295 lines
36 KiB
XML
295 lines
36 KiB
XML
<document id="5363DA5CBAACDD525A1E6348E1427A37" ID-DOI="10.5281/zenodo.170190" ID-GBIF-Dataset="0a606273-f8f3-4c9e-87c6-76f5f96a221c" ID-ISSN="11755326" ID-Zenodo-Dep="170190" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459751819524" checkinUser="plazi" docAuthor="Robertson, Desiree R. & Holzenthal, Ralph W." docDate="2005" docId="03FA87D68206FF8AFEF37D26FBFEFB61" docLanguage="en" docName="zt01063p068.pdf" docOrigin="Zootaxa 1063" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Tolhuaca cupulifera Schmid" docType="treatment" docVersion="9" lastPageNumber="62" masterDocId="FFC3FFAE8202FF83FFFB7850FFD2FFAF" masterDocTitle="The Neotropical caddisfly genus To l h u a c a (Trichoptera: Glossosomatidae)" masterLastPageNumber="68" masterPageNumber="53" pageNumber="57" updateTime="1698211932345" updateUser="plazi">
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<mods:titleInfo id="E02FD09ABD6DDB8F0B0A2EBF436B591A">
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<mods:title id="C5CA477F4FB01F3E7288275696EB660A">The Neotropical caddisfly genus To l h u a c a (Trichoptera: Glossosomatidae)</mods:title>
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<mods:name id="BFC6689DEDB7119AFD42595670C9D9B6" type="personal">
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<mods:namePart id="0B422F2667C5F532BCC52190046E3616">Robertson, Desiree R.</mods:namePart>
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<mods:name id="E689564944F55F4919220DC1337EF8CA" type="personal">
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<mods:namePart id="913046E89ACDCEEC46429016C3141E97">Holzenthal, Ralph W.</mods:namePart>
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<mods:typeOfResource id="EA6C4558FCF19E830EC624AC73226027">text</mods:typeOfResource>
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<mods:title id="E99B55262BA090D05738312E25836884">Zootaxa</mods:title>
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<mods:part id="92BC3103BA424EDF8FB58A55B8454DEE">
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<mods:date id="9A5806953B6BB17FC11CBA267A7D68C4">2005</mods:date>
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<mods:detail id="78DA7644D77566596496C054F2C6F1D0" type="volume">
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<mods:number id="69BDAA88E16940F5339C490FB9241F6A">1063</mods:number>
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<mods:classification id="30994D95C0CD9A11B161B48D27E8B06C">journal article</mods:classification>
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<mods:identifier id="350B24E0F021E5FB7D3D5B667C4829FD" type="DOI">10.5281/zenodo.170190</mods:identifier>
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<treatment id="03FA87D68206FF8AFEF37D26FBFEFB61" ID-DOI="http://doi.org/10.5281/zenodo.6265889" ID-GBIF-Taxon="119337914" ID-Zenodo-Dep="6265889" LSID="urn:lsid:plazi:treatment:03FA87D68206FF8AFEF37D26FBFEFB61" httpUri="http://treatment.plazi.org/id/03FA87D68206FF8AFEF37D26FBFEFB61" lastPageId="9" lastPageNumber="62" pageId="4" pageNumber="57">
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<subSubSection id="C349654B8206FF87FEF37D26FE7CFA1C" pageId="4" pageNumber="57" type="nomenclature">
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<paragraph id="8BEC36C08206FF87FEF37D26FDB1FA3F" blockId="4.[264,611,1398,1459]" box="[264,611,1398,1424]" pageId="4" pageNumber="57">
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<heading id="D0A481AC8206FF87FEF37D26FDB1FA3F" bold="true" box="[264,611,1398,1424]" fontSize="11" level="1" pageId="4" pageNumber="57" reason="1">
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<taxonomicName id="4C534D438206FF87FEF37D26FDB1FA3F" ID-CoL="57BG4" authority="Schmid" authorityName="Schmid" box="[264,611,1398,1424]" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="cupulifera">
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<emphasis id="B927EAD28206FF87FEF37D26FDB1FA3F" bold="true" box="[264,611,1398,1424]" pageId="4" pageNumber="57">
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<emphasis id="B927EAD28206FF87FEF37D26FE2FFA3F" bold="true" box="[264,509,1398,1424]" italics="true" pageId="4" pageNumber="57">Tolhuaca cupulifera</emphasis>
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Schmid
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BEC36C08206FF87FEF37DC9FE7CFA1C" blockId="4.[264,611,1398,1459]" box="[264,430,1433,1459]" pageId="4" pageNumber="57">
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<figureCitation id="13682A458206FF87FEF37DC9FEA7FA1C" box="[264,373,1433,1459]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 3" captionStartId-0="5.[264,368,1369,1393]" captionStartId-1="6.[264,368,1778,1802]" captionTargetBox-0="[264,1323,291,1318]" captionTargetId-0="figure@5.[264,1323,291,1318]" captionTargetPageId-0="5" captionText-0="FIGURE 1. Tolhuaca cupulifera. Adult: dorsal view of head and thorax. FIGURE 2. Tolhuaca cupulifera. Adult: A –– foretibial spur; B –– mesotibial spurs, at same scale for comparison." captionText-1="FIGURE 3. Tolhuaca cupulifera Schmid. Wings: A –– forewing; B –– hind wing." httpUri-0="https://zenodo.org/record/170191/files/figure.png" pageId="4" pageNumber="57">Figs. 1–3</figureCitation>
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,
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<figureCitation id="13682A458206FF87FE797DC9FE40FA1C" box="[386,402,1433,1459]" captionStart="FIGURE 5" captionStartId="7.[264,368,1783,1807]" captionTargetBox="[298,1289,321,1756]" captionTargetId="figure@7.[298,1289,321,1757]" captionTargetPageId="7" captionText="FIGURE 5. Tolhuaca cupulifera Schmid. Male genitalia: A –– lateral; B –– dorsal; C –– process of sternum VI; D –– endotheca fully everted as observed in holotype; E –– detail of endothecal thornlike spines; F –– ventral view of phallobase." httpUri="https://zenodo.org/record/170193/files/figure.png" pageId="4" pageNumber="57">5</figureCitation>
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,
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<figureCitation id="13682A458206FF87FE5B7DC9FE7CFA1C" box="[416,430,1433,1459]" captionStart="FIGURE 6" captionStartId="8.[264,368,1581,1605]" captionTargetBox="[290,1294,287,1549]" captionTargetId="figure@8.[290,1294,287,1550]" captionTargetPageId="8" captionText="FIGURE 6. Tolhuaca cupulifera Schmid. Female genitalia: A –– lateral; B –– dorsal; C –– ventral." httpUri="https://zenodo.org/record/170194/files/figure.png" pageId="4" pageNumber="57">6</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C349654B8206FF87FEF37DBAFD4BF8FC" pageId="4" pageNumber="57" type="reference_group">
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||
<paragraph id="8BEC36C08206FF87FEF37DBAFC6CF9EF" blockId="4.[264,1323,1513,1600]" pageId="4" pageNumber="57">
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||
<treatmentCitationGroup id="AB4311EE8206FF87FEF37DBAFC6CF9EF" pageId="4" pageNumber="57">
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||
<taxonomicName id="4C534D438206FF87FEF37DBAFD61F9AF" ID-CoL="57BG4" authority="Schmid, 1964: 337" authorityName="Schmid" authorityPageNumber="337" authorityYear="1964" box="[264,691,1513,1537]" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="cupulifera">
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<emphasis id="B927EAD28206FF87FEF37DBAFE32F9AE" box="[264,480,1514,1537]" italics="true" pageId="4" pageNumber="57">Tolhuaca cupulifera</emphasis>
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<treatmentCitation id="0AF210D18206FF87FE1C7DB9FD61F9AF" author="Schmid" box="[487,691,1513,1536]" page="337" pageId="4" pageNumber="57" year="1964">
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<bibRefCitation id="EFC24B318206FF87FE1C7DB9FDADF9AF" author="Schmid" box="[487,639,1513,1536]" pageId="4" pageNumber="68" refString="Schmid, F. (1964) Contribution a l'etude des Trichopteres neotropicaux V. Tijdschrift voor Entomologie, 107, 307 - 339." type="journal article" year="1964">Schmid, 1964</bibRefCitation>
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: 337
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</treatmentCitation>
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</taxonomicName>
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[
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<typeStatus id="54E888628206FF87FD3B7DB9FD2AF9AF" box="[704,760,1513,1536]" pageId="4" pageNumber="57">Type</typeStatus>
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locality:
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<collectingCountry id="F34476508206FF87FCA57DB9FC48F9AF" box="[862,922,1513,1536]" name="Chile" pageId="4" pageNumber="57">Chile</collectingCountry>
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, Pichinahuel, Arauco; NMNH; male; in
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<taxonomicName id="4C534D438206FF87FEAC7E59FDC7F98F" ID-CoL="G8F" box="[343,533,1545,1568]" class="Insecta" family="Sericostomatidae" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="family">Sericostomatidae</taxonomicName>
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]. –
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<treatmentCitation id="0AF210D18206FF87FDCE7E59FD1BF98F" author="Flint" box="[565,713,1545,1568]" page="52" pageId="4" pageNumber="57" year="1967">
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<bibRefCitation id="EFC24B318206FF87FDCE7E59FD7AF98F" author="Flint" box="[565,680,1545,1568]" pageId="4" pageNumber="67" refString="Flint, O. S., Jr. (1967) Studies of Neotropical caddis flies, II: Trichoptera collected by Prof. Dr. J. Illies in the Chilean Subregion. Beitrage zur Neotropischen Fauna, 5, 45 - 68." type="journal article" year="1967">Flint 1967</bibRefCitation>
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:52
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</treatmentCitation>
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[correction of transposition of wing figures in original description; to
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<taxonomicName id="4C534D438206FF87FE237E79FD43F9EF" ID-CoL="623KK" box="[472,657,1577,1600]" class="Insecta" family="Glossosomatidae" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="family">Glossosomatidae</taxonomicName>
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,
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<taxonomicName id="4C534D438206FF87FD607E79FCFAF9EF" ID-CoL="K7S" box="[667,808,1577,1600]" class="Insecta" family="Glossosomatidae" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="subFamily" subFamily="Protoptilinae">Protoptilinae</taxonomicName>
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; distribution].
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BEC36C08206FF87FEF37E21FD4BF8FC" blockId="4.[264,1323,1649,1875]" pageId="4" pageNumber="57">
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<taxonomicName id="4C534D438206FF87FEF37E21FE2EF925" ID-CoL="57BG4" box="[264,508,1649,1674]" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="cupulifera">
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<emphasis id="B927EAD28206FF87FEF37E21FE2EF925" box="[264,508,1649,1674]" italics="true" pageId="4" pageNumber="57">Tolhuaca cupulifera</emphasis>
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</taxonomicName>
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can be distinguished from
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<taxonomicName id="4C534D438206FF87FC937E21FBC6F925" box="[872,1044,1649,1674]" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="brasiliensis" status="sp. nov.">
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<emphasis id="B927EAD28206FF87FC937E21FBC6F925" box="[872,1044,1649,1674]" italics="true" pageId="4" pageNumber="57">T. brasiliensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21457A98206FF87FBD77E21FB16F924" box="[1068,1220,1649,1675]" pageId="4" pageNumber="57" rank="species">new species</taxonomicNameLabel>
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, by the presence of sclerotized, conical, thornlike spines in the endotheca. Tergum X is also much more excavate apicomesally, while it is shallowly excavate in the new species,
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<taxonomicName id="4C534D438206FF87FAEE7E91FE5DF8AD" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="brasiliensis">
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<emphasis id="B927EAD28206FF87FAEE7E91FE5DF8AD" italics="true" pageId="4" pageNumber="57">T. brasiliensis</emphasis>
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</taxonomicName>
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. Tergum X is triangular in lateral view in
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<taxonomicName id="4C534D438206FF87FC517EB9FB91F8AD" ID-CoL="57BG4" box="[938,1091,1769,1794]" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="cupulifera">
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<emphasis id="B927EAD28206FF87FC517EB9FB91F8AD" box="[938,1091,1769,1794]" italics="true" pageId="4" pageNumber="57">T. cupulifera</emphasis>
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</taxonomicName>
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and quadrate in
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<taxonomicName id="4C534D438206FF87FAEE7EB9FE5DF885" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="brasiliensis">
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<emphasis id="B927EAD28206FF87FAEE7EB9FE5DF885" italics="true" pageId="4" pageNumber="57">T. brasiliensis</emphasis>
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</taxonomicName>
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. The wings of
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<taxonomicName id="4C534D438206FF87FDB37F41FD0CF885" ID-CoL="57BG4" box="[584,734,1809,1834]" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="cupulifera">
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<emphasis id="B927EAD28206FF87FDB37F41FD0CF885" box="[584,734,1809,1834]" italics="true" pageId="4" pageNumber="57">T. cupulifera</emphasis>
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</taxonomicName>
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are broader than the wings of
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<taxonomicName id="4C534D438206FF87FBB87F41FB35F885" box="[1091,1255,1809,1834]" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="brasiliensis">
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<emphasis id="B927EAD28206FF87FBB87F41FB35F885" box="[1091,1255,1809,1834]" italics="true" pageId="4" pageNumber="57">T. brasiliensis</emphasis>
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</taxonomicName>
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. The species is known only from
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<collectingCountry id="F34476508206FF87FDAA7F69FD47F8FC" box="[593,661,1849,1875]" name="Chile" pageId="4" pageNumber="57">Chile</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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||
<caption id="DF2C66488207FF86FEF37D09FE5EFA15" httpUri="https://zenodo.org/record/170191/files/figure.png" pageId="5" pageNumber="58" targetBox="[264,1323,291,1318]" targetPageId="5">
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||
<paragraph id="8BEC36C08207FF86FEF37D09FE5EFA15" blockId="5.[264,1323,1369,1466]" pageId="5" pageNumber="58">
|
||
<emphasis id="B927EAD28207FF86FEF37D09FE58FADE" bold="true" box="[264,394,1369,1393]" pageId="5" pageNumber="58">FIGURE 1.</emphasis>
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<emphasis id="B927EAD28207FF86FE6A7D09FDBDFADF" box="[401,623,1369,1392]" italics="true" pageId="5" pageNumber="58">
|
||
<taxonomicName id="4C534D438207FF86FE6A7D09FDB9FADF" box="[401,619,1369,1392]" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="cupulifera">Tolhuaca cupulifera</taxonomicName>
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||
.
|
||
</emphasis>
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Adult: dorsal view of head and thorax.
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<emphasis id="B927EAD28207FF86FEF37D2BFE58FA3C" bold="true" box="[264,394,1403,1427]" pageId="5" pageNumber="58">FIGURE 2.</emphasis>
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||
<emphasis id="B927EAD28207FF86FE6A7D2CFDBDFA3C" box="[401,623,1404,1427]" italics="true" pageId="5" pageNumber="58">
|
||
<taxonomicName id="4C534D438207FF86FE6A7D2CFDB8FA3C" box="[401,618,1404,1427]" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="cupulifera">Tolhuaca cupulifera</taxonomicName>
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||
.
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</emphasis>
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Adult: A––foretibial spur; B––mesotibial spurs, at same scale for comparison.
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</paragraph>
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</caption>
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<subSubSection id="C349654B8207FF8AFEC37DA1FBFEFB61" lastPageId="9" lastPageNumber="62" pageId="5" pageNumber="58" type="description">
|
||
<paragraph id="8BEC36C08207FF86FEC37DA1FC2FF8E4" blockId="5.[264,1323,1521,1867]" pageId="5" pageNumber="58">
|
||
Adult. Length of forewing: male 4.7–5.0 mm (n=5); female 5.0–
|
||
<quantity id="4CAB9B258207FF86FB8C7DA1FB04F9A4" box="[1143,1238,1521,1547]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="58" unit="mm" value="5.3">5.3 mm</quantity>
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(n=3). Body, wings, and appendages nearly uniformly fuscous, tibia and tarsi yellowish brown. Forewing (
|
||
<figureCitation id="13682A458207FF86FE747E11FE0BF9F4" box="[399,473,1601,1627]" captionStart="FIGURE 3" captionStartId="6.[264,368,1778,1802]" captionText="FIGURE 3. Tolhuaca cupulifera Schmid. Wings: A –– forewing; B –– hind wing." pageId="5" pageNumber="58">Fig. 3</figureCitation>
|
||
A) relatively broad, apex blunt; with retrorse setae along veins; Sc incomplete, not reaching anterior margin; fork I emerging at or immediately beyond cord; fork II emerging anteriorly to cord; forks III and IV longer than their stems; A3 forming short loop, intersecting A2 basally; crossveins
|
||
<emphasis id="B927EAD28207FF86FCB77EE9FC84F97D" box="[844,854,1721,1746]" italics="true" pageId="5" pageNumber="58">r</emphasis>
|
||
,
|
||
<emphasis id="B927EAD28207FF86FC907EE9FCA8F97D" box="[875,890,1721,1746]" italics="true" pageId="5" pageNumber="58">d</emphasis>
|
||
,
|
||
<emphasis id="B927EAD28207FF86FC767EE9FC64F97D" box="[909,950,1721,1746]" italics="true" pageId="5" pageNumber="58">rm</emphasis>
|
||
, and
|
||
<emphasis id="B927EAD28207FF86FBF97EE9FBEFF97D" box="[1026,1085,1721,1746]" italics="true" pageId="5" pageNumber="58">mcu</emphasis>
|
||
present, forming a relatively straight line, but with
|
||
<emphasis id="B927EAD28207FF86FD867EB1FD6AF955" box="[637,696,1761,1786]" italics="true" pageId="5" pageNumber="58">mcu</emphasis>
|
||
slightly basal to cord.
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<collectingCountry id="F34476508207FF86FC3C7EB1FBD0F954" box="[967,1026,1761,1787]" name="India" pageId="5" pageNumber="58">Hind</collectingCountry>
|
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wing (
|
||
<figureCitation id="13682A458207FF86FBA87EB1FB45F954" box="[1107,1175,1761,1787]" captionStart="FIGURE 3" captionStartId="6.[264,368,1778,1802]" captionText="FIGURE 3. Tolhuaca cupulifera Schmid. Wings: A –– forewing; B –– hind wing." pageId="5" pageNumber="58">Fig. 3</figureCitation>
|
||
B) relatively broad; Sc converging with R1 near wing margin; fork II about 4 times longer than its stem; fork III longer than its stem; crossveins
|
||
<emphasis id="B927EAD28207FF86FD277F61FD34F8E5" box="[732,742,1841,1866]" italics="true" pageId="5" pageNumber="58">r</emphasis>
|
||
,
|
||
<emphasis id="B927EAD28207FF86FD0F7F61FCCFF8E5" box="[756,797,1841,1866]" italics="true" pageId="5" pageNumber="58">rm</emphasis>
|
||
, and
|
||
<emphasis id="B927EAD28207FF86FCA47F61FC48F8E5" box="[863,922,1841,1866]" italics="true" pageId="5" pageNumber="58">mcu</emphasis>
|
||
present.
|
||
</paragraph>
|
||
<caption id="DF2C66488204FF85FEF37EA2FB89F8A6" box="[264,1115,1778,1802]" pageId="6" pageNumber="59">
|
||
<paragraph id="8BEC36C08204FF85FEF37EA2FB89F8A6" blockId="6.[264,1182,1778,1838]" box="[264,1115,1778,1802]" pageId="6" pageNumber="59">
|
||
<emphasis id="B927EAD28204FF85FEF37EA2FE58F8A5" bold="true" box="[264,394,1778,1802]" pageId="6" pageNumber="59">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C534D438204FF85FE6A7EA2FD15F8A6" authority="Schmid." authorityName="Schmid." box="[401,711,1778,1801]" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="59" phylum="Arthropoda" rank="species" species="cupulifera">
|
||
<emphasis id="B927EAD28204FF85FE6A7EA2FDBAF8A6" box="[401,616,1778,1801]" italics="true" pageId="6" pageNumber="59">Tolhuaca cupulifera</emphasis>
|
||
Schmid.
|
||
</taxonomicName>
|
||
Wings: A––forewing; B––hind wing.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2C66488204FF85FEF37F46FB4CF882" box="[264,1182,1814,1838]" httpUri="https://zenodo.org/record/170192/files/figure.png" pageId="6" pageNumber="59" targetBox="[273,1310,324,1740]" targetPageId="6">
|
||
<paragraph id="8BEC36C08204FF85FEF37F46FB4CF882" blockId="6.[264,1182,1778,1838]" box="[264,1182,1814,1838]" pageId="6" pageNumber="59">
|
||
<emphasis id="B927EAD28204FF85FEF37F46FE58F881" bold="true" box="[264,394,1814,1838]" pageId="6" pageNumber="59">FIGURE 4.</emphasis>
|
||
<emphasis id="B927EAD28204FF85FE6A7F47FDA9F881" box="[401,635,1815,1838]" italics="true" pageId="6" pageNumber="59">
|
||
<taxonomicName id="4C534D438204FF85FE6A7F47FDA5F881" box="[401,631,1815,1838]" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="59" phylum="Arthropoda" rank="species" species="brasiliensis" status="sp. nov.">Tolhuaca brasiliensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A21457A98204FF85FD797F46FCD5F882" box="[642,775,1814,1837]" pageId="6" pageNumber="59" rank="species">new species</taxonomicNameLabel>
|
||
. Wings: A––forewing; B––hind wing.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2C66488205FF84FEF37EA7FD00F8FC" httpUri="https://zenodo.org/record/170193/files/figure.png" pageId="7" pageNumber="60" targetBox="[298,1289,321,1756]" targetPageId="7">
|
||
<paragraph id="8BEC36C08205FF84FEF37EA7FD00F8FC" blockId="7.[264,1323,1783,1875]" pageId="7" pageNumber="60">
|
||
<emphasis id="B927EAD28205FF84FEF37EA7FE5FF8A0" bold="true" box="[264,397,1783,1807]" pageId="7" pageNumber="60">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C534D438205FF84FE637EA7FD07F8A1" authority="Schmid." authorityName="Schmid." box="[408,725,1783,1806]" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="cupulifera">
|
||
<emphasis id="B927EAD28205FF84FE637EA7FDA0F8A1" box="[408,626,1783,1806]" italics="true" pageId="7" pageNumber="60">Tolhuaca cupulifera</emphasis>
|
||
Schmid.
|
||
</taxonomicName>
|
||
Male genitalia: A––lateral; B––dorsal; C––process of sternum VI; D––endotheca fully everted as observed in holotype; E––detail of endothecal thornlike spines; F––ventral view of phallobase.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2C6648820AFF8BFEF37E7DFAD6F9EB" box="[264,1284,1581,1605]" httpUri="https://zenodo.org/record/170194/files/figure.png" pageId="8" pageNumber="61" targetBox="[290,1294,287,1549]" targetPageId="8">
|
||
<paragraph id="8BEC36C0820AFF8BFEF37E7DFAD6F9EB" blockId="8.[264,1284,1581,1605]" box="[264,1284,1581,1605]" pageId="8" pageNumber="61">
|
||
<emphasis id="B927EAD2820AFF8BFEF37E7DFE59F9EA" bold="true" box="[264,395,1581,1605]" pageId="8" pageNumber="61">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C534D43820AFF8BFE6A7E7EFD15F9EB" authority="Schmid." authorityName="Schmid." box="[401,711,1581,1605]" class="Insecta" family="Glossosomatidae" genus="Tolhuaca" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="cupulifera">
|
||
<emphasis id="B927EAD2820AFF8BFE6A7E7EFDBAF9EA" box="[401,616,1582,1605]" italics="true" pageId="8" pageNumber="61">Tolhuaca cupulifera</emphasis>
|
||
Schmid.
|
||
</taxonomicName>
|
||
Female genitalia: A––lateral; B––dorsal; C––ventral.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEC36C0820AFF8AFEC37E2CFD5BFDE1" blockId="8.[264,1323,1660,1886]" lastBlockId="9.[264,1325,284,1230]" lastPageId="9" lastPageNumber="62" pageId="8" pageNumber="61">
|
||
Male genitalia (
|
||
<figureCitation id="13682A45820AFF8BFE067E2CFD98F939" box="[509,586,1660,1686]" captionStart="FIGURE 5" captionStartId="7.[264,368,1783,1807]" captionTargetBox="[298,1289,321,1756]" captionTargetId="figure@7.[298,1289,321,1757]" captionTargetPageId="7" captionText="FIGURE 5. Tolhuaca cupulifera Schmid. Male genitalia: A –– lateral; B –– dorsal; C –– process of sternum VI; D –– endotheca fully everted as observed in holotype; E –– detail of endothecal thornlike spines; F –– ventral view of phallobase." httpUri="https://zenodo.org/record/170193/files/figure.png" pageId="8" pageNumber="61">Fig. 5</figureCitation>
|
||
). Sternum VI (
|
||
<figureCitation id="13682A45820AFF8BFCE17E2CFCB7F939" box="[794,869,1660,1686]" captionStart="FIGURE 5" captionStartId="7.[264,368,1783,1807]" captionTargetBox="[298,1289,321,1756]" captionTargetId="figure@7.[298,1289,321,1757]" captionTargetPageId="7" captionText="FIGURE 5. Tolhuaca cupulifera Schmid. Male genitalia: A –– lateral; B –– dorsal; C –– process of sternum VI; D –– endotheca fully everted as observed in holotype; E –– detail of endothecal thornlike spines; F –– ventral view of phallobase." httpUri="https://zenodo.org/record/170193/files/figure.png" pageId="8" pageNumber="61">Fig. 5</figureCitation>
|
||
C) with thin, digitate mesal process, projecting caudally. Abdominal segment IX (
|
||
<figureCitation id="13682A45820AFF8BFCB47EF4FC49F911" box="[847,923,1700,1726]" captionStart="FIGURE 5" captionStartId="7.[264,368,1783,1807]" captionTargetBox="[298,1289,321,1756]" captionTargetId="figure@7.[298,1289,321,1757]" captionTargetPageId="7" captionText="FIGURE 5. Tolhuaca cupulifera Schmid. Male genitalia: A –– lateral; B –– dorsal; C –– process of sternum VI; D –– endotheca fully everted as observed in holotype; E –– detail of endothecal thornlike spines; F –– ventral view of phallobase." httpUri="https://zenodo.org/record/170193/files/figure.png" pageId="8" pageNumber="61">Fig. 5</figureCitation>
|
||
A) well developed dorsally and laterally, but extremely reduced ventrally, forming very thin sclerotized strap; anterior margin slightly rounded; tergum IX, in dorsal view (
|
||
<figureCitation id="13682A45820AFF8BFC7B7EA4FC1AF8A1" box="[896,968,1780,1806]" captionStart="FIGURE 5" captionStartId="7.[264,368,1783,1807]" captionTargetBox="[298,1289,321,1756]" captionTargetId="figure@7.[298,1289,321,1757]" captionTargetPageId="7" captionText="FIGURE 5. Tolhuaca cupulifera Schmid. Male genitalia: A –– lateral; B –– dorsal; C –– process of sternum VI; D –– endotheca fully everted as observed in holotype; E –– detail of endothecal thornlike spines; F –– ventral view of phallobase." httpUri="https://zenodo.org/record/170193/files/figure.png" pageId="8" pageNumber="61">Fig. 5</figureCitation>
|
||
B), with posteromesal margin triangular, covered with fine microtrichia; membranous connection between segments IX and X distinct. Tergum X (
|
||
<figureCitation id="13682A45820AFF8BFDAE7F14FD75F8F1" box="[597,679,1860,1886]" captionStart="FIGURE 5" captionStartId="7.[264,368,1783,1807]" captionTargetBox="[298,1289,321,1756]" captionTargetId="figure@7.[298,1289,321,1757]" captionTargetPageId="7" captionText="FIGURE 5. Tolhuaca cupulifera Schmid. Male genitalia: A –– lateral; B –– dorsal; C –– process of sternum VI; D –– endotheca fully everted as observed in holotype; E –– detail of endothecal thornlike spines; F –– ventral view of phallobase." httpUri="https://zenodo.org/record/170193/files/figure.png" pageId="8" pageNumber="61">Figs. 5</figureCitation>
|
||
A, 5B) covered with fine microtrichia; in lateral view (
|
||
<figureCitation id="13682A45820BFF8AFEEA794CFE8BFE99" box="[273,345,284,310]" captionStart="FIGURE 5" captionStartId="7.[264,368,1783,1807]" captionTargetBox="[298,1289,321,1756]" captionTargetId="figure@7.[298,1289,321,1757]" captionTargetPageId="7" captionText="FIGURE 5. Tolhuaca cupulifera Schmid. Male genitalia: A –– lateral; B –– dorsal; C –– process of sternum VI; D –– endotheca fully everted as observed in holotype; E –– detail of endothecal thornlike spines; F –– ventral view of phallobase." httpUri="https://zenodo.org/record/170193/files/figure.png" pageId="9" pageNumber="62">Fig. 5</figureCitation>
|
||
A), triangular, apex rounded, setose dorsally to ventrally, with acute ventromesal process; in dorsal view (
|
||
<figureCitation id="13682A45820BFF8AFDC27914FD56FEF1" box="[569,644,324,350]" captionStart="FIGURE 5" captionStartId="7.[264,368,1783,1807]" captionTargetBox="[298,1289,321,1756]" captionTargetId="figure@7.[298,1289,321,1757]" captionTargetPageId="7" captionText="FIGURE 5. Tolhuaca cupulifera Schmid. Male genitalia: A –– lateral; B –– dorsal; C –– process of sternum VI; D –– endotheca fully everted as observed in holotype; E –– detail of endothecal thornlike spines; F –– ventral view of phallobase." httpUri="https://zenodo.org/record/170193/files/figure.png" pageId="9" pageNumber="62">Fig. 5</figureCitation>
|
||
B), with lateral margins slightly sinuous; apex strongly excavate medially. Phallobase (
|
||
<figureCitation id="13682A45820BFF8AFD79793CFD01FE29" box="[642,723,364,390]" captionStart="FIGURE 5" captionStartId="7.[264,368,1783,1807]" captionTargetBox="[298,1289,321,1756]" captionTargetId="figure@7.[298,1289,321,1757]" captionTargetPageId="7" captionText="FIGURE 5. Tolhuaca cupulifera Schmid. Male genitalia: A –– lateral; B –– dorsal; C –– process of sternum VI; D –– endotheca fully everted as observed in holotype; E –– detail of endothecal thornlike spines; F –– ventral view of phallobase." httpUri="https://zenodo.org/record/170193/files/figure.png" pageId="9" pageNumber="62">Figs. 5</figureCitation>
|
||
A, 5F) large and bulbous, projecting apicodorsally, lightly sclerotized, with small, stout setae at posterior end, running transversally from dorsum to venter; endotheca (
|
||
<figureCitation id="13682A45820BFF8AFD8E79ECFD18FE79" box="[629,714,444,470]" captionStart="FIGURE 5" captionStartId="7.[264,368,1783,1807]" captionTargetBox="[298,1289,321,1756]" captionTargetId="figure@7.[298,1289,321,1757]" captionTargetPageId="7" captionText="FIGURE 5. Tolhuaca cupulifera Schmid. Male genitalia: A –– lateral; B –– dorsal; C –– process of sternum VI; D –– endotheca fully everted as observed in holotype; E –– detail of endothecal thornlike spines; F –– ventral view of phallobase." httpUri="https://zenodo.org/record/170193/files/figure.png" pageId="9" pageNumber="62">Figs. 5</figureCitation>
|
||
A, 5D) membranous except for lightly sclerotized subbasal ring, greatly enlarged and tubular when evaginated, with 12 or 13 heavily sclerotized conical, thornlike spines with enlarged bases, each sitting within membranous ovoid cupule (
|
||
<figureCitation id="13682A45820BFF8AFE4B7A64FDD3FDE1" box="[432,513,564,590]" captionStart="FIGURE 5" captionStartId="7.[264,368,1783,1807]" captionTargetBox="[298,1289,321,1756]" captionTargetId="figure@7.[298,1289,321,1757]" captionTargetPageId="7" captionText="FIGURE 5. Tolhuaca cupulifera Schmid. Male genitalia: A –– lateral; B –– dorsal; C –– process of sternum VI; D –– endotheca fully everted as observed in holotype; E –– detail of endothecal thornlike spines; F –– ventral view of phallobase." httpUri="https://zenodo.org/record/170193/files/figure.png" pageId="9" pageNumber="62">Figs. 5</figureCitation>
|
||
A, 5D, 5E).
|
||
</paragraph>
|
||
<paragraph id="8BEC36C0820BFF8AFEC37A0CFE7AFCC9" blockId="9.[264,1325,284,1230]" pageId="9" pageNumber="62">
|
||
Female genitalia (
|
||
<figureCitation id="13682A45820BFF8AFDE97A0CFD8CFDD9" box="[530,606,604,630]" captionStart="FIGURE 6" captionStartId="8.[264,368,1581,1605]" captionTargetBox="[290,1294,287,1549]" captionTargetId="figure@8.[290,1294,287,1550]" captionTargetPageId="8" captionText="FIGURE 6. Tolhuaca cupulifera Schmid. Female genitalia: A –– lateral; B –– dorsal; C –– ventral." httpUri="https://zenodo.org/record/170194/files/figure.png" pageId="9" pageNumber="62">Fig. 6</figureCitation>
|
||
). Sternum V with slightly raised, sclerotized linear mesal ridge running parallel to anterior and posterior margins. Sternum VI process associated with strongly oblique apodeme. Segment VII normally developed. Abdominal segment VIII synscleritous, more distinctly sclerotized along anterior margin, venter lightly sclerotized, becoming membranous posteroventrally. Tergum IX lightly sclerotized. Segment X, in lateral view, elongate, digitate; in dorsal and ventral views, bulbous, bearing cerci.
|
||
</paragraph>
|
||
<paragraph id="8BEC36C0820BFF8AFEC37B24FEAFFBD1" blockId="9.[264,1325,284,1230]" pageId="9" pageNumber="62">
|
||
Material examined:
|
||
<emphasis id="B927EAD2820BFF8AFDBA7B24FCFAFC21" bold="true" box="[577,808,884,910]" pageId="9" pageNumber="62">
|
||
<collectingCountry id="F3447650820BFF8AFDBA7B24FD76FC21" box="[577,676,884,910]" name="Chile" pageId="9" pageNumber="62">CHILE</collectingCountry>
|
||
: BíoBío:
|
||
</emphasis>
|
||
Arauco, Pichinahuel, [ca.
|
||
<geoCoordinate id="EE675007820BFF8AFB607B24FAF7FC21" box="[1179,1317,884,910]" direction="south" orientation="latitude" pageId="9" pageNumber="62" precision="15" value="-37.79778">37º47’52”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE675007820BFF8AFEF37BCCFE7DFC19" box="[264,431,924,950]" direction="west" orientation="longitude" pageId="9" pageNumber="62" precision="15" value="-73.04361">073º02’37”W</geoCoordinate>
|
||
],
|
||
<date id="FFED1000820BFF8AFE337BCCFD84FC19" box="[456,598,924,950]" pageId="9" pageNumber="62" value="1959-01-01" valueMax="1959-01-30" valueMin="1959-01-01">1–30.i.1959</date>
|
||
, Peña —
|
||
<specimenCount id="9D55FD49820BFF8AFD227BCCFCFCFC19" box="[729,814,924,950]" pageId="9" pageNumber="62" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus id="54E88862820BFF8AFCB87BCCFC7FFC19" box="[835,941,924,950]" pageId="9" pageNumber="62" type="holotype">holotype</typeStatus>
|
||
) (UMSP000115157) (NMNH);
|
||
<emphasis id="B927EAD2820BFF8AFEF37B94FEA2FC71" bold="true" box="[264,368,964,990]" pageId="9" pageNumber="62">BíoBío:</emphasis>
|
||
Estero Pichinahuel,
|
||
<quantity id="4CAB9B25820BFF8AFD9D7B94FD71FC71" box="[614,675,964,990]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="62" unit="km" value="2.0">2 km</quantity>
|
||
W Parque Nacional Nahuelbuta entrance,
|
||
<geoCoordinate id="EE675007820BFF8AFB677B94FAF6FC71" box="[1180,1316,964,990]" direction="south" orientation="latitude" pageId="9" pageNumber="62" precision="15" value="-37.79778">37°47'52"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE675007820BFF8AFEF37BBCFE75FBA9" box="[264,423,1004,1030]" direction="west" orientation="longitude" pageId="9" pageNumber="62" precision="15" value="-73.04361">073°02'37"W</geoCoordinate>
|
||
,
|
||
<quantity id="4CAB9B25820BFF8AFE427BBCFDCEFBA9" box="[441,540,1004,1030]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="9" pageNumber="62" unit="m" value="1070.0">1070 m</quantity>
|
||
,
|
||
<date id="FFED1000820BFF8AFDD57BBCFD03FBA9" box="[558,721,1004,1030]" pageId="9" pageNumber="62" value="2005-02-11" valueMax="2005-02-12" valueMin="2005-02-11">11–12.ii.2005</date>
|
||
, Holzenthal, Blahnik, Chamorro —
|
||
<specimenCount id="9D55FD49820BFF8AFB607BBCFAD8FBA9" box="[1179,1290,1004,1030]" pageId="9" pageNumber="62" type="female">1 female</specimenCount>
|
||
,
|
||
<specimenCount id="9D55FD49820BFF8AFAE77BBCFE9EFB81" pageId="9" pageNumber="62" type="male">4 males</specimenCount>
|
||
(UMSP);
|
||
<emphasis id="B927EAD2820BFF8AFE327C44FDE4FB81" bold="true" box="[457,566,1044,1070]" pageId="9" pageNumber="62">Malleco:</emphasis>
|
||
Cord. Nahuelbuta, Cabreria, [
|
||
<geoCoordinate id="EE675007820BFF8AFC607C44FBFAFB81" box="[923,1064,1044,1070]" direction="south" orientation="latitude" pageId="9" pageNumber="62" precision="15" value="-37.825">37º49’30”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE675007820BFF8AFBCD7C44FB0FFB81" box="[1078,1245,1044,1070]" direction="west" orientation="longitude" pageId="9" pageNumber="62" precision="15" value="-73.01667">073º01’00”W</geoCoordinate>
|
||
],
|
||
<quantity id="4CAB9B25820BFF8AFB0A7C44FEF3FBF9" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="62" unit="m" value="1100.0">1100 m</quantity>
|
||
,
|
||
<date id="FFED1000820BFF8AFECB7C6CFE6CFBF9" box="[304,446,1084,1110]" pageId="9" pageNumber="62" value="1977-01-09" valueMax="1977-01-15" valueMin="1977-01-09">9–15.i.1977</date>
|
||
, Peña —
|
||
<specimenCount id="9D55FD49820BFF8AFDCC7C6CFD4DFBF9" box="[567,671,1084,1110]" pageId="9" pageNumber="62" type="female">1 female</specimenCount>
|
||
(NMNH); same except
|
||
<date id="FFED1000820BFF8AFC3F7C6CFBB2FBF9" box="[964,1120,1084,1110]" pageId="9" pageNumber="62" value="1977-01-15" valueMax="1977-01-20" valueMin="1977-01-15">15–20.i.1977</date>
|
||
, Peña —
|
||
<specimenCount id="9D55FD49820BFF8AFB227C6CFAF9FBF9" box="[1241,1323,1084,1110]" pageId="9" pageNumber="62" type="male">1 male</specimenCount>
|
||
(NMNH).
|
||
</paragraph>
|
||
<paragraph id="8BEC36C0820BFF8AFEC37CDCFBFEFB61" blockId="9.[264,1325,284,1230]" pageId="9" pageNumber="62">
|
||
Distribution.
|
||
<collectingCountry id="F3447650820BFF8AFE1D7CDCFDFBFB09" box="[486,553,1164,1190]" name="Chile" pageId="9" pageNumber="62">Chile</collectingCountry>
|
||
. This species occurs in the Cordillera Nahuelbuta region and extends south to Valdivia based on records provided by
|
||
<bibRefCitation id="EFC24B31820BFF8AFC6C7CE4FBFAFB61" author="Flint" box="[919,1064,1204,1230]" pageId="9" pageNumber="67" refString="Flint, O. S., Jr. (1967) Studies of Neotropical caddis flies, II: Trichoptera collected by Prof. Dr. J. Illies in the Chilean Subregion. Beitrage zur Neotropischen Fauna, 5, 45 - 68." type="journal article" year="1967">Flint (1967)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |