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<document id="D2915E73E5D56325A124DC0450E5F8E6" ID-CLB-Dataset="68519" ID-DOI="10.11646/zootaxa.5150.2.5" ID-GBIF-Dataset="e099b088-6bbc-42eb-a57b-84bdfe32f64e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6621301" ID-ZooBank="D15CA296-D186-43EA-90D3-4979596A46F0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654625951008" checkinUser="plazi" docAuthor="Rojas-Buffet, Carolina, Bouzan, Rodrigo Salvador, Sierwald, Petra, Brescovit, Antonio Do- Mingos &amp; Simó, Miguel" docDate="2022" docId="03FA87BBFFDCFFB9D6B93E4205950028" docLanguage="en" docName="zootaxa.5150.2.5.pdf" docOrigin="Zootaxa 5150 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sandalodesmus Silvestri 1902" docType="key" docVersion="7" lastPageNumber="266" masterDocId="FFC3FFC3FFDAFFBFD62E3D250009074D" masterDocTitle="Sandalodesmus joachimadisi n. sp., the first Chelodesmidae member from Uruguay with an updated key to species of the genus (Diplopoda: Polydesmida)" masterLastPageNumber="268" masterPageNumber="260" pageNumber="266" updateTime="1699339059308" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="060E2C422EC00088D3ACB33EECB9CBC2">Sandalodesmus joachimadisi n. sp., the first Chelodesmidae member from Uruguay with an updated key to species of the genus (Diplopoda: Polydesmida)</mods:title>
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<mods:namePart id="3377F58147AC192067B162DD5A2B8155">Rojas-Buffet, Carolina</mods:namePart>
<mods:affiliation id="D7B9A75955655F26AFC368A9A0E069E5">Sección Entomología, Facultad de Ciencias, Universidad de la República, Montevideo, Uruguay. &amp; Departamento de Ecología y Biología Evolutiva, Instituto de Investigaciones Biológicas Clemente Estable, Ministerio de Educación y Cultura, Montevideo, Uruguay.</mods:affiliation>
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<mods:namePart id="1A8158E980B0F6D6C61064641806E511">Bouzan, Rodrigo Salvador</mods:namePart>
<mods:nameIdentifier id="B6BF3AA54700E66CEE06D61DEC908F8D" type="ORCID">0000-0002-5331-7031</mods:nameIdentifier>
<mods:affiliation id="9546559554284733CF5110DFD3846A3E">Laboratório de Coleções Zoológicas, Instituto Butantan, São Paulo, Brazil. &amp; Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. &amp; rodrigobouzan @ outlook. com; https: // orcid. org / 0000 - 0002 - 5331 - 7031</mods:affiliation>
<mods:nameIdentifier id="F7B3D059C3AB1F3E6AAFE91F0270F1DD" type="email">rodrigobouzan@outlook.com</mods:nameIdentifier>
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<mods:namePart id="C2B4EC3533DAD1E7BE298C4E4DFBE43B">Sierwald, Petra</mods:namePart>
<mods:nameIdentifier id="1A05C78DE135EBD1516C22617C12A2C7" type="ORCID">0000-0003-2592-1298</mods:nameIdentifier>
<mods:affiliation id="71310F3EE7183A4384B0068D60300EB7">Insect Division, Field Museum of Natural History, Chicago, IL. USA. &amp; psierwald @ fieldmuseum. org; https: // orcid. org / 0000 - 0003 - 2592 - 1298</mods:affiliation>
<mods:nameIdentifier id="87D3213B3177E5A29BD7E9BA35DA8F3F" type="email">psierwald@fieldmuseum.org</mods:nameIdentifier>
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<mods:namePart id="2238619947079355ACD0FFD1B76E5FAA">Brescovit, Antonio Do- Mingos</mods:namePart>
<mods:nameIdentifier id="A5D769F52EABD2A170DF3302B56A5DA9" type="ORCID">0000-0002-1511-5324</mods:nameIdentifier>
<mods:affiliation id="548D5DC028634AE6BA388DC3F5B96651">Laboratório de Coleções Zoológicas, Instituto Butantan, São Paulo, Brazil. &amp; antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324</mods:affiliation>
<mods:nameIdentifier id="CC2B3559CC2169E83B5E4BC3B69AF761" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<mods:namePart id="38D056E9EF53CEA213C661B5364BA55B">Simó, Miguel</mods:namePart>
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<mods:affiliation id="CA54DDB9828F544A12B5FE6A1971CAB2">Sección Entomología, Facultad de Ciencias, Universidad de la República, Montevideo, Uruguay. &amp; simo @ fcien. edu. uy; https: // orcid. org / 0000 - 0002 - 9477 - 5880</mods:affiliation>
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<mods:title id="E18A0802E5CF7C3490EC0D49ECB348AC">Zootaxa</mods:title>
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<mods:date id="431E9E0AB7A6BF073A88005401499CD2">2022</mods:date>
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<treatment id="03FA87BBFFDCFFB9D6B93E4205950028" ID-DOI="http://doi.org/10.5281/zenodo.6628478" ID-GBIF-Taxon="195827681" ID-Zenodo-Dep="6628478" LSID="urn:lsid:plazi:treatment:03FA87BBFFDCFFB9D6B93E4205950028" httpUri="http://treatment.plazi.org/id/03FA87BBFFDCFFB9D6B93E4205950028" lastPageNumber="266" pageId="6" pageNumber="266">
<subSubSection id="C3496526FFDCFFB9D6B93E42039604CD" box="[151,927,870,897]" pageId="6" pageNumber="266" type="nomenclature">
<paragraph id="8BEC36ADFFDCFFB9D6B93E42039604CD" blockId="6.[151,927,870,897]" box="[151,927,870,897]" pageId="6" pageNumber="266">
<heading id="D0A481C1FFDCFFB9D6B93E42039604CD" bold="true" box="[151,927,870,897]" fontSize="11" level="1" pageId="6" pageNumber="266" reason="1">
<emphasis id="B927EABFFFDCFFB9D6B93E42039604CD" bold="true" box="[151,927,870,897]" pageId="6" pageNumber="266">
Key to males of
<taxonomicName id="4C534D2EFFDCFFB9D74E3E43021404CC" ID-CoL="7PYLQ" authorityName="Silvestri" authorityYear="1902" box="[352,541,870,897]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="genus">
<emphasis id="B927EABFFFDCFFB9D74E3E43021404CC" bold="true" box="[352,541,870,897]" italics="true" pageId="6" pageNumber="266">Sandalodesmus</emphasis>
</taxonomicName>
(based on gonopod structures)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3496526FFDCFFB9D6B93E8005950028" pageId="6" pageNumber="266" type="key">
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<paragraph id="8BEC36ADFFDCFFB9D6B93E80059504F7" blockId="6.[151,1437,933,1894]" box="[151,1436,933,955]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93E80059504F7" box="[151,1436,933,955]" pageId="6" pageNumber="266">1 Acropodite process with a prominent, subtriangular secondary process on the lateral side............................ 2</keyLead>
</paragraph>
<paragraph id="8BEC36ADFFDCFFB9D6B93EE60595049A" blockId="6.[151,1437,933,1894]" box="[151,1436,962,984]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93EE60595049A" box="[151,1436,962,984]" pageId="6" pageNumber="266">- Acropodite process without a secondary process............................................................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E08FFDCFFB9D6B93EC50595037D" pageId="6" pageNumber="266">
<paragraph id="8BEC36ADFFDCFFB9D6B93EC5059504BB" blockId="6.[151,1437,933,1894]" box="[151,1436,992,1014]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93EC5059504BB" box="[151,1436,992,1014]" pageId="6" pageNumber="266">
2 (1) Acropodite process rounded, curved mesad (see
<bibRefCitation id="EFC24B5CFFDCFFB9D49A3EC5034804BB" author="Schubart, O." box="[692,833,992,1014]" pageId="6" pageNumber="266" pagination="165 - 202" refId="ref6122" refString="Schubart, O. (1946) Contribuicao ao conhecimento do genero Leptodesmus (Familia Leptodesmidae, Diplopoda). Anais da Academia Brasileira de Ciencias, 18, 165 - 202." type="journal article" year="1946">Schubart 1946</bibRefCitation>
, fig. 4).....................
<taxonomicName id="4C534D2EFFDCFFB9D2B83EC5059504BB" ID-CoL="79HJ9" authority="(Schubart, 1946)" baseAuthorityName="Schubart" baseAuthorityYear="1946" box="[1174,1436,992,1014]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="araujoi">
<emphasis id="B927EABFFFDCFFB9D2B83EC504FC04B8" box="[1174,1269,992,1013]" italics="true" pageId="6" pageNumber="266">S. araujoi</emphasis>
(
<bibRefCitation id="EFC24B5CFFDCFFB9D32F3EC5059C04BB" author="Schubart, O." box="[1281,1429,992,1014]" pageId="6" pageNumber="266" pagination="165 - 202" refId="ref6122" refString="Schubart, O. (1946) Contribuicao ao conhecimento do genero Leptodesmus (Familia Leptodesmidae, Diplopoda). Anais da Academia Brasileira de Ciencias, 18, 165 - 202." type="journal article" year="1946">Schubart, 1946</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEC36ADFFDCFFB9D6B93ED80595037D" blockId="6.[151,1437,933,1894]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93ED80595037D" pageId="6" pageNumber="266">
- Acropodite process subtriangular, projected up (see
<bibRefCitation id="EFC24B5CFFDCFFB9D4FB3ED80354035E" author="Silvestri, F." box="[725,861,1021,1043]" pageId="6" pageNumber="266" pagination="1 - 12" refId="ref6358" refString="Silvestri, F. (1895) Viaggio del dottor Alfredo Borelli nella Repubblica Argentina e nel Paraguay. XIV. Chilopodi e Diplopodi. Bollettino del musei di zoologia e di anatomia comparata della Reale Universita di Torino, 10 (203), 1 - 12. https: // doi. org / 10.5962 / bhl. part. 8048" type="journal article" year="1895">Silvestri 1895</bibRefCitation>
, fig. 3;
<bibRefCitation id="EFC24B5CFFDCFFB9D5893ED8043C035F" author="Hoffman, R. L." box="[935,1077,1021,1043]" pageId="6" pageNumber="266" pagination="247 - 259" refId="ref5695" refString="Hoffman, R. L. (1982) Chelodesmid studies. XVIII. A synopsis of the genus Sandalodesmus Silvestri, 1902, and proposal of the new tribe Sandalodesmini (Polydesmida: Chelodesmidae). Spixiana, 5 (3), 247 - 259. https: // www. biodiversitylibrary. org / page / 28194567" type="journal article" year="1982">Hoffman 1982</bibRefCitation>
, figs 1, 2).................................................................................................
<taxonomicName id="4C534D2EFFDCFFB9D2AF393E0595037D" ID-CoL="79HLG" authority="(Silvestri, 1895)" baseAuthorityName="Silvestri" baseAuthorityYear="1895" box="[1153,1436,1050,1072]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="salvadorii">
<emphasis id="B927EABFFFDCFFB9D2AF393E04F30362" box="[1153,1274,1050,1072]" italics="true" pageId="6" pageNumber="266">S. salvadorii</emphasis>
(
<bibRefCitation id="EFC24B5CFFDCFFB9D329393F059C037D" author="Silvestri, F." box="[1287,1429,1050,1072]" pageId="6" pageNumber="266" pagination="1 - 12" refId="ref6358" refString="Silvestri, F. (1895) Viaggio del dottor Alfredo Borelli nella Repubblica Argentina e nel Paraguay. XIV. Chilopodi e Diplopodi. Bollettino del musei di zoologia e di anatomia comparata della Reale Universita di Torino, 10 (203), 1 - 12. https: // doi. org / 10.5962 / bhl. part. 8048" type="journal article" year="1895">Silvestri, 1895</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E08FFDCFFB9D6B9391D059503CA" pageId="6" pageNumber="266">
<paragraph id="8BEC36ADFFDCFFB9D6B9391D05950300" blockId="6.[151,1437,933,1894]" box="[151,1436,1080,1102]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B9391D05950300" box="[151,1436,1080,1102]" pageId="6" pageNumber="266">3 (1) Prefemur short, less than half of the total length of telopodite, not extending distad beyond base of the prefemoral process.. 4</keyLead>
</paragraph>
<paragraph id="8BEC36ADFFDCFFB9D6B93970059503CA" blockId="6.[151,1437,933,1894]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93970059503CA" pageId="6" pageNumber="266">- Prefemur longer, more than half of the total length of telopodite, extending distad to about midlength of prefemoral process.. ................................................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E08FFDCFFB9D6B939B5059503AD" pageId="6" pageNumber="266">
<paragraph id="8BEC36ADFFDCFFB9D6B939B50595038E" blockId="6.[151,1437,933,1894]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B939B50595038E" pageId="6" pageNumber="266">
4 (3) Acropodite short, scarcely longer than prefemoral process, its distal edge exceeded by solenomere (see
<bibRefCitation id="EFC24B5CFFDCFFB9D29839B5054703E8" author="Schubart, O." box="[1206,1358,1167,1190]" pageId="6" pageNumber="266" pagination="77 - 132" refId="ref6155" refString="Schubart, O. (1954 a) Sobre os Diplopoda dos estados do Parana e Santa Catarina. I. Proterospermophora. Arquivos do Museu Paranaense, 10, 77 - 132." type="journal article" year="1954">Schubart 1954a</bibRefCitation>
, fig. 16) ….........................................................................
<taxonomicName id="4C534D2EFFDCFFB9D25B39880595038E" ID-CoL="9368X" authority="(Schubart, 1954)" baseAuthorityName="Schubart" baseAuthorityYear="1954" box="[1141,1436,1197,1219]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="camellatus">
<emphasis id="B927EABFFFDCFFB9D25B398804FC038F" box="[1141,1269,1197,1218]" italics="true" pageId="6" pageNumber="266">S. camellatus</emphasis>
(Schubart, 1954)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEC36ADFFDCFFB9D6B939EE059503AD" blockId="6.[151,1437,933,1894]" box="[151,1436,1226,1248]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B939EE059503AD" box="[151,1436,1226,1248]" pageId="6" pageNumber="266">- Acropodite long, forming a concave shield that contains the solenomere......................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E08FFDCFFB9D6B939C205950275" pageId="6" pageNumber="266">
<paragraph id="8BEC36ADFFDCFFB9D6B939C2059503B3" blockId="6.[151,1437,933,1894]" box="[151,1436,1255,1278]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B939C2059503B3" box="[151,1436,1255,1278]" pageId="6" pageNumber="266">5 (4) Prefemoral process shorter than acropodite................................................................. 6</keyLead>
</paragraph>
<paragraph id="8BEC36ADFFDCFFB9D6B9382005950275" blockId="6.[151,1437,933,1894]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B9382005950275" pageId="6" pageNumber="266">
- Prefemoral process as long as acropodite, apically broad and rounded in mesal aspect, without a basal lobe; acropodite with a prominent spiculate lobe on the lateral side (see
<bibRefCitation id="EFC24B5CFFDCFFB9D49C380703360275" author="Schubart, O." box="[690,831,1314,1336]" pageId="6" pageNumber="266" pagination="23 - 32" refId="ref6273" refString="Schubart, O. (1958) &quot; Leptodesmidae &quot; Brasileiras. VI: Especies novas do Estado do Rio Grande do Sul (Diplopoda, Proterosper- mophora). Revista brasileira de biologia, 18 (1), 23 - 32." type="journal article" year="1958">Schubart 1958</bibRefCitation>
, fig. 4)....................
<taxonomicName id="4C534D2EFFDCFFB9D2AA380605950275" ID-CoL="6XHLG" authority="(Schubart, 1958)" authorityName="" baseAuthorityName="Schubart" baseAuthorityYear="1958" box="[1156,1436,1314,1336]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="repandus">
<emphasis id="B927EABFFFDCFFB9D2AA380604FD027A" box="[1156,1268,1314,1336]" italics="true" pageId="6" pageNumber="266">S. repandus</emphasis>
(
<bibRefCitation id="EFC24B5CFFDCFFB9D32F3807059C0275" author="Schubart, O." box="[1281,1429,1314,1336]" pageId="6" pageNumber="266" pagination="23 - 32" refId="ref6273" refString="Schubart, O. (1958) &quot; Leptodesmidae &quot; Brasileiras. VI: Especies novas do Estado do Rio Grande do Sul (Diplopoda, Proterosper- mophora). Revista brasileira de biologia, 18 (1), 23 - 32." type="journal article" year="1958">Schubart, 1958</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E08FFDCFFB9D6B938650595023F" pageId="6" pageNumber="266">
<paragraph id="8BEC36ADFFDCFFB9D6B9386505950218" blockId="6.[151,1437,933,1894]" box="[151,1436,1343,1366]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B9386505950218" box="[151,1436,1343,1366]" pageId="6" pageNumber="266">6 (5) Nontorsate prefemoral process.......................................................................... 7</keyLead>
</paragraph>
<paragraph id="8BEC36ADFFDCFFB9D6B938780595023F" blockId="6.[151,1437,933,1894]" box="[151,1436,1373,1395]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B938780595023F" box="[151,1436,1373,1395]" pageId="6" pageNumber="266">- Torsate prefemoral process............................................................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E08FFDCFFB9D6B9385E059502E0" pageId="6" pageNumber="266">
<paragraph id="8BEC36ADFFDCFFB9D6B9385E059502C2" blockId="6.[151,1437,933,1894]" box="[151,1436,1402,1424]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B9385E059502C2" box="[151,1436,1402,1424]" pageId="6" pageNumber="266">
7 (6) Solenomere massive and apically reflexed (see
<bibRefCitation id="EFC24B5CFFDCFFB9D484385F033102C2" author="Hoffman, R. L." box="[682,824,1402,1424]" pageId="6" pageNumber="266" pagination="247 - 259" refId="ref5695" refString="Hoffman, R. L. (1982) Chelodesmid studies. XVIII. A synopsis of the genus Sandalodesmus Silvestri, 1902, and proposal of the new tribe Sandalodesmini (Polydesmida: Chelodesmidae). Spixiana, 5 (3), 247 - 259. https: // www. biodiversitylibrary. org / page / 28194567" type="journal article" year="1982">Hoffman 1982</bibRefCitation>
, fig. 5)......................
<taxonomicName id="4C534D2EFFDCFFB9D2B9385E059502C2" ID-CoL="6XHJB" authority="Hoffman, 1982" authorityName="Hoffman" authorityYear="1982" box="[1175,1436,1402,1424]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="avilectus">
<emphasis id="B927EABFFFDCFFB9D2B9385E050D02C2" box="[1175,1284,1402,1424]" italics="true" pageId="6" pageNumber="266">S. avilectus</emphasis>
<bibRefCitation id="EFC24B5CFFDCFFB9D324385F059502C2" author="Hoffman, R. L." box="[1290,1436,1402,1424]" pageId="6" pageNumber="266" pagination="247 - 259" refId="ref5695" refString="Hoffman, R. L. (1982) Chelodesmid studies. XVIII. A synopsis of the genus Sandalodesmus Silvestri, 1902, and proposal of the new tribe Sandalodesmini (Polydesmida: Chelodesmidae). Spixiana, 5 (3), 247 - 259. https: // www. biodiversitylibrary. org / page / 28194567" type="journal article" year="1982">Hoffman, 1982</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEC36ADFFDCFFB9D6B938BD059502E0" blockId="6.[151,1437,933,1894]" box="[151,1436,1431,1454]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B938BD059502E0" box="[151,1436,1431,1454]" pageId="6" pageNumber="266">
- Solenomere not massive (see
<bibRefCitation id="EFC24B5CFFDCFFB9D7D138BD029102E0" author="Schubart, O." box="[511,664,1431,1454]" pageId="6" pageNumber="266" pagination="113 - 146" refId="ref6189" refString="Schubart, O. (1954 b) Diplopodos Argentinos del Museo de la Ciudad Eva Peron. I. Familia Leptodesmidae. Ministerio de educacion de la nacion. Universidad nacional de Eva Peron. Notas del Museo, 17, 113 - 146." type="journal article" year="1954">Schubart 1954b</bibRefCitation>
, fig. 14)...............................
<taxonomicName id="4C534D2EFFDCFFB9D24338BD059502E0" ID-CoL="79HLH" authority="(Schubart, 1954)" baseAuthorityName="Schubart" baseAuthorityYear="1954" box="[1133,1436,1431,1454]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="iguazuensis">
<emphasis id="B927EABFFFDCFFB9D24338BD04FC02E0" box="[1133,1269,1432,1453]" italics="true" pageId="6" pageNumber="266">S. iguazuensis</emphasis>
(Schubart, 1954)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E08FFDCFFB9D6B9389005940148" pageId="6" pageNumber="266">
<paragraph id="8BEC36ADFFDCFFB9D6B93890059502AA" blockId="6.[151,1437,933,1894]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93890059502AA" pageId="6" pageNumber="266">
8 (6) Prefemoral process large, concave, excavated to accommodate the acropodite (see
<bibRefCitation id="EFC24B5CFFDCFFB9D5D2389004830287" author="Hoffman, R. L." box="[1020,1162,1461,1483]" pageId="6" pageNumber="266" pagination="247 - 259" refId="ref5695" refString="Hoffman, R. L. (1982) Chelodesmid studies. XVIII. A synopsis of the genus Sandalodesmus Silvestri, 1902, and proposal of the new tribe Sandalodesmini (Polydesmida: Chelodesmidae). Spixiana, 5 (3), 247 - 259. https: // www. biodiversitylibrary. org / page / 28194567" type="journal article" year="1982">Hoffman 1982</bibRefCitation>
, figs 3, 4)............................................................................................
<taxonomicName id="4C534D2EFFDCFFB9D2B338F6059502AA" ID-CoL="6XHLF" authority="Hoffman, 1982" authorityName="Hoffman" authorityYear="1982" box="[1181,1436,1490,1512]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="librellus">
<emphasis id="B927EABFFFDCFFB9D2B338F6050D02AA" box="[1181,1284,1490,1512]" italics="true" pageId="6" pageNumber="266">S. librellus</emphasis>
<bibRefCitation id="EFC24B5CFFDCFFB9D32438F7059502AA" author="Hoffman, R. L." box="[1290,1436,1490,1512]" pageId="6" pageNumber="266" pagination="247 - 259" refId="ref5695" refString="Hoffman, R. L. (1982) Chelodesmid studies. XVIII. A synopsis of the genus Sandalodesmus Silvestri, 1902, and proposal of the new tribe Sandalodesmini (Polydesmida: Chelodesmidae). Spixiana, 5 (3), 247 - 259. https: // www. biodiversitylibrary. org / page / 28194567" type="journal article" year="1982">Hoffman, 1982</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEC36ADFFDCFFB9D6B938D505940148" blockId="6.[151,1437,933,1894]" box="[151,1437,1520,1542]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B938D505940148" box="[151,1437,1520,1542]" pageId="6" pageNumber="266">
- Prefemoral process small, Z-shaped (
<figureCitation id="13682A28FFDCFFB9D46838D5029F0148" box="[582,662,1520,1542]" captionStart="FIGURE 3" captionStartId="4.[151,250,1347,1371]" captionTargetBox="[151,1436,181,1323]" captionTargetId="figure-17@4.[151,1436,181,1323]" captionTargetPageId="4" captionText="FIGURE 3. Sexual characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), left gonopod, mesal view. B. Holotype ♂ (FCE-Myr 0664), left gonopod, lateral view. C. Holotype ♂ (FCE-Myr 0664), left gonopod, ventral view. D. Paratype ♀ (FCE-Myr 0664), left vulva, ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Ev: external valve; Iv: internal valve; Op: operculum; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm (A–C), 0.2 mm (D). Photographed by: Damián Hagopián." figureDoi="http://doi.org/10.5281/zenodo.6621311" httpUri="https://zenodo.org/record/6621311/files/figure.png" pageId="6" pageNumber="266">Figs 3A</figureCitation>
,
<figureCitation id="13682A28FFDCFFB9D48E38D502B6014B" box="[672,703,1520,1542]" captionStart="FIGURE 4" captionStartId="5.[151,250,697,721]" captionTargetBox="[155,1429,187,670]" captionTargetId="figure-20@5.[151,1436,181,673]" captionTargetPageId="5" captionText="FIGURE 4. Gonopod drawings of Sandalodesmus joachimadisi n. sp., holotype ♂ (FCE-Myr 0664), left gonopod. A. Mesal view. B. Lateral view. C. Ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6621313" httpUri="https://zenodo.org/record/6621313/files/figure.png" pageId="6" pageNumber="266">4A</figureCitation>
)..............................
<taxonomicName id="4C534D2EFFDCFFB9D26738D505690148" ID-CoL="993J7" authority="Rojas-Buffet &amp; Bouzan &amp; Sierwald &amp; Brescovit &amp; Simó, 2022" authorityName="Rojas-Buffet &amp; Bouzan &amp; Sierwald &amp; Brescovit &amp; Simó" authorityYear="2022" box="[1097,1376,1520,1541]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="joachimadisi" status="sp. nov.">
<emphasis id="B927EABFFFDCFFB9D26738D505690148" box="[1097,1376,1520,1541]" italics="true" pageId="6" pageNumber="266">Sandalodesmus joachimadisi</emphasis>
</taxonomicName>
<emphasis id="B927EABFFFDCFFB9D34838D505940148" bold="true" box="[1382,1437,1520,1541]" pageId="6" pageNumber="266">
<taxonomicNameLabel id="A21457C4FFDCFFB9D34838D505940148" box="[1382,1437,1520,1541]" pageId="6" pageNumber="266" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E08FFDCFFB9D6B93B2805950110" pageId="6" pageNumber="266">
<paragraph id="8BEC36ADFFDCFFB9D6B93B280595016F" blockId="6.[151,1437,933,1894]" box="[151,1436,1549,1571]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93B280595016F" box="[151,1436,1549,1571]" pageId="6" pageNumber="266">9 (3) Acropodite with a prominent laminate deltoid lobe on the mesal side; prefemoral process straight.................... 10</keyLead>
</paragraph>
<paragraph id="8BEC36ADFFDCFFB9D6B93B0E05950110" blockId="6.[151,1437,933,1894]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93B0E05950110" pageId="6" pageNumber="266">- Acropodite with a lobe on the mesal side; prefemoral process rotated 180° so that the distal half is on the opposite side from the basal half.......................................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E08FFDCFFB9D6B93B400595019E" pageId="6" pageNumber="266">
<paragraph id="8BEC36ADFFDCFFB9D6B93B40059501D5" blockId="6.[151,1437,933,1894]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93B40059501D5" pageId="6" pageNumber="266">
10 (9) Mesal deltoid lobe of acropodite placed apically and concealing solenomere, latter extending beyond end of acropodite (see
<bibRefCitation id="EFC24B5CFFDCFFB9D6CC3BA7016601D5" author="Schubart, O." box="[226,367,1666,1688]" pageId="6" pageNumber="266" pagination="355 - 367" refId="ref6235" refString="Schubart, O. (1956) &quot; Leptodesmidae &quot; Brasileiras. III: Especies de Minas Gerais (Diplopoda, Proterospermophora). Revista brasileira de biologia, 16 (3), 355 - 367." type="journal article" year="1956">Schubart 1956</bibRefCitation>
, fig. 3).........................................................
<taxonomicName id="4C534D2EFFDCFFB9D25A3BA6059501D5" ID-CoL="6XHLL" authority="(Schubart, 1956)" baseAuthorityName="Schubart" baseAuthorityYear="1956" box="[1140,1436,1666,1688]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="stramineus">
<emphasis id="B927EABFFFDCFFB9D25A3BA604FC01D5" box="[1140,1269,1667,1688]" italics="true" pageId="6" pageNumber="266">S. stramineus</emphasis>
(
<bibRefCitation id="EFC24B5CFFDCFFB9D32F3BA7059C01D5" author="Schubart, O." box="[1281,1429,1666,1688]" pageId="6" pageNumber="266" pagination="355 - 367" refId="ref6235" refString="Schubart, O. (1956) &quot; Leptodesmidae &quot; Brasileiras. III: Especies de Minas Gerais (Diplopoda, Proterospermophora). Revista brasileira de biologia, 16 (3), 355 - 367." type="journal article" year="1956">Schubart, 1956</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEC36ADFFDCFFB9D6B93B850595019E" blockId="6.[151,1437,933,1894]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93B850595019E" pageId="6" pageNumber="266">
- Mesal deltoid lobe of acropodite placed subapically, solenomere entirely visible and extending beyond end of acropodite (see
<bibRefCitation id="EFC24B5CFFDCFFB9D6CC3B980179019E" author="Hoffman, R. L." box="[226,368,1725,1747]" pageId="6" pageNumber="266" pagination="33 - 42" refId="ref5609" refString="Hoffman, R. L. (1967) Chelodesmid Studies III. Notes on the status of Gonioleptodesmus, the description of a new species from Minas Gerais, and a key to the presently known members of the genus (Diplopoda, Polydesmida). Papeis Avulsos do Departamento de Zoologia, 21 (4), 33 - 42." type="journal article" year="1967">Hoffman 1967</bibRefCitation>
, figs 3, 4;
<bibRefCitation id="EFC24B5CFFDCFFB9D7FB3B98026A019F" author="Hoffman, R. L." box="[469,611,1725,1747]" pageId="6" pageNumber="266" pagination="247 - 259" refId="ref5695" refString="Hoffman, R. L. (1982) Chelodesmid studies. XVIII. A synopsis of the genus Sandalodesmus Silvestri, 1902, and proposal of the new tribe Sandalodesmini (Polydesmida: Chelodesmidae). Spixiana, 5 (3), 247 - 259. https: // www. biodiversitylibrary. org / page / 28194567" type="journal article" year="1982">Hoffman 1982</bibRefCitation>
, fig. 11).....................................
<taxonomicName id="4C534D2EFFDCFFB9D2AF3B980595019E" ID-CoL="9949J" authority="(Hoffman, 1967)" baseAuthorityName="Hoffman" baseAuthorityYear="1967" box="[1153,1436,1725,1747]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="schubarti">
<emphasis id="B927EABFFFDCFFB9D2AF3B9804FD019F" box="[1153,1268,1725,1746]" italics="true" pageId="6" pageNumber="266">S. schubarti</emphasis>
(
<bibRefCitation id="EFC24B5CFFDCFFB9D32E3B98059C019E" author="Hoffman, R. L." box="[1280,1429,1725,1747]" pageId="6" pageNumber="266" pagination="33 - 42" refId="ref5609" refString="Hoffman, R. L. (1967) Chelodesmid Studies III. Notes on the status of Gonioleptodesmus, the description of a new species from Minas Gerais, and a key to the presently known members of the genus (Diplopoda, Polydesmida). Papeis Avulsos do Departamento de Zoologia, 21 (4), 33 - 42." type="journal article" year="1967">Hoffman, 1967</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E08FFDCFFB9D6B93BFF05950040" pageId="6" pageNumber="266">
<paragraph id="8BEC36ADFFDCFFB9D6B93BFF059501BD" blockId="6.[151,1437,933,1894]" box="[151,1436,1754,1776]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93BFF059501BD" box="[151,1436,1754,1776]" pageId="6" pageNumber="266">
11 (9) Outer edge of solenomere entire, smooth (see
<bibRefCitation id="EFC24B5CFFDCFFB9D4B23BFF032301A2" author="Schubart, O." box="[668,810,1754,1776]" pageId="6" pageNumber="266" pagination="321 - 440" refId="ref6100" refString="Schubart, O. (1944) Os Diplopodos de Pirassununga. Acta Zoologica Lilloana, 2, 321 - 440." type="journal article" year="1944">Schubart 1944</bibRefCitation>
, fig. 22)....................
<taxonomicName id="4C534D2EFFDCFFB9D2AA3BFE059501BD" ID-CoL="79HJC" authority="(Schubart, 1944)" baseAuthorityName="Schubart" baseAuthorityYear="1944" box="[1156,1436,1754,1776]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="gasparae">
<emphasis id="B927EABFFFDCFFB9D2AA3BFE04FC01BD" box="[1156,1269,1755,1776]" italics="true" pageId="6" pageNumber="266">S. gasparae</emphasis>
(
<bibRefCitation id="EFC24B5CFFDCFFB9D32F3BFF059C01BD" author="Schubart, O." box="[1281,1429,1754,1776]" pageId="6" pageNumber="266" pagination="321 - 440" refId="ref6100" refString="Schubart, O. (1944) Os Diplopodos de Pirassununga. Acta Zoologica Lilloana, 2, 321 - 440." type="journal article" year="1944">Schubart, 1944</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEC36ADFFDCFFB9D6B93BDD05950040" blockId="6.[151,1437,933,1894]" box="[151,1436,1784,1806]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93BDD05950040" box="[151,1436,1784,1806]" pageId="6" pageNumber="266">- Outer edge of solenomere denticulate..................................................................… 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A72E08FFDCFFB9D6B93A3005950028" pageId="6" pageNumber="266">
<paragraph id="8BEC36ADFFDCFFB9D6B93A3005950005" blockId="6.[151,1437,933,1894]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93A3005950005" pageId="6" pageNumber="266">
12 (11) Solenomere elongate, the seminal groove sinuately curved (see
<bibRefCitation id="EFC24B5CFFDCFFB9D56B3A3003DB0067" author="Schubart, O." box="[837,978,1813,1835]" pageId="6" pageNumber="266" pagination="321 - 440" refId="ref6100" refString="Schubart, O. (1944) Os Diplopodos de Pirassununga. Acta Zoologica Lilloana, 2, 321 - 440." type="journal article" year="1944">Schubart 1944</bibRefCitation>
, figs 25, 26)....................................................................................................
<taxonomicName id="4C534D2EFFDCFFB9D2733A1605950005" ID-CoL="6XHLH" authority="(Schubart, 1944)" baseAuthorityName="Schubart" baseAuthorityYear="1944" box="[1117,1436,1842,1864]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="hippocampus">
<emphasis id="B927EABFFFDCFFB9D2733A1604FC000A" box="[1117,1269,1842,1864]" italics="true" pageId="6" pageNumber="266">S. hippocampus</emphasis>
(
<bibRefCitation id="EFC24B5CFFDCFFB9D32F3A17059C0005" author="Schubart, O." box="[1281,1429,1842,1864]" pageId="6" pageNumber="266" pagination="321 - 440" refId="ref6100" refString="Schubart, O. (1944) Os Diplopodos de Pirassununga. Acta Zoologica Lilloana, 2, 321 - 440." type="journal article" year="1944">Schubart, 1944</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEC36ADFFDCFFB9D6B93A7505950028" blockId="6.[151,1437,933,1894]" box="[151,1436,1871,1894]" pageId="6" pageNumber="266">
<keyLead id="30A29598FFDCFFB9D6B93A7505950028" box="[151,1436,1871,1894]" pageId="6" pageNumber="266">
- Solenomere short, the seminal groove arcuately curved (see
<bibRefCitation id="EFC24B5CFFDCFFB9D50B3A7503B40028" author="Schubart, O." box="[805,957,1871,1894]" pageId="6" pageNumber="266" pagination="77 - 132" refId="ref6155" refString="Schubart, O. (1954 a) Sobre os Diplopoda dos estados do Parana e Santa Catarina. I. Proterospermophora. Arquivos do Museu Paranaense, 10, 77 - 132." type="journal article" year="1954">Schubart 1954a</bibRefCitation>
, fig. 14)......
<taxonomicName id="4C534D2EFFDCFFB9D2463A7505950028" ID-CoL="79HLF" authority="(Schubart, 1954)" baseAuthorityName="Schubart" baseAuthorityYear="1954" box="[1128,1436,1871,1894]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="paranaensis">
<emphasis id="B927EABFFFDCFFB9D2463A7504FC0028" box="[1128,1269,1872,1893]" italics="true" pageId="6" pageNumber="266">S. paranaensis</emphasis>
(Schubart, 1954)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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