706]" italics="true" pageId="5" pageNumber="176">Ehecatusa</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="DA974D00FFA8FFC5CC9398F5FBC55BE4" authorityName="Villalobos-Hiriart & Moreno-Juárez & Álvarez" authorityYear="2024" box="[1189,1436,1680,1706]" class="Malacostraca" family="Pseudothelphusidae" genus="Xoconochcothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="2FE3EA91FFA8FFC5CC9398F5FBC55BE4" box="[1189,1436,1680,1706]" italics="true" pageId="5" pageNumber="176">Xoconochcothelphusa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="34D057EAFFA8FFC5C8A198D1FEBF5B81" box="[151,230,1716,1743]" pageId="5" pageNumber="176" rank="genus">
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<emphasis id="2FE3EA91FFA8FFC5C8A198D1FE865B80" bold="true" box="[151,223,1716,1742]" pageId="5" pageNumber="176">n. gen</emphasis>
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.
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</taxonomicNameLabel>
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and
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<taxonomicName id="DA974D00FFA8FFC5C92A98D1FF9C5B80" authorityName="Pretzmann" authorityYear="1965" box="[284,453,1716,1742]" class="Malacostraca" family="Pseudothelphusidae" genus="Spirothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
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<emphasis id="2FE3EA91FFA8FFC5C92A98D1FF9C5B80" box="[284,453,1716,1742]" italics="true" pageId="5" pageNumber="176">Spirothelphusa</emphasis>
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</taxonomicName>
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(see diagnosis of those genera), the latter one with a wide rounded lobe on distal half of principal axis and one accessory subapical spine on the caudolateral border (
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<figureCitation id="85AC2A06FFA8FFC5CBCF98BDFA115BBC" box="[1017,1096,1752,1778]" captionStart="FIGURE 3" captionStartId="10.[152,255,1086,1110]" captionTargetBox="[444,1143,237,1063]" captionTargetId="figure-57@10.[444,1144,237,1063]" captionTargetPageId="10" captionText="FIGURE 3. Ehecatusa mixtepensis (Rodríguez & Smalley, in Smalley, 1970), male holotype (cl 16.8 mm, cw 26.9 mm) (CNCR 309), distal portion of left G1. A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, distal view. Abbreviations: Ac = apical cavity; Apclb = accessory prominences of caudolateral border; Ccac = central crest apical cavity; Ccclb = crenate crest of caudo-lateral border; Cmp = caudo-marginal projection; Cslp = conical spines of lateral process; Dlcmp = distal lobe of caudo-marginal projection; Map = marginal plate; Mas = marginal suture; Mep = mesial process; Sfac = setae field of apical cavity; Spch = spermatic channel." figureDoi="http://doi.org/10.5281/zenodo.13934119" httpUri="https://zenodo.org/record/13934119/files/figure.png" pageId="5" pageNumber="176">Figs. 3</figureCitation>
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,
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<figureCitation id="85AC2A06FFA8FFC5CC6098BDFA3F5BBD" box="[1110,1126,1752,1779]" captionStart="FIGURE 5" captionStartId="12.[152,255,1199,1223]" captionTargetBox="[451,1137,345,1176]" captionTargetId="figure-106@12.[451,1137,345,1176]" captionTargetPageId="12" captionText="FIGURE 5. Xoconochcothelphusa chiapensis (Rodríguez & Smalley, 1972) n. comb., male holotype (cl 15.3 mm, cw 26.5 mm) (CNCR 310), distal portion of left G1. A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, distal view. Abbreviations: Ac = apical cavity; Ccac = central crest apical cavity; Cmp = caudo-marginal projection; Cs = caudal surface; Dccs = distal crest of caudal surface; Dlcmp = distal lobe of caudo-marginal projection; Lp = lateral process; Lsh = lateral shoulder; Map = marginal plate; Mas = marginal suture; Mep = mesial process; Sf = spine field of apical cavity; Spch = spermatic channel." figureDoi="http://doi.org/10.5281/zenodo.13934124" httpUri="https://zenodo.org/record/13934124/files/figure.png" pageId="5" pageNumber="176">5</figureCitation>
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,
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<figureCitation id="85AC2A06FFA8FFC5CC4498BDFADB5BBC" box="[1138,1154,1752,1778]" captionStart="FIGURE 7" captionStartId="16.[152,255,1214,1238]" captionTargetBox="[518,1069,237,1191]" captionTargetId="figure-55@16.[518,1069,237,1191]" captionTargetPageId="16" captionText="FIGURE 7. Spirothelphusa verticalis Rathbun, 1893, male (cl 20.7 mm, cw 33.2 mm) (CNCR 25444), distal portion of left G1. A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, distal view. Abbreviations: Ac = apical cavity; Apls = apical plate of lateral surface; Asus = accessory subapical spine; Cmp = caudo-marginal projection; Dlcmp = distal lobe of caudo-marginal projection; Las = lateral suture; Llopa = lateral lobe of principal axis; Map = marginal plate; Mas = marginal suture; Mep = mesial process." figureDoi="http://doi.org/10.5281/zenodo.13934128" httpUri="https://zenodo.org/record/13934128/files/figure.png" pageId="5" pageNumber="176">7</figureCitation>
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). Mesodistal lobe can be present (
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<taxonomicName id="DA974D00FFA8FFC5C8C19899FF0F5A58" box="[247,342,1788,1814]" class="Malacostraca" family="Pseudothelphusidae" genus="Tehuana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
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<emphasis id="2FE3EA91FFA8FFC5C8C19899FF0F5A58" box="[247,342,1788,1814]" italics="true" pageId="5" pageNumber="176">Tehuana</emphasis>
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</taxonomicName>
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). Mesial process as triangular, subrectangular or as irregular plate or ax-shaped laminar expansion, commonly in longitudinal position relative to principal axis of gonopod and closing the apical cavity, rarely reduced (some species of
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<taxonomicName id="DA974D00FFA8FFC5C96E9921FFEE5A10" box="[344,439,1860,1886]" class="Malacostraca" family="Pseudothelphusidae" genus="Tehuana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
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<emphasis id="2FE3EA91FFA8FFC5C96E9921FFEE5A10" box="[344,439,1860,1886]" italics="true" pageId="5" pageNumber="176">Tehuana</emphasis>
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</taxonomicName>
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). Apical cavity oblique or transversal to principal axis of gonopod, u-shaped (
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<taxonomicName id="DA974D00FFA8FFC5CD169921FBCC5A10" box="[1312,1429,1860,1886]" class="Malacostraca" family="Pseudothelphusidae" genus="Alvarezius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
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<emphasis id="2FE3EA91FFA8FFC5CD169921FBCC5A10" box="[1312,1429,1860,1886]" italics="true" pageId="5" pageNumber="176">Alvarezius</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="DA974D00FFA8FFC5C8A1990DFF075ACC" authorityName="Moreno, Villalobos & Alvarez" authorityYear="2022" box="[151,350,1896,1922]" class="Malacostraca" family="Pseudothelphusidae" genus="Mokayathelphusa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
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<emphasis id="2FE3EA91FFA8FFC5C8A1990DFF075ACC" box="[151,350,1896,1922]" italics="true" pageId="5" pageNumber="176">Mokayathelphusa</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="DA974D00FFA8FFC5C95B990DFC755ACC" authorityName="de Saussure" authorityYear="1857" box="[365,556,1896,1922]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
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<emphasis id="2FE3EA91FFA8FFC5C95B990DFC755ACC" box="[365,556,1896,1922]" italics="true" pageId="5" pageNumber="176">Pseudothelphusa</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="DA974D00FFA8FFC5CA0C990DFCF55ACC" authorityName="Alvarez" authorityYear="1989" box="[570,684,1896,1922]" class="Malacostraca" family="Pseudothelphusidae" genus="Smalleyus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
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<emphasis id="2FE3EA91FFA8FFC5CA0C990DFCF55ACC" box="[570,684,1896,1922]" italics="true" pageId="5" pageNumber="176">Smalleyus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="DA974D00FFA8FFC5CA8C990DFD405ACC" box="[698,793,1896,1922]" class="Malacostraca" family="Pseudothelphusidae" genus="Tehuana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
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<emphasis id="2FE3EA91FFA8FFC5CA8C990DFD405ACC" box="[698,793,1896,1922]" italics="true" pageId="5" pageNumber="176">Tehuana</emphasis>
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</taxonomicName>
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), closed cephalically by internal angle of mesial process, or rounded (
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<taxonomicName id="DA974D00FFA8FFC5C91A99E9FFB75AE8" authorityName="Smalley & Adkison" authorityYear="1984" box="[300,494,1932,1958]" class="Malacostraca" family="Pseudothelphusidae" genus="Disparithelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
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<emphasis id="2FE3EA91FFA8FFC5C91A99E9FFB75AE8" box="[300,494,1932,1958]" italics="true" pageId="5" pageNumber="176">Disparithelphusa</emphasis>
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</taxonomicName>
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,
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|
<taxonomicName id="DA974D00FFA8FFC5CA3799E9FC2F5AE8" authorityName="Ng & Low" authorityYear="2010" box="[513,630,1932,1958]" class="Malacostraca" family="Pseudothelphusidae" genus="Ehecatusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2FE3EA91FFA8FFC5CA3799E9FC2F5AE8" box="[513,630,1932,1958]" italics="true" pageId="5" pageNumber="176">Ehecatusa</emphasis>
|
|
</taxonomicName>
|
|
), or elongated and narrow (
|
|
<taxonomicName id="DA974D00FFA8FFC5CBF199E9FA295AE8" authorityName="Pretzmann" authorityYear="1965" box="[967,1136,1932,1958]" class="Malacostraca" family="Pseudothelphusidae" genus="Spirothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2FE3EA91FFA8FFC5CBF199E9FA295AE8" box="[967,1136,1932,1958]" italics="true" pageId="5" pageNumber="176">Spirothelphusa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="DA974D00FFA8FFC5CCB499E9FB205AE8" authorityName="Villalobos-Hiriart & Moreno-Juárez & Álvarez" authorityYear="2024" box="[1154,1401,1932,1958]" class="Malacostraca" family="Pseudothelphusidae" genus="Xoconochcothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|
<emphasis id="2FE3EA91FFA8FFC5CCB499E9FB205AE8" box="[1154,1401,1932,1958]" italics="true" pageId="5" pageNumber="176">Xoconochcothelphusa</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="2FE3EA91FFA8FFC5CDB099E9FE915A84" bold="true" pageId="5" pageNumber="176">
|
|
<taxonomicNameLabel id="34D057EAFFA8FFC5CDB099E9FE915A84" pageId="5" pageNumber="176" rank="genus">n. gen.</taxonomicNameLabel>
|
|
</emphasis>
|
|
); field of spines distally located, irregularly disposed, with variable number of terminal-pore setae; opening of spermatic channel in caudal position, median crest present or absent.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="558D6508FFABFFC6C8F19EF3FD3A5C0E" pageId="6" pageNumber="177" type="type_taxon">
|
|
<paragraph id="1D283683FFABFFC6C8F19EF3FCAD5DFF" blockId="6.[151,1437,150,644]" box="[199,756,150,177]" pageId="6" pageNumber="177">
|
|
<emphasis id="2FE3EA91FFABFFC6C8F19EF3FF145DFE" bold="true" box="[199,333,150,176]" pageId="6" pageNumber="177">
|
|
<typeStatus id="C22C8821FFABFFC6C8F19EF3FF5B5DFE" box="[199,258,150,176]" pageId="6" pageNumber="177">Type</typeStatus>
|
|
genus
|
|
</emphasis>
|
|
.
|
|
<taxonomicName id="DA974D00FFABFFC6C96D9EF3FCA95DFF" ID-CoL="7PWRV" authority="de Saussure, 1857" authorityName="de Saussure" authorityYear="1857" box="[347,752,150,177]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2FE3EA91FFABFFC6C96D9EF3FC435DFE" box="[347,538,150,176]" italics="true" pageId="6" pageNumber="177">Pseudothelphusa</emphasis>
|
|
<bibRefCitation id="79064B72FFABFFC6CA149EF3FCA95DFF" author="De Saussure, H." box="[546,752,150,177]" pageId="6" pageNumber="177" pagination="304 - 306" refId="ref14140" refString="De Saussure, H. (1857) Diagnosis de quelques Crustaces nouveaux des Antilles et du Mexique. Revue et Magasin de Zoologie Pure et Appliquee, Series 2, 9, 304 - 306." type="journal article" year="1857">de Saussure, 1857</bibRefCitation>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="1D283683FFABFFC6C8F19EDFFD3A5C0E" blockId="6.[151,1437,150,644]" pageId="6" pageNumber="177">
|
|
<emphasis id="2FE3EA91FFABFFC6C8F19EDFFFD55D9A" bold="true" box="[199,396,186,212]" pageId="6" pageNumber="177">Genera included</emphasis>
|
|
.
|
|
<taxonomicName id="DA974D00FFABFFC6C9AF9EDFFD5A5D9A" ID-CoL="BBPFS" authority="Moreno & Villalobos" authorityName="Moreno & Villalobos" box="[409,771,186,213]" class="Malacostraca" family="Pseudothelphusidae" genus="Alvarezius" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2FE3EA91FFABFFC6C9AF9EDFFC575D9A" box="[409,526,186,212]" italics="true" pageId="6" pageNumber="177">Alvarezius</emphasis>
|
|
Moreno & Villalobos
|
|
</taxonomicName>
|
|
, in Moreno, Villalobos & Álvarez, 2022b;
|
|
<taxonomicName id="DA974D00FFABFFC6CCEC9EDFFF995DB6" ID-CoL="8FXNZ" authority="Smalley & Adkison, 1984" authorityName="Smalley & Adkison" authorityYear="1984" class="Malacostraca" family="Pseudothelphusidae" genus="Disparithelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2FE3EA91FFABFFC6CCEC9EDFFBC55D9A" box="[1242,1436,186,212]" italics="true" pageId="6" pageNumber="177">Disparithelphusa</emphasis>
|
|
<bibRefCitation id="79064B72FFABFFC6C8A19EBBFF995DB6" author="Smalley, A. & Adkison, D. L." box="[151,448,222,248]" pageId="6" pageNumber="177" pagination="127 - 133" refId="ref16210" refString="Smalley, A. & Adkison, D. L. (1984) Disparithelphusa pecki, a new genus and species of fresh-water crab from Mexico (Brachyura: Pseudothelphusidae). Journal of Crustacean Biology, 4 (1), 127 - 133. https: // doi. org / 10.2307 / 1547901" type="journal article" year="1984">Smalley & Adkison, 1984</bibRefCitation>
|
|
</taxonomicName>
|
|
;
|
|
<taxonomicName id="DA974D00FFABFFC6C9FA9EBBFD545DB6" ID-CoL="7PCVS" authority="Ng & Low, 2010" authorityName="Ng & Low" authorityYear="2010" box="[460,781,222,249]" class="Malacostraca" family="Pseudothelphusidae" genus="Ehecatusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2FE3EA91FFABFFC6C9FA9EBBFC185DB6" box="[460,577,222,248]" italics="true" pageId="6" pageNumber="177">Ehecatusa</emphasis>
|
|
<bibRefCitation id="79064B72FFABFFC6CA7F9EBAFD545DB6" author="Ng, P. K. L. & Low, M. E. Y." box="[585,781,222,249]" pageId="6" pageNumber="177" pagination="34 - 46" refId="ref15233" refString="Ng, P. K. L. & Low, M. E. Y. (2010) On the generic nomenclature of nine brachyuran names, with four replacement names and two nomina protecta (Crustacea: Decapoda). Zootaxa, 2489 (1), 34 - 46. https: // doi. org / 10.11646 / zootaxa. 2489.1.2" type="journal article" year="2010">Ng & Low, 2010</bibRefCitation>
|
|
</taxonomicName>
|
|
;
|
|
<taxonomicName id="DA974D00FFABFFC6CB2F9EBBFBC15DB6" ID-CoL="BBPYW" authority="Moreno, Villalobos & Alvarez, 2022 b" authorityName="Moreno, Villalobos & Alvarez" authorityYear="2022" box="[793,1432,222,251]" class="Malacostraca" family="Pseudothelphusidae" genus="Mokayathelphusa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2FE3EA91FFABFFC6CB2F9EBBFDB95DB6" box="[793,992,222,248]" italics="true" pageId="6" pageNumber="177">Mokayathelphusa</emphasis>
|
|
Moreno, Villalobos & Álvarez, 2022b
|
|
</taxonomicName>
|
|
;
|
|
<taxonomicName id="DA974D00FFABFFC6C8A19F67FC7C5C53" ID-CoL="7PWRV" authority="de Saussure, 1857" authorityName="de Saussure" authorityYear="1857" box="[151,549,258,285]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2FE3EA91FFABFFC6C8A19F67FF0F5C52" box="[151,342,258,284]" italics="true" pageId="6" pageNumber="177">Pseudothelphusa</emphasis>
|
|
<bibRefCitation id="79064B72FFABFFC6C96D9F67FC7C5C53" author="De Saussure, H." box="[347,549,258,285]" pageId="6" pageNumber="177" pagination="304 - 306" refId="ref14140" refString="De Saussure, H. (1857) Diagnosis de quelques Crustaces nouveaux des Antilles et du Mexique. Revue et Magasin de Zoologie Pure et Appliquee, Series 2, 9, 304 - 306." type="journal article" year="1857">de Saussure, 1857</bibRefCitation>
|
|
</taxonomicName>
|
|
;
|
|
<taxonomicName id="DA974D00FFABFFC6CA199F67FD1D5C52" ID-CoL="7Q22V" authority="Alvarez, 1989" authorityName="Alvarez" authorityYear="1989" box="[559,836,258,284]" class="Malacostraca" family="Pseudothelphusidae" genus="Smalleyus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2FE3EA91FFABFFC6CA199F67FCF85C52" box="[559,673,258,284]" italics="true" pageId="6" pageNumber="177">Smalleyus</emphasis>
|
|
<bibRefCitation id="79064B72FFABFFC6CA929F67FD1D5C52" author="Alvarez, F." box="[676,836,258,284]" pageId="6" pageNumber="177" pagination="45 - 49" refId="ref13433" refString="Alvarez, F. (1989) Smalleyus tricristatus, new genus, new species, and Pseudothelphusa parabelliana, new species (Brachyura: Pseudothelphusidae) from Los Tuxtlas, Veracruz, Mexico. Proceedings of the Biological Society of Washington, 102 (1), 45 - 49." type="journal article" year="1989">Alvarez, 1989</bibRefCitation>
|
|
</taxonomicName>
|
|
;
|
|
<taxonomicName id="DA974D00FFABFFC6CB7B9F67FAE05C53" ID-CoL="7Q2MP" authority="Pretzmann, 1965" authorityName="Pretzmann" authorityYear="1965" box="[845,1209,258,285]" class="Malacostraca" family="Pseudothelphusidae" genus="Spirothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2FE3EA91FFABFFC6CB7B9F67FDAF5C52" box="[845,1014,258,284]" italics="true" pageId="6" pageNumber="177">Spirothelphusa</emphasis>
|
|
<bibRefCitation id="79064B72FFABFFC6CBCD9F66FAE05C53" author="Pretzmann, G." box="[1019,1209,258,285]" pageId="6" pageNumber="177" pagination="1 - 10" refId="ref15525" refString="Pretzmann, G. (1965) Vorlaufiger Bericht uber die Familie Pseudothelphusidae. Sitzungsberichte der Mathematisch-naturwissenschaftlichen Klasse der Osterreichischen Akademie der Wissenschaften, Wien, 1, 1 - 10." type="journal article" year="1965">Pretzmann, 1965</bibRefCitation>
|
|
</taxonomicName>
|
|
;
|
|
<taxonomicName id="DA974D00FFABFFC6CCF59F67FF485C0E" ID-CoL="7Q3V3" authority="Rodriguez & Smalley" authorityName="Rodriguez & Smalley" class="Malacostraca" family="Pseudothelphusidae" genus="Tehuana" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2FE3EA91FFABFFC6CCF59F67FB7B5C52" box="[1219,1314,258,284]" italics="true" pageId="6" pageNumber="177">Tehuana</emphasis>
|
|
Rodríguez & Smalley
|
|
</taxonomicName>
|
|
, in
|
|
<bibRefCitation id="79064B72FFABFFC6C90F9F43FF855C0E" author="Smalley, A. E." box="[313,476,294,320]" pageId="6" pageNumber="177" pagination="96 - 106" refId="ref16153" refString="Smalley, A. E. (1970) A new genus of freshwater crabs from Guatemala, with a key to the Middle American genera (Crustacea, Decapoda, Pseudothelphusidae). American Midland Naturalist, 83 (1), 96 - 106. https: // doi. org / 10.2307 / 2424009" type="journal article" year="1970">Smalley, 1970</bibRefCitation>
|
|
, and
|
|
<taxonomicName id="DA974D00FFABFFC6CA209F43FD545C0E" authorityName="Villalobos-Hiriart & Moreno-Juárez & Álvarez" authorityYear="2024" box="[534,781,294,320]" class="Malacostraca" family="Pseudothelphusidae" genus="Xoconochcothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|
<emphasis id="2FE3EA91FFABFFC6CA209F43FD545C0E" box="[534,781,294,320]" italics="true" pageId="6" pageNumber="177">Xoconochcothelphusa</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="2FE3EA91FFABFFC6CB229F43FD3A5C0E" bold="true" box="[788,867,294,320]" pageId="6" pageNumber="177">
|
|
<taxonomicNameLabel id="34D057EAFFABFFC6CB229F43FD3A5C0E" box="[788,867,294,320]" pageId="6" pageNumber="177" rank="genus">n. gen.</taxonomicNameLabel>
|
|
</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="558D6508FFABFFC6C8F19F2FFE865FCA" pageId="6" pageNumber="177" type="discussion">
|
|
<paragraph id="1D283683FFABFFC6C8F19F2FFE865FCA" blockId="6.[151,1437,150,644]" pageId="6" pageNumber="177">
|
|
<emphasis id="2FE3EA91FFABFFC6C8F19F2FFF6D5C2A" bold="true" box="[199,308,330,356]" pageId="6" pageNumber="177">Remarks</emphasis>
|
|
. The subfamily
|
|
<taxonomicName id="DA974D00FFABFFC6C9D09F2FFC9D5C2A" authorityName="Ortmann" authorityYear="1893" box="[486,708,330,356]" class="Malacostraca" family="Pseudothelphusidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="subFamily" subFamily="Pseudothelphusinae">Pseudothelphusinae</taxonomicName>
|
|
is distributed exclusively in
|
|
<collectingCountry id="65807613FFABFFC6CBC89F2FFA0E5C2A" box="[1022,1111,330,356]" name="Mexico" pageId="6" pageNumber="177">Mexico</collectingCountry>
|
|
. The genus
|
|
<taxonomicName id="DA974D00FFABFFC6CCE09F2FFBCC5C2A" authorityName="de Saussure" authorityYear="1857" box="[1238,1429,330,356]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2FE3EA91FFABFFC6CCE09F2FFBCC5C2A" box="[1238,1429,330,356]" italics="true" pageId="6" pageNumber="177">Pseudothelphusa</emphasis>
|
|
</taxonomicName>
|
|
, occupies a large area in
|
|
<collectingCountry id="65807613FFABFFC6C9989F0BFC5F5CC6" box="[430,518,366,392]" name="Mexico" pageId="6" pageNumber="177">Mexico</collectingCountry>
|
|
, along the Pacific slope from the southern of
|
|
<collectingRegion id="DF53F861FFABFFC6CC229F0BFA3F5CC6" box="[1044,1126,366,392]" country="Mexico" name="Sonora" pageId="6" pageNumber="177">Sonora</collectingRegion>
|
|
, which is the northernmost limit of the superfamily
|
|
<taxonomicName id="DA974D00FFABFFC6C99E9FF7FCCE5CE2" authorityName="Ortmann" authorityYear="1893" box="[424,663,402,428]" pageId="6" pageNumber="177" rank="superFamily" superFamily="Pseudothelphusoidea">Pseudothelphusoidea</taxonomicName>
|
|
, to the south reaching coastal rivers of
|
|
<collectingRegion id="DF53F861FFABFFC6CC799FF7FAFE5CE2" box="[1103,1191,402,428]" country="Mexico" name="Oaxaca" pageId="6" pageNumber="177">Oaxaca</collectingRegion>
|
|
; in the central portion of the country, it extends its presence through the Transmexican Volcanic Belt from the west in
|
|
<collectingRegion id="DF53F861FFABFFC6CCF89FD3FB7B5C9E" box="[1230,1314,438,464]" country="Mexico" name="Nayarit" pageId="6" pageNumber="177">Nayarit</collectingRegion>
|
|
to the east in
|
|
<collectingRegion id="DF53F861FFABFFC6C8819FBFFF5F5CBA" box="[183,262,474,500]" country="Mexico" name="Puebla" pageId="6" pageNumber="177">Puebla</collectingRegion>
|
|
, with one extra record of
|
|
<taxonomicName id="DA974D00FFABFFC6CA009FBEFDD25CBA" authority="Alvarez 1989" authorityName="Alvarez" authorityYear="1989" box="[566,907,474,500]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="parabelliana">
|
|
<emphasis id="2FE3EA91FFABFFC6CA009FBEFCBF5CBA" box="[566,742,474,500]" italics="true" pageId="6" pageNumber="177">P. parabelliana</emphasis>
|
|
<bibRefCitation id="79064B72FFABFFC6CAD89FBFFDD25CBA" author="Alvarez, F." box="[750,907,474,500]" pageId="6" pageNumber="177" pagination="45 - 49" refId="ref13433" refString="Alvarez, F. (1989) Smalleyus tricristatus, new genus, new species, and Pseudothelphusa parabelliana, new species (Brachyura: Pseudothelphusidae) from Los Tuxtlas, Veracruz, Mexico. Proceedings of the Biological Society of Washington, 102 (1), 45 - 49." type="journal article" year="1989">Alvarez 1989</bibRefCitation>
|
|
</taxonomicName>
|
|
, in Los Tuxtlas region in
|
|
<collectingRegion id="DF53F861FFABFFC6CC8F9FBEFB795CBB" box="[1209,1312,475,501]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="177">Veracruz</collectingRegion>
|
|
. The other eight genera in the subfamily occur in southern
|
|
<collectingCountry id="65807613FFABFFC6CA8D9F9BFD4A5F56" box="[699,787,510,536]" name="Mexico" pageId="6" pageNumber="177">Mexico</collectingCountry>
|
|
, in the states of
|
|
<collectingRegion id="DF53F861FFABFFC6CBE69F9AFA6E5F57" box="[976,1079,511,537]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="177">Veracruz</collectingRegion>
|
|
,
|
|
<collectingRegion id="DF53F861FFABFFC6CC759F9BFAC35F56" box="[1091,1178,510,536]" country="Mexico" name="Oaxaca" pageId="6" pageNumber="177">Oaxaca</collectingRegion>
|
|
,
|
|
<collectingRegion id="DF53F861FFABFFC6CC909F9BFB585F56" box="[1190,1281,510,536]" country="Mexico" name="Tabasco" pageId="6" pageNumber="177">Tabasco</collectingRegion>
|
|
and
|
|
<collectingRegion id="DF53F861FFABFFC6CD0D9F9BFBC15F56" box="[1339,1432,510,536]" country="Mexico" name="Chiapas" pageId="6" pageNumber="177">Chiapas</collectingRegion>
|
|
, in the latter one a new region is added that includes the Pacific slope from the town of Tonalá, to the border with
|
|
<collectingCountry id="65807613FFABFFC6C8A19C23FF4A5F2E" box="[151,275,582,608]" name="Guatemala" pageId="6" pageNumber="177">Guatemala</collectingCountry>
|
|
, known as the Soconusco region, which is the distribution area of
|
|
<taxonomicName id="DA974D00FFABFFC6CC3E9C23FB245F2E" authorityName="Villalobos-Hiriart & Moreno-Juárez & Álvarez" authorityYear="2024" baseAuthorityName="Rodriguez & Smalley" baseAuthorityYear="1972" box="[1032,1405,582,608]" class="Malacostraca" family="Pseudothelphusidae" genus="Xoconochcothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="chiapensis" status="comb. nov.">
|
|
<emphasis id="2FE3EA91FFABFFC6CC3E9C23FB245F2E" box="[1032,1405,582,608]" italics="true" pageId="6" pageNumber="177">Xoconochcothelphusa chiapensis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="2FE3EA91FFABFFC6CDB09C23FE865FCA" bold="true" pageId="6" pageNumber="177">
|
|
<taxonomicNameLabel id="34D057EAFFABFFC6CDB09C23FE865FCA" pageId="6" pageNumber="177" rank="species">n. comb.</taxonomicNameLabel>
|
|
</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |