260 lines
35 KiB
XML
260 lines
35 KiB
XML
<document id="0F0CEF4DCEDADD2D2BCCBDD38877915A" ID-DOI="10.5281/zenodo.5160535" ID-ISSN="1942-1354" ID-Zenodo-Dep="5160535" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628108913532" checkinUser="felipe" docAuthor="Sokolov, Igor M." docDate="2011" docId="03FA87B0FF957735A7862C70FAE7FEAE" docLanguage="en" docName="InsectaMundi.0164.1-14.pdf" docOrigin="Insecta Mundi 2011 (164)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Anillinus smokiensis Sokolov 2011, new species" docType="treatment" docVersion="3" lastPageNumber="11" masterDocId="FFC3FFC8FF9F7739A7462A52FFF2FFF5" masterDocTitle="Five new species of Anillinus Casey from the Southern Appalachian Mountains and the Piedmont Plateau of eastern U. S. A. (Coleoptera: Carabidae: Trechinae: Bembidiini)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="9" updateTime="1699034592004" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="17C2EB06E6CBC0C5691CE16D222937DE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A7E98B99F7DA9D67C3C824B8079974CD">
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<mods:title id="17C8DBF08417D2464EA408715297023C">Five new species of Anillinus Casey from the Southern Appalachian Mountains and the Piedmont Plateau of eastern U. S. A. (Coleoptera: Carabidae: Trechinae: Bembidiini)</mods:title>
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</mods:titleInfo>
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<mods:name id="B89D049737BF467DA9A071DEB9A6F9A7" type="personal">
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<mods:role id="79F8AB431A1BF5063099B323B573D263">
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<mods:roleTerm id="932382133B056AB8E8B578928240E071">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B073EB912AF637FE9EBD8E0B0EBA6089">Sokolov, Igor M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="937876AFACAF0075CC21F1D9DEEA0AB7">text</mods:typeOfResource>
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<mods:titleInfo id="F23E1606DD2D2ABD190DB89D292B2A98">
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<mods:title id="A66C9C2B5A26C23D55CCFB2186EC3574">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="C010D770230C991FC8FCB8C8DBBDB9DB">
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<mods:date id="720E58804E04924AF4F662E01FE84D08">2011</mods:date>
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<mods:detail id="9E8A766D1284D7D62904123FC7D42188" type="pubDate">
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<mods:number id="11948AF16E211A9B9486DC28D93F1DAE">2011-04-15</mods:number>
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</mods:detail>
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<mods:detail id="0C0721B3E8CF292A830330C067763B08" type="volume">
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<mods:number id="7EF3962FDADD9A819C28B4A564671E59">2011</mods:number>
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</mods:detail>
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<mods:detail id="570EC9E2C85EC117B5C2D9A3B9EB93C2" type="issue">
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<mods:number id="DEFB52D97EC65AFFA13654860BA59F0D">164</mods:number>
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</mods:detail>
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<mods:extent id="8E79C0D5A2DE7C4061117088573E1C47" unit="page">
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<mods:start id="93B25D84466800B95EA42AF4E5FA812A">1</mods:start>
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<mods:end id="034C1AA1EE0D6B168AD0F3D8A7D32BF6">14</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="E2E936E8296D76ACD06DEAF51DA4D0B8">journal article</mods:classification>
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<mods:identifier id="31428132806BCA56E1F7723F6D10022D" type="DOI">10.5281/zenodo.5160535</mods:identifier>
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<mods:identifier id="A602073002162A9D376CF9A6162D87B2" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="230E6B8B32B941BC0483F9F4081C2DF7" type="Zenodo-Dep">5160535</mods:identifier>
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</mods:mods>
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<treatment id="03FA87B0FF957735A7862C70FAE7FEAE" LSID="urn:lsid:plazi:treatment:03FA87B0FF957735A7862C70FAE7FEAE" httpUri="http://treatment.plazi.org/id/03FA87B0FF957735A7862C70FAE7FEAE" lastPageId="12" lastPageNumber="11" pageId="10" pageNumber="9">
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<subSubSection id="C349652DFF957733A7862C70FD0CF9CE" box="[192,766,1570,1595]" pageId="10" pageNumber="9" type="nomenclature">
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<paragraph id="8BEC36A6FF957733A7862C70FD0CF9CE" blockId="10.[192,766,1570,1627]" box="[192,766,1570,1595]" pageId="10" pageNumber="9">
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<heading id="D0A481CAFF957733A7862C70FD0CF9CE" bold="true" box="[192,766,1570,1595]" fontSize="10" level="2" pageId="10" pageNumber="9" reason="3">
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<taxonomicName id="4C534D25FF957733A7862C70FDBCF9CE" ID-CoL="6727G" authority="Sokolov" authorityName="Sokolov" authorityYear="2011" box="[192,590,1570,1595]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="smokiensis" status="sp. nov.">
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<emphasis id="B927EAB4FF957733A7862C70FDBCF9CE" bold="true" box="[192,590,1570,1595]" pageId="10" pageNumber="9">
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<emphasis id="B927EAB4FF957733A7862C70FE29F9CE" bold="true" box="[192,475,1570,1595]" italics="true" pageId="10" pageNumber="9">Anillinus smokiensis</emphasis>
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Sokolov
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</emphasis>
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</taxonomicName>
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,
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<emphasis id="B927EAB4FF957733A51A2C71FD0CF9CE" bold="true" box="[604,766,1571,1595]" pageId="10" pageNumber="9">
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<taxonomicNameLabel id="A21457CFFF957733A51A2C71FD0CF9CE" box="[604,766,1571,1595]" pageId="10" pageNumber="9" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C349652DFF957733A7862C11FE6CF9AE" box="[192,414,1603,1627]" pageId="10" pageNumber="9" type="description">
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<paragraph id="8BEC36A6FF957733A7862C11FE6CF9AE" blockId="10.[192,766,1570,1627]" box="[192,414,1603,1627]" pageId="10" pageNumber="9">
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<figureCitation id="13682A23FF957733A7862C11FED9F9AE" box="[192,299,1603,1627]" captionStart="Figure 1-5" captionStartId="3.[192,276,1475,1497]" captionTargetBox="[197,1384,196,1453]" captionTargetId="figure-160@3.[192,1440,192,1458]" captionTargetPageId="3" captionText="Figure 1-5. Habitus images. 1) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.), holotype. 2) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.), paratype. 3) A. chilhowee (TN, Polk Co., Chilhowee Mtn.), holotype. 4) A. smokiensis (TN, Blount Co., GSMNP), paratype. 5) A. chandleri (SC, Edgefield Co., Sumter National Forest), holotype." figureDoi="http://doi.org/10.5281/zenodo.5160539" httpUri="https://zenodo.org/record/5160539/files/figure.png" pageId="10" pageNumber="9">Figure 4</figureCitation>
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,
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<figureCitation id="13682A23FF957733A6712C11FEBAF9AE" box="[311,328,1603,1627]" captionStart="Figure 6-10" captionStartId="4.[192,276,877,899]" captionTargetBox="[192,1440,193,852]" captionTargetId="figure-262@4.[192,1440,192,852]" captionTargetPageId="4" captionText="Figure 6-10. Pronotum images. 6) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.). 7) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.). 8) A. chilhowee (TN, Polk Co., Chilhowee Mtn.). 9) A. smokiensis (TN, Blount Co., GSMNP). 10) A. chandleri (SC, Edgefield Co., Sumter National Forest)." figureDoi="http://doi.org/10.5281/zenodo.5160541" httpUri="https://zenodo.org/record/5160541/files/figure.png" pageId="10" pageNumber="9">9</figureCitation>
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,
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<figureCitation id="13682A23FF957733A6122C11FE86F9AE" box="[340,372,1603,1627]" captionStart="Figure 11-15" captionStartId="5.[192,276,1691,1713]" captionTargetBox="[202,1419,207,1664]" captionTargetId="figure-14@5.[192,1440,192,1696]" captionTargetPageId="5" captionText="Figure 11-15. Illustrations of male aedeagus. 11) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.). 12) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.). 13) A. chilhowee (TN, Polk Co., Chilhowee Mtn.). 14) A. smokiensis (TN, Blount Co., GSMNP). 15) A. chandleri (SC, Edgefield Co., Sumter National Forest). a-Median lobe, right lateral aspect; b-Left paramere, left lateral aspect; c-Right paramere, right lateral aspect. Scale = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5160545" httpUri="https://zenodo.org/record/5160545/files/figure.png" pageId="10" pageNumber="9">14</figureCitation>
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,
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<figureCitation id="13682A23FF957733A6C62C11FE6CF9AE" box="[384,414,1603,1627]" captionStart="Figure 17" captionStartId="7.[192,275,991,1013]" captionTargetBox="[192,1440,192,969]" captionTargetId="figure-273@7.[192,1440,192,969]" captionTargetPageId="7" captionText="Figure 17. Locality records for the valentinei-group and cornelli-group species. Anillinus valentinei – red triangle; A. smokiensis – red quadrangle; A. chandleri – red star; A. cornelli – red circles." figureDoi="http://doi.org/10.5281/zenodo.5160551" httpUri="https://zenodo.org/record/5160551/files/figure.png" pageId="10" pageNumber="9">17</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C349652DFF957732A7862CD1FC9DFECE" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9" type="materials_examined">
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<paragraph id="8BEC36A6FF957733A7862CD1FC88F90E" blockId="10.[192,1440,1667,1787]" pageId="10" pageNumber="9">
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<materialsCitation id="3B3B3CFBFF957733A7862CD1FC2BF94E" collectingDate="2006-04-14" collectorName="Litter & A. K. Tishechkin" county="Blount Co." elevation="605" latitude="35.609833" location="Gregory cave" longLatPrecision="11" longitude="-83.80583" municipality="Male" pageId="10" pageNumber="9" specimenCount="605" stateProvince="Tennessee" typeStatus="holotype">
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<emphasis id="B927EAB4FF957733A7862CD1FEB5F96E" bold="true" box="[192,327,1667,1691]" pageId="10" pageNumber="9">
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<typeStatus id="54E88804FF957733A7862CD1FEB3F96E" box="[192,321,1667,1691]" pageId="10" pageNumber="9" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingMunicipality id="6B88ACDCFF957733A6092CD1FE7FF96E" box="[335,397,1667,1691]" pageId="10" pageNumber="9">Male</collectingMunicipality>
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labeled /
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<collectingRegion id="4997F844FF957733A5472CD1FD42F96E" box="[513,688,1667,1691]" country="United States of America" name="Tennessee" pageId="10" pageNumber="9">TENNESSEE</collectingRegion>
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:
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<collectingCounty id="628D4E2AFF957733A5FA2CD1FCB6F96E" box="[700,836,1667,1691]" pageId="10" pageNumber="9">Blount Co.</collectingCounty>
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, GSMNP,
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<location id="8E8C607DFF957733A48F2CD1FB82F96E" LSID="urn:lsid:plazi:treatment:03FA87B0FF957735A7862C70FAE7FEAE:8E8C607DFF957733A48F2CD1FB82F96E" box="[969,1136,1667,1691]" county="Blount Co." latitude="35.609833" longLatPrecision="11" longitude="-83.80583" municipality="Male" name="Gregory cave" pageId="10" pageNumber="9" stateProvince="Tennessee">Gregory cave</location>
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,
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<geoCoordinate id="EE675061FF957733A33B2CD1FAF1F96E" box="[1149,1283,1667,1691]" degrees="35" direction="north" minutes="36.59" orientation="latitude" pageId="10" pageNumber="9" precision="9" value="35.609833">35°36.59’N</geoCoordinate>
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<geoCoordinate id="EE675061FF957733A24B2CD1FA69F96E" box="[1293,1435,1667,1691]" degrees="83" direction="west" minutes="48.35" orientation="longitude" pageId="10" pageNumber="9" precision="9" value="-83.80583">83°48.35’W</geoCoordinate>
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,
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<specimenCount id="9D55FD2FFF957733A7862CF1FEFEF94E" box="[192,268,1699,1723]" pageId="10" pageNumber="9" type="generic">
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<quantity id="4CAB9B43FF957733A7862CF1FEFAF94E" box="[192,264,1699,1723]" metricMagnitude="2" metricUnit="m" metricValue="6.05" pageId="10" pageNumber="9" unit="m" value="605.0">
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<elevation id="007ED195FF957733A7862CF1FEFAF94E" box="[192,264,1699,1723]" metricMagnitude="2" metricUnit="m" metricValue="6.05" pageId="10" pageNumber="9" unit="m" value="605.0">605m</elevation>
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</quantity>
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.
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</specimenCount>
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<collectorName id="26A65370FF957733A6532CF1FEACF94E" box="[277,350,1699,1723]" pageId="10" pageNumber="9">Litter</collectorName>
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sifting at entrance.
