372 lines
46 KiB
XML
372 lines
46 KiB
XML
<document id="5B7A4006F594A0F3CBE03C7FB8FDF8C2" ID="10.11646/zootaxa.4250.4.2SLASH10731" ID-CLB-Dataset="33291" ID-DOI="10.11646/zootaxa.4250.4.2" ID-GBIF-Dataset="ce4e62c5-1d2a-478e-a0f3-1d2637e76f8e" ID-ISSN="1175-5326" ID-Zenodo-Dep="495277" ID-ZooBank="CE45B4DB-641B-4D37-AD35-93CCB384BE1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1491462921247" checkinUser="plazi" docAuthor="Kompier, Tom" docDate="2017" docId="03FAA703ED384634FF5FF05BA035FC17" docLanguage="en" docName="zootaxa.4250.4.2SLASH10731.pdf" docOrigin="Zootaxa 4250 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lyriothemis kameliyae Kompier, 2017, spec. nov." docType="treatment" docVersion="9" lastPageNumber="320" masterDocId="FFC3DF7BED394631FFC8F307A445FFFA" masterDocTitle="The riddle of Lyriothemis bivittata (Rambur, 1842): Lyriothemis kameliyae spec. nov. (Odonata: Libellulidae)" masterLastPageNumber="326" masterPageNumber="315" pageNumber="316" updateTime="1698438726893" updateUser="ExternalLinkService">
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<mods:titleInfo id="93F69FD1FFD5F483DA870704BC9950F9">
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<mods:title id="5D0543F7FB4FBC1AD74BD89AF726909E">The riddle of Lyriothemis bivittata (Rambur, 1842): Lyriothemis kameliyae spec. nov. (Odonata: Libellulidae)</mods:title>
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<mods:namePart id="E2AF17F7DB25D6D562FA9859C46B2B6C">Kompier, Tom</mods:namePart>
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<mods:typeOfResource id="81197282ADEF4EAE53935479D0415538">text</mods:typeOfResource>
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<mods:title id="1633BDE60DABF4443A3E9D421FD7BC04">Zootaxa</mods:title>
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<mods:part id="F6654AB3990E51802EE3727B80678EB7">
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<mods:date id="AE4FB8E7DB697DA02A9DDCDEBBDD7E2F">2017</mods:date>
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<mods:number id="8F590E09FAA74529FADDC86079C6C23B">4250</mods:number>
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<mods:identifier id="6FF617FD2BAF88A0157B60A3C19E65AD" type="DOI">10.11646/zootaxa.4250.4.2</mods:identifier>
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<treatment id="03FAA703ED384634FF5FF05BA035FC17" ID-DOI="http://doi.org/10.5281/zenodo.5689898" ID-GBIF-Taxon="128343991" ID-Zenodo-Dep="5689898" LSID="urn:lsid:plazi:treatment:03FAA703ED384634FF5FF05BA035FC17" httpUri="http://treatment.plazi.org/id/03FAA703ED384634FF5FF05BA035FC17" lastPageId="5" lastPageNumber="320" pageId="1" pageNumber="316">
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<subSubSection id="C349459EED384630FF5FF05BA601FC8F" box="[151,580,859,886]" pageId="1" pageNumber="316" type="nomenclature">
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<paragraph id="8BEC1615ED384630FF5FF05BA601FC8F" blockId="1.[151,627,859,919]" box="[151,580,859,886]" pageId="1" pageNumber="316">
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<heading id="D0A4A179ED384630FF5FF05BA601FC8F" bold="true" box="[151,580,859,886]" fontSize="11" level="1" pageId="1" pageNumber="316" reason="1">
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<emphasis id="B927CA07ED384630FF5FF05BA601FC8F" bold="true" box="[151,580,859,886]" pageId="1" pageNumber="316">
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1.
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<taxonomicName id="4C536D96ED384630FF7CF05CA580FC8F" ID-CoL="9X3FH" box="[180,453,859,885]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="kameliyae" status="sp. nov.">
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<emphasis id="B927CA07ED384630FF7CF05CA580FC8F" bold="true" box="[180,453,859,885]" italics="true" pageId="1" pageNumber="316">Lyriothemis kameliyae</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A214777CED384630FE04F05BA601FC8F" box="[460,580,859,886]" pageId="1" pageNumber="316" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C349459EED384630FF5FF079A636FC6D" box="[151,627,894,919]" pageId="1" pageNumber="316" type="description">
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<paragraph id="8BEC1615ED384630FF5FF079A636FC6D" blockId="1.[151,627,859,919]" box="[151,627,894,919]" pageId="1" pageNumber="316">
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(
|
||
<figureCitation id="13680A90ED384630FF57F079A4ACFC6D" box="[159,233,894,919]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="1" pageNumber="316">Figs. 1</figureCitation>
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B, 1E, 2B–C, 2F, 3, 4A-B, 5C, 6B)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C349459EED384630FF5FF0C1A696FB95" pageId="1" pageNumber="316" type="materials_examined">
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<paragraph id="8BEC1615ED384630FF5FF0C1A696FB95" blockId="1.[151,1437,966,2035]" pageId="1" pageNumber="316">
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<materialsCitation id="3B3B1C48ED384630FF5FF0C1A7DEFBF9" ID-GBIF-Occurrence="1455704846" collectingDate="2015-06-29" collectionCode="RMNH" collectorName="T. Kompier" country="Vietnam" latitude="21.159" location="Type" longLatPrecision="75" longitude="104.926" municipality="Xuan Son National Park" pageId="1" pageNumber="316" specimenCount="2" stateProvince="Phu Tho Province" typeStatus="holotype">
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||
<emphasis id="B927CA07ED384630FF5FF0C1A513FC25" bold="true" box="[151,342,966,991]" pageId="1" pageNumber="316">
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<typeStatus id="54E8A8B7ED384630FF5FF0C1A497FC25" box="[151,210,966,991]" pageId="1" pageNumber="316">
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<location id="8E8C40CEED384630FF5FF0C1A497FC25" LSID="urn:lsid:plazi:treatment:03FAA703ED384634FF5FF05BA035FC17:8E8C40CEED384630FF5FF0C1A497FC25" box="[151,210,966,991]" country="Vietnam" latitude="21.159" longLatPrecision="75" longitude="104.926" municipality="Xuan Son National Park" name="Type" pageId="1" pageNumber="316" stateProvince="Phu Tho Province">Type</location>
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||
</typeStatus>
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||
specimens
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||
</emphasis>
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||
.
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||
<emphasis id="B927CA07ED384630FEA2F0C1A592FC25" bold="true" box="[362,471,966,991]" pageId="1" pageNumber="316">
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||
<typeStatus id="54E8A8B7ED384630FEA2F0C1A592FC25" box="[362,471,966,991]" pageId="1" pageNumber="316" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
male:
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<collectingMunicipality id="6B888C6FED384630FDE5F0C0A715FC25" box="[557,848,966,991]" pageId="1" pageNumber="316">Xuan Son National Park</collectingMunicipality>
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||
,
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||
<collectingRegion id="4997D8F7ED384630FCA9F0C1A07CFC25" box="[865,1081,966,991]" country="Vietnam" name="Phu Tho" pageId="1" pageNumber="316">
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||
Phu
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||
<collectingRegion id="4997D8F7ED384630FC50F0C1A07CFC25" box="[920,1081,966,991]" inferredBySuffix="true" pageId="1" pageNumber="316">Tho Province</collectingRegion>
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||
</collectingRegion>
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, northern
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||
<collectingCountry id="F3445685ED384630FB7BF0C1A157FC25" box="[1203,1298,966,991]" name="Vietnam" pageId="1" pageNumber="316">Vietnam</collectingCountry>
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(
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<geoCoordinate id="EE6770D2ED384630FAEEF0C1A1DDFC24" box="[1318,1432,966,991]" direction="north" orientation="latitude" pageId="1" pageNumber="316" precision="55" value="21.159">21.159 N</geoCoordinate>
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,
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<geoCoordinate id="EE6770D2ED384630FF5FF0EDA556FBF9" box="[151,275,1002,1027]" direction="east" orientation="longitude" pageId="1" pageNumber="316" precision="55" value="104.926">104.926 E</geoCoordinate>
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),
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||
<date id="FFED30D5ED384630FEE6F0EDA5F4FBF9" box="[302,433,1002,1027]" pageId="1" pageNumber="316" value="2015-06-29">
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||
<collectingDate id="EFA9C93DED384630FEE6F0EDA5F4FBF9" box="[302,433,1002,1027]" pageId="1" pageNumber="316" value="2015-06-29">29.VI.2015</collectingDate>
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||
</date>
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||
,
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<collectorName id="26A673C3ED384630FE0BF0ECA60FFBF9" box="[451,586,1002,1027]" pageId="1" pageNumber="316">T. Kompier</collectorName>
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leg, (deposited at
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||
<collectionCode id="ED428ED0ED384630FCFBF0ECA7CAFBF9" box="[819,911,1003,1027]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="316">RMNH</collectionCode>
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||
).
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B3B1C48ED384630FC64F0EDA4B8FBDD" ID-GBIF-Occurrence="1455704844" collectingDate="2015-06-29" collectionCode="RMNH" collectorName="T. Kompier" country="Vietnam" latitude="21.159" location="Type" longLatPrecision="75" longitude="104.926" municipality="Xuan Son National Park" pageId="1" pageNumber="316" specimenCount="1" stateProvince="Phu Tho Province" typeStatus="allotype">
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<emphasis id="B927CA07ED384630FC64F0EDA054FBF9" bold="true" box="[940,1041,1002,1027]" pageId="1" pageNumber="316">
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||
<typeStatus id="54E8A8B7ED384630FC64F0EDA054FBF9" box="[940,1041,1002,1027]" pageId="1" pageNumber="316" type="allotype">Allotype</typeStatus>
|
||
</emphasis>
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||
female: same location, date and collector
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B3B1C48ED384630FEC5F709A03AFBDD" ID-GBIF-Occurrence="1455704848" box="[269,1151,1038,1063]" collectingDate="2016-08-01" collectionCode="RMNH" collectorName="T. Kompier" country="Vietnam" latitude="21.159" location="Type" longLatPrecision="75" longitude="104.926" municipality="Xuan Son National Park" pageId="1" pageNumber="316" specimenCount="1" specimenCount-male="1" stateProvince="Phu Tho Province" typeStatus="paratype">
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||
<emphasis id="B927CA07ED384630FEC5F709A53FFBDD" bold="true" box="[269,378,1038,1063]" pageId="1" pageNumber="316">
|
||
<typeStatus id="54E8A8B7ED384630FEC5F709A53FFBDD" box="[269,378,1038,1063]" pageId="1" pageNumber="316" type="paratype">Paratype</typeStatus>
|
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</emphasis>
|
||
<specimenCount id="9D55DD9CED384630FE4EF708A5E9FBDD" box="[390,428,1039,1063]" pageId="1" pageNumber="316" type="male">1♂</specimenCount>
|
||
: same location and collector as
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||
<typeStatus id="54E8A8B7ED384630FCF9F709A7D6FBDD" box="[817,915,1038,1063]" pageId="1" pageNumber="316" type="holotype">holotype</typeStatus>
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||
male,
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||
<date id="FFED30D5ED384630FC21F709A03AFBDD" box="[1001,1151,1038,1063]" pageId="1" pageNumber="316" value="2016-08-01">
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||
<collectingDate id="EFA9C93DED384630FC21F709A03AFBDD" box="[1001,1151,1038,1063]" pageId="1" pageNumber="316" value="2016-08-01">01.VIII.2016</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B3B1C48ED384630FB58F708A635FBB1" ID-GBIF-Occurrence="1455704847" collectingDate="2016-07-31" collectionCode="RMNH" collectorName="T. Kompier" country="Vietnam" latitude="21.159" location="Type" longLatPrecision="75" longitude="104.926" municipality="Xuan Son National Park" pageId="1" pageNumber="316" specimenCount="1" specimenCount-female="1" stateProvince="Phu Tho Province" typeStatus="allotype">
|
||
<specimenCount id="9D55DD9CED384630FB58F708A0F0FBDD" box="[1168,1205,1039,1063]" pageId="1" pageNumber="316" type="female">1♀</specimenCount>
|
||
: same location and collector as
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||
<typeStatus id="54E8A8B7ED384630FEEBF735A538FBB1" box="[291,381,1074,1099]" pageId="1" pageNumber="316" type="allotype">allotype</typeStatus>
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||
female,
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||
<date id="FFED30D5ED384630FE2CF735A635FBB1" box="[484,624,1074,1099]" pageId="1" pageNumber="316" value="2016-07-31">
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||
<collectingDate id="EFA9C93DED384630FE2CF735A635FBB1" box="[484,624,1074,1099]" pageId="1" pageNumber="316" value="2016-07-31">31.VII.2016</collectingDate>
|
||
</date>
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B3B1C48ED384630FDB6F734A696FB95" ID-GBIF-Occurrence="1455704845" collectingDate="2014-05-24" collectorName="T. Kompier" country="Vietnam" latitude="21.708" location="Huu Lien Nature Reserve" longLatPrecision="75" longitude="106.374" pageId="1" pageNumber="316" specimenCount="1" specimenCount-male="1" stateProvince="Lang Son Province" typeStatus="allotype">
|
||
<specimenCount id="9D55DD9CED384630FDB6F734A6E6FBB1" box="[638,675,1075,1099]" pageId="1" pageNumber="316" type="male">1♂</specimenCount>
|
||
:
|
||
<location id="8E8C40CEED384630FD7AF734A79EFBB0" LSID="urn:lsid:plazi:treatment:03FAA703ED384634FF5FF05BA035FC17:8E8C40CEED384630FD7AF734A79EFBB0" box="[690,987,1074,1099]" country="Vietnam" latitude="21.708" longLatPrecision="75" longitude="106.374" name="Huu Lien Nature Reserve" pageId="1" pageNumber="316" stateProvince="Lang Son Province">Huu Lien Nature Reserve</location>
|
||
,
|
||
<collectingRegion id="4997D8F7ED384630FC21F734A082FBB1" box="[1001,1223,1074,1099]" country="Vietnam" name="Lang Son" pageId="1" pageNumber="316">
|
||
Lang
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||
<collectingRegion id="4997D8F7ED384630FBE4F734A082FBB1" box="[1068,1223,1074,1099]" inferredBySuffix="true" pageId="1" pageNumber="316">Son Province</collectingRegion>
|
||
</collectingRegion>
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||
, northern
|
||
<collectingCountry id="F3445685ED384630FAF5F735A1D9FBB1" box="[1341,1436,1074,1099]" name="Vietnam" pageId="1" pageNumber="316">Vietnam</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE6770D2ED384630FF56F750A54EFB94" box="[158,267,1111,1135]" direction="north" orientation="latitude" pageId="1" pageNumber="316" precision="55" value="21.708">21.708 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE6770D2ED384630FEDEF751A5C9FB95" box="[278,396,1110,1135]" direction="east" orientation="longitude" pageId="1" pageNumber="316" precision="55" value="106.374">106.374 E</geoCoordinate>
|
||
),
|
||
<date id="FFED30D5ED384630FE69F750A653FB95" box="[417,534,1111,1135]" pageId="1" pageNumber="316" value="2014-05-24">
|
||
<collectingDate id="EFA9C93DED384630FE69F750A653FB95" box="[417,534,1111,1135]" pageId="1" pageNumber="316" value="2014-05-24">24.V.2014</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26A673C3ED384630FDE9F750A6E7FB95" box="[545,674,1110,1135]" pageId="1" pageNumber="316">T. Kompier</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349459EED384630FF0FF77DA778FB20" pageId="1" pageNumber="316" type="etymology">
|
||
<paragraph id="8BEC1615ED384630FF0FF77DA778FB20" blockId="1.[151,1437,966,2035]" pageId="1" pageNumber="316">
|
||
<emphasis id="B927CA07ED384630FF0FF77DA503FB69" bold="true" box="[199,326,1146,1171]" pageId="1" pageNumber="316">Etymology</emphasis>
|
||
. The species is named after Kameliya Petrova in gratitude for her assistance with my book “Dragonflies and Damselflies of the Serra dos Orgaos” and unrelenting support for my continuing research on
|
||
<taxonomicName id="4C536D96ED384630FF5FF7C5A4B3FB21" box="[151,246,1218,1243]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="316" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
|
||
in
|
||
<collectingCountry id="F3445685ED384630FED1F7C5A539FB21" box="[281,380,1218,1243]" name="Vietnam" pageId="1" pageNumber="316">Vietnam</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C536D96ED384630FE4EF7C4A5BBFB21" box="[390,510,1219,1243]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="326" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED384630FE4EF7C4A5BBFB21" box="[390,510,1219,1243]" italics="true" pageId="1" pageNumber="316">Kameliyae</emphasis>
|
||
</taxonomicName>
|
||
, a noun in the genitive case.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349459EED384634FF0FF7E1A035FC17" lastPageId="5" lastPageNumber="320" pageId="1" pageNumber="316" type="description">
|
||
<paragraph id="8BEC1615ED384630FF0FF7E1A027FABD" blockId="1.[151,1437,966,2035]" pageId="1" pageNumber="316">
|
||
<emphasis id="B927CA07ED384630FF0FF7E1A65BFB05" bold="true" box="[199,542,1254,1279]" pageId="1" pageNumber="316">
|
||
Description of male
|
||
<typeStatus id="54E8A8B7ED384630FE70F7E1A65BFB05" box="[440,542,1254,1279]" pageId="1" pageNumber="316" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. (
|
||
<figureCitation id="13680A90ED384630FDFDF7E1A630FB05" box="[565,629,1254,1279]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="1" pageNumber="316">Fig. 1</figureCitation>
|
||
B)
|
||
<emphasis id="B927CA07ED384630FD53F7E1A69DFB04" box="[667,728,1254,1278]" italics="true" pageId="1" pageNumber="316">Head</emphasis>
|
||
. (cf.
|
||
<figureCitation id="13680A90ED384630FCDBF7E1A70EFB05" box="[787,843,1254,1279]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="1" pageNumber="316">Fig 1</figureCitation>
|
||
F). Labium black. Labrum, anteclypeus, postclypeus and antefrons white, postfrons metallic dark green, squarish. Vertex metallic dark green, with two small tubercles, antennae black, occiput black. Compound eyes blackish grey, chestnut brown dorsally.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED384630FF0FF654A5D5FA91" blockId="1.[151,1437,966,2035]" box="[199,400,1362,1387]" pageId="1" pageNumber="316">
|
||
<emphasis id="B927CA07ED384630FF0FF654A57CFA91" box="[199,313,1363,1387]" italics="true" pageId="1" pageNumber="316">Prothorax</emphasis>
|
||
. Black.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED384630FF0FF671A0CEF955" blockId="1.[151,1437,966,2035]" pageId="1" pageNumber="316">
|
||
<emphasis id="B927CA07ED384630FF0FF671A505FA74" box="[199,320,1398,1422]" italics="true" pageId="1" pageNumber="316">Synthorax.</emphasis>
|
||
(
|
||
<figureCitation id="13680A90ED384630FE98F671A5D5FA75" box="[336,400,1398,1423]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="1" pageNumber="316">Fig. 1</figureCitation>
|
||
B) Blackish brown with markings as follows: a vague brown dorsal stripe on either side of dorsal carina, mesepimeron with a broad oblong yellow stripe, and metepimeron almost completely yellow.
|
||
<emphasis id="B927CA07ED384630FAAFF69CA1D9FA48" box="[1383,1436,1435,1458]" italics="true" pageId="1" pageNumber="316">Legs</emphasis>
|
||
black. Femora with short spines growing progressively longer distally, tibiae with longer black spines.
|
||
<emphasis id="B927CA07ED384630FAD2F6B8A11BFA2C" box="[1306,1374,1471,1494]" italics="true" pageId="1" pageNumber="316">Wings</emphasis>
|
||
(
|
||
<figureCitation id="13680A90ED384630FAA6F6B9A4E7FA01" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="1" pageNumber="316">Fig. 1</figureCitation>
|
||
B) hyaline. Two brown streaks above and below median space in both fore- and hindwing as follows: first two cells between subcosta and radius blackish brown, third cell tinted brown and cubito-anal space blackish brown until first crossvein. Veins black. Membranule brown. Anal loop 20–21 celled. 2 bridge crossveins in all wings, last antenodal crossvein complete. Forewing with 1 cubito-anal crossvein, triangle 2-celled, subtriangle 3-celled. Hindwing with 2 cubito-anal crossveins, triangle 2-celled. Nodal index in forewing 12–17:18–11 and in hindwing 12–14:14–13. Pterostigma black, covering 2.5 cells in all wings, 4 times longer than wide.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED384630FF0FF5BCA76EF809" blockId="1.[151,1437,966,2035]" pageId="1" pageNumber="316">
|
||
<emphasis id="B927CA07ED384630FF0FF5BCA57DF929" box="[199,312,1723,1747]" italics="true" pageId="1" pageNumber="316">Abdomen.</emphasis>
|
||
(
|
||
<figureCitation id="13680A90ED384630FE8FF5BDA5C2F929" box="[327,391,1722,1747]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="1" pageNumber="316">Fig. 1</figureCitation>
|
||
B) As broad as thorax in dorsal view, S3 broadest. S1 yellow with black dorsum, S2 dorsally bright red to somewhat above lateral carina, lower part yellow with some blackish blotches, lateral carina black. Hamulus (cf.
|
||
<figureCitation id="13680A90ED384630FEF3F405A5C7F8E1" box="[315,386,1794,1819]" captionStart="FIGURE 5" captionStartId="6.[151,250,1662,1684]" captionTargetBox="[159,1412,1204,1644]" captionTargetId="figure@6.[159,1412,1204,1645]" captionTargetPageId="6" captionText="FIGURE 5. Hamuli in anteroventral view: (A) L. tricolor, Taiwan, courtesy Wen-Chi Yeh; (B) L. bivittata, Huu Lien NR, 23. VII. 2016; (C) L. kameliyae, paratype, Huu Lien NR, 24. V. 2014." httpUri="https://zenodo.org/record/495283/files/figure.png" pageId="1" pageNumber="316">Fig. 5</figureCitation>
|
||
C) black, large, shaped like a rectangular dish, with a backward-directed spine on inner posterior corner and a large v-shaped incision approximately in middle of inner edge. Pale yellow genital lobe very slender, pointed and about 4 times as long as wide. S3 dorsally bright red to black lateral carina, dorsal carina black, ventrally yellow. S4–7 bright red dorsally, with black lateral, dorsal and posterior carina, ventrally yellow. S8 bright red, with broad blackish borders along dorsal and posterior carina. Lateral carina and venter black. S9–10 black.
|
||
<emphasis id="B927CA07ED384630FF2CF4B1A5ECF834" box="[228,425,1974,1998]" italics="true" pageId="1" pageNumber="316">Anal appendages</emphasis>
|
||
. Cerci black, twice length of S10, extending just beyond epiproct, pointed, subapically with 5 small ventral teeth. Epiproct of normal
|
||
<typeStatus id="54E8A8B7ED384630FD56F4DCA694F809" box="[670,721,2011,2035]" pageId="1" pageNumber="316">type</typeStatus>
|
||
, brown.
|
||
</paragraph>
|
||
<caption id="DF2C469DED3B4633FF5FF496A7D4F81F" httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="2" pageNumber="317" targetBox="[216,1358,229,1915]" targetPageId="2">
|
||
<paragraph id="8BEC1615ED3B4633FF5FF496A7D4F81F" blockId="2.[151,1437,1937,2021]" pageId="2" pageNumber="317">
|
||
<emphasis id="B927CA07ED3B4633FF5FF496A550F85C" bold="true" box="[151,277,1937,1959]" pageId="2" pageNumber="317">FIGURE 1.</emphasis>
|
||
Lateral scans: (A) ♂
|
||
<taxonomicName id="4C536D96ED3B4633FE31F496A77BF85D" authority="Huu Lien" authorityName="Huu Lien" box="[505,830,1937,1959]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="317" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3B4633FE31F496A688F85D" box="[505,717,1937,1959]" italics="true" pageId="2" pageNumber="317">Lyriothemis bivittata</emphasis>
|
||
, Huu Lien
|
||
</taxonomicName>
|
||
NR, 23.VII.2016; (B) ♂
|
||
<taxonomicName id="4C536D96ED3B4633FB81F495A08EF85D" box="[1097,1227,1937,1959]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="317" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED3B4633FB81F495A08EF85D" box="[1097,1227,1937,1959]" italics="true" pageId="2" pageNumber="317">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
, holotype; (C) ♂
|
||
<emphasis id="B927CA07ED3B4633FA4FF495A4ADF83C" italics="true" pageId="2" pageNumber="317">
|
||
<taxonomicName id="4C536D96ED3B4633FA4FF495A4A1F83C" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="317" phylum="Arthropoda" rank="species" species="tricolor">L. tricolor</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
Taiwan (courtesy of Wen-Chi Yeh); (D) ♀
|
||
<taxonomicName id="4C536D96ED3B4633FD55F4B6A739F83C" authority="Xuan Son" authorityName="Xuan Son" box="[669,892,1968,1990]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="317" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3B4633FD55F4B6A757F83C" box="[669,786,1968,1990]" italics="true" pageId="2" pageNumber="317">L. bivittata,</emphasis>
|
||
Xuan Son
|
||
</taxonomicName>
|
||
NP, 31.VII.2016; (E) ♀
|
||
<taxonomicName id="4C536D96ED3B4633FBBCF4B6A0B0F83C" box="[1140,1269,1968,1990]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="317" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED3B4633FBBCF4B6A0B0F83C" box="[1140,1269,1968,1990]" italics="true" pageId="2" pageNumber="317">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Xuan Son NP, 31.VII.2016; (F) ♀
|
||
<emphasis id="B927CA07ED3B4633FE7EF4D7A667F81F" box="[438,546,1999,2021]" italics="true" pageId="2" pageNumber="317">
|
||
<taxonomicName id="4C536D96ED3B4633FE7EF4D7A65BF81F" box="[438,542,1999,2021]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="317" phylum="Arthropoda" rank="species" species="tricolor">L. tricolor</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
Taiwan (courtesy of Wen-Chi Yeh).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEC1615ED3A4632FF0FF39FA0C0FF4A" blockId="3.[151,1437,151,608]" box="[199,1157,151,176]" pageId="3" pageNumber="318">Measurements (mm): Abdomen incl. appendages 26.5, hindwing 35, pterostigma 3.5.</paragraph>
|
||
<paragraph id="8BEC1615ED3A4632FF0FF3BBA670FE57" blockId="3.[151,1437,151,608]" pageId="3" pageNumber="318">
|
||
<emphasis id="B927CA07ED3A4632FF0FF3BBA593FF2F" bold="true" box="[199,470,188,213]" pageId="3" pageNumber="318">
|
||
Variation in
|
||
<typeStatus id="54E8A8B7ED3A4632FEA8F3BBA593FF2F" box="[352,470,188,213]" pageId="3" pageNumber="318" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
. Male from Huu Lien (
|
||
<figureCitation id="13680A90ED3A4632FD2EF3BBA773FF2E" box="[742,822,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1818,1840]" captionTargetBox="[185,1417,665,1777]" captionTargetId="figure@3.[174,1417,665,1780]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal scans: (A) ♀ Lyriothemis bivittata, Xuan Son NP, 31. VII. 2016; (B) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (C) ♂ L. kameliyae, paratype, Huu Lien NR, 24. V. 2014. Photos in natural habitat: (D) ♀ L. bivittata (not collected), Tam Dao NP, 18. VIII. 2013; (E) ♀ L. sp., Quang Binh Province, 20. V. 2016; (F) ♂ (not collected) L. kameliyae, Huu Lien NR, 21. IV. 2014." httpUri="https://zenodo.org/record/495280/files/figure.png" pageId="3" pageNumber="318">Figs. 2</figureCitation>
|
||
C, 3D, 5C) with much reduced yellow stripe (only on ventral half) on mesepimeron. S1–2 with extensive black on ventral half, including genital lobe. One cercus with only 4 small ventral teeth. Nodal index in forewing 10–19:17–10 and in hindwing 10–14:13–9, one forewing with 3 bridge crossveins. Anal loop 19–20 celled. Male from Xuan Son (
|
||
<figureCitation id="13680A90ED3A4632FC59F220A796FEBA" box="[913,979,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,993,1015]" captionTargetBox="[148,1429,197,954]" captionTargetId="figure@4.[148,1430,197,954]" captionTargetPageId="4" captionText="FIGURE 3. Variation in ♂ L. kameliyae: (A) Paratype, Xuan Son NP, 1. VIII. 2016; (B) Huu Lien NR, 21. IV. 2014 (not collected); (C) Xuan Son NP, 18. IV. 2015 (not collected); (D) Paratype, Huu Lien NR, 24. V. 2014." httpUri="https://zenodo.org/record/495281/files/figure.png" pageId="3" pageNumber="318">Fig. 3</figureCitation>
|
||
A) with somewhat reduced yellow stripe on mesepimeron, constricted dorsally. Wings lightly tinted amber; brown streaks at base not extending past 2nd cell between subcosta and radius. Nodal index in forewing 9–19:18–10, in hindwing 10–14:
|
||
<date id="FFED30D5ED3A4632FBB2F277A095FE72" box="[1146,1232,368,392]" pageId="3" pageNumber="318" value="2014-04-09">14–9. 4</date>
|
||
bridge crossveins in all wings. Anal loop 20–24 celled.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED3A4632FF0FF2BFA0A5FE2A" blockId="3.[151,1437,151,608]" box="[199,1248,439,464]" pageId="3" pageNumber="318">Measurements (mm): Abdomen incl. appendages 28–29, hindwing 33–34, pterostigma 3–3.5.</paragraph>
|
||
<paragraph id="8BEC1615ED3A4632FF0FF2DBA725FD9A" blockId="3.[151,1437,151,608]" pageId="3" pageNumber="318">
|
||
<emphasis id="B927CA07ED3A4632FF0FF2DBA66EFE0F" bold="true" box="[199,555,476,501]" pageId="3" pageNumber="318">
|
||
Description of female
|
||
<typeStatus id="54E8A8B7ED3A4632FE04F2DBA66EFE0F" box="[460,555,476,501]" pageId="3" pageNumber="318" type="allotype">allotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13680A90ED3A4632FDF1F2DBA6CDFE0E" box="[569,648,476,501]" captionStart="FIGURE 4" captionStartId="4.[151,250,1902,1924]" captionTargetBox="[154,1418,1098,1857]" captionTargetId="figure@4.[154,1418,1098,1857]" captionTargetPageId="4" captionText="FIGURE 4. Lyriothemis ♀: (A) L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (B) idem, allotype; (C) L. bivittata, Xuan Son NP, 31. VII. 2016; (D) L. sp., Quang Binh Province, 20. V. 2016." httpUri="https://zenodo.org/record/495282/files/figure.png" pageId="3" pageNumber="318">Figs. 4</figureCitation>
|
||
B, 6B).
|
||
<emphasis id="B927CA07ED3A4632FD16F2DBA75EFE0E" box="[734,795,476,500]" italics="true" pageId="3" pageNumber="318">Head</emphasis>
|
||
. Labium and labrum black. Anteclypeus brownish white, postclypeus dark brown with off-white dorsolateral corners. Antefrons brownish white in middle, off-white towards sides, postfrons metallic dark green, rounded. Vertex metallic dark green, with two small tubercles, antennae black, occiput black. Compound eyes blackish brown.
|
||
</paragraph>
|
||
<caption id="DF2C469DED3A4632FF5FF41DA537F876" httpUri="https://zenodo.org/record/495280/files/figure.png" pageId="3" pageNumber="318" targetBox="[185,1417,665,1777]" targetPageId="3">
|
||
<paragraph id="8BEC1615ED3A4632FF5FF41DA537F876" blockId="3.[151,1436,1818,1932]" pageId="3" pageNumber="318">
|
||
<emphasis id="B927CA07ED3A4632FF5FF41DA54BF8D5" bold="true" box="[151,270,1818,1840]" pageId="3" pageNumber="318">FIGURE 2</emphasis>
|
||
. Dorsal scans: (A) ♀
|
||
<taxonomicName id="4C536D96ED3A4632FE3BF41DA779F8CA" authority="Xuan Son" authorityName="Xuan Son" box="[499,828,1818,1840]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="318" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3A4632FE3BF41DA68BF8CA" box="[499,718,1818,1840]" italics="true" pageId="3" pageNumber="318">Lyriothemis bivittata,</emphasis>
|
||
Xuan Son
|
||
</taxonomicName>
|
||
NP, 31.VII.2016; (B) ♀
|
||
<taxonomicName id="4C536D96ED3A4632FBF7F41CA084F8CA" box="[1087,1217,1818,1840]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="318" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED3A4632FBF7F41CA084F8CA" box="[1087,1217,1818,1840]" italics="true" pageId="3" pageNumber="318">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Xuan Son NP, 31.VII.2016; (C) ♂
|
||
<taxonomicName id="4C536D96ED3A4632FE5DF43DA65DF8B5" box="[405,536,1849,1871]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="318" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED3A4632FE5DF43DA65DF8B5" box="[405,536,1849,1871]" italics="true" pageId="3" pageNumber="318">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Huu Lien NR, 24.V.2014. Photos in natural habitat: (D) ♀
|
||
<taxonomicName id="4C536D96ED3A4632FB31F43DA12EF8B5" box="[1273,1387,1849,1871]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="318" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3A4632FB31F43DA12EF8B5" box="[1273,1387,1849,1871]" italics="true" pageId="3" pageNumber="318">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
(not collected), Tam Dao NP, 18.VIII.2013; (E) ♀
|
||
<emphasis id="B927CA07ED3A4632FDAFF45EA6E5F894" box="[615,672,1881,1902]" italics="true" pageId="3" pageNumber="318">L. sp.</emphasis>
|
||
, Quang Binh Province, 20.V.2016; (F) ♂ (not collected)
|
||
<taxonomicName id="4C536D96ED3A4632FB2AF45EA480F876" authority="Huu Lien" authorityName="Huu Lien" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="318" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED3A4632FB2AF45EA121F894" box="[1250,1380,1880,1902]" italics="true" pageId="3" pageNumber="318">L. kameliyae</emphasis>
|
||
, Huu Lien
|
||
</taxonomicName>
|
||
NR, 21.IV.2014.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2C469DED3D4635FF5FF0E6A02AFBEF" httpUri="https://zenodo.org/record/495281/files/figure.png" pageId="4" pageNumber="319" targetBox="[148,1429,197,954]" targetPageId="4">
|
||
<paragraph id="8BEC1615ED3D4635FF5FF0E6A02AFBEF" blockId="4.[151,1436,993,1045]" pageId="4" pageNumber="319">
|
||
<emphasis id="B927CA07ED3D4635FF5FF0E6A552FC0C" bold="true" box="[151,279,993,1015]" pageId="4" pageNumber="319">FIGURE 3.</emphasis>
|
||
Variation in ♂
|
||
<emphasis id="B927CA07ED3D4635FE0FF0E5A613FC0D" box="[455,598,993,1015]" italics="true" pageId="4" pageNumber="319">
|
||
<taxonomicName id="4C536D96ED3D4635FE0FF0E5A614FC0D" box="[455,593,993,1015]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="kameliyae">L. kameliyae</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
(A) Paratype, Xuan Son NP, 1.VIII.2016; (B) Huu Lien NR, 21.IV.2014 (not collected); (C) Xuan Son NP, 18.IV.2015 (not collected); (D) Paratype, Huu Lien NR, 24.V.2014.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2C469DED3D4635FF5FF469A779F859" httpUri="https://zenodo.org/record/495282/files/figure.png" pageId="4" pageNumber="319" targetBox="[154,1418,1098,1857]" targetPageId="4">
|
||
<paragraph id="8BEC1615ED3D4635FF5FF469A779F859" blockId="4.[151,1436,1902,1955]" pageId="4" pageNumber="319">
|
||
<emphasis id="B927CA07ED3D4635FF5FF469A556F87E" bold="true" box="[151,275,1902,1924]" pageId="4" pageNumber="319">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C536D96ED3D4635FED2F469A5D7F87E" box="[282,402,1902,1924]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="319" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B927CA07ED3D4635FED2F469A5D7F87E" box="[282,402,1902,1924]" italics="true" pageId="4" pageNumber="319">Lyriothemis</emphasis>
|
||
</taxonomicName>
|
||
♀: (A)
|
||
<taxonomicName id="4C536D96ED3D4635FE2CF477A623F87E" box="[484,614,1902,1925]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED3D4635FE2CF477A623F87E" box="[484,614,1902,1925]" italics="true" pageId="4" pageNumber="319">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Xuan Son NP, 31.VII.2016; (B) idem, allotype; (C)
|
||
<taxonomicName id="4C536D96ED3D4635FB20F477A4FBF859" authority="Xuan Son" authorityName="Xuan Son" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3D4635FB20F477A11BF87E" box="[1256,1374,1902,1925]" italics="true" pageId="4" pageNumber="319">L. bivittata,</emphasis>
|
||
Xuan Son
|
||
</taxonomicName>
|
||
NP, 31.VII.2016; (D)
|
||
<emphasis id="B927CA07ED3D4635FE69F489A59CF859" box="[417,473,1934,1955]" italics="true" pageId="4" pageNumber="319">L. sp.</emphasis>
|
||
, Quang Binh Province, 20.V.2016.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEC1615ED3C4634FF0FF39FA7F4FF4A" blockId="5.[151,1437,151,1005]" box="[199,945,151,176]" pageId="5" pageNumber="320">
|
||
<emphasis id="B927CA07ED3C4634FF0FF39FA57CFF4A" box="[199,313,152,176]" italics="true" pageId="5" pageNumber="320">Prothorax</emphasis>
|
||
. Black, but anterior lobe with anterior edge pale yellow.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED3C4634FF0FF3BBA03AFDE2" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="320">
|
||
<emphasis id="B927CA07ED3C4634FF0FF3BBA505FF2E" box="[199,320,188,212]" italics="true" pageId="5" pageNumber="320">Synthorax.</emphasis>
|
||
(
|
||
<figureCitation id="13680A90ED3C4634FE99F3BBA5D3FF2E" box="[337,406,188,213]" captionStart="FIGURE 4" captionStartId="4.[151,250,1902,1924]" captionTargetBox="[154,1418,1098,1857]" captionTargetId="figure@4.[154,1418,1098,1857]" captionTargetPageId="4" captionText="FIGURE 4. Lyriothemis ♀: (A) L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (B) idem, allotype; (C) L. bivittata, Xuan Son NP, 31. VII. 2016; (D) L. sp., Quang Binh Province, 20. V. 2016." httpUri="https://zenodo.org/record/495282/files/figure.png" pageId="5" pageNumber="320">Fig. 4</figureCitation>
|
||
B) Blackish brown with markings as follows: indistinct pale brown dorsal stripe on either side of dorsal carina, mesepimeron with a broad oblong pale yellow stripe, somewhat narrower in dorsal half, metepimeron almost completely pale yellow.
|
||
<emphasis id="B927CA07ED3C4634FD55F202A697FEE6" box="[669,722,261,284]" italics="true" pageId="5" pageNumber="320">Legs</emphasis>
|
||
black. Femora with short spines growing progressively longer distally, tibiae with longer black spines.
|
||
<emphasis id="B927CA07ED3C4634FD90F22FA6DEFEC5" box="[600,667,296,319]" italics="true" pageId="5" pageNumber="320">Wings</emphasis>
|
||
(cf.
|
||
<figureCitation id="13680A90ED3C4634FD18F220A75DFEBA" box="[720,792,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="5" pageNumber="320">Figs. 1</figureCitation>
|
||
E, 2B) hyaline, slightly tinted amber, apices darkest. Two brown streaks above and below median space in both fore- and hindwing as follows: first three cells between subcosta and radius blackish brown, fourth cell tinted brown, cubito-anal space blackish brown until first crossvein. Veins black. Membranule brown. Anal loop 23 celled. 2 bridge crossveins in left wings,
|
||
<quantity id="4CABBBF0ED3C4634FB69F293A08AFE57" box="[1185,1231,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="320" unit="in" value="3.0">3 in</quantity>
|
||
right wings. Last antenodal crossvein complete. Forewing with 1 cubito-anal crossvein, triangle 2-celled, subtriangle 3-celled. Hindwing with 2 cubito-anal crossveins, triangle 2-celled. Nodal index in forewing 11–18:19–11 and in hindwing 12–14:15–11. Pterostigma brown, covering 3 cells in all wings, 4 times longer than wide.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED3C4634FF0FF123A134FD0A" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="320">
|
||
<emphasis id="B927CA07ED3C4634FF0FF123A57DFDC6" box="[199,312,548,572]" italics="true" pageId="5" pageNumber="320">Abdomen.</emphasis>
|
||
(
|
||
<figureCitation id="13680A90ED3C4634FE80F123A5C8FDC6" box="[328,397,548,573]" captionStart="FIGURE 4" captionStartId="4.[151,250,1902,1924]" captionTargetBox="[154,1418,1098,1857]" captionTargetId="figure@4.[154,1418,1098,1857]" captionTargetPageId="4" captionText="FIGURE 4. Lyriothemis ♀: (A) L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (B) idem, allotype; (C) L. bivittata, Xuan Son NP, 31. VII. 2016; (D) L. sp., Quang Binh Province, 20. V. 2016." httpUri="https://zenodo.org/record/495282/files/figure.png" pageId="5" pageNumber="320">Fig. 4</figureCitation>
|
||
B) S1 blackish brown, yellow towards venter. S2 pale yellow, with black carinas and broad black line running parallel to the dorsal carina halfway down towards the lateral carina. S3–6 pale orange with black carinas, a broader black line over dorsal carina and another black line parallel to it halfway down towards lateral carina. Ventrally yellow. S7 with more extensive black, but ventrally yellow. S8 virtually black with only a tiny pale orange spot towards anterior margin and ventrally black, S9–10 and conical cerci all black, cerci slightly longer than S10. S8 somewhat dilated.
|
||
<emphasis id="B927CA07ED3C4634FD85F1DFA6A9FD0A" box="[589,748,728,752]" italics="true" pageId="5" pageNumber="320">Vulvar lamina</emphasis>
|
||
(
|
||
<figureCitation id="13680A90ED3C4634FD34F1D0A77BFD0A" box="[764,830,727,752]" captionStart="FIGURE 6" captionStartId="8.[151,250,737,759]" captionTargetBox="[165,1417,196,718]" captionTargetId="figure@8.[165,1417,196,718]" captionTargetPageId="8" captionText="FIGURE 6. Vulvar lamina: (A) L. bivittata, Xuan Son NP, 31. VII. 2016; (B) L. kameliyae allotype." httpUri="https://zenodo.org/record/495284/files/figure.png" pageId="5" pageNumber="320">Fig. 6</figureCitation>
|
||
B) short, slightly curled, relatively widely incised.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED3C4634FF0FF1FAA02AFCEE" blockId="5.[151,1437,151,1005]" box="[199,1135,764,789]" pageId="5" pageNumber="320">Measurements (mm): Abdomen incl. appendages 27.5, hindwing 37, pterostigma 4.</paragraph>
|
||
<paragraph id="8BEC1615ED3C4634FF0FF018A05AFC32" blockId="5.[151,1437,151,1005]" pageId="5" pageNumber="320">
|
||
<emphasis id="B927CA07ED3C4634FF0FF018A58DFCC2" bold="true" box="[199,456,799,824]" pageId="5" pageNumber="320">
|
||
Variation in
|
||
<typeStatus id="54E8A8B7ED3C4634FE94F018A58DFCC2" box="[348,456,799,824]" pageId="5" pageNumber="320" type="paratype">paratype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13680A90ED3C4634FE1FF018A664FCC2" box="[471,545,799,824]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="5" pageNumber="320">Figs. 1</figureCitation>
|
||
E, 2B, 4A). Dorsal stripes on synthorax paler and more distinct. Abdomen with reduced blackish striping, S2–7 brownish orange dorsally, carinas only thinly black. S8–10 and cerci completely black. Streaks at wing bases reduced, only covering first 2 cells between subcosta and radius and only half of first cell in cubito-anal space. Wings only slightly tinted amber. 3–5 bridge crossveins, anal loop 25–26 celled. 12– 18:18–12 ante- and postnodal crossveins in forewing, 12–12:12–
|
||
<quantity id="4CABBBF0ED3C4634FCA5F0B7A7E3FC32" box="[877,934,943,968]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="5" pageNumber="320" unit="in" value="12.0">12 in</quantity>
|
||
hindwing.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED3C4634FF0FF0D2A035FC17" blockId="5.[151,1437,151,1005]" box="[199,1136,980,1005]" pageId="5" pageNumber="320">Measurements (mm): Abdomen incl. appendages 27, hindwing 36, pterostigma 3.5.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |