269 lines
36 KiB
XML
269 lines
36 KiB
XML
<document id="7F8372EBD764E50978F2CC501E41ABED" ID-DOI="10.5281/zenodo.5160535" ID-ISSN="1942-1354" ID-Zenodo-Dep="5160535" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628108913532" checkinUser="felipe" docAuthor="Sokolov, Igor M." docDate="2011" docId="03FA87B0FF967733A7862E10FA52FA0E" docLanguage="en" docName="InsectaMundi.0164.1-14.pdf" docOrigin="Insecta Mundi 2011 (164)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Anillinus chilhowee Sokolov 2011, new species" docType="treatment" docVersion="3" lastPageNumber="9" masterDocId="FFC3FFC8FF9F7739A7462A52FFF2FFF5" masterDocTitle="Five new species of Anillinus Casey from the Southern Appalachian Mountains and the Piedmont Plateau of eastern U. S. A. (Coleoptera: Carabidae: Trechinae: Bembidiini)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="8" updateTime="1699034592004" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="D58FA7017F89D8C7F6211522A26175D2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0F35F32973FEEB1B38ABA2992C0C87F4">
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<mods:title id="7CC7A140B20E3E330291F9DD929A59CF">Five new species of Anillinus Casey from the Southern Appalachian Mountains and the Piedmont Plateau of eastern U. S. A. (Coleoptera: Carabidae: Trechinae: Bembidiini)</mods:title>
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</mods:titleInfo>
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<mods:name id="AC630A9D03C1326741B1589BCECB3144" type="personal">
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<mods:role id="D9E32BB1E3CF6530D0512D9F46D2DACF">
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<mods:roleTerm id="16FED94467852BB28102E464992E14E1">Author</mods:roleTerm>
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<mods:namePart id="46BC1699A99F4F15F78CF506B9018397">Sokolov, Igor M.</mods:namePart>
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<mods:typeOfResource id="9E72F1EABA78C0DF9BCE34270BE4B646">text</mods:typeOfResource>
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<mods:titleInfo id="57595E5BA2A816EA649187A232004C7B">
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<mods:title id="6F6F2BAFAC377D340C0A382D7B4B2959">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="747058BD19E103A9F28D2CE766F90B2E">
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<mods:date id="0B371AC524310AD2FC7CE0D6DE423810">2011</mods:date>
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<mods:detail id="F7B64B0F6BE1B049E0C74188D33420CA" type="pubDate">
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<mods:number id="3618275D62442169BBF731959145207D">2011-04-15</mods:number>
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</mods:detail>
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<mods:detail id="30C0E5C466DE219B796E592B6C2FB29C" type="volume">
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<mods:number id="9EB581C239BC14D68D0E04848C57A6FE">2011</mods:number>
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</mods:detail>
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<mods:detail id="BD495E87FB629F1AD2BF2A7985325FEF" type="issue">
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<mods:number id="66FC18B974777202EC681772B767F276">164</mods:number>
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<mods:extent id="00ED282AC87831BB2256A37E3DA0153E" unit="page">
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<mods:start id="575EC05B081B72EF9D324C50484D26DB">1</mods:start>
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<mods:end id="4919B4AA5130B37FCBF4066259D33856">14</mods:end>
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<mods:classification id="90BC03934B25D810EB7C1A82C9D1F53B">journal article</mods:classification>
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<mods:identifier id="12371263D87728CDF0BCEF8A5392768B" type="DOI">10.5281/zenodo.5160535</mods:identifier>
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<mods:identifier id="D20A913FC6C6FFE08995320C38C78A58" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="23057564203F5C8583E26620AF19BDEA" type="Zenodo-Dep">5160535</mods:identifier>
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</mods:mods>
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<treatment id="03FA87B0FF967733A7862E10FA52FA0E" LSID="urn:lsid:plazi:treatment:03FA87B0FF967733A7862E10FA52FA0E" httpUri="http://treatment.plazi.org/id/03FA87B0FF967733A7862E10FA52FA0E" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
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<subSubSection id="C349652DFF967730A7862E10FD0AFBAE" box="[192,760,1090,1115]" pageId="9" pageNumber="8" type="nomenclature">
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<paragraph id="8BEC36A6FF967730A7862E10FD0AFBAE" blockId="9.[192,760,1090,1147]" box="[192,760,1090,1115]" pageId="9" pageNumber="8">
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<heading id="D0A481CAFF967730A7862E10FD0AFBAE" bold="true" box="[192,760,1090,1115]" fontSize="10" level="2" pageId="9" pageNumber="8" reason="3">
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<emphasis id="B927EAB4FF967730A7862E10FD0AFBAE" bold="true" box="[192,760,1090,1115]" pageId="9" pageNumber="8">
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<taxonomicName id="4C534D25FF967730A7862E10FDBAFBAE" ID-CoL="66PVW" authority="Sokolov" authorityName="Sokolov" authorityYear="2011" box="[192,584,1090,1115]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="chilhowee" status="sp. nov.">
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<emphasis id="B927EAB4FF967730A7862E10FE20FBAE" bold="true" box="[192,466,1090,1115]" italics="true" pageId="9" pageNumber="8">Anillinus chilhowee</emphasis>
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Sokolov
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21457CFFF967730A5122E11FD0AFBAE" box="[596,760,1091,1115]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C349652DFF967730A7862E31FE6CFB8E" box="[192,414,1123,1147]" pageId="9" pageNumber="8" type="description">
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<paragraph id="8BEC36A6FF967730A7862E31FE6CFB8E" blockId="9.[192,760,1090,1147]" box="[192,414,1123,1147]" pageId="9" pageNumber="8">
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<figureCitation id="13682A23FF967730A7862E31FED8FB8E" box="[192,298,1123,1147]" captionStart="Figure 1-5" captionStartId="3.[192,276,1475,1497]" captionTargetBox="[197,1384,196,1453]" captionTargetId="figure-160@3.[192,1440,192,1458]" captionTargetPageId="3" captionText="Figure 1-5. Habitus images. 1) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.), holotype. 2) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.), paratype. 3) A. chilhowee (TN, Polk Co., Chilhowee Mtn.), holotype. 4) A. smokiensis (TN, Blount Co., GSMNP), paratype. 5) A. chandleri (SC, Edgefield Co., Sumter National Forest), holotype." figureDoi="http://doi.org/10.5281/zenodo.5160539" httpUri="https://zenodo.org/record/5160539/files/figure.png" pageId="9" pageNumber="8">Figure 3</figureCitation>
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,
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<figureCitation id="13682A23FF967730A6712E31FEB5FB8E" box="[311,327,1123,1147]" captionStart="Figure 6-10" captionStartId="4.[192,276,877,899]" captionTargetBox="[192,1440,193,852]" captionTargetId="figure-262@4.[192,1440,192,852]" captionTargetPageId="4" captionText="Figure 6-10. Pronotum images. 6) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.). 7) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.). 8) A. chilhowee (TN, Polk Co., Chilhowee Mtn.). 9) A. smokiensis (TN, Blount Co., GSMNP). 10) A. chandleri (SC, Edgefield Co., Sumter National Forest)." figureDoi="http://doi.org/10.5281/zenodo.5160541" httpUri="https://zenodo.org/record/5160541/files/figure.png" pageId="9" pageNumber="8">8</figureCitation>
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,
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<figureCitation id="13682A23FF967730A6122E31FE86FB8E" box="[340,372,1123,1147]" captionStart="Figure 11-15" captionStartId="5.[192,276,1691,1713]" captionTargetBox="[202,1419,207,1664]" captionTargetId="figure-14@5.[192,1440,192,1696]" captionTargetPageId="5" captionText="Figure 11-15. Illustrations of male aedeagus. 11) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.). 12) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.). 13) A. chilhowee (TN, Polk Co., Chilhowee Mtn.). 14) A. smokiensis (TN, Blount Co., GSMNP). 15) A. chandleri (SC, Edgefield Co., Sumter National Forest). a-Median lobe, right lateral aspect; b-Left paramere, left lateral aspect; c-Right paramere, right lateral aspect. Scale = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5160545" httpUri="https://zenodo.org/record/5160545/files/figure.png" pageId="9" pageNumber="8">13</figureCitation>
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,
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<figureCitation id="13682A23FF967730A6C62E31FE6CFB8E" box="[384,414,1123,1147]" captionStart="Figure 16" captionStartId="6.[192,276,1017,1039]" captionTargetBox="[192,1440,192,996]" captionTargetId="figure-321@6.[192,1440,192,996]" captionTargetPageId="6" captionText="Figure 16. Locality records for the moseleyae-group (red color) and loweae-group species from Chilhowee and Starr Mountains (yellow color). Anillinus moseleyae – red quadrangles; A. carltoni, new species – red circle; A. unicoi – red star; A. juliae – yellow circle; A. chilhowee – yellow star." figureDoi="http://doi.org/10.5281/zenodo.5160547" httpUri="https://zenodo.org/record/5160547/files/figure.png" pageId="9" pageNumber="8">16</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C349652DFF967730A7862EF1FEA0FA8E" pageId="9" pageNumber="8" type="materials_examined">
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<paragraph id="8BEC36A6FF967730A7862EF1FAC6FAEE" blockId="9.[192,1440,1187,1307]" pageId="9" pageNumber="8">
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<materialsCitation id="3B3B3CFBFF967730A7862EF1FF1EFB0E" collectingDate="2005-09-18" collectorName="Forest & A. K. Tishechkin" county="Polk Co." determinerName="S. and C. A. Tishechkin" elevation="715" latitude="35.116665" location="Cherokee Natl. Forest" longLatPrecision="92" longitude="-84.624" municipality="Male" pageId="9" pageNumber="8" specimenCount="715" stateProvince="Tennessee" typeStatus="holotype">
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<emphasis id="B927EAB4FF967730A7862EF1FEB5FB4E" bold="true" box="[192,327,1187,1211]" pageId="9" pageNumber="8">
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<typeStatus id="54E88804FF967730A7862EF1FEB3FB4E" box="[192,321,1187,1211]" pageId="9" pageNumber="8" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingMunicipality id="6B88ACDCFF967730A6082EF1FE7EFB4E" box="[334,396,1187,1211]" pageId="9" pageNumber="8">Male</collectingMunicipality>
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labeled /
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<collectingRegion id="4997F844FF967730A5462EF1FD5DFB4E" box="[512,687,1187,1211]" country="United States of America" name="Tennessee" pageId="9" pageNumber="8">TENNESSEE</collectingRegion>
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:
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<collectingCounty id="628D4E2AFF967730A5FC2EF1FCD7FB4E" box="[698,805,1187,1211]" pageId="9" pageNumber="8">Polk Co.</collectingCounty>
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,
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<location id="8E8C607DFF967730A4742EF1FBB8FB4E" LSID="urn:lsid:plazi:treatment:03FA87B0FF967733A7862E10FA52FA0E:8E8C607DFF967730A4742EF1FBB8FB4E" box="[818,1098,1187,1211]" county="Polk Co." latitude="35.116665" longLatPrecision="92" longitude="-84.624" municipality="Male" name="Cherokee Natl. Forest" pageId="9" pageNumber="8" stateProvince="Tennessee">Cherokee Natl. Forest</location>
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||
,
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<location id="8E8C607DFF967730A3112EF1FAFFFB4E" LSID="urn:lsid:plazi:treatment:03FA87B0FF967733A7862E10FA52FA0E:8E8C607DFF967730A3112EF1FAFFFB4E" box="[1111,1293,1187,1211]" county="Polk Co." latitude="35.116665" longLatPrecision="92" longitude="-84.624" municipality="Male" name="Chilhowee Mt." pageId="9" pageNumber="8" stateProvince="Tennessee">Chilhowee Mt.</location>
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at
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<geoCoordinate id="EE675061FF967730A2702EF1FA52FB4E" box="[1334,1440,1187,1211]" degrees="35" direction="north" minutes="7.0" orientation="latitude" pageId="9" pageNumber="8" precision="92" value="35.116665">35°7.0’N</geoCoordinate>
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||
<geoCoordinate id="EE675061FF967730A7862E91FEBCFB2E" box="[192,334,1219,1243]" degrees="84" direction="west" minutes="37.44" orientation="longitude" pageId="9" pageNumber="8" precision="9" value="-84.624">84°37.44’W</geoCoordinate>
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,
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<specimenCount id="9D55FD2FFF967730A61E2E91FE56FB2E" box="[344,420,1219,1243]" pageId="9" pageNumber="8" type="generic">
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||
<quantity id="4CAB9B43FF967730A61E2E91FE52FB2E" box="[344,416,1219,1243]" metricMagnitude="2" metricUnit="m" metricValue="7.15" pageId="9" pageNumber="8" unit="m" value="715.0">
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||
<elevation id="007ED195FF967730A61E2E91FE52FB2E" box="[344,416,1219,1243]" metricMagnitude="2" metricUnit="m" metricValue="7.15" pageId="9" pageNumber="8" unit="m" value="715.0">715m</elevation>
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</quantity>
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||
.
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</specimenCount>
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<date id="FFED1066FF967730A6EF2E91FDBBFB2E" box="[425,585,1219,1243]" pageId="9" pageNumber="8" value="2005-09-18">
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||
<collectingDate id="EFA9E98EFF967730A6EF2E91FDBBFB2E" box="[425,585,1219,1243]" pageId="9" pageNumber="8" value="2005-09-18">18 Sept 2005</collectingDate>
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</date>
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.
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<collectorName id="26A65370FF967730A5152E96FD50FB2E" box="[595,674,1220,1243]" pageId="9" pageNumber="8">Forest</collectorName>
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litter.
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<collectorName id="26A65370FF967730A5B22E91FC45FB2E" box="[756,951,1219,1243]" pageId="9" pageNumber="8">A.K. Tishechkin</collectorName>
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/
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<taxonomicName id="4C534D25FF967730A4802E91FB8CFB2E" authorityName="Sokolov & Carlton" authorityYear="2004" box="[966,1150,1219,1243]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="barri">
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<emphasis id="B927EAB4FF967730A4802E91FB8CFB2E" box="[966,1150,1219,1243]" italics="true" pageId="9" pageNumber="8">Anillinus barri</emphasis>
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</taxonomicName>
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<determinerName id="7E30DA62FF967730A3C22E91FA52FB2E" box="[1156,1440,1219,1243]" pageId="9" pageNumber="8">S. and C. A. Tishechkin</determinerName>
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det.
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</materialsCitation>
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/
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<materialsCitation id="3B3B3CFBFF967730A7BD2EB1FAC2FAEE" collectionCode="USNM" pageId="9" pageNumber="8" specimenCount="2" stateProvince="Tennessee" typeStatus="holotype">
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<typeStatus id="54E88804FF967730A7BD2EB1FE6BFB0E" box="[251,409,1251,1275]" pageId="9" pageNumber="8" type="holotype">HOLOTYPE</typeStatus>
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,
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<taxonomicName id="4C534D25FF967730A6E22EB1FD09FB0E" authority="Sokolov" authorityName="Sokolov" authorityYear="2011" box="[420,763,1251,1275]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="chilhowee">
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<emphasis id="B927EAB4FF967730A6E22EB1FD66FB0E" box="[420,660,1251,1275]" italics="true" pageId="9" pageNumber="8">Anillinus chilhowee</emphasis>
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Sokolov
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</taxonomicName>
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, des. 2009/. The
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<typeStatus id="54E88804FF967730A4862EB1FBDBFB0E" box="[960,1065,1251,1275]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
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is dissected and bears a plastic rectangle with genitalia mounted in dimethylhydantoin formaldehyde resin. Deposited
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<collectionCode id="ED42AE63FF967730A3912F51FAC2FAEE" box="[1239,1328,1283,1307]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="8" type="Museum">USNM</collectionCode>
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</materialsCitation>
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||
.
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</paragraph>
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<paragraph id="8BEC36A6FF967730A7862F11FEA0FA8E" blockId="9.[192,1440,1347,1403]" pageId="9" pageNumber="8">
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<emphasis id="B927EAB4FF967730A7862F11FE87FAAE" bold="true" box="[192,373,1347,1371]" pageId="9" pageNumber="8">Type locality.</emphasis>
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<collectingCountry id="F3447636FF967730A63C2F11FE3AFAAE" box="[378,456,1347,1371]" name="United States of America" pageId="9" pageNumber="8">U.S.A.</collectingCountry>
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<collectingRegion id="4997F844FF967730A68B2F16FDBFFAAE" box="[461,589,1348,1371]" country="United States of America" name="Tennessee" pageId="9" pageNumber="8">Tennessee</collectingRegion>
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, Polk County, Cherokee National Forest, Chilhowee Mountain,
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<geoCoordinate id="EE675061FF967730A27C2F11FA52FAAE" box="[1338,1440,1347,1371]" degrees="35" direction="north" minutes="7.0" orientation="latitude" pageId="9" pageNumber="8" precision="92" value="35.116665">35°7.0’N</geoCoordinate>
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<geoCoordinate id="EE675061FF967730A7862F31FEBDFA8E" box="[192,335,1379,1403]" degrees="84" direction="west" minutes="37.44" orientation="longitude" pageId="9" pageNumber="8" precision="9" value="-84.624">84°37.44’W</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C349652DFF967730A7862FF1FAEDFA4E" box="[192,1311,1443,1467]" pageId="9" pageNumber="8" type="etymology">
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<paragraph id="8BEC36A6FF967730A7862FF1FAEDFA4E" blockId="9.[192,1311,1443,1467]" box="[192,1311,1443,1467]" pageId="9" pageNumber="8">
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||
<emphasis id="B927EAB4FF967730A7862FF1FEAFFA4E" bold="true" box="[192,349,1443,1467]" pageId="9" pageNumber="8">Etymology.</emphasis>
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The name of this species is based on the name of the mountain where it occurs.
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</paragraph>
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</subSubSection>
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<subSubSection id="C349652DFF967733A7862FB1FB0EFE2E" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="description">
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||
<paragraph id="8BEC36A6FF967730A7862FB1FB5EF98E" blockId="9.[192,1440,1507,1915]" pageId="9" pageNumber="8">
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||
<emphasis id="B927EAB4FF967730A7862FB1FE96FA0E" bold="true" box="[192,356,1507,1531]" pageId="9" pageNumber="8">Description</emphasis>
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||
. Small for genus (ABL =
|
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<quantity id="4CAB9B43FF967730A5D52FB1FCF3FA0E" box="[659,769,1507,1531]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" pageId="9" pageNumber="8" unit="mm" value="1.64">1.64 mm</quantity>
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||
). Habitus (
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<figureCitation id="13682A23FF967730A4C32FB1FC3BFA0E" box="[901,969,1507,1531]" captionStart="Figure 1-5" captionStartId="3.[192,276,1475,1497]" captionTargetBox="[197,1384,196,1453]" captionTargetId="figure-160@3.[192,1440,192,1458]" captionTargetPageId="3" captionText="Figure 1-5. Habitus images. 1) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.), holotype. 2) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.), paratype. 3) A. chilhowee (TN, Polk Co., Chilhowee Mtn.), holotype. 4) A. smokiensis (TN, Blount Co., GSMNP), paratype. 5) A. chandleri (SC, Edgefield Co., Sumter National Forest), holotype." figureDoi="http://doi.org/10.5281/zenodo.5160539" httpUri="https://zenodo.org/record/5160539/files/figure.png" pageId="9" pageNumber="8">Fig. 3</figureCitation>
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) moderately convex, ovoidal (WE/ABL 0.38), head of normal proportions for the genus (WH/WPm 0.71), pronotum narrow compared to elytra (WPm/WE 0.78). Body color rufobrunneus, appendages testaceous. Dorsal microsculpture well-developed; covering all head, except two paramedial patches without microsculpture on vertex, and all pronotum except medial part of the disc. Elytra with well-developed polygonal microsculpture.
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</paragraph>
|
||
<paragraph id="8BEC36A6FF967730A7B62CD6FDBCF90E" blockId="9.[192,1440,1507,1915]" pageId="9" pageNumber="8">
|
||
Pronotum (
|
||
<figureCitation id="13682A23FF967730A6382CD1FE37F96E" box="[382,453,1667,1691]" captionStart="Figure 6-10" captionStartId="4.[192,276,877,899]" captionTargetBox="[192,1440,193,852]" captionTargetId="figure-262@4.[192,1440,192,852]" captionTargetPageId="4" captionText="Figure 6-10. Pronotum images. 6) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.). 7) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.). 8) A. chilhowee (TN, Polk Co., Chilhowee Mtn.). 9) A. smokiensis (TN, Blount Co., GSMNP). 10) A. chandleri (SC, Edgefield Co., Sumter National Forest)." figureDoi="http://doi.org/10.5281/zenodo.5160541" httpUri="https://zenodo.org/record/5160541/files/figure.png" pageId="9" pageNumber="8">Fig. 8</figureCitation>
|
||
) moderately convex and of normal proportions for genus (WPm/LP 1.26), with margins rectilinear and moderately constricted posteriad (WPm/WPp 1.26). Anterior angles evident, very slightly prominent. Posterior angles obtuse (115°). Width between posterior angles greater than between anterior angles (WPa/WPp 0.92).
|
||
</paragraph>
|
||
<paragraph id="8BEC36A6FF967730A7B62D51FBA4F88E" blockId="9.[192,1440,1507,1915]" pageId="9" pageNumber="8">Elytra moderately convex, depressed along suture, slightly elongated (LE/ABL 0.58), with traces of 2 interneurs. Humeri rectangular and rounded. Lateral margins subparallel, slightly divergent in basal forth, evenly rounded to apex in apical third, maximal width of elytra at midpoint. Elytra without subapical sinuation. Vestiture of elytra short (lesser than one-third of discal setae).</paragraph>
|
||
<paragraph id="8BEC36A6FF957733A7B62A91FD03FF0E" blockId="10.[192,1440,195,475]" pageId="10" pageNumber="9">Prothoracic leg of males with moderately dilated tarsomere 1. Profemur moderately swollen. Metafemora unmodified. Sternum VII of males unmodified.</paragraph>
|
||
<paragraph id="8BEC36A6FF957733A7B62B51FB0EFE2E" blockId="10.[192,1440,195,475]" pageId="10" pageNumber="9">
|
||
Median lobe of aedeagus (
|
||
<figureCitation id="13682A23FF957733A5662B51FD8DFEEE" box="[544,639,259,283]" captionStart="Figure 11-15" captionStartId="5.[192,276,1691,1713]" captionTargetBox="[202,1419,207,1664]" captionTargetId="figure-14@5.[192,1440,192,1696]" captionTargetPageId="5" captionText="Figure 11-15. Illustrations of male aedeagus. 11) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.). 12) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.). 13) A. chilhowee (TN, Polk Co., Chilhowee Mtn.). 14) A. smokiensis (TN, Blount Co., GSMNP). 15) A. chandleri (SC, Edgefield Co., Sumter National Forest). a-Median lobe, right lateral aspect; b-Left paramere, left lateral aspect; c-Right paramere, right lateral aspect. Scale = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5160545" httpUri="https://zenodo.org/record/5160545/files/figure.png" pageId="10" pageNumber="9">Fig. 13a</figureCitation>
|
||
) evenly arcuate and twisted, with apex greatly enlarged and widely rounded. Ventral margin of median lobe enlarged bearing numerous poriferous canals, which are also present on walls of median lobe across its lower half. Dorsal copulatory sclerites forming a short, curled blade-like structure with long basal prolongations. Ventral sclerite and spines of internal sac absent. Left paramere (
|
||
<figureCitation id="13682A23FF957733A60E2BD1FE5EFE6E" box="[328,428,387,411]" captionStart="Figure 11-15" captionStartId="5.[192,276,1691,1713]" captionTargetBox="[202,1419,207,1664]" captionTargetId="figure-14@5.[192,1440,192,1696]" captionTargetPageId="5" captionText="Figure 11-15. Illustrations of male aedeagus. 11) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.). 12) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.). 13) A. chilhowee (TN, Polk Co., Chilhowee Mtn.). 14) A. smokiensis (TN, Blount Co., GSMNP). 15) A. chandleri (SC, Edgefield Co., Sumter National Forest). a-Median lobe, right lateral aspect; b-Left paramere, left lateral aspect; c-Right paramere, right lateral aspect. Scale = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5160545" httpUri="https://zenodo.org/record/5160545/files/figure.png" pageId="10" pageNumber="9">Fig. 13b</figureCitation>
|
||
) not enlarged, paramere apex with four poriferous canals, bearing one long (more distal) and one short setae in apical position. Right paramere (
|
||
<figureCitation id="13682A23FF957733A4EC2BF1FBFAFE4E" box="[938,1032,419,443]" captionStart="Figure 11-15" captionStartId="5.[192,276,1691,1713]" captionTargetBox="[202,1419,207,1664]" captionTargetId="figure-14@5.[192,1440,192,1696]" captionTargetPageId="5" captionText="Figure 11-15. Illustrations of male aedeagus. 11) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.). 12) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.). 13) A. chilhowee (TN, Polk Co., Chilhowee Mtn.). 14) A. smokiensis (TN, Blount Co., GSMNP). 15) A. chandleri (SC, Edgefield Co., Sumter National Forest). a-Median lobe, right lateral aspect; b-Left paramere, left lateral aspect; c-Right paramere, right lateral aspect. Scale = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5160545" httpUri="https://zenodo.org/record/5160545/files/figure.png" pageId="10" pageNumber="9">Fig. 13c</figureCitation>
|
||
) elongate, with subparallel apical portion, bearing four long setae that are slightly shorter than the apical part of paramere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349652DFF957733A7862851FC66FDAE" pageId="10" pageNumber="9" type="distribution">
|
||
<paragraph id="8BEC36A6FF957733A7862851FAD4FDEE" blockId="10.[192,1318,515,539]" box="[192,1318,515,539]" pageId="10" pageNumber="9">
|
||
<emphasis id="B927EAB4FF957733A7862851FE87FDEE" bold="true" box="[192,373,515,539]" pageId="10" pageNumber="9">Distribution.</emphasis>
|
||
Known only from the Chilhowee Mountain, Polk County,
|
||
<collectingRegion id="4997F844FF957733A3742856FB41FDEE" box="[1074,1203,516,539]" country="United States of America" name="Tennessee" pageId="10" pageNumber="9">Tennessee</collectingRegion>
|
||
(
|
||
<figureCitation id="13682A23FF957733A3842851FAEAFDEE" box="[1218,1304,515,539]" captionStart="Figure 16" captionStartId="6.[192,276,1017,1039]" captionTargetBox="[192,1440,192,996]" captionTargetId="figure-321@6.[192,1440,192,996]" captionTargetPageId="6" captionText="Figure 16. Locality records for the moseleyae-group (red color) and loweae-group species from Chilhowee and Starr Mountains (yellow color). Anillinus moseleyae – red quadrangles; A. carltoni, new species – red circle; A. unicoi – red star; A. juliae – yellow circle; A. chilhowee – yellow star." figureDoi="http://doi.org/10.5281/zenodo.5160547" httpUri="https://zenodo.org/record/5160547/files/figure.png" pageId="10" pageNumber="9">Fig. 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEC36A6FF957733A7862811FC66FDAE" blockId="10.[192,916,579,603]" box="[192,916,579,603]" pageId="10" pageNumber="9">
|
||
<emphasis id="B927EAB4FF957733A7862811FEC0FDAE" bold="true" box="[192,306,579,603]" pageId="10" pageNumber="9">Habitat.</emphasis>
|
||
A single male was collected by sifting forest litter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349652DFF957733A78628D1FA52FA0E" pageId="10" pageNumber="9" type="diagnosis">
|
||
<paragraph id="8BEC36A6FF957733A78628D1FD08FD0E" blockId="10.[192,1440,643,1051]" pageId="10" pageNumber="9">
|
||
<emphasis id="B927EAB4FF957733A78628D1FE1FFD6E" bold="true" box="[192,493,643,667]" pageId="10" pageNumber="9">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C534D25FF957733A6B728D1FD2FFD6E" authorityName="Sokolov" authorityYear="2011" box="[497,733,643,667]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="chilhowee">
|
||
<emphasis id="B927EAB4FF957733A6B728D1FD2FFD6E" box="[497,733,643,667]" italics="true" pageId="10" pageNumber="9">Anillinus chilhowee</emphasis>
|
||
</taxonomicName>
|
||
belongs to the loweae-group of species (
|
||
<bibRefCitation id="EFC24B57FF957733A3EB28D1FF0DFD4E" author="Sokolov, I. M. & C. E. Carlton" pageId="10" pageNumber="9" pagination="1 - 23" refId="ref9274" refString="Sokolov, I. M., and C. E. Carlton. 2010. New species of Anillinus Casey (Carabidae: Trechinae: Bembidiini) from the Southern Appalachians and phylogeography of the A. loweae species group. Zootaxa 2502: 1 - 23." type="journal article" year="2010">Sokolov and Carlton 2010</bibRefCitation>
|
||
), which is characterized by two paramedial patches of smoothed microsculpture on the head. It is distinguished from all other species in this group by the form of the median lobe and armature of the internal sac, and especially by the smaller size.
|
||
</paragraph>
|
||
<paragraph id="8BEC36A6FF957733A7B62951FD5AFC2E" blockId="10.[192,1440,643,1051]" pageId="10" pageNumber="9">
|
||
<taxonomicName id="4C534D25FF957733A7B62951FE13FCEE" authorityName="Sokolov" authorityYear="2011" box="[240,481,771,795]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="chilhowee">
|
||
<emphasis id="B927EAB4FF957733A7B62951FE13FCEE" box="[240,481,771,795]" italics="true" pageId="10" pageNumber="9">Anillinus chilhowee</emphasis>
|
||
</taxonomicName>
|
||
occurs sympatrically with one or two species of
|
||
<taxonomicName id="4C534D25FF957733A3592951FB60FCEE" authorityName="Casey" authorityYear="1918" box="[1055,1170,771,795]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B927EAB4FF957733A3592951FB60FCEE" box="[1055,1170,771,795]" italics="true" pageId="10" pageNumber="9">Anillinus</emphasis>
|
||
</taxonomicName>
|
||
, and undescribed species from the langdoni
|
||
<emphasis id="B927EAB4FF957733A6892976FE2AFCCE" box="[463,472,804,827]" italics="true" pageId="10" pageNumber="9">-</emphasis>
|
||
group (
|
||
<bibRefCitation id="EFC24B57FF957733A56B2971FCE4FCCE" author="Sokolov, I. M. & Y. Y. Sokolova & C. E. Carlton" box="[557,790,803,827]" pageId="10" pageNumber="9" pagination="1 - 20" refId="ref9158" refString="Sokolov, I. M., Y. Y. Sokolova, and C. E. Carlton. 2007. New species of Anillinus Casey (Carabidae: Trechinae: Bembidiini) from Great Smoky Mountains National Park, U. S. A. and phylogeography of the A. langdoni species group. Zootaxa 1542: 1 - 20." type="journal article" year="2007">Sokolov et al. 2007</bibRefCitation>
|
||
), and, possibly,
|
||
<taxonomicName id="4C534D25FF957733A4902976FB44FCCE" authority="Barr. It" authorityName="Barr. It" authorityYear="1995" box="[982,1206,804,827]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="steevesi">
|
||
<emphasis id="B927EAB4FF957733A4902976FBA1FCCE" box="[982,1107,804,827]" italics="true" pageId="10" pageNumber="9">A. steevesi</emphasis>
|
||
Barr. It
|
||
</taxonomicName>
|
||
is also presumably sympatric with one or two species of
|
||
<taxonomicName id="4C534D25FF957733A5382911FCF9FCAE" box="[638,779,835,859]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B927EAB4FF957733A5382911FCF9FCAE" box="[638,779,835,859]" italics="true" pageId="10" pageNumber="9">Serranillus</emphasis>
|
||
</taxonomicName>
|
||
. Externally, it can be distinguished from all species of
|
||
<taxonomicName id="4C534D25FF957733A7862931FEBAFC8E" box="[192,328,867,891]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B927EAB4FF957733A7862931FEBAFC8E" box="[192,328,867,891]" italics="true" pageId="10" pageNumber="9">Serranillus</emphasis>
|
||
</taxonomicName>
|
||
by the presence of long discal elytral setae. From the species of the langdoni-group
|
||
<taxonomicName id="4C534D25FF957733A24B2936FA52FC8E" authorityName="Sokolov" authorityYear="2011" box="[1293,1440,867,891]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="chilhowee">
|
||
<emphasis id="B927EAB4FF957733A24B2936FA52FC8E" box="[1293,1440,867,891]" italics="true" pageId="10" pageNumber="9">A. chilhowee</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by the presence of areas with smoothed microsculpture on the head and pronotum, and from
|
||
<taxonomicName id="4C534D25FF957733A67229F6FE42FC4E" authorityName="Barr. It" authorityYear="1995" box="[308,432,932,955]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="steevesi">
|
||
<emphasis id="B927EAB4FF957733A67229F6FE42FC4E" box="[308,432,932,955]" italics="true" pageId="10" pageNumber="9">A. steevesi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C534D25FF957733A6FB29F6FDA4FC4E" authorityName="Sokolov" authorityYear="2011" box="[445,598,931,955]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="chilhowee">
|
||
<emphasis id="B927EAB4FF957733A6FB29F6FDA4FC4E" box="[445,598,931,955]" italics="true" pageId="10" pageNumber="9">A. chilhowee</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by the smaller size (1.64 versus
|
||
<quantity id="4CAB9B43FF957733A39829F1FA76FC4E" box="[1246,1412,931,955]" metricMagnitude="-3" metricUnit="m" metricValue="1.795" metricValueMax="1.92" metricValueMin="1.67" pageId="10" pageNumber="9" unit="mm" value="1.795" valueMax="1.92" valueMin="1.67">1.67-1.92 mm</quantity>
|
||
of
|
||
<taxonomicName id="4C534D25FF957733A7862996FECEFC2E" authorityName="Barr. It" authorityYear="1995" box="[192,316,964,987]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="steevesi">
|
||
<emphasis id="B927EAB4FF957733A7862996FECEFC2E" box="[192,316,964,987]" italics="true" pageId="10" pageNumber="9">A. steevesi</emphasis>
|
||
</taxonomicName>
|
||
) and form of the median lobe.
|
||
</paragraph>
|
||
<paragraph id="8BEC36A6FF957733A7B629B1FE02FBEE" blockId="10.[192,1440,643,1051]" pageId="10" pageNumber="9">
|
||
The key to the loweae-group species (
|
||
<bibRefCitation id="EFC24B57FF957733A5EB29B1FC14FC0E" author="Sokolov, I. M. & C. E. Carlton" box="[685,998,995,1019]" pageId="10" pageNumber="9" pagination="1 - 23" refId="ref9274" refString="Sokolov, I. M., and C. E. Carlton. 2010. New species of Anillinus Casey (Carabidae: Trechinae: Bembidiini) from the Southern Appalachians and phylogeography of the A. loweae species group. Zootaxa 2502: 1 - 23." type="journal article" year="2010">Sokolov and Carlton 2010</bibRefCitation>
|
||
) is modified from couplet 8 to accommodate the
|
||
<taxonomicName id="4C534D25FF957733A6092E56FE1AFBEE" authorityName="Sokolov" authorityYear="2011" box="[335,488,1027,1051]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="chilhowee">
|
||
<emphasis id="B927EAB4FF957733A6092E56FE1AFBEE" box="[335,488,1027,1051]" italics="true" pageId="10" pageNumber="9">A. chilhowee</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
</paragraph>
|
||
<paragraph id="8BEC36A6FF957733A7862E11FA52FB4E" blockId="10.[192,1440,1091,1275]" pageId="10" pageNumber="9">
|
||
8. Male median lobe with dorsal sclerites very long, forming filament-like structures with greatly elongated basal processes. Setae of right paramere long, much longer than the length of paramere itself. Beetles from the limestone area at the eastern part of Rich Mountain ridge in Great Smoky Mountains National Park ....................................
|
||
<taxonomicName id="4C534D25FF957733A4AA2EF1FA52FB4E" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" box="[1004,1440,1186,1211]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="gimmeli">
|
||
<emphasis id="B927EAB4FF957733A4AA2EF1FA52FB4E" bold="true" box="[1004,1440,1186,1211]" pageId="10" pageNumber="9">
|
||
<emphasis id="B927EAB4FF957733A4AA2EF1FB8DFB4E" bold="true" box="[1004,1151,1186,1211]" italics="true" pageId="10" pageNumber="9">A. gimmeli</emphasis>
|
||
Sokolov and Carlton
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BEC36A6FF957733A7862E96FA52FB0E" blockId="10.[192,1440,1091,1275]" pageId="10" pageNumber="9">
|
||
— Male median lobe with dorsal sclerites shorter, forming blade-like structures. Setae of right paramere shorter than the length of paramere itself ...........................................................
|
||
<emphasis id="B927EAB4FF957733A23A2EB1FA52FB0E" bold="true" box="[1404,1440,1251,1275]" pageId="10" pageNumber="9">8A</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BEC36A6FF957733A7862F71FA52FA8E" blockId="10.[192,1440,1315,1531]" pageId="10" pageNumber="9">
|
||
8A(8). Median lobe with strongly enlarged and symmetrically rounded apex, and without a patch of spines near apical orifice. Beetles from Chilhowee Mountain (Polk County, TN) to the south of the Hiwassee River ................................................................................
|
||
<emphasis id="B927EAB4FF957733A3CC2F31FA52FA8E" bold="true" box="[1162,1440,1378,1403]" pageId="10" pageNumber="9">
|
||
<taxonomicName id="4C534D25FF957733A3CC2F31FAC5FA8E" authority="Sokolov, 2011" authorityName="Sokolov" authorityYear="2011" box="[1162,1335,1378,1403]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="chilhowee" status="sp. nov.">
|
||
<emphasis id="B927EAB4FF957733A3CC2F31FAC5FA8E" bold="true" box="[1162,1335,1378,1403]" italics="true" pageId="10" pageNumber="9">A. chilhowee</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21457CFFF957733A2792F31FA52FA8E" box="[1343,1440,1379,1403]" pageId="10" pageNumber="9" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BEC36A6FF957733A7862FD6FA52FA0E" blockId="10.[192,1440,1315,1531]" pageId="10" pageNumber="9">
|
||
— Apex of median lobe EITHER with sinuation on dorsal side, OR noticeably asymmetrical, OR not enlarged (its width much lesser than half width of median lobe itself). Beetles from eastern portions of southern Appalachian Mountains to the north and east of the Hiwassee River ...... .........................................................................................................................
|
||
<emphasis id="B927EAB4FF957733A3942FB1FB13FA0E" bold="true" box="[1234,1249,1507,1531]" pageId="10" pageNumber="9">9</emphasis>
|
||
(follow the key)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |