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<document id="009C8C18F3B57DA50E13603BE7AC983A" ID-DOI="10.11646/zootaxa.3990.2.2" ID-GBIF-Dataset="1740ec81-c55e-44ce-9a6d-cd62acd5d898" ID-ISSN="1175-5326" ID-Zenodo-Dep="242947" ID-ZooBank="BDF1BC11-4848-4858-99DC-0336A5FBEE2B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461267666231" checkinUser="plazi" docAuthor="Helwerda, Renate A." docDate="2015" docId="03FA878C9514FE4EFF26E6F1027DCD90" docLanguage="en" docName="zt03990p220.pdf" docOrigin="Zootaxa 3990 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bathyacteon valdesi" docType="treatment" docVersion="9" lastPageNumber="201" masterDocId="FFC3FFF49510FE4AFFB1E574036DCA1D" masterDocTitle="Acteonidae, Bullinidae and Ringiculidae (Gastropoda: Heterobranchia) from the Plio-Pleistocene of the Philippines" masterLastPageNumber="220" masterPageNumber="197" pageNumber="201" updateTime="1698610896101" updateUser="plazi">
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<mods:title id="92783D90286518E455D3812BE0109A26">Acteonidae, Bullinidae and Ringiculidae (Gastropoda: Heterobranchia) from the Plio-Pleistocene of the Philippines</mods:title>
</mods:titleInfo>
<mods:name id="CCA4C19D57BBE5462A2E5387B897028F" type="personal">
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<mods:namePart id="3E7FD49953F967385D737791C788346F">Helwerda, Renate A.</mods:namePart>
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<mods:typeOfResource id="6618844283BD55B43E37674ACFF37ACD">text</mods:typeOfResource>
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<mods:title id="9F53670E7F050E79F4174AF278D9A8C4">Zootaxa</mods:title>
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<mods:part id="8C4A8AE0FB246E5B267D8F805A564E70">
<mods:date id="654A1B2817FF781496FC9A976E852E6D">2015</mods:date>
<mods:detail id="5A9125C5D8FCBA1EC23BB7B55B4608BE" type="volume">
<mods:number id="A258D7B95CDA3BC450B719A0A43DF944">3990</mods:number>
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<mods:classification id="7B422D9CFC0E4779B092BB4DFCFE1F67">journal article</mods:classification>
<mods:identifier id="B1D0AFEEBF47E81499A837E9C8691DAF" type="DOI">10.11646/zootaxa.3990.2.2</mods:identifier>
<mods:identifier id="74121930AAC3AD85E0F31EAD2BBFD8FD" type="GBIF-Dataset">1740ec81-c55e-44ce-9a6d-cd62acd5d898</mods:identifier>
<mods:identifier id="1F22D85E927E0F402B58342EB77CE1A0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="879F96AFC103AF60C89B5A09BA01AB09" type="Zenodo-Dep">242947</mods:identifier>
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<treatment id="03FA878C9514FE4EFF26E6F1027DCD90" ID-DOI="http://doi.org/10.5281/zenodo.6116497" ID-GBIF-Taxon="119638240" ID-Zenodo-Dep="6116497" LSID="urn:lsid:plazi:treatment:03FA878C9514FE4EFF26E6F1027DCD90" httpUri="http://treatment.plazi.org/id/03FA878C9514FE4EFF26E6F1027DCD90" lastPageNumber="201" pageId="4" pageNumber="201">
<subSubSection id="C34965119514FE4EFF26E6F10243C9DD" pageId="4" pageNumber="201" type="nomenclature">
<paragraph id="8BEC369A9514FE4EFF26E6F1016AC982" blockId="4.[151,519,901,960]" box="[151,519,901,927]" pageId="4" pageNumber="201">
<heading id="D0A481F69514FE4EFF26E6F1016AC982" bold="true" box="[151,519,901,927]" fontSize="11" level="1" pageId="4" pageNumber="201" reason="1">
<emphasis id="B927EA889514FE4EFF26E6F1016AC982" bold="true" box="[151,519,901,927]" pageId="4" pageNumber="201">
<taxonomicName id="4C534D199514FE4EFF26E6F102E5C982" ID-CoL="7QB4R" box="[151,392,901,927]" class="Gastropoda" family="Acteonidae" genus="Bathyacteon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="201" phylum="Mollusca" rank="species" species="valdesi">
<emphasis id="B927EA889514FE4EFF26E6F102E5C982" bold="true" box="[151,392,901,927]" italics="true" pageId="4" pageNumber="201">Bathyacteon valdesi</emphasis>
</taxonomicName>
nov. spec.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEC369A9514FE4EFF26E6D30243C9DD" blockId="4.[151,519,901,960]" box="[151,302,935,960]" pageId="4" pageNumber="201">
(
<figureCitation id="13682A1F9514FE4EFF2FE6D3024BC9DD" box="[158,294,935,960]" captionStart="FIGURE 1 – 7. 1" captionStartId="3.[151,250,1768,1790]" captionTargetBox="[247,1326,195,1729]" captionTargetId="figure@3.[247,1326,195,1729]" captionTargetPageId="3" captionText="FIGURE 1 – 7. 1. Japonacteon sieboldii (Reeve, 1842). RGM 961.831. Locality Anda 4. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. For these and remaining figures, all material derives from the Piazencian or Gelasian of the Santa Cruz Formation of Cabarruyan Island or nearby Luzon Island, Philippines. Scale bars = 1 mm. 2. Japonacteon sieboldii (Reeve, 1842). RGM 961.830. Locality Roxas. (a) apertural view. 3. Japonacteon sieboldii (Reeve, 1842). RGM 961.739. Locality Anda 4. (a) apertural view, (b) apical view. 4. Bathyacteon valdesi nov. spec .. Paratype RGM 961.746. Locality AndaDeVos. (a) apertural view, (b) apical view. 5. Bathyacteon valdesi nov. spec .. Holotype RGM 961.786. Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 6. Acteon spec 1. RGM 961.768. Locality Anda 4. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 7. Acteon spec 1. RGM 961.741. Locality Anda 4. (a) apertural view, (b) apical view." httpUri="https://zenodo.org/record/242948/files/figure.png" pageId="4" pageNumber="201">Figures 4–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34965119514FE4EFF26E69B0297CE85" pageId="4" pageNumber="201" type="materials_examined">
<paragraph id="8BEC369A9514FE4EFF26E69B0281CE30" blockId="4.[151,1437,1007,1933]" pageId="4" pageNumber="201">
<emphasis id="B927EA889514FE4EFF26E69B022BCE15" bold="true" box="[151,326,1007,1032]" pageId="4" pageNumber="201">
<typeStatus id="54E888389514FE4EFF26E69B03BFCE15" box="[151,210,1007,1032]" pageId="4" pageNumber="201">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54E888389514FE4EFEFEE69B02DACE15" box="[335,439,1007,1032]" pageId="4" pageNumber="201" type="holotype">Holotype</typeStatus>
<collectionCode id="ED42AE5F9514FE4EFE71E684016DCE15" box="[448,512,1008,1032]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="201">RGM</collectionCode>
961.786 (
<typeStatus id="54E888389514FE4EFDC2E68401C0CE15" box="[627,685,1008,1032]" pageId="4" pageNumber="201">Type</typeStatus>
locality).
<typeStatus id="54E888389514FE4EFC92E68400E8CE15" box="[803,901,1008,1032]" pageId="4" pageNumber="201" type="paratype">Paratype</typeStatus>
<collectionCode id="ED42AE5F9514FE4EFC3CE68400A0CE15" box="[909,973,1008,1032]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="201">RGM</collectionCode>
961.746 (
<typeStatus id="54E888389514FE4EFBF0E6840716CE15" box="[1089,1147,1008,1032]" pageId="4" pageNumber="201">Type</typeStatus>
locality) and 7
<typeStatus id="54E888389514FE4EFA80E68406F1CE15" box="[1329,1436,1008,1032]" pageId="4" pageNumber="201" type="paratype">paratypes</typeStatus>
<collectionCode id="ED42AE5F9514FE4EFF26E16003BACE31" box="[151,215,1044,1068]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="201">RGM</collectionCode>
961.848 (
<typeStatus id="54E888389514FE4EFEF6E16002EDCE31" box="[327,384,1044,1068]" pageId="4" pageNumber="201">Type</typeStatus>
locality).
</paragraph>
<paragraph id="8BEC369A9514FE4EFF76E1430067CE68" blockId="4.[151,1437,1007,1933]" pageId="4" pageNumber="201">
<emphasis id="B927EA889514FE4EFF76E14302E5CE4D" bold="true" box="[199,392,1079,1104]" pageId="4" pageNumber="201">Other material.</emphasis>
Anda3 (1)
<collectionCode id="ED42AE5F9514FE4EFDADE14C0131CE4D" box="[540,604,1080,1104]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="201">RGM</collectionCode>
961.781, Anda6 (8)
<collectionCode id="ED42AE5F9514FE4EFCEEE14C00F2CE4D" box="[863,927,1080,1104]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="201">RGM</collectionCode>
961.782, AndaDeVos (65)
<collectionCode id="ED42AE5F9514FE4EFB5CE14C0640CE4D" box="[1261,1325,1080,1104]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="201">RGM</collectionCode>
961.783, AndaClif3 (3)
<collectionCode id="ED42AE5F9514FE4EFE8DE1280211CE69" box="[316,380,1116,1140]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="201">RGM</collectionCode>
961.784, Roxas (6)
<collectionCode id="ED42AE5F9514FE4EFDD1E12801CDCE69" box="[608,672,1116,1140]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="201">RGM</collectionCode>
961.785.
</paragraph>
<paragraph id="8BEC369A9514FE4EFF76E10B0297CE85" blockId="4.[151,1437,1007,1933]" box="[199,506,1151,1176]" pageId="4" pageNumber="201">
<emphasis id="B927EA889514FE4EFF76E10B0208CE85" bold="true" box="[199,357,1151,1176]" pageId="4" pageNumber="201">
<typeStatus id="54E888389514FE4EFF76E10B026FCE85" box="[199,258,1151,1176]" pageId="4" pageNumber="201">Type</typeStatus>
locality.
</emphasis>
AndaDeVos.
</paragraph>
</subSubSection>
<subSubSection id="C34965119514FE4EFF76E1D001BBCEFD" pageId="4" pageNumber="201" type="etymology">
<paragraph id="8BEC369A9514FE4EFF76E1D001BBCEFD" blockId="4.[151,1437,1007,1933]" pageId="4" pageNumber="201">
<emphasis id="B927EA889514FE4EFF76E1D002C5CEA0" bold="true" box="[199,424,1188,1213]" pageId="4" pageNumber="201">Derivatio nominis.</emphasis>
Named after Dr. Ángel Valdés, who made a major contribution to our knowledge on
<taxonomicName id="4C534D199514FE4EFF26E1B3027ACEFD" box="[151,279,1223,1248]" class="Gastropoda" family="Acteonidae" kingdom="Animalia" pageId="4" pageNumber="201" phylum="Mollusca" rank="family">Acteonidae</taxonomicName>
and other “cephalaspid” heterobranchs.
</paragraph>
</subSubSection>
<subSubSection id="C34965119514FE4EFF76E198009BCF50" pageId="4" pageNumber="201" type="diagnosis">
<paragraph id="8BEC369A9514FE4EFF76E198009BCF50" blockId="4.[151,1437,1007,1933]" pageId="4" pageNumber="201">
<emphasis id="B927EA889514FE4EFF76E198022CCF18" bold="true" box="[199,321,1260,1285]" pageId="4" pageNumber="201">Diagnosis.</emphasis>
Shell wide, thick; H
<quantity id="4CAB9B7F9514FE4EFD9FE19801FECF19" box="[558,659,1260,1284]" metricMagnitude="-3" metricUnit="m" metricValue="2.44" pageId="4" pageNumber="201" unit="mm" value="2.44">2.44 mm</quantity>
,
<geoCoordinate id="EE67505D9514FE4EFD2EE199019CCF19" box="[671,753,1260,1284]" direction="west" orientation="longitude" pageId="4" pageNumber="201" precision="555" value="-1.78">
W
<quantity id="4CAB9B7F9514FE4EFD71E198019CCF19" box="[704,753,1260,1284]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="4" pageNumber="201" unit="mm" value="1.78">1.78</quantity>
</geoCoordinate>
mm; protoconch large, low, 1.2 whorl, DP
<quantity id="4CAB9B7F9514FE4EFB64E1980656CF19" box="[1237,1339,1260,1284]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="201" unit="mm" value="0.8">0.80 mm</quantity>
; nucleus large, DN
<quantity id="4CAB9B7F9514FE4EFEBAE064021CCF3A" box="[267,369,1296,1320]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="201" unit="mm" value="0.38">0.38 mm</quantity>
; teleoconch weakly shouldered, smooth except for growth lines and few spiral grooves on base; aperture wide; inner lip and columella covered with callus; umbilicus closed.
</paragraph>
</subSubSection>
<subSubSection id="C34965119514FE4EFF76E0230787CCE1" pageId="4" pageNumber="201" type="description">
<paragraph id="8BEC369A9514FE4EFF76E023038BCC38" blockId="4.[151,1437,1007,1933]" pageId="4" pageNumber="201">
<emphasis id="B927EA889514FE4EFF76E0230234CF6D" bold="true" box="[199,345,1367,1392]" pageId="4" pageNumber="201">Description.</emphasis>
Shell small, relatively wide for a species of
<taxonomicName id="4C534D199514FE4EFCDCE0230082CF6D" box="[877,1007,1367,1392]" class="Gastropoda" family="Acteonidae" kingdom="Animalia" pageId="4" pageNumber="201" phylum="Mollusca" rank="family">Acteonidae</taxonomicName>
, rather thick-shelled, with a rounded shoulder and convex sides. Largest specimen has a total of 2.8 whorls, including a protoconch of 1.2 whorl. Protoconch smooth, shiny, margin indistinct. Teleoconch glossy, smooth except for weak prosocyrt growth lines and 3–5 weak spiral grooves without punctuations on base. Aperture wide and rather short, teardrop-shaped. Parietal callus present, thickest on columella. One weak fold present on columella in large specimens. Umbilicus closed.
</paragraph>
<paragraph id="8BEC369A9514FE4EFF76E35B0787CCE1" blockId="4.[151,1437,1007,1933]" pageId="4" pageNumber="201">
<emphasis id="B927EA889514FE4EFF76E35B0212CC55" bold="true" box="[199,383,1583,1608]" pageId="4" pageNumber="201">Differentiation.</emphasis>
<taxonomicName id="4C534D199514FE4EFE3BE3440008CC55" authority="Thiele, 1925" authorityName="Thiele" authorityYear="1925" box="[394,869,1583,1608]" class="Gastropoda" family="Acteonidae" genus="Bathyacteon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="201" phylum="Mollusca" rank="species" species="aequatorialis">
<emphasis id="B927EA889514FE4EFE3BE34401DBCC55" box="[394,694,1584,1608]" italics="true" pageId="4" pageNumber="201">Bathyacteon aequatorialis</emphasis>
(Thiele, 1925)
</taxonomicName>
is similar to
<taxonomicName id="4C534D199514FE4EFBB9E3450717CC55" box="[1032,1146,1584,1608]" class="Gastropoda" family="Acteonidae" genus="Bathyacteon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="201" phylum="Mollusca" rank="species" species="valdesi">
<emphasis id="B927EA889514FE4EFBB9E3450717CC55" box="[1032,1146,1584,1608]" italics="true" pageId="4" pageNumber="201">B. valdesi</emphasis>
</taxonomicName>
nov. spec. in its general shape and size, but
<taxonomicName id="4C534D199514FE4EFEDFE32101A5CC70" authority="Thiele, 1925" authorityName="Thiele" authorityYear="1925" box="[366,712,1620,1645]" class="Gastropoda" family="Acteonidae" genus="Bathyacteon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="201" phylum="Mollusca" rank="species" species="aequatorialis">
<emphasis id="B927EA889514FE4EFEDFE321014FCC71" box="[366,546,1620,1644]" italics="true" pageId="4" pageNumber="201">B. aequatorialis</emphasis>
(Thiele, 1925)
</taxonomicName>
has a smaller protoconch, which is usually corroded, but appears to have 0.8 whorl and a diameter of 500 µm. Furthermore, there are weak, punctuated spiral grooves, with large gaps in between, present on most of the teleoconch whorl in
<taxonomicName id="4C534D199514FE4EFCFFE3E80648CCA8" authority="Thiele, 1925" authorityName="Thiele" authorityYear="1925" box="[846,1317,1692,1717]" class="Gastropoda" family="Acteonidae" genus="Bathyacteon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="201" phylum="Mollusca" rank="species" species="aequatorialis">
<emphasis id="B927EA889514FE4EFCFFE3E8071ACCA9" box="[846,1143,1692,1716]" italics="true" pageId="4" pageNumber="201">Bathyacteon aequatorialis</emphasis>
(Thiele, 1925)
</taxonomicName>
. A lack of punctuated spiral grooves is rare within
<taxonomicName id="4C534D199514FE4EFDC1E3CB019ECCC5" box="[624,755,1727,1752]" class="Gastropoda" family="Acteonidae" kingdom="Animalia" pageId="4" pageNumber="201" phylum="Mollusca" rank="family">Acteonidae</taxonomicName>
. It also occurs in
<taxonomicName id="4C534D199514FE4EFC7BE3B50658CCC5" authority="Lamarck, 1816" authorityName="Lamarck" authorityYear="1816" box="[970,1333,1727,1752]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="201" phylum="Mollusca" rank="species" species="nitidula">
<emphasis id="B927EA889514FE4EFC7BE3B5070ACCC5" box="[970,1127,1728,1752]" italics="true" pageId="4" pageNumber="201">Pupa nitidula</emphasis>
(Lamarck, 1816)
</taxonomicName>
, but this species has a coeloconoid spire, a strong, double columellar fold and a lamella on the parietal area.
</paragraph>
</subSubSection>
<subSubSection id="C34965119514FE4EFF76E273027DCD90" pageId="4" pageNumber="201" type="discussion">
<paragraph id="8BEC369A9514FE4EFF76E273027DCD90" blockId="4.[151,1437,1007,1933]" pageId="4" pageNumber="201">
<emphasis id="B927EA889514FE4EFF76E2730256CD3D" bold="true" box="[199,315,1799,1824]" pageId="4" pageNumber="201">Remarks.</emphasis>
This species is tentatively placed in genus
<taxonomicName id="4C534D199514FE4EFC82E27C00D2CD3D" box="[819,959,1800,1824]" class="Gastropoda" family="Acteonidae" genus="Bathyacteon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="201" phylum="Mollusca" rank="genus">
<emphasis id="B927EA889514FE4EFC82E27C00D2CD3D" box="[819,959,1800,1824]" italics="true" pageId="4" pageNumber="201">Bathyacteon</emphasis>
</taxonomicName>
, because it resembles the
<typeStatus id="54E888389514FE4EFB5EE27D0672CD3C" box="[1263,1311,1801,1825]" pageId="4" pageNumber="201">type</typeStatus>
species
<taxonomicName id="4C534D199514FE4EFA35E27D02B4CD58" authority="Thiele, 1925" authorityName="Thiele" authorityYear="1925" class="Gastropoda" family="Acteonidae" genus="Bathyacteon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="201" phylum="Mollusca" rank="species" species="aequatorialis">
<emphasis id="B927EA889514FE4EFA35E27D0241CD59" italics="true" pageId="4" pageNumber="201">B. aequatorialis</emphasis>
(Thiele, 1925)
</taxonomicName>
much more strongly than any other
<taxonomicName id="4C534D199514FE4EFC32E2580768CD58" box="[899,1029,1836,1861]" class="Gastropoda" family="Acteonidae" kingdom="Animalia" pageId="4" pageNumber="201" phylum="Mollusca" rank="family">Acteonidae</taxonomicName>
, with respect to its shell shape and sculpture. This placement is tentative, because
<taxonomicName id="4C534D199514FE4EFD1CE2240054CD75" box="[685,825,1872,1896]" class="Gastropoda" family="Acteonidae" genus="Bathyacteon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="4" pageNumber="201" phylum="Mollusca" rank="genus">
<emphasis id="B927EA889514FE4EFD1CE2240054CD75" box="[685,825,1872,1896]" italics="true" pageId="4" pageNumber="201">Bathyacteon</emphasis>
</taxonomicName>
is delimited from other acteonid genera using radular characters.
</paragraph>
</subSubSection>
</treatment>
</document>
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