147 lines
17 KiB
XML
147 lines
17 KiB
XML
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<mods:title id="7187F108C4238717D502D4F73AA9C368">An inordinate fondness for Osedax (Siboglinidae: Annelida): Fourteen new species of bone worms from California</mods:title>
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<mods:namePart id="B8EFB256AA236C276A5365608AF252C3">Rouse, Greg W.</mods:namePart>
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<mods:namePart id="86FA7050CCBE3488733C9F199AE7AEC7">Goffredi, Shana K.</mods:namePart>
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<mods:namePart id="11C5454B6722E26DFB0B5163F0935E13">Johnson, Shannon B.</mods:namePart>
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<mods:namePart id="A1BFB034030BED5F73F06490A7D652E3">Vrijenhoek, Robert C.</mods:namePart>
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<mods:title id="73D99C68E00551F251CB3E05D33B03E7">Zootaxa</mods:title>
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<mods:date id="2AE924585C68A1BF6CEDBA4906E372FE">2018</mods:date>
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<mods:number id="F8A8029C7F9C627AD1EF4D3A51D45617">2018-02-05</mods:number>
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<mods:number id="05F95C7FF158735AA540D4D8AE4B3752">4377</mods:number>
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<treatment id="03FC87E5FFB161591CB1FF3B2DC2FC77" ID-DOI="http://doi.org/10.5281/zenodo.5978272" ID-GBIF-Taxon="141129090" ID-Zenodo-Dep="5978272" LSID="urn:lsid:plazi:treatment:03FC87E5FFB161591CB1FF3B2DC2FC77" httpUri="http://treatment.plazi.org/id/03FC87E5FFB161591CB1FF3B2DC2FC77" lastPageNumber="459" pageId="8" pageNumber="459">
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<subSubSection id="C34F6578FFB161591CB1FF3B2FE5FF67" box="[151,494,152,178]" pageId="8" pageNumber="459" type="nomenclature">
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<paragraph id="8BEA36F3FFB161591CB1FF3B2FE5FF67" blockId="8.[151,494,152,178]" box="[151,494,152,178]" pageId="8" pageNumber="459">
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<heading id="D0A2819FFFB161591CB1FF3B2FE5FF67" bold="true" box="[151,494,152,178]" fontSize="11" level="1" pageId="8" pageNumber="459" reason="1">
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<taxonomicName id="4C554D70FFB161591CB1FF3B2FE5FF67" ID-CoL="84KRZ" authority="Caullery, 1914" authorityName="Caullery" authorityYear="1914" box="[151,494,152,178]" class="Polychaeta" family="Siboglinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sabellida" pageId="8" pageNumber="459" phylum="Annelida" rank="family">
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<emphasis id="B921EAE1FFB161591CB1FF3B2FE5FF67" bold="true" box="[151,494,152,178]" pageId="8" pageNumber="459">
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Siboglinidae
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<bibRefCitation id="EFC44B02FFB161591D1FFF3B2FE5FF67" author="Caullery, M." box="[313,494,152,178]" journalOrPublisher="Bulletin de la Societe Zoologique de France (Evolution et Zoologie)" pageId="8" pageNumber="459" pagination="350 - 353" part="39" refString="Caullery, M. (1914) Sur les Siboglinidae, type nouveau d'invertebres receuillis par l'expedition du Siboga. Bulletin de la Societe Zoologique de France (Evolution et Zoologie), 39, 350 - 353." title="Sur les Siboglinidae, type nouveau d'invertebres receuillis par l'expedition du Siboga" type="journal article" year="1914">Caullery, 1914</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34F6578FFB161591CB1FF7D2CB9FF2D" box="[151,690,222,248]" pageId="8" pageNumber="459" type="reference_group">
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<paragraph id="8BEA36F3FFB161591CB1FF7D2CB9FF2D" blockId="8.[151,690,222,248]" box="[151,690,222,248]" pageId="8" pageNumber="459">
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<heading id="D0A2819FFFB161591CB1FF7D2CB9FF2D" bold="true" box="[151,690,222,248]" fontSize="11" level="1" pageId="8" pageNumber="459" reason="1">
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<taxonomicName id="4C554D70FFB161591CB1FF7D2CB9FF2D" ID-CoL="84S22" authority="Rouse, Goffredi & Vrijenhoek 2004" authorityName="Rouse, Goffredi & Vrijenhoek" authorityYear="2004" box="[151,690,222,248]" class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sabellida" pageId="8" pageNumber="459" phylum="Annelida" rank="genus">
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<emphasis id="B921EAE1FFB161591CB1FF7D2CB9FF2D" bold="true" box="[151,690,222,248]" pageId="8" pageNumber="459">
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<emphasis id="B921EAE1FFB161591CB1FF7D2EFAFF2D" bold="true" box="[151,241,222,248]" italics="true" pageId="8" pageNumber="459">Osedax</emphasis>
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<bibRefCitation id="EFC44B02FFB161591CDEFF7D2CB9FF2D" author="Rouse, G. W. & Goffredi, S. K. & Vrijenhoek, R. C." box="[248,690,222,248]" journalOrPublisher="Science" pageId="8" pageNumber="459" pagination="668 - 671" part="305" publicationUrl="https://doi.org/10.1126/science.1098650" refString="Rouse, G. W., Goffredi, S. K. & Vrijenhoek, R. C. (2004) Osedax: Bone-eating marine worms with dwarf males. Science, 305, 668 - 671. https: // doi. org / 10.1126 / science. 1098650" title="Osedax: Bone-eating marine worms with dwarf males" type="journal article" year="2004">Rouse, Goffredi & Vrijenhoek 2004</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34F6578FFB161591CB1FE862F5EFD3B" pageId="8" pageNumber="459" type="diagnosis">
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<paragraph id="8BEA36F3FFB161591CB1FE862F5EFD3B" blockId="8.[151,1437,293,930]" pageId="8" pageNumber="459">
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<emphasis id="B921EAE1FFB161591CB1FE862F1AFEEB" bold="true" box="[151,273,293,318]" pageId="8" pageNumber="459">Diagnosis.</emphasis>
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Siboglinids with females having a contractile trunk, bulbous ovisac, and branching ‘roots’. A crown of palps usually present, with or without pinnules. Trunk lies within transparent tube emergent from bone surface. Mostly males are dwarfs resembling larvae, exceptionally having a crown, contractile trunk, bulbous testis sac, and branching ‘roots’, as in
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<taxonomicName id="4C554D70FFB161591D8DFE312C2FFE7C" box="[427,548,402,425]" class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sabellida" pageId="8" pageNumber="459" phylum="Annelida" rank="species" species="priapus">
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<emphasis id="B921EAE1FFB161591D8DFE312C2FFE7C" box="[427,548,402,425]" italics="true" pageId="8" pageNumber="459">O. priapus</emphasis>
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</taxonomicName>
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. Crown in females, when present, comprised of cylindrical oviduct plus four palps.
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<taxonomicName id="4C554D70FFB161591CC6FE162F84FE18" box="[224,399,437,462]" class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sabellida" pageId="8" pageNumber="459" phylum="Annelida" rank="species" species="priapus">
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<emphasis id="B921EAE1FFB161591CC6FE162F84FE18" box="[224,399,437,462]" italics="true" pageId="8" pageNumber="459">Osedax priapus</emphasis>
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</taxonomicName>
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, the only species known to produce adult bone-eating males, has crown with only two palps. No mouth or obvious gut. Cylindrical trunk comprised mostly of longitudinal muscles and glands, with large dorsal and ventral blood vessels present. Oviduct or sperm duct runs dorsally along trunk surface into posterior ovisac, or testis sac. Ovisac or testis sac enclosed by epidermis and trophosome with bacteriocytes, which also extends outwards as vascularized ‘roots’. No chaetae or segmentation apparent in females or bone-eating males. In most species, paedomorphic dwarf males cluster around oviduct in gelatinous tube surrounding trunk of female. The dwarf males possess anterior prototroch and posterior hooked chaetae arranged on two segments. Hooks, lacking rostrum, comprise capitium with curved teeth over subrostral process. Internally, males contain spermatids and sperm anteriorly.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34F6578FFB161591CE1FD5A2DC2FC77" pageId="8" pageNumber="459" type="discussion">
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<paragraph id="8BEA36F3FFB161591CE1FD5A2DC2FC77" blockId="8.[151,1437,293,930]" pageId="8" pageNumber="459">
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<emphasis id="B921EAE1FFB161591CE1FD5A2F3FFCC7" bold="true" box="[199,308,761,786]" pageId="8" pageNumber="459">Remarks</emphasis>
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. This diagnosis is revised from that in
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<bibRefCitation id="EFC44B02FFB161591ED7FD592DC0FCC7" author="Rouse, G. W. & Goffredi, S. K. & Vrijenhoek, R. C." box="[753,971,761,786]" journalOrPublisher="Science" pageId="8" pageNumber="459" pagination="668 - 671" part="305" publicationUrl="https://doi.org/10.1126/science.1098650" refString="Rouse, G. W., Goffredi, S. K. & Vrijenhoek, R. C. (2004) Osedax: Bone-eating marine worms with dwarf males. Science, 305, 668 - 671. https: // doi. org / 10.1126 / science. 1098650" title="Osedax: Bone-eating marine worms with dwarf males" type="journal article" year="2004">
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Rouse
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<emphasis id="B921EAE1FFB161591F19FD592D73FCC7" box="[831,888,761,786]" italics="true" pageId="8" pageNumber="459">et al.</emphasis>
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(2004)
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</bibRefCitation>
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to accommodate the diversity of
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<taxonomicName id="4C554D70FFB16159196EFD5A2B97FCC7" authorityName="Rouse, Goffredi & Vrijenhoek" authorityYear="2004" box="[1352,1436,761,786]" class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sabellida" pageId="8" pageNumber="459" phylum="Annelida" rank="genus">
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<emphasis id="B921EAE1FFB16159196EFD5A2B97FCC7" box="[1352,1436,761,786]" italics="true" pageId="8" pageNumber="459">Osedax</emphasis>
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</taxonomicName>
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forms. These include
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<taxonomicName id="4C554D70FFB161591DB4FCBE2C24FCE3" authority="Rouse & Goffredi & Johnson & Vrijenhoek, 2018" authorityName="Rouse & Goffredi & Johnson & Vrijenhoek" authorityYear="2018" box="[402,559,797,822]" class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sabellida" pageId="8" pageNumber="459" phylum="Annelida" rank="species" species="jabba" status="sp. nov.">
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<emphasis id="B921EAE1FFB161591DB4FCBE2C24FCE3" box="[402,559,797,822]" italics="true" pageId="8" pageNumber="459">Osedax jabba</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A212579AFFB161591E1FFCBE2C70FCE3" box="[569,635,797,822]" pageId="8" pageNumber="459" rank="species">
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<emphasis id="B921EAE1FFB161591E1FFCBE2C70FCE3" bold="true" box="[569,635,797,822]" pageId="8" pageNumber="459">n. sp.</emphasis>
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</taxonomicNameLabel>
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where the females lack palps entirely and
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<taxonomicName id="4C554D70FFB161591848FCBE2B2AFCE0" box="[1134,1313,797,822]" class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sabellida" pageId="8" pageNumber="459" phylum="Annelida" rank="species" species="priapus">
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<emphasis id="B921EAE1FFB161591848FCBE2B2AFCE0" box="[1134,1313,797,822]" italics="true" pageId="8" pageNumber="459">Osedax priapus</emphasis>
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</taxonomicName>
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where the males are not paedomorphic dwarfs, but also consume bone and have similar anatomy to females. Many
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<taxonomicName id="4C554D70FFB16159196EFCE22B97FC8F" authorityName="Rouse, Goffredi & Vrijenhoek" authorityYear="2004" box="[1352,1436,833,858]" class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sabellida" pageId="8" pageNumber="459" phylum="Annelida" rank="genus">
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<emphasis id="B921EAE1FFB16159196EFCE22B97FC8F" box="[1352,1436,833,858]" italics="true" pageId="8" pageNumber="459">Osedax</emphasis>
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</taxonomicName>
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species have palps that lack obvious pinnules. The dorsal placement of the oviduct reflects the reorientation of
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<taxonomicName id="4C554D70FFB161591CB1FC2A2EE0FC77" authorityName="Rouse, Goffredi & Vrijenhoek" authorityYear="2004" box="[151,235,905,930]" class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sabellida" pageId="8" pageNumber="459" phylum="Annelida" rank="genus">
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<emphasis id="B921EAE1FFB161591CB1FC2A2EE0FC77" box="[151,235,905,930]" italics="true" pageId="8" pageNumber="459">Osedax</emphasis>
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</taxonomicName>
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as reported in
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<bibRefCitation id="EFC44B02FFB161591DB7FC2A2C90FC77" author="Huusgaard, R. S. & Vismann, B. & Kuhl, M. & Macnaugton, M. & Colmander, V & Rouse, G. W." box="[401,667,905,930]" journalOrPublisher="The potent respiratory system of Osedax mucofloris (Siboglinidae, Annelida) - a prerequisite for the origin of bone-eating Osedax? PLoS ONE" pageId="8" pageNumber="459" pagination="35975" part="7" publicationUrl="https://doi.org/10.1371/journal.pone.0035975" refString="Huusgaard, R. S., Vismann, B., Kuhl, M., Macnaugton, M., Colmander, V., Rouse, G. W., Glover, A. G., Dahlgren, T. & Worsaae, K. (2012) The potent respiratory system of Osedax mucofloris (Siboglinidae, Annelida) - a prerequisite for the origin of bone-eating Osedax? PLoS ONE, 7, e 35975. https: // doi. org / 10.1371 / journal. pone. 0035975" title="Glover, A. G., Dahlgren, T. & Worsaae, K" type="journal article" year="2012">
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Huusgaard
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<emphasis id="B921EAE1FFB161591E37FC292C42FC77" box="[529,585,905,930]" italics="true" pageId="8" pageNumber="459">et al.</emphasis>
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(2012)
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</bibRefCitation>
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and
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<bibRefCitation id="EFC44B02FFB161591EF4FC292DCEFC77" author="Worsaae, K. & Rimskaya-Korsakova, N. N. & Rouse, G. W." box="[722,965,905,930]" journalOrPublisher="BMC Evolutionary Biology" pageId="8" pageNumber="459" pagination="83" part="16" publicationUrl="https://doi.org/10.1186/s12862-016-0639-7" refString="Worsaae, K., Rimskaya-Korsakova, N. N. & Rouse, G. W. (2016) Neural reconstruction of bone-eating Osedax spp. (Annelida) and evolution of the siboglinid nervous system. BMC Evolutionary Biology, 16, 83. https: // doi. org / 10.1186 / s 12862 - 016 - 0639 - 7" title="Neural reconstruction of bone-eating Osedax spp. (Annelida) and evolution of the siboglinid nervous system" type="journal article" year="2016">
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Worsaae
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<emphasis id="B921EAE1FFB161591F1EFC292D62FC77" box="[824,873,905,930]" italics="true" pageId="8" pageNumber="459">et al</emphasis>
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. (2016)
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</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |