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<document id="5FD8265B5EA6458EBD44CF55EC0489E2" ID-DOI="10.11646/zootaxa.5343.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8324250" ID-ZooBank="20BF9AB4-F86D-42EA-AEAA-7C03EBA44EB6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1694069790509" checkinUser="plazi" docAuthor="Yamasaki, Takeshi, Hashimoto, Yoshiaki, Endo, Tomoji, Hyodo, Fujio, Itioka, Takao, Mohamed, Maryati &amp; Meleng, Paulus" docDate="2023" docId="03FA87E7FFBCFFD6FF7D3E82FDCE8A93" docLanguage="en" docName="zootaxa.5343.1.3.pdf" docOrigin="Zootaxa 5343 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5343.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Utivarachna fukasawana Kishida 1940" docType="treatment" docVersion="2" lastPageNumber="68" masterDocId="FFC3FF9FFFB7FFDBFFEA3E15FFEC8816" masterDocTitle="Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species" masterLastPageNumber="73" masterPageNumber="55" pageNumber="66" updateTime="1694071302809" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="36BDE1680A3B888C1C90BF0D6B20844B">Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species</mods:title>
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<mods:affiliation id="9697DEF118E740208D743282AB3DD516">Institute of Natural and Environmental Sciences, University of Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan &amp; Museum of Nature and Human Activities, Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan &amp; k 0468874 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2419 - 188 X</mods:affiliation>
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<mods:namePart id="8751F80EBDDD68CD3D04E47A17A08BA9">Hashimoto, Yoshiaki</mods:namePart>
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<mods:affiliation id="C301C9FBD6DEB33B874291896E5C6676">Institute of Natural and Environmental Sciences, University of Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan &amp; Museum of Nature and Human Activities, Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan &amp; aenictus @ mac. com; https: // orcid. org / 0000 - 0002 - 6696 - 5011</mods:affiliation>
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<mods:affiliation id="5FC291C9BB5548D6AE13F5BDE5E3D8D4">School of Human Sciences, Kobe College, Okadayama 4 - 1, Nishinomiya, Hyogo, 662 - 8505, Japan &amp; endo @ mail. kobe-c. ac. jp; https: // orcid. org / 0009 - 0007 - 8279 - 9897</mods:affiliation>
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<mods:affiliation id="0E908F9DF2A0C6AF4CE8BC5976A5F635">Research Core for Interdisciplinary Sciences, Okayama University, 3 - 1 - 1, Tsushimanaka, Okayama, 700 - 8530, Japan &amp; fhyodo @ cc. okayama-u. ac. jp; https: // orcid. org / 0000 - 0001 - 8344 - 9353</mods:affiliation>
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<mods:namePart id="EE9891B2F0B0922171F7541DCEAFC3FF">Itioka, Takao</mods:namePart>
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<mods:affiliation id="4B32BA4014C56A0E66446BFDB5F063B4">Graduate School of Human and Environmental Studies, Kyoto University, Yoshida-nihonmatsu, Kyoto 606 - 8501, Japan &amp; ichioka. takao. 5 m @ kyoto-u. ac. jp; https: // orcid. org / 0000 - 0002 - 8295 - 5657</mods:affiliation>
<mods:nameIdentifier id="C0327A4AD18EBC5B048F4032DBD153C1" type="email">ichioka.takao.5m@kyoto-u.ac.jp</mods:nameIdentifier>
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<mods:namePart id="D25FDE85BEC6D5C85DFB6437B62E1BD7">Mohamed, Maryati</mods:namePart>
<mods:affiliation id="BCEDF76B86B201A03FE1D8ED955A6945">Tun Hussein Onn Malaysia, Hab Pendidikan Tinggi Pagoh, KM 1, Jalan Panchor, 84600 Panchor, Johor, Malaysia &amp; maryatihayati 52 @ gmail. com</mods:affiliation>
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<mods:namePart id="72A3181AC7CF7B4C9D4D335EABE369FC">Meleng, Paulus</mods:namePart>
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<mods:affiliation id="9606A165D18672FEC89064FAC72F5CE3">Research Development and Innovation Division, Forest Department Sarawak, Kuching, KM 10, Jalan Datuk Amar Stephen Kalong Ningkan, 93250 Kuching, Sarawak, Malaysia &amp; paulusm @ sarawak. gov. my; https: // orcid. org / 0000 - 0003 - 3777 - 0597</mods:affiliation>
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<treatment id="03FA87E7FFBCFFD6FF7D3E82FDCE8A93" LSID="urn:lsid:plazi:treatment:03FA87E7FFBCFFD6FF7D3E82FDCE8A93" httpUri="http://treatment.plazi.org/id/03FA87E7FFBCFFD6FF7D3E82FDCE8A93" lastPageId="13" lastPageNumber="68" pageId="11" pageNumber="66">
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<heading id="D0A4819DFFBCFFD0FF7D3E82FD9088A7" bold="true" box="[151,636,151,178]" fontSize="11" level="1" pageId="11" pageNumber="66" reason="1">
<taxonomicName id="4C534D72FFBCFFD0FF7D3E82FD9088A7" ID-CoL="7F64H" authority="Kishida, 1940" authorityName="Kishida" authorityYear="1940" box="[151,636,151,178]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFBCFFD0FF7D3E82FD9088A7" bold="true" box="[151,636,151,178]" pageId="11" pageNumber="66">
<emphasis id="B927EAE3FFBCFFD0FF7D3E82FE2588A4" bold="true" box="[151,457,151,178]" italics="true" pageId="11" pageNumber="66">Utivarachna fukasawana</emphasis>
<bibRefCitation id="EFC24B00FFBCFFD0FE3A3E82FD9088A7" author="Kishida, K." box="[464,636,151,177]" pageId="11" pageNumber="66" pagination="138 - 145" refId="ref9036" refString="Kishida, K. (1940) Notes on two species of spiders, Doosia japonica and Utivarachna fukasawana. Acta Arachnologica, 5, 138 - 145. https: // doi. org / 10.2476 / asjaa. 5.138" type="journal article" year="1940">Kishida, 1940</bibRefCitation>
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<figureCitation id="13682A74FFBCFFD0FF7D3EAEFEF888C3" box="[151,276,187,213]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStart-3="FIGURE 19" captionStartId-0="11.[151,250,1119,1143]" captionStartId-1="11.[151,250,1708,1732]" captionStartId-2="12.[151,250,909,933]" captionStartId-3="12.[151,250,1336,1360]" captionTargetBox-0="[151,1435,478,1095]" captionTargetBox-1="[151,1433,1204,1685]" captionTargetBox-2="[151,1436,181,886]" captionTargetBox-3="[151,1435,1030,1313]" captionTargetId-0="figure-81@11.[151,1436,478,1095]" captionTargetId-1="figure-117@11.[151,1435,1203,1685]" captionTargetId-2="figure-17@12.[151,1436,181,886]" captionTargetId-3="figure-60@12.[151,1435,1030,1313]" captionTargetPageId-0="11" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="12" captionText-0="FIGURE 16. Utivarachna fukasawana Kishida, 1940, male (LCo20070821_O1). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view. Scale bars: 1 mm." captionText-1="FIGURE 17. Utivarachna fukasawana, male (LCo20070821_O1). A left palp, prolateral view; B same, ventral view; C same, retrolateral view; D same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum." captionText-2="FIGURE 18. Utivarachna fukasawana Kishida, 1940, female (RNMH.ARA.15300). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label; F label, opposite side. Scale bars A–D: 1 mm." captionText-3="FIGURE 19. Utivarachna fukasawana, female (RNMH.ARA.15300). A epigyne, ventral view; B internal organs, ventral view; C same, dorsal view. Scale bars: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.8324294" figureDoi-1="http://doi.org/10.5281/zenodo.8324296" figureDoi-2="http://doi.org/10.5281/zenodo.8324300" figureDoi-3="http://doi.org/10.5281/zenodo.8324302" httpUri-0="https://zenodo.org/record/8324294/files/figure.png" httpUri-1="https://zenodo.org/record/8324296/files/figure.png" httpUri-2="https://zenodo.org/record/8324300/files/figure.png" httpUri-3="https://zenodo.org/record/8324302/files/figure.png" pageId="11" pageNumber="66">Figs 16–19</figureCitation>
</paragraph>
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<treatmentCitation id="0AF210E0FFBCFFD0FF7D3F15FD92890E" author="Kishida, K." box="[151,638,256,280]" page="142" pageId="11" pageNumber="66" year="1940">
<taxonomicName id="4C534D72FFBCFFD0FF7D3F15FD92890E" ID-CoL="7F64H" authority="Kishida, 1940: 142" authorityName="Kishida" authorityPageNumber="142" authorityYear="1940" box="[151,638,256,280]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFBCFFD0FF7D3F15FE72890E" box="[151,414,256,280]" italics="true" pageId="11" pageNumber="66">Utivarachna fukasawana</emphasis>
<bibRefCitation id="EFC24B00FFBCFFD0FE413F15FD92890E" author="Kishida, K." box="[427,638,256,280]" pageId="11" pageNumber="66" pagination="138 - 145" refId="ref9036" refString="Kishida, K. (1940) Notes on two species of spiders, Doosia japonica and Utivarachna fukasawana. Acta Arachnologica, 5, 138 - 145. https: // doi. org / 10.2476 / asjaa. 5.138" type="journal article" year="1940">Kishida, 1940: 142</bibRefCitation>
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;
<treatmentCitation id="0AF210E0FFBCFFD0FD7A3F15FC33890E" author="Deeleman-Reinhold, C. L." box="[656,991,256,280]" page="373" pageId="11" pageNumber="66" year="2001">
<bibRefCitation id="EFC24B00FFBCFFD0FD7A3F15FC33890E" author="Deeleman-Reinhold, C. L." box="[656,991,256,280]" pageId="11" pageNumber="66" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001: 373</bibRefCitation>
</treatmentCitation>
, figs 575–580;
<treatmentCitation id="0AF210E0FFBCFFD0FB783F15FE4A8920" author="Dankittipakul, P. &amp; Tavano, M. &amp; Singtripop, T." page="137" pageId="11" pageNumber="66" year="2011">
<bibRefCitation id="EFC24B00FFBCFFD0FB783F15FE4A8920" author="Dankittipakul, P. &amp; Tavano, M. &amp; Singtripop, T." pageId="11" pageNumber="66" pagination="133 - 146" refId="ref8829" refString="Dankittipakul, P., Tavano, M. &amp; Singtripop, T. (2011) Two new species of the spider genus Utivarachna Kishida, 1940 from Southwest Asia (Araneae, Corinnidae). Annali del Museo Civico di Storia Naturale Giacomo Doria, 103, 133 - 146." type="journal article" year="2011">Dankittipakul, Tavano &amp; Singtripop, 2011: 137</bibRefCitation>
</treatmentCitation>
, fig. 6.
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</paragraph>
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<emphasis id="B927EAE3FFBCFFD0FF7D3F71FE6F8968" bold="true" box="[151,387,356,382]" pageId="11" pageNumber="66">Material examined.</emphasis>
<materialsCitation id="3B3B3CACFFBCFFD0FE613F70FCF28968" box="[395,798,356,383]" collectionCode="RMNH" country="Brunei" location="Brunei" pageId="11" pageNumber="66" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D55FD78FFBCFFD0FE613F70FE5B8969" box="[395,439,357,383]" pageId="11" pageNumber="66" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED42AE34FFBCFFD0FE223F70FDCD8969" box="[456,545,357,383]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="66" type="Museum">RMNH</collectionCode>
.ARA. 15300),
<collectingCountry id="F3447661FFBCFFD0FD253F71FCF28968" box="[719,798,356,382]" name="Brunei" pageId="11" pageNumber="66">Brunei</collectingCountry>
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,
<materialsCitation id="3B3B3CACFFBCFFD0FCC13F71FDD789D0" collectingDate="1992-02-08" collectionCode="R, MNHAH" country="Malaysia" location="West Ridge" municipality="Lambir Hills National Park" pageId="11" pageNumber="66" specimenCount="1" typeStatus="neotype">
Kuala Belalong,
<location id="8E8C602AFFBCFFD0FC013F70FB9E8968" LSID="urn:lsid:plazi:treatment:03FA87E7FFBCFFD6FF7D3E82FDCE8A93:8E8C602AFFBCFFD0FC013F70FB9E8968" box="[1003,1138,356,383]" country="Malaysia" municipality="Lambir Hills National Park" name="West Ridge" pageId="11" pageNumber="66">West Ridge</location>
,
<date id="FFED1031FFBCFFD0FB943F70FB0B8969" box="[1150,1255,357,383]" pageId="11" pageNumber="66" value="1992-02-08">
<collectingDate id="EFA9E9D9FFBCFFD0FB943F70FB0B8969" box="[1150,1255,357,383]" pageId="11" pageNumber="66" value="1992-02-08">8.II.1992</collectingDate>
</date>
, leg.
<collectionCode id="ED42AE34FFBCFFD0FACD3F70FAD18969" box="[1319,1341,357,383]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="66">R</collectionCode>
. Snazell (label data shown in
<figureCitation id="13682A74FFBCFFD0FE603F9DFDD489B5" box="[394,568,392,419]" captionStart="FIGURE 18" captionStartId="12.[151,250,909,933]" captionTargetBox="[151,1436,181,886]" captionTargetId="figure-17@12.[151,1436,181,886]" captionTargetPageId="12" captionText="FIGURE 18. Utivarachna fukasawana Kishida, 1940, female (RNMH.ARA.15300). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label; F label, opposite side. Scale bars A–D: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324300" httpUri="https://zenodo.org/record/8324300/files/figure.png" pageId="11" pageNumber="66">Fig. 18E and F</figureCitation>
are the same as those of the
<typeStatus id="54E88853FFBCFFD0FC673F9CFC0789B5" box="[909,1003,393,419]" pageId="11" pageNumber="66">neotype</typeStatus>
). 1 ♁ (LCo20070821_O1;
<collectionCode id="ED42AE34FFBCFFD0FAC83F9CFA7C89B5" box="[1314,1424,393,419]" pageId="11" pageNumber="66">MNHAH</collectionCode>
),
<collectingMunicipality id="6B88AC8BFFBCFFD0FF7D3FB9FE2089D0" box="[151,460,428,454]" pageId="11" pageNumber="66">Lambir Hills National Park</collectingMunicipality>
,
<collectingCountry id="F3447661FFBCFFD0FE3D3FB9FDD789D0" box="[471,571,428,454]" name="Malaysia" pageId="11" pageNumber="66">Sarawak</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF2C6679FFBCFFD0FF7D3A4AFBD68C8C" ID-DOI="http://doi.org/10.5281/zenodo.8324294" ID-Zenodo-Dep="8324294" httpUri="https://zenodo.org/record/8324294/files/figure.png" pageId="11" pageNumber="66" startId="11.[151,250,1119,1143]" targetBox="[151,1435,478,1095]" targetPageId="11" targetType="figure">
<paragraph id="8BEC36F1FFBCFFD0FF7D3A4AFBD68C8C" blockId="11.[151,1436,1119,1179]" pageId="11" pageNumber="66">
<emphasis id="B927EAE3FFBCFFD0FF7D3A4AFEF38C61" bold="true" box="[151,287,1119,1143]" pageId="11" pageNumber="66">FIGURE 16.</emphasis>
<taxonomicName id="4C534D72FFBCFFD0FECE3A4AFD548C61" authority="Kishida, 1940" authorityName="Kishida" authorityYear="1940" box="[292,696,1119,1143]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFBCFFD0FECE3A4AFDCF8C61" box="[292,547,1119,1143]" italics="true" pageId="11" pageNumber="66">Utivarachna fukasawana</emphasis>
<bibRefCitation id="EFC24B00FFBCFFD0FDC33A4AFD548C61" author="Kishida, K." box="[553,696,1119,1143]" pageId="11" pageNumber="66" pagination="138 - 145" refId="ref9036" refString="Kishida, K. (1940) Notes on two species of spiders, Doosia japonica and Utivarachna fukasawana. Acta Arachnologica, 5, 138 - 145. https: // doi. org / 10.2476 / asjaa. 5.138" type="journal article" year="1940">Kishida, 1940</bibRefCitation>
</taxonomicName>
, male (LCo20070821_O1).
<emphasis id="B927EAE3FFBCFFD0FC393A4AFC098C61" bold="true" box="[979,997,1119,1143]" pageId="11" pageNumber="66">A</emphasis>
habitus, dorsal view;
<emphasis id="B927EAE3FFBCFFD0FB283A4AFB3F8C61" bold="true" box="[1218,1235,1119,1143]" pageId="11" pageNumber="66">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFBCFFD0FF7D3A96FF458C8D" bold="true" box="[151,169,1155,1179]" pageId="11" pageNumber="66">C</emphasis>
cephalic part and chelicerae, frontal view;
<emphasis id="B927EAE3FFBCFFD0FDB53A96FD9D8C8D" bold="true" box="[607,625,1155,1179]" pageId="11" pageNumber="66">D</emphasis>
leg I, left, prolateral view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF2C6679FFBCFFD0FF7D38B9FB058EFF" ID-DOI="http://doi.org/10.5281/zenodo.8324296" ID-Zenodo-Dep="8324296" httpUri="https://zenodo.org/record/8324296/files/figure.png" pageId="11" pageNumber="66" startId="11.[151,250,1708,1732]" targetBox="[151,1433,1204,1685]" targetPageId="11" targetType="figure">
<paragraph id="8BEC36F1FFBCFFD0FF7D38B9FB058EFF" blockId="11.[151,1436,1708,1769]" pageId="11" pageNumber="66">
<emphasis id="B927EAE3FFBCFFD0FF7D38B9FECC8ED3" bold="true" box="[151,288,1708,1733]" pageId="11" pageNumber="66">FIGURE 17.</emphasis>
<taxonomicName id="4C534D72FFBCFFD0FECC38B8FDCA8ED3" authorityName="Kishida" authorityYear="1940" box="[294,550,1709,1733]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFBCFFD0FECC38B8FDCA8ED3" box="[294,550,1709,1733]" italics="true" pageId="11" pageNumber="66">Utivarachna fukasawana</emphasis>
</taxonomicName>
, male (LCo20070821_O1).
<emphasis id="B927EAE3FFBCFFD0FCAC38B9FCB48ED2" bold="true" box="[838,856,1708,1732]" pageId="11" pageNumber="66">A</emphasis>
left palp, prolateral view;
<emphasis id="B927EAE3FFBCFFD0FB8F38B8FB9A8ED3" bold="true" box="[1125,1142,1709,1733]" pageId="11" pageNumber="66">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFBCFFD0FAA338B9FAB78ED2" bold="true" box="[1353,1371,1708,1732]" pageId="11" pageNumber="66">C</emphasis>
same, retrolateral view;
<emphasis id="B927EAE3FFBCFFD0FEA138C4FEB18EFF" bold="true" box="[331,349,1745,1769]" pageId="11" pageNumber="66">D</emphasis>
same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum.
</paragraph>
</caption>
<subSubSection id="C349657AFFBCFFD0FF2D392EFE538FD7" pageId="11" pageNumber="66" type="diagnosis">
<paragraph id="8BEC36F1FFBCFFD0FF2D392EFE538FD7" blockId="11.[151,1436,1851,1985]" pageId="11" pageNumber="66">
<emphasis id="B927EAE3FFBCFFD0FF2D392EFEAE8F43" bold="true" box="[199,322,1851,1877]" pageId="11" pageNumber="66">Diagnosis.</emphasis>
In the
<taxonomicName id="4C534D72FFBCFFD0FE72392EFDF28F43" authorityName="Kishida" authorityYear="1940" box="[408,542,1851,1877]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFBCFFD0FE72392EFDF28F43" box="[408,542,1851,1877]" italics="true" pageId="11" pageNumber="66">fukasawana</emphasis>
</taxonomicName>
-group,
<taxonomicName id="4C534D72FFBCFFD0FD933929FCC88F43" authorityName="Kishida" authorityYear="1940" box="[633,804,1851,1877]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFBCFFD0FD933929FCC88F43" box="[633,804,1851,1877]" italics="true" pageId="11" pageNumber="66">U. fukasawana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D72FFBCFFD0FC8A3929FBEC8F43" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[864,1024,1851,1877]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="ichneumon">
<emphasis id="B927EAE3FFBCFFD0FC8A3929FBEC8F43" box="[864,1024,1851,1877]" italics="true" pageId="11" pageNumber="66">U. ichneumon</emphasis>
</taxonomicName>
are easily recognizable by the long stalk projection of the posterior carapace, especially in males. In males,
<taxonomicName id="4C534D72FFBCFFD0FC2D3975FB9C8F6F" authorityName="Kishida" authorityYear="1940" box="[967,1136,1887,1913]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFBCFFD0FC2D3975FB9C8F6F" box="[967,1136,1887,1913]" italics="true" pageId="11" pageNumber="66">U. fukasawana</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C534D72FFBCFFD0FF7D3991FED98F8B" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[151,309,1923,1949]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="ichneumon">
<emphasis id="B927EAE3FFBCFFD0FF7D3991FED98F8B" box="[151,309,1923,1949]" italics="true" pageId="11" pageNumber="66">U. ichneumon</emphasis>
</taxonomicName>
by the wide cymbium and bulb. In females,
<taxonomicName id="4C534D72FFBCFFD0FCC03991FC3E8F8B" authorityName="Kishida" authorityYear="1940" box="[810,978,1923,1949]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFBCFFD0FCC03991FC3E8F8B" box="[810,978,1923,1949]" italics="true" pageId="11" pageNumber="66">U. fukasawana</emphasis>
</taxonomicName>
has short bursae, whereas
<taxonomicName id="4C534D72FFBCFFD0FB153991FA708F8B" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[1279,1436,1923,1949]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="ichneumon">
<emphasis id="B927EAE3FFBCFFD0FB153991FA708F8B" box="[1279,1436,1923,1949]" italics="true" pageId="11" pageNumber="66">U. ichneumon</emphasis>
</taxonomicName>
has long, tube-like bursae.
</paragraph>
</subSubSection>
<caption id="DF2C6679FFBBFFD7FF7D3D98FF358BFB" ID-DOI="http://doi.org/10.5281/zenodo.8324300" ID-Zenodo-Dep="8324300" httpUri="https://zenodo.org/record/8324300/files/figure.png" pageId="12" pageNumber="67" startId="12.[151,250,909,933]" targetBox="[151,1436,181,886]" targetPageId="12" targetType="figure">
<paragraph id="8BEC36F1FFBBFFD7FF7D3D98FF358BFB" blockId="12.[151,1437,909,1006]" pageId="12" pageNumber="67">
<emphasis id="B927EAE3FFBBFFD7FF7D3D98FECD8BB3" bold="true" box="[151,289,909,933]" pageId="12" pageNumber="67">FIGURE 18.</emphasis>
<taxonomicName id="4C534D72FFBBFFD7FEC23D9BFD2E8BB0" authority="Kishida, 1940" authorityName="Kishida" authorityYear="1940" box="[296,706,910,934]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="67" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFBBFFD7FEC23D9BFDC58BB0" box="[296,553,910,934]" italics="true" pageId="12" pageNumber="67">Utivarachna fukasawana</emphasis>
<bibRefCitation id="EFC24B00FFBBFFD7FDDA3D9BFD2E8BB0" author="Kishida, K." box="[560,706,910,934]" pageId="12" pageNumber="67" pagination="138 - 145" refId="ref9036" refString="Kishida, K. (1940) Notes on two species of spiders, Doosia japonica and Utivarachna fukasawana. Acta Arachnologica, 5, 138 - 145. https: // doi. org / 10.2476 / asjaa. 5.138" type="journal article" year="1940">Kishida, 1940</bibRefCitation>
</taxonomicName>
, female (RNMH.ARA.15300).
<emphasis id="B927EAE3FFBBFFD7FBEC3D98FBF48BB3" bold="true" box="[1030,1048,909,933]" pageId="12" pageNumber="67">A</emphasis>
habitus, dorsal view;
<emphasis id="B927EAE3FFBBFFD7FB163D9BFAE18BB0" bold="true" box="[1276,1293,910,934]" pageId="12" pageNumber="67">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFBBFFD7FF3F3DA4FF0B8BDF" bold="true" box="[213,231,945,969]" pageId="12" pageNumber="67">C</emphasis>
cephalic part and chelicerae, frontal view;
<emphasis id="B927EAE3FFBBFFD7FD703DA7FD408BDC" bold="true" box="[666,684,946,970]" pageId="12" pageNumber="67">D</emphasis>
leg I, left, prolateral view;
<emphasis id="B927EAE3FFBBFFD7FC2A3DA7FC3D8BDC" bold="true" box="[960,977,946,970]" pageId="12" pageNumber="67">E</emphasis>
label;
<emphasis id="B927EAE3FFBBFFD7FBFF3DA7FBC88BDC" bold="true" box="[1045,1060,946,970]" pageId="12" pageNumber="67">F</emphasis>
label, opposite side. Scale bars A–D: 1 mm.
</paragraph>
</caption>
<caption id="DF2C6679FFBBFFD7FF7D3B2DFBB58D62" ID-DOI="http://doi.org/10.5281/zenodo.8324302" ID-Zenodo-Dep="8324302" httpUri="https://zenodo.org/record/8324302/files/figure.png" pageId="12" pageNumber="67" startId="12.[151,250,1336,1360]" targetBox="[151,1435,1030,1313]" targetPageId="12" targetType="figure">
<paragraph id="8BEC36F1FFBBFFD7FF7D3B2DFBB58D62" blockId="12.[151,1436,1336,1396]" pageId="12" pageNumber="67">
<emphasis id="B927EAE3FFBBFFD7FF7D3B2DFEF38D46" bold="true" box="[151,287,1336,1360]" pageId="12" pageNumber="67">FIGURE 19.</emphasis>
<taxonomicName id="4C534D72FFBBFFD7FECE3B2DFDCF8D46" authorityName="Kishida" authorityYear="1940" box="[292,547,1336,1360]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="67" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFBBFFD7FECE3B2DFDCF8D46" box="[292,547,1336,1360]" italics="true" pageId="12" pageNumber="67">Utivarachna fukasawana</emphasis>
</taxonomicName>
, female (RNMH.ARA.15300).
<emphasis id="B927EAE3FFBBFFD7FC893B2DFC998D46" bold="true" box="[867,885,1336,1360]" pageId="12" pageNumber="67">A</emphasis>
epigyne, ventral view;
<emphasis id="B927EAE3FFBBFFD7FB8A3B2DFB9D8D46" bold="true" box="[1120,1137,1336,1360]" pageId="12" pageNumber="67">B</emphasis>
internal organs, ventral view;
<emphasis id="B927EAE3FFBBFFD7FF7D3B49FF458D62" bold="true" box="[151,169,1372,1396]" pageId="12" pageNumber="67">C</emphasis>
same, dorsal view. Scale bars: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca.
</paragraph>
</caption>
<subSubSection id="C349657AFFBBFFD6FF2D3BD2FC678A77" lastPageId="13" lastPageNumber="68" pageId="12" pageNumber="67" type="description">
<paragraph id="8BEC36F1FFBBFFD7FF2D3BD2FDB48ECB" blockId="12.[151,1437,1478,2009]" pageId="12" pageNumber="67">
<emphasis id="B927EAE3FFBBFFD7FF2D3BD2FE9A8DF7" bold="true" box="[199,374,1479,1505]" pageId="12" pageNumber="67">Measurements</emphasis>
(♁/
<specimenCount id="9D55FD78FFBBFFD7FE4F3BDDFE5B8DF5" box="[421,439,1480,1507]" pageId="12" pageNumber="67" type="female">♀</specimenCount>
). Carapace length 3.35/2.86; cephalic width 1.45/1.33; thoracic width 1.77/1.62. Diameter of eyes: AME 0.18/0.17; ALE 0.17/0.17; PME 0.15/0.13; PLE 0.13/0.16. Interdistances between eyes: AME–AME 0.06/0.05; AME–ALE 0.07/0.06; ALE–ALE 0.58/0.50; PME–PME 0.17/0.16; PME–PLE 0.35/0.27; PLE–PLE 1.11/0.97; ALE–PLE 0.22/0.20; AME–PME 0.07/0.05. MOA: length 0.35/0.28; anterior width 0.41/0.36; posterior width 0.46/0.43. Clypeus height 0.40/0.33. Abdomen length 2.76/2.73; width 1.27/1.76. Legs: I 10.56 (2.80, 1.10, 2.70, 2.80, 1.16)/8.73 (2.45, 0.86, 2.25, 2.15, 1.02); II 8.92 (2.46, 0.90, 2.20, 2.33, 1.03)/7.90 (2.25, 0.73, 2.02, 1.97, 0.93); III 5.36 (1.50, 0.62, 1.17, 1.50, 0.57)/5.04 (1.43, 0.58, 1.11, 1.35, 0.57); IV 7.15 (2.00, 0.70, 1.57, 2.12, 0.76)/7.13 (2.00, 0.63, 1.65, 2.10, 0.75).
</paragraph>
<paragraph id="8BEC36F1FFBBFFD7FF2D38F2FD018FA3" blockId="12.[151,1437,1478,2009]" pageId="12" pageNumber="67">
<emphasis id="B927EAE3FFBBFFD7FF2D38F2FEE88F17" bold="true" box="[199,260,1767,1793]" pageId="12" pageNumber="67">Male</emphasis>
(
<figureCitation id="13682A74FFBBFFD7FEFE38F3FE4C8F16" box="[276,416,1766,1792]" captionStart="FIGURE 16" captionStartId="11.[151,250,1119,1143]" captionTargetBox="[151,1435,478,1095]" captionTargetId="figure-81@11.[151,1436,478,1095]" captionTargetPageId="11" captionText="FIGURE 16. Utivarachna fukasawana Kishida, 1940, male (LCo20070821_O1). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324294" httpUri="https://zenodo.org/record/8324294/files/figure.png" pageId="12" pageNumber="67">Fig. 16A–D</figureCitation>
). Carapace oval, entirely granulated; posterior stalk almost as long as 1/4 carapace length; PER slightly recurved in dorsal view; ocular tubercle accompanied with PLE; thoracic lateral margins undulating. Chelicerae granulated, with five prolateral and five retrolateral teeth. Ventral clypeus margin with small wedge-shaped projection between chelicerae. Pedicel short. Abdomen elongate oval; anterior two-thirds with weakly sclerotized dorsal scutum. Leg I and II elongated, covered with fine spherical granulation; ventral sides of tibia, metatarsus, and tarsus bearing small tooth-like cusps.
</paragraph>
<paragraph id="8BEC36F1FFBBFFD6FF2D39ABFC5E890B" blockId="12.[151,1437,1478,2009]" lastBlockId="13.[151,1437,151,645]" lastPageId="13" lastPageNumber="68" pageId="12" pageNumber="67">
Palp (
<figureCitation id="13682A74FFBBFFD7FEE739ABFE718FCF" box="[269,413,1982,2009]" captionStart="FIGURE 17" captionStartId="11.[151,250,1708,1732]" captionTargetBox="[151,1433,1204,1685]" captionTargetId="figure-117@11.[151,1435,1203,1685]" captionTargetPageId="11" captionText="FIGURE 17. Utivarachna fukasawana, male (LCo20070821_O1). A left palp, prolateral view; B same, ventral view; C same, retrolateral view; D same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum." figureDoi="http://doi.org/10.5281/zenodo.8324296" httpUri="https://zenodo.org/record/8324296/files/figure.png" pageId="12" pageNumber="67">Fig. 17A–D</figureCitation>
). Cymbium strongly convex in retrolateral margin. Bulb inverted pear-shaped, tapering posteriorly; anterior half membranous, and prolateral half weakly sclerotized with subtegulum. Embolus coiled in horizontal plane, inclining dorsally, and plane completely covered by cymbium. Sperm duct U-curved in posterior part of bulb. RTA strongly extended, apically developed into semi-transparent plate with apical claw; base of claw well-sclerotized and developed; claw slender, strongly curved dorsally.
</paragraph>
<paragraph id="8BEC36F1FFBAFFD6FF2D3F32FED2899F" blockId="13.[151,1437,151,645]" pageId="13" pageNumber="68">
Coloration and setation (
<figureCitation id="13682A74FFBAFFD6FE033F32FD948957" box="[489,632,295,321]" captionStart="FIGURE 16" captionStartId="11.[151,250,1119,1143]" captionTargetBox="[151,1435,478,1095]" captionTargetId="figure-81@11.[151,1436,478,1095]" captionTargetPageId="11" captionText="FIGURE 16. Utivarachna fukasawana Kishida, 1940, male (LCo20070821_O1). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324294" httpUri="https://zenodo.org/record/8324294/files/figure.png" pageId="13" pageNumber="68">Fig. 16A–D</figureCitation>
). Carapace and chelicerae brown, covered with fine setae. Abdomen cream, entirely covered with short setae. All legs brownish cream, except for femora I–II and trochanters I–II brown to light brown.
</paragraph>
<paragraph id="8BEC36F1FFBAFFD6FF2D3F86FCAF89E3" blockId="13.[151,1437,151,645]" pageId="13" pageNumber="68">
<emphasis id="B927EAE3FFBAFFD6FF2D3F86FEF289BB" bold="true" box="[199,286,403,429]" pageId="13" pageNumber="68">Female</emphasis>
(
<figureCitation id="13682A74FFBAFFD6FEDA3F86FE5389BB" box="[304,447,403,429]" captionStart="FIGURE 18" captionStartId="12.[151,250,909,933]" captionTargetBox="[151,1436,181,886]" captionTargetId="figure-17@12.[151,1436,181,886]" captionTargetPageId="12" captionText="FIGURE 18. Utivarachna fukasawana Kishida, 1940, female (RNMH.ARA.15300). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label; F label, opposite side. Scale bars A–D: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324300" httpUri="https://zenodo.org/record/8324300/files/figure.png" pageId="13" pageNumber="68">Fig. 18A–D</figureCitation>
). Somatic characters almost identical to males, except for abdomen shape and shorter posterior stalk; posterior stalk almost as long as 1/5 carapace length. Chelicerae granulated, with six prolateral and five retrolateral teeth. Abdomen oval, without dorsal scutum.
</paragraph>
<paragraph id="8BEC36F1FFBAFFD6FF2D3FEAFAF28A2B" blockId="13.[151,1437,151,645]" pageId="13" pageNumber="68">
Copulatory organs (
<figureCitation id="13682A74FFBAFFD6FE4D3FEAFDDD8A0F" box="[423,561,511,537]" captionStart="FIGURE 19" captionStartId="12.[151,250,1336,1360]" captionTargetBox="[151,1435,1030,1313]" captionTargetId="figure-60@12.[151,1435,1030,1313]" captionTargetPageId="12" captionText="FIGURE 19. Utivarachna fukasawana, female (RNMH.ARA.15300). A epigyne, ventral view; B internal organs, ventral view; C same, dorsal view. Scale bars: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8324302" httpUri="https://zenodo.org/record/8324302/files/figure.png" pageId="13" pageNumber="68">Fig. 19A–C</figureCitation>
). Copulatory atrium square. Copulatory openings located at anterior corner of atrium, extending anteriorly with several coils and connected to slender bursae. Spermathecae globular.
</paragraph>
<paragraph id="8BEC36F1FFBAFFD6FF2D3C52FC678A77" blockId="13.[151,1437,151,645]" box="[199,907,583,609]" pageId="13" pageNumber="68">
Coloration and setation (
<figureCitation id="13682A74FFBAFFD6FE363C52FD848A77" box="[476,616,583,609]" captionStart="FIGURE 18" captionStartId="12.[151,250,909,933]" captionTargetBox="[151,1436,181,886]" captionTargetId="figure-17@12.[151,1436,181,886]" captionTargetPageId="12" captionText="FIGURE 18. Utivarachna fukasawana Kishida, 1940, female (RNMH.ARA.15300). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label; F label, opposite side. Scale bars A–D: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324300" httpUri="https://zenodo.org/record/8324300/files/figure.png" pageId="13" pageNumber="68">Fig. 18A–D</figureCitation>
). Almost same as in male.
</paragraph>
</subSubSection>
<subSubSection id="C349657AFFBAFFD6FF2D3C7FFDCE8A93" box="[199,546,618,645]" pageId="13" pageNumber="68" type="distribution">
<paragraph id="8BEC36F1FFBAFFD6FF2D3C7FFDCE8A93" blockId="13.[151,1437,151,645]" box="[199,546,618,645]" pageId="13" pageNumber="68">
<emphasis id="B927EAE3FFBAFFD6FF2D3C7FFE8D8A92" bold="true" box="[199,353,618,644]" pageId="13" pageNumber="68">Distribution.</emphasis>
<collectingCountry id="F3447661FFBAFFD6FE8D3C7EFE5A8A93" box="[359,438,619,645]" name="Brunei" pageId="13" pageNumber="68">Brunei</collectingCountry>
,
<collectingCountry id="F3447661FFBAFFD6FE283C7EFDCE8A93" box="[450,546,619,645]" name="Malaysia" pageId="13" pageNumber="68">Sarawak</collectingCountry>
</paragraph>
</subSubSection>
</treatment>
</document>
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