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<document ID-DOI="http://doi.org/10.5281/zenodo.5720677" ID-GBIF-Dataset="b231b31b-455c-43ff-80a5-80e9d0874636" ID-ISBN="978-84-96553-77-4" ID-Zenodo-Dep="5720677" approvalRequired="1" approvalRequired_for_taxonomicNames="1" checkinTime="1633653942716" checkinUser="conny" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2011" docId="9525582DFF8F2603F16EFD09F94AFCF0" docLanguage="en" docName="hbmw_2_Procaviidae_0028.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 2 Hoofed Mammals, Barcelona: Lynx Edicions" docTitle="Dendrohyrax arboreus" docType="treatment" docVersion="8" lastPageNumber="47" masterDocId="691C2055FF892605F116FF9DFFB9FFE3" masterDocTitle="Procaviidae" masterLastPageNumber="47" masterPageNumber="41" pageNumber="47" updateTime="1653418574080" updateUser="tatiana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Procaviidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2011</mods:dateIssued>
<mods:dateOther type="pubDate">2011-08-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 2 Hoofed Mammals</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>41</mods:start>
<mods:end>47</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.5720677</mods:identifier>
<mods:identifier type="GBIF-Dataset">b231b31b-455c-43ff-80a5-80e9d0874636</mods:identifier>
<mods:identifier type="ISBN">978-84-96553-77-4</mods:identifier>
<mods:identifier type="Zenodo-Dep">5720677</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5720697" ID-GBIF-Taxon="190577662" ID-Zenodo-Dep="5720697" LSID="urn:lsid:plazi:treatment:9525582DFF8F2603F16EFD09F94AFCF0" httpUri="http://treatment.plazi.org/id/9525582DFF8F2603F16EFD09F94AFCF0" lastPageNumber="47" pageId="6" pageNumber="47">
<subSubSection box="[120,152,660,706]" pageId="6" pageNumber="47" type="multiple">
<paragraph blockId="6.[117,1152,660,784]" box="[120,152,660,706]" pageId="6" pageNumber="47">
<heading box="[120,152,660,706]" pageId="6" pageNumber="47">
<figureCitation box="[120,152,660,706]" captionStartId="2.[129,161,3439,3460]" captionTargetBox="[12,2784,13,3636]" captionTargetPageId="1" captionText="1. Rock Hyrax (Procavia capensis), 2. Bush Hyrax (Heterohyrax brucei), 3. Western Tree Hyrax (Dendrohyrax dorsalis), 4. Southern Tree Hyrax (Dendrohyrax arboreus), 5. Eastern Tree Hyrax (Dendrohyrax validus)" figureDoi="http://doi.org/10.5281/zenodo.6514167" httpUri="https://zenodo.org/record/6514167/files/figure.png" pageId="6" pageNumber="47">4.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[170,611,660,706]" pageId="6" pageNumber="47" type="vernacular_names">
<paragraph blockId="6.[117,1152,660,784]" box="[170,611,660,706]" pageId="6" pageNumber="47">
<heading box="[170,611,660,706]" pageId="6" pageNumber="47">
<vernacularName box="[170,611,660,706]" pageId="6" pageNumber="47">Southern Tree Hyrax</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[659,1069,660,706]" pageId="6" pageNumber="47" type="nomenclature">
<paragraph blockId="6.[117,1152,660,784]" box="[659,1069,660,706]" pageId="6" pageNumber="47">
<heading box="[659,1069,660,706]" pageId="6" pageNumber="47">
<taxonomicName baseAuthorityName="A. Smith" baseAuthorityYear="1827" box="[659,1069,660,706]" class="Mammalia" family="Procaviidae" genus="Dendrohyrax" kingdom="Animalia" order="Hyracoidea" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="arboreus">
<emphasis box="[659,1069,660,706]" italics="true" pageId="6" pageNumber="47">Dendrohyrax arboreus</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="47" type="vernacular_names">
<paragraph blockId="6.[117,1152,660,784]" box="[119,1151,724,745]" pageId="6" pageNumber="47">
<heading box="[119,1151,724,745]" pageId="6" pageNumber="47">
<emphasis bold="true" box="[119,196,724,745]" pageId="6" pageNumber="47">French:</emphasis>
<vernacularName box="[206,387,724,745]" pageId="6" pageNumber="47">Daman des arbres</vernacularName>
/
<emphasis bold="true" box="[408,499,724,745]" pageId="6" pageNumber="47">German:</emphasis>
<vernacularName box="[509,752,724,745]" pageId="6" pageNumber="47">Sidlicher Baumschliefer</vernacularName>
/
<emphasis bold="true" box="[772,863,724,745]" pageId="6" pageNumber="47">Spanish:</emphasis>
<vernacularName box="[873,1151,724,745]" pageId="6" pageNumber="47">Damanarboricola meridional</vernacularName>
</heading>
</paragraph>
<paragraph blockId="6.[117,1152,660,784]" box="[119,490,763,784]" pageId="6" pageNumber="47">
<heading box="[119,490,763,784]" pageId="6" pageNumber="47">
<emphasis bold="true" box="[119,365,763,784]" pageId="6" pageNumber="47">Other common names:</emphasis>
<vernacularName box="[374,490,763,784]" pageId="6" pageNumber="47">Tree Dassie</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="47" type="materials_examined">
<paragraph blockId="6.[733,1319,831,1257]" box="[734,1317,831,864]" pageId="6" pageNumber="47">
<emphasis bold="true" box="[734,889,831,864]" pageId="6" pageNumber="47">Taxonomy.</emphasis>
<taxonomicName authority="A. Smith, 1827" authorityName="A. Smith" authorityYear="1827" box="[902,1313,831,864]" class="Mammalia" family="Procaviidae" genus="Hyrax" kingdom="Animalia" order="Hyracoidea" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="arboreus">Hyrax arboreus A. Smith, 1827</taxonomicName>
,
</paragraph>
<paragraph blockId="6.[733,1319,831,1257]" pageId="6" pageNumber="47">
<materialsCitation ID-GBIF-Occurrence="3761283304" pageId="6" pageNumber="47">
<collectingCountry box="[735,914,878,903]" name="South Africa" pageId="6" pageNumber="47">South Africa</collectingCountry>
,
<collectingRegion box="[931,1207,878,903]" country="South Africa" name="Western Cape" pageId="6" pageNumber="47">Western Cape Prov.</collectingRegion>
forests of
<collectingRegion box="[772,842,909,942]" country="South Africa" name="Western Cape" pageId="6" pageNumber="47">Cape</collectingRegion>
of Good Hope.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="47" type="discussion">
<paragraph blockId="6.[733,1319,831,1257]" pageId="6" pageNumber="47">Seven subspecies have been described, but taxonomic boundaries are poorly defined.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="47" type="distribution">
<paragraph blockId="6.[733,1319,831,1257]" box="[734,1133,1067,1100]" pageId="6" pageNumber="47">
<emphasis bold="true" box="[734,1133,1067,1100]" pageId="6" pageNumber="47">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph blockId="6.[733,1319,831,1257]" pageId="6" pageNumber="47">
<taxonomicName authority="A. Smith, 1827" authorityName="A. Smith" authorityYear="1827" box="[733,1180,1110,1139]" class="Mammalia" family="Procaviidae" genus="Dendrohyrax" kingdom="Animalia" order="Hyracoidea" pageId="6" pageNumber="47" phylum="Chordata" rank="subSpecies" species="arboreus" subSpecies="arboreus">
<collectingCountry name="South Africa" pageId="6" pageNumber="47">D.a.arboreusA.Smith,1827—SouthAfrica(EasternCape&amp;KwaZulu-Natalprovinces).</collectingCountry>
</taxonomicName>
</paragraph>
<paragraph blockId="6.[733,1319,831,1257]" pageId="6" pageNumber="47">
<taxonomicName authority="Brauer, 1913" authorityName="Brauer" authorityYear="1913" box="[733,1193,1224,1257]" class="Mammalia" family="Procaviidae" genus="Dendrohyrax" kingdom="Animalia" order="Hyracoidea" pageId="6" pageNumber="47" phylum="Chordata" rank="subSpecies" species="arboreus" subSpecies="adolfifriederici">
<collectingCountry name="Democratic Republic of the Congo" pageId="6" pageNumber="47">D.a.adolfifriedericiBrauer,1913—EDRCongo,SWUganda,Rwanda,andBurundi.</collectingCountry>
</taxonomicName>
</paragraph>
<paragraph blockId="6.[117,772,1308,1341]" box="[117,772,1308,1341]" pageId="6" pageNumber="47">
<taxonomicName authority="Thomas &amp; Schwann, 1904" box="[117,646,1308,1341]" class="Mammalia" family="Procaviidae" genus="Dendrohyrax" kingdom="Animalia" order="Hyracoidea" pageId="6" pageNumber="47" phylum="Chordata" rank="subSpecies" species="arboreus" subSpecies="bettoni ">D. a. bettoni Thomas &amp;Schwann, 1904</taxonomicName>
— Kenya.
</paragraph>
<paragraph blockId="6.[117,1012,1348,1374]" box="[117,1012,1348,1374]" pageId="6" pageNumber="47">
<taxonomicName authority="Hahn, 1933" box="[117,452,1348,1374]" class="Mammalia" family="Procaviidae" genus="Dendrohyrax" kingdom="Animalia" order="Hyracoidea" pageId="6" pageNumber="47" phylum="Chordata" rank="subSpecies" species="arboreus" subSpecies="bettoni ">D. a. braueri Hahn, 1933</taxonomicName>
- NE Amzola. S DR Gonzo and Zambia.
</paragraph>
<paragraph blockId="6.[116,676,1378,1420]" box="[116,676,1378,1420]" pageId="6" pageNumber="47">
<taxonomicName authority="Thomas, 1900" box="[116,520,1378,1420]" class="Mammalia" family="Procaviidae" genus="Dendrohyrax" kingdom="Animalia" order="Hyracoidea" pageId="6" pageNumber="47" phylum="Chordata" rank="subSpecies" species="crawshayi " subSpecies="bettoni ">D.a. crawshayi Thomas, 1900</taxonomicName>
- C Kenya.
</paragraph>
<paragraph box="[116,1045,1426,1459]" pageId="6" pageNumber="47">
<taxonomicName authority=" Neumann, 1902" box="[116,557,1426,1455]" class="Mammalia" family="Procaviidae" genus="Dendrohyrax" kingdom="Animalia" order="Hyracoidea" pageId="6" pageNumber="47" phylum="Chordata" rank="subSpecies" species="ruwenzorii" subSpecies="bettoni ">D. a. ruwenzorii Neumann, 1902</taxonomicName>
- NE DR Congo (Ruwenzori Mts).
</paragraph>
<paragraph blockId="6.[117,1316,1456,1536]" pageId="6" pageNumber="47">
<taxonomicName authority="Matschie, 1892" box="[117,569,1466,1494]" class="Mammalia" family="Procaviidae" genus="Dendrohyrax" kingdom="Animalia" order="Hyracoidea" pageId="6" pageNumber="47" phylum="Chordata" rank="subSpecies" species="stuhlmanni " subSpecies="bettoni ">D. a. stuhlmanni Matschie, 1892</taxonomicName>
- SW KenyaîTanzania, SE DR VCongo, Malawi, and Mozambique.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="47" type="description">
<paragraph pageId="6" pageNumber="47">
<emphasis bold="true" box="[116,369,1538,1571]" pageId="6" pageNumber="47">Descriptive notes.</emphasis>
Head-body 32-60 cm; weight 1.7-4.
<quantity box="[909,976,1538,1571]" metricMagnitude="0" metricUnit="kg" metricValue="5.0" pageId="6" pageNumber="47" unit="kg" value="5.0">5 kg</quantity>
. Males and females are approximately the same size. General coloration dark brown to gray, but very variable; may appear grizzled due to buff band below black tips of guard hairs. Individuals from high rainfall areas have darker pelage. Head often darker than rest of body and ventral pelage pale, creamy, or white. Dark yellow 23-30 mm long erectile hairs in the middle of back surrounding the dorsal gland. Four digits on forefeet, three on hindfeet, with nails rather than claws, inner digit on hindfoot curved for grooming. Soles of feet are padded pink, but in ruwenzorii black in color. Cheek teeth hypsodont. Number of mammae variable. Subspecies ruwenzorii and stuhlmanni have four mammae, two front and two back. In males the anus-preputial opening distance is 1-7 cm. The penisis short, simply built, and slightly curved. Longevity over ten years.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="47" type="biology_ecology">
<paragraph pageId="6" pageNumber="47">
<emphasis bold="true" box="[116,226,1971,2004]" pageId="6" pageNumber="47">Habitat.</emphasis>
Afro-montane forest from sea level to subalpine areas up to about
<quantity box="[1167,1271,1971,2004]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="47" unit="m" value="4500.0">4500 m</quantity>
. In the Ruwenzori they also live among rock boulders. In central
<collectingCountry box="[992,1174,2010,2043]" name="Mozambique" pageId="6" pageNumber="47">Mozambique</collectingCountry>
Southern Tree Hyraxes occur in lowland evergreen forests as well as in the evergreen forests bordering the Save River. In East Africa they occur in drier Acacia woodland and evergreen riverine forests, mainly in fig trees. In southern and eastern Africa, Southern Tree Hyraxes are dependent on cavities oftrees, epiphytes, or dense matted forest vegetation for shelter. In fragmented afro-montane forest in
<collectingCountry box="[933,1110,2211,2240]" name="South Africa" pageId="6" pageNumber="47">South Africa</collectingCountry>
they persist as a dynamic mainland-island metapopulation. They have the ability to live in small forest patches, under moderate disturbance, and in a significant human-altered matrix. Therefore, all forest patches should be conserved and human disturbance reduced. Known predators include hawk-eagles (Hieraeutus sp.), crowned eagles (Stephanoaetus coronatus), Leopards (Panthera pardus), Golden Cats (Profelis aurata), genets (Genetta spp.), Servals (Leptailurus serval), Caracals (Caracal caracal), and larger owls and snakes. Tree hyraxes have a number of species of biting lice as parasites.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="47" type="food_feeding">
<paragraph pageId="6" pageNumber="47">
<emphasis bold="true" box="[115,373,2530,2555]" pageId="6" pageNumber="47">Food and Feeding.</emphasis>
Predominantly herbivorous, browsing leaves, buds, twigs, and fruits from forbs and trees year-round. Important dietary species in East Africa include Podocarpus latifolius, Schefflera volkensii, Ilex mitis, andJuniperus procera, and in southern Africa P. falcatus, Schotia laitfolia, Cassine aethiopica, Euclea natalensis, and
<taxonomicName authorityName="Pallas" authorityYear="1766" box="[1047,1267,2643,2672]" class="Mammalia" family="Procaviidae" genus="Eugenia" kingdom="Animalia" order="Hyracoidea" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="capensis">Eugenia capensis</taxonomicName>
. In the Virunga Volcanoes, leaves of Hagenia abyssinica, Hypericum revolutum, Senecio maranguensis, Galium ruwenzoriense, and Pleoppeltis excavata formed the bulk of their diet. In the Ngorongoro Crater,
<collectingCountry box="[496,624,2757,2790]" name="Tanzania" pageId="6" pageNumber="47">Tanzania</collectingCountry>
, they browse almost exclusively on Ficus natalensis and Acacia albida trees.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="47" type="breeding">
<paragraph pageId="6" pageNumber="47">
<emphasis bold="true" box="[114,248,2835,2868]" pageId="6" pageNumber="47">Breeding.</emphasis>
Reproductive biology unknown. Gestation period 220-240 days; 1-2 young per female.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="47" type="activity">
<paragraph pageId="6" pageNumber="47">
<emphasis bold="true" box="[112,346,2914,2947]" pageId="6" pageNumber="47">Activity patterns.</emphasis>
Tree hyraxes are nocturnal, with activity peaks after sunset and in the early morning before sunrise.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="47" type="biology_ecology">
<paragraph lastBlockId="6.[1391,2595,287,788]" pageId="6" pageNumber="47">
<emphasis bold="true" box="[114,809,2993,3026]" pageId="6" pageNumber="47">Movements, Home range and Social organization.</emphasis>
Primarily solitary, but groups of two and three can be found (likely mother and subadult young). Occasionally seen as family groups, several females with single male. Population densities and social organization are poorly known. The calls start with several cracking sounds, which are followed by a loud scream, repeated several times. Several short cracking sounds follow immediately after each of the first few screams. Later in the sequence the scream occurs alone. Mostly males but also females produce these calls. Each animal call is so distinctive it allows individual recognition by the human observer.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="47" type="conservation">
<paragraph blockId="6.[1391,2595,287,788]" pageId="6" pageNumber="47">
<emphasis bold="true" box="[1392,1742,401,434]" pageId="6" pageNumber="47">Status and Conservation.</emphasis>
Currently classified as Least Concern on The IUCN Red List. Rare and officially protected in
<collectingCountry box="[1860,2041,440,473]" name="South Africa" pageId="6" pageNumber="47">South Africa</collectingCountry>
. Several populations are probably endangered due to deforestation. Certain African tribes hunt tree hyraxes as a source of food, to collect skins, and as medicine. They are also important in the spiritual traditions of some tribes.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="47" type="bibRefCitation_list">
<paragraph blockId="6.[1391,2595,287,788]" pageId="6" pageNumber="47">
<emphasis bold="true" box="[1392,1545,605,630]" pageId="6" pageNumber="47">Bibliography.</emphasis>
Bothma (1966, 1971), Castely &amp; Kerley (1993), Coetzee (1966), Dobson (1876), Eley (1994), EIlerman et al. (1953), Estes (1991), Fischer (1992), Gaylard (1994), Gaylard &amp; Kerley (1997, 2001), Hahn (1934), Hoeck (1978c), Kingdon (1971, 1997), Meester et al. (1986), Meyer (1978), Milner (1994), Milner &amp; Harris (19993, 1999b), O'Donoghue (1963), Roberts (1951), Roche (1962, 1972), Rudnai (1984), Sale (1969), Schlitter (1993), Skinner &amp; Chimimba (2005).
</paragraph>
</subSubSection>
</treatment>
</document>