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<date id="FFED1066FF957733A51B2CF1FCF5F94E" box="[605,775,1699,1723]" pageId="10" pageNumber="9" value="2006-04-14">
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<collectingDate id="EFA9E98EFF957733A51B2CF1FCF5F94E" box="[605,775,1699,1723]" pageId="10" pageNumber="9" value="2006-04-14">14 April 2006</collectingDate>
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</date>
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<collectorName id="26A65370FF957733A4492CF1FC2BF94E" box="[783,985,1699,1723]" pageId="10" pageNumber="9">A.K. Tishechkin</collectorName>
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</materialsCitation>
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/
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<materialsCitation id="3B3B3CFBFF957733A4B72CF1FC84F90E" collectionCode="USNM" pageId="10" pageNumber="9" specimenCount="2" stateProvince="Tennessee" typeStatus="holotype">
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<typeStatus id="54E88804FF957733A4B72CF1FB62F94E" box="[1009,1168,1699,1723]" pageId="10" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
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,
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<taxonomicName id="4C534D25FF957733A3DA2CF1FED1F92E" authority="Sokolov" authorityName="Sokolov" authorityYear="2011" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="smokiensis">
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<emphasis id="B927EAB4FF957733A3DA2CF1FA52F94E" box="[1180,1440,1699,1723]" italics="true" pageId="10" pageNumber="9">Anillinus smokiensis</emphasis>
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Sokolov
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</taxonomicName>
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, des. 2009/. The
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<typeStatus id="54E88804FF957733A6A92C91FDABF92E" box="[495,601,1731,1755]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
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is dissected and bears a plastic rectangle with genitalia mounted in dimethylhydantoin formaldehyde resin. Deposited
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<collectionCode id="ED42AE63FF957733A4582CB1FC84F90E" box="[798,886,1763,1787]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="9" type="Museum">USNM</collectionCode>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BEC36A6FF957733A7862D71FBECF8AE" blockId="10.[192,1440,1827,1883]" pageId="10" pageNumber="9">
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<emphasis id="B927EAB4FF957733A7862D71FE8CF8CE" bold="true" box="[192,382,1827,1851]" pageId="10" pageNumber="9">Type locality.</emphasis>
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<collectingCountry id="F3447636FF957733A6C12D71FE2AF8CE" box="[391,472,1827,1851]" name="United States of America" pageId="10" pageNumber="9">U.S.A.</collectingCountry>
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<collectingRegion id="4997F844FF957733A6A72D76FD94F8CE" box="[481,614,1828,1851]" country="United States of America" name="Tennessee" pageId="10" pageNumber="9">Tennessee</collectingRegion>
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, Blount County, Great Smoky Mountain National Park, southern slopes of Rich Mountain ridge, Gregory Cave at
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<geoCoordinate id="EE675061FF957733A4462D11FC74F8AE" box="[768,902,1859,1883]" degrees="35" direction="north" minutes="36.59" orientation="latitude" pageId="10" pageNumber="9" precision="9" value="35.609833">35°36.59’N</geoCoordinate>
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<geoCoordinate id="EE675061FF957733A4CA2D11FBE8F8AE" box="[908,1050,1859,1883]" degrees="83" direction="west" minutes="48.35" orientation="longitude" pageId="10" pageNumber="9" precision="9" value="-83.80583">83°48.35’W</geoCoordinate>
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.
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</paragraph>
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<paragraph id="8BEC36A6FF947732A7862A91FC9DFECE" blockId="11.[192,1440,195,315]" pageId="11" pageNumber="10">
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<emphasis id="B927EAB4FF947732A7862A91FEBCFF2E" bold="true" box="[192,334,195,219]" pageId="11" pageNumber="10">
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<typeStatus id="54E88804FF947732A7862A91FEBCFF2E" box="[192,334,195,219]" pageId="11" pageNumber="10" type="paratype">Paratypes</typeStatus>
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</emphasis>
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(7). One male with the same data as
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<typeStatus id="54E88804FF947732A4502A91FC76FF2E" box="[790,900,195,219]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
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. Two males and
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<specimenCount id="9D55FD2FFF947732A3132A91FB1CFF2E" box="[1109,1262,195,219]" pageId="11" pageNumber="10" type="female">four females</specimenCount>
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labeled /TEN- NESSEE: Blount Co., GSMNP, Gregory cave,
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<geoCoordinate id="EE675061FF947732A5AB2AB1FC81FF0E" box="[749,883,227,251]" degrees="35" direction="north" minutes="36.59" orientation="latitude" pageId="11" pageNumber="10" precision="9" value="35.609833">35°36.59’N</geoCoordinate>
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<geoCoordinate id="EE675061FF947732A43F2AB1FBF5FF0E" box="[889,1031,227,251]" degrees="83" direction="west" minutes="48.35" orientation="longitude" pageId="11" pageNumber="10" precision="9" value="-83.80583">83°48.35’W</geoCoordinate>
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. Litter sifting at entrance.
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<date id="FFED1066FF947732A20D2AB1FF09FEEE" pageId="11" pageNumber="10" value="2004-07-28">28.VII. 2004</date>
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<collectionCode id="ED42AE63FF947732A6462B51FEE4FEEE" box="[256,278,259,283]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="10" type="Museum">V</collectionCode>
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.Bayless C.Carlton A. Tishechkin/ and
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<specimenCount id="9D55FD2FFF947732A5A22B56FCA2FEEE" box="[740,848,259,283]" pageId="11" pageNumber="10" type="male">one male</specimenCount>
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and
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<specimenCount id="9D55FD2FFF947732A4CF2B56FBF9FEEE" box="[905,1035,259,283]" pageId="11" pageNumber="10" type="female">one female</specimenCount>
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from those bear labels /Molecular voucher # 93/ and /Molecular voucher # 94/, respectively.
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</paragraph>
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</subSubSection>
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<subSubSection id="C349652DFF947732A7862B31FDEAFE6E" pageId="11" pageNumber="10" type="etymology">
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<paragraph id="8BEC36A6FF947732A7862B31FDEAFE6E" blockId="11.[192,1440,355,411]" pageId="11" pageNumber="10">
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<emphasis id="B927EAB4FF947732A7862B31FEA9FE8E" bold="true" box="[192,347,355,379]" pageId="11" pageNumber="10">Etymology.</emphasis>
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The name of this species is a Latinized adjective based on the name of the mountain region in which this species occurs.
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</paragraph>
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</subSubSection>
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<subSubSection id="C349652DFF947732A7862B91FC36FAAE" pageId="11" pageNumber="10" type="description">
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||
<paragraph id="8BEC36A6FF947732A7862B91FBF1FD6E" blockId="11.[192,1441,451,1371]" pageId="11" pageNumber="10">
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<emphasis id="B927EAB4FF947732A7862B91FE96FE2E" bold="true" box="[192,356,451,475]" pageId="11" pageNumber="10">Description</emphasis>
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. Large-sized for genus (ABL range
|
||
<quantity id="4CAB9B43FF947732A4672B91FC3FFE2E" box="[801,973,451,475]" metricMagnitude="-3" metricUnit="m" metricValue="1.895" metricValueMax="1.93" metricValueMin="1.86" pageId="11" pageNumber="10" unit="mm" value="1.895" valueMax="1.93" valueMin="1.86">1.86-1.93 mm</quantity>
|
||
, mean 1.90±
|
||
<quantity id="4CAB9B43FF947732A32E2B91FB1AFE2E" box="[1128,1256,451,475]" metricMagnitude="-5" metricUnit="m" metricValue="2.9" pageId="11" pageNumber="10" unit="mm" value="0.029">0.029 mm</quantity>
|
||
, n=7). Habitus (
|
||
<figureCitation id="13682A23FF947732A78F2BB1FEFFFE0E" box="[201,269,483,507]" captionStart="Figure 1-5" captionStartId="3.[192,276,1475,1497]" captionTargetBox="[197,1384,196,1453]" captionTargetId="figure-160@3.[192,1440,192,1458]" captionTargetPageId="3" captionText="Figure 1-5. Habitus images. 1) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.), holotype. 2) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.), paratype. 3) A. chilhowee (TN, Polk Co., Chilhowee Mtn.), holotype. 4) A. smokiensis (TN, Blount Co., GSMNP), paratype. 5) A. chandleri (SC, Edgefield Co., Sumter National Forest), holotype." figureDoi="http://doi.org/10.5281/zenodo.5160539" httpUri="https://zenodo.org/record/5160539/files/figure.png" pageId="11" pageNumber="10">Fig. 4</figureCitation>
|
||
) moderately convex, subparallel (WE/ABL 0.37±0.012), head of normal proportions for the genus (WH/WPm 0.74±0.019), pronotum narrow compared to elytra (WPm/WE 0.81±0.017). Body color rufotestaceous, appendages testaceous. Dorsal microsculpture mostly effaced, polygonal microsculpture present on a small triangular area at middle of vertex and in frontal furrows on head, and also on base of pronotum; most parts of vertex, lateral parts of head, disc and front part of pronotum with effaced microsculpture. Elytra with well-developed polygonal microsculpture.
|
||
</paragraph>
|
||
<paragraph id="8BEC36A6FF947732A7B628F6FBFBFCEE" blockId="11.[192,1441,451,1371]" pageId="11" pageNumber="10">
|
||
Pronotum (
|
||
<figureCitation id="13682A23FF947732A63828F1FE34FD4E" box="[382,454,675,699]" captionStart="Figure 6-10" captionStartId="4.[192,276,877,899]" captionTargetBox="[192,1440,193,852]" captionTargetId="figure-262@4.[192,1440,192,852]" captionTargetPageId="4" captionText="Figure 6-10. Pronotum images. 6) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.). 7) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.). 8) A. chilhowee (TN, Polk Co., Chilhowee Mtn.). 9) A. smokiensis (TN, Blount Co., GSMNP). 10) A. chandleri (SC, Edgefield Co., Sumter National Forest)." figureDoi="http://doi.org/10.5281/zenodo.5160541" httpUri="https://zenodo.org/record/5160541/files/figure.png" pageId="11" pageNumber="10">Fig. 9</figureCitation>
|
||
) moderately convex and comparatively transverse (WPm/LP 1.33±0.025), with margins rectilinear and moderately constricted posteriad (WPm/WPp 1.31±0.023). Anterior angles indistinct, only slightly prominent. Posterior angles obtuse (100-120°). Width between posterior angles only slightly greater than between anterior angles (WPa/WPp 0.97±0.020).
|
||
</paragraph>
|
||
<paragraph id="8BEC36A6FF947732A7B62971FEFEFC4E" blockId="11.[192,1441,451,1371]" pageId="11" pageNumber="10">Elytra moderately convex, slightly depressed along suture, of normal length for genus (LE/ABL 0.56±0.009), with traces of 3-4 interneurs. Humeri rectangular, moderately rounded. Margins subparallel, slightly divergent in basal forth, evenly rounded to apex in apical third, maximal width of elytra at midpoint. Elytra without subapical sinuation. Vestiture of elytra short (lesser than one-third of discal setae).</paragraph>
|
||
<paragraph id="8BEC36A6FF947732A7B62991FD03FC0E" blockId="11.[192,1441,451,1371]" pageId="11" pageNumber="10">Prothoracic leg of males with moderately dilated tarsomere 1. Profemur moderately swollen. Metafemora unmodified. Sternum VII of males unmodified.</paragraph>
|
||
<paragraph id="8BEC36A6FF947732A7B62E51FC36FAAE" blockId="11.[192,1441,451,1371]" pageId="11" pageNumber="10">
|
||
Median lobe of aedeagus (
|
||
<figureCitation id="13682A23FF947732A56C2E51FD7EFBEE" box="[554,652,1027,1051]" captionStart="Figure 11-15" captionStartId="5.[192,276,1691,1713]" captionTargetBox="[202,1419,207,1664]" captionTargetId="figure-14@5.[192,1440,192,1696]" captionTargetPageId="5" captionText="Figure 11-15. Illustrations of male aedeagus. 11) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.). 12) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.). 13) A. chilhowee (TN, Polk Co., Chilhowee Mtn.). 14) A. smokiensis (TN, Blount Co., GSMNP). 15) A. chandleri (SC, Edgefield Co., Sumter National Forest). a-Median lobe, right lateral aspect; b-Left paramere, left lateral aspect; c-Right paramere, right lateral aspect. Scale = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5160545" httpUri="https://zenodo.org/record/5160545/files/figure.png" pageId="11" pageNumber="10">Fig. 14a</figureCitation>
|
||
) evenly arcuate and twisted, with pointed apex slightly tapering and shortly rounded at tip. The wall of median lobe forming evident incision and characteristic beak like fold above the apex. Ventral margin of median lobe not enlarged, bearing numerous poriferous canals scattered across the ventral side of the lobe. Dorsal copulatory sclerites forming a curled blade-like structure with characteristic basal prolongations. Ventral sclerite of internal sac absent. Spines represented by three groups: 7-8 small spines are nearby the apical part of dorsal sclerites, group of about 20 long spines occupy the space near apex of median lobe, and two spines above the preceding group of spines. Also a small band with minute sclerotized setose structures extends above the end of dorsal sclerites in ostial area. Left paramere (
|
||
<figureCitation id="13682A23FF947732A6F82F51FDD2FAEE" box="[446,544,1283,1307]" captionStart="Figure 11-15" captionStartId="5.[192,276,1691,1713]" captionTargetBox="[202,1419,207,1664]" captionTargetId="figure-14@5.[192,1440,192,1696]" captionTargetPageId="5" captionText="Figure 11-15. Illustrations of male aedeagus. 11) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.). 12) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.). 13) A. chilhowee (TN, Polk Co., Chilhowee Mtn.). 14) A. smokiensis (TN, Blount Co., GSMNP). 15) A. chandleri (SC, Edgefield Co., Sumter National Forest). a-Median lobe, right lateral aspect; b-Left paramere, left lateral aspect; c-Right paramere, right lateral aspect. Scale = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5160545" httpUri="https://zenodo.org/record/5160545/files/figure.png" pageId="11" pageNumber="10">Fig. 14b</figureCitation>
|
||
) not enlarged, paramere apex with four poriferous canals, but bearing only two small setae in distal positions. Right paramere (
|
||
<figureCitation id="13682A23FF947732A4142F71FC47FACE" box="[850,949,1315,1339]" captionStart="Figure 11-15" captionStartId="5.[192,276,1691,1713]" captionTargetBox="[202,1419,207,1664]" captionTargetId="figure-14@5.[192,1440,192,1696]" captionTargetPageId="5" captionText="Figure 11-15. Illustrations of male aedeagus. 11) Anillinus unicoi (NC, Graham Co., Unicoi Mtns.). 12) A. carltoni (TN, Blount Co., GSMNP, Thunderhead Mtn.). 13) A. chilhowee (TN, Polk Co., Chilhowee Mtn.). 14) A. smokiensis (TN, Blount Co., GSMNP). 15) A. chandleri (SC, Edgefield Co., Sumter National Forest). a-Median lobe, right lateral aspect; b-Left paramere, left lateral aspect; c-Right paramere, right lateral aspect. Scale = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5160545" httpUri="https://zenodo.org/record/5160545/files/figure.png" pageId="11" pageNumber="10">Fig. 14c</figureCitation>
|
||
) short, with subparallel apical portion, bearing four long setae that are longer than the paramere itself.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349652DFF947732A7862FD1FC98F9CE" pageId="11" pageNumber="10" type="distribution">
|
||
<paragraph id="8BEC36A6FF947732A7862FD1FC96FA4E" blockId="11.[192,1440,1411,1467]" pageId="11" pageNumber="10">
|
||
<emphasis id="B927EAB4FF947732A7862FD1FE87FA6E" bold="true" box="[192,373,1411,1435]" pageId="11" pageNumber="10">Distribution.</emphasis>
|
||
Known only from Blount County, TN, where it was collected in a cave on the southern slopes of Rich Mountain ridge, within GSMNP (
|
||
<figureCitation id="13682A23FF947732A4462FF1FCA4FA4E" box="[768,854,1443,1467]" captionStart="Figure 17" captionStartId="7.[192,275,991,1013]" captionTargetBox="[192,1440,192,969]" captionTargetId="figure-273@7.[192,1440,192,969]" captionTargetPageId="7" captionText="Figure 17. Locality records for the valentinei-group and cornelli-group species. Anillinus valentinei – red triangle; A. smokiensis – red quadrangle; A. chandleri – red star; A. cornelli – red circles." figureDoi="http://doi.org/10.5281/zenodo.5160551" httpUri="https://zenodo.org/record/5160551/files/figure.png" pageId="11" pageNumber="10">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEC36A6FF947732A7862FB1FC98F9CE" blockId="11.[192,1440,1507,1595]" pageId="11" pageNumber="10">
|
||
<emphasis id="B927EAB4FF947732A7862FB1FEC3FA0E" bold="true" box="[192,305,1507,1531]" pageId="11" pageNumber="10">Habitat.</emphasis>
|
||
The series of specimens was collected by sifting a thick, wet layer of leaves along the dry bed of an ephemeral stream near the entrance of the cave. The locality is situated in a limestone area in pinemixed hardwood forest at relatively low altitude (
|
||
<quantity id="4CAB9B43FF947732A4562C71FCAFF9CE" box="[784,861,1571,1595]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="10" unit="m" value="600.0">600 m</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349652DFF947735A7862C31FAE7FEAE" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10" type="diagnosis">
|
||
<paragraph id="8BEC36A6FF947732A7862C31FD2EF8AE" blockId="11.[192,1440,1635,1883]" pageId="11" pageNumber="10">
|
||
<emphasis id="B927EAB4FF947732A7862C31FE07F98E" bold="true" box="[192,501,1635,1659]" pageId="11" pageNumber="10">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C534D25FF947732A6BC2C31FD08F98E" authorityName="Sokolov" authorityYear="2011" box="[506,762,1635,1659]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="smokiensis">
|
||
<emphasis id="B927EAB4FF947732A6BC2C31FD08F98E" box="[506,762,1635,1659]" italics="true" pageId="11" pageNumber="10">Anillinus smokiensis</emphasis>
|
||
</taxonomicName>
|
||
belongs to the troglobitic valentinei-group of species. In addition to
|
||
<taxonomicName id="4C534D25FF947732A60A2CD6FDAEF96E" authority="(Jeanell)" baseAuthorityName="Jeanell" box="[332,604,1667,1691]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="valentinei">
|
||
<emphasis id="B927EAB4FF947732A60A2CD6FE15F96E" box="[332,487,1667,1691]" italics="true" pageId="11" pageNumber="10">A. valentinei</emphasis>
|
||
(Jeanell)
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="54E88804FF947732A5D02CD6FD3EF969" box="[662,716,1668,1692]" pageId="11" pageNumber="10">type</typeStatus>
|
||
species of the older genus
|
||
<taxonomicName id="4C534D25FF947732A3492CD1FAE7F96E" authority="Jeannel" authorityName="Jeannel" authorityYear="1963" box="[1039,1301,1667,1691]" class="Insecta" family="Carabidae" genus="Troglanillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B927EAB4FF947732A3492CD1FB58F96E" box="[1039,1194,1667,1691]" italics="true" pageId="11" pageNumber="10">Troglanillus</emphasis>
|
||
Jeannel
|
||
</taxonomicName>
|
||
, which was synonymized with
|
||
<taxonomicName id="4C534D25FF947732A6D92CF1FDE2F94E" authorityName="Casey" authorityYear="1918" box="[415,528,1699,1723]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B927EAB4FF947732A6D92CF1FDE2F94E" box="[415,528,1699,1723]" italics="true" pageId="11" pageNumber="10">Anillinus</emphasis>
|
||
</taxonomicName>
|
||
by Barr (
|
||
<bibRefCitation id="EFC24B57FF947732A5382CF1FD0BF94E" author="Barr, T. C." box="[638,761,1699,1723]" pageId="11" pageNumber="10" pagination="239 - 248" refId="ref8335" refString="Barr, T. C. 1995. Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from southeastern United States, with description of a new genus and two new species. Special Bulletin Japanese Society of Coleopterology 4: 239 - 248." type="journal article" year="1995">Barr 1995</bibRefCitation>
|
||
), the group includes several undescribed species from the caves of
|
||
<collectingRegion id="4997F844FF947732A6632C91FE6AF92E" box="[293,408,1731,1755]" country="United States of America" name="Alabama" pageId="11" pageNumber="10">Alabama</collectingRegion>
|
||
. Besides the habitat, this group is characterized by the combination of comparatively large size, mostly reduced microsculpture on foreparts, and the presence of rows of large spines in the internal sac of median lobe.
|
||
<taxonomicName id="4C534D25FF947732A5512D51FCEAF8EE" authorityName="Sokolov" authorityYear="2011" box="[535,792,1795,1819]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="smokiensis">
|
||
<emphasis id="B927EAB4FF947732A5512D51FCEAF8EE" box="[535,792,1795,1819]" italics="true" pageId="11" pageNumber="10">Anillinus smokiensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C534D25FF947732A36E2D56FB36F8EE" box="[1064,1220,1795,1819]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="valentinei">
|
||
<emphasis id="B927EAB4FF947732A36E2D56FB36F8EE" box="[1064,1220,1795,1819]" italics="true" pageId="11" pageNumber="10">A. valentinei</emphasis>
|
||
</taxonomicName>
|
||
by the form of the median lobe and armature of the internal sac (cf.
|
||
<bibRefCitation id="EFC24B57FF947732A4722D71FC2CF8CE" author="Jeannel, R." box="[820,990,1827,1851]" pageId="11" pageNumber="10" pagination="145 - 152" refId="ref8636" refString="Jeannel, R. 1963. Supplement a la monographie des Anillini (1). Sur quelques especes nouvelles de l'Amerique du Nord. Revue Francaise d'Entomologie 30: 145 - 152." type="journal article" year="1963">Jeannel 1963</bibRefCitation>
|
||
: p.147, fig. 2), and especially by the smaller size (cf.
|
||
<quantity id="4CAB9B43FF947732A6C22D11FDFCF8AE" box="[388,526,1859,1883]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="11" pageNumber="10" unit="mm" value="2.5" valueMax="2.7" valueMin="2.3">2.3-2.7 mm</quantity>
|
||
of
|
||
<taxonomicName id="4C534D25FF947732A5762D16FD3EF8AE" box="[560,716,1859,1883]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="valentinei">
|
||
<emphasis id="B927EAB4FF947732A5762D16FD3EF8AE" box="[560,716,1859,1883]" italics="true" pageId="11" pageNumber="10">A. valentinei</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEC36A6FF937735A7B62A91FAE7FEAE" blockId="12.[192,1441,195,347]" pageId="12" pageNumber="11">
|
||
<taxonomicName id="4C534D25FF937735A7B62A91FE01FF2E" authorityName="Sokolov" authorityYear="2011" box="[240,499,195,219]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="smokiensis">
|
||
<emphasis id="B927EAB4FF937735A7B62A91FE01FF2E" box="[240,499,195,219]" italics="true" pageId="12" pageNumber="11">Anillinus smokiensis</emphasis>
|
||
</taxonomicName>
|
||
apparently is the only cavernicolous anilline species in the Smokies. Many species may be found incidentally in caves, especially near entrances, but are also found in epigean leaf litter not associated with caves. Externally,
|
||
<taxonomicName id="4C534D25FF937735A5F02B56FCA6FEEE" authorityName="Sokolov" authorityYear="2011" box="[694,852,259,283]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="smokiensis">
|
||
<emphasis id="B927EAB4FF937735A5F02B56FCA6FEEE" box="[694,852,259,283]" italics="true" pageId="12" pageNumber="11">A. smokiensis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all species of
|
||
<taxonomicName id="4C534D25FF937735A25D2B51FA52FEEE" box="[1307,1440,259,283]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B927EAB4FF937735A25D2B51FA52FEEE" box="[1307,1440,259,283]" italics="true" pageId="12" pageNumber="11">Serranillus</emphasis>
|
||
</taxonomicName>
|
||
by the presence of long discal elytral setae. From
|
||
<taxonomicName id="4C534D25FF937735A4542B71FC10FECE" box="[786,994,291,315]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B927EAB4FF937735A4542B71FC77FECE" box="[786,901,291,315]" italics="true" pageId="12" pageNumber="11">Anillinus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
inhabiting low and mid elevations of GSMNP,
|
||
<taxonomicName id="4C534D25FF937735A6722B16FE2BFEAE" authorityName="Sokolov" authorityYear="2011" box="[308,473,323,347]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="smokiensis">
|
||
<emphasis id="B927EAB4FF937735A6722B16FE2BFEAE" box="[308,473,323,347]" italics="true" pageId="12" pageNumber="11">A. smokiensis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by the effaced microsculpture of the pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |