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<document id="B6AB50A18F90B85485A7F86CED8CEECF" ID-DOI="10.5281/zenodo.5174200" ID-ISSN="1942-1354" ID-Zenodo-Dep="5174200" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628335400619" checkinUser="felipe" docAuthor="Blahnik, Roger J., Arefina-Armitage, I. &amp; J, Brian" docDate="2012" docId="03FA8783FF98FF945CF0F9592ED7F145" docLanguage="en" docName="InsectaMundi.0229.1-25.pdf" docOrigin="Insecta Mundi 2012 (229)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Chimarra aculeata Blahnik &amp; Arefina-Armitage &amp; J 2012, sp. n." docType="treatment" docVersion="3" lastPageNumber="6" masterDocId="FFC3FFFBFF9DFF935C30FFB12F7FF47E" masterDocTitle="The Genus Chimarra Stephens (Trichoptera: Philopotamidae) in Vietnam" masterLastPageNumber="25" masterPageNumber="1" pageNumber="4" updateTime="1699045144500" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="D55FE31FB49C5BB43010BF1A513B6A6B">
<mods:title id="664028F6E2932638BF808E198B539FF7">The Genus Chimarra Stephens (Trichoptera: Philopotamidae) in Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="0136473D6CF724FC8E0C42A15CDE8C50" type="personal">
<mods:role id="0C7D9E01172A5B250FF04A781F36D0EB">
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<mods:namePart id="1EC76E2C86E0246EBBFF68C6D43A6433">Blahnik, Roger J.</mods:namePart>
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<mods:role id="67A7B9C24912F26A5A1A70FB9C3A2248">
<mods:roleTerm id="692C5D089FE33D3BF47044DE4A59E00A">Author</mods:roleTerm>
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<mods:namePart id="DB77CCE052DD96B76D24AD9320397E96">Arefina-Armitage, I.</mods:namePart>
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<mods:namePart id="D43EA7C9C4A8E049A382B6EDED2932CB">J, Brian</mods:namePart>
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<mods:title id="798221E7B31DAFFB63319ED179BEAC15">Insecta Mundi</mods:title>
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<mods:part id="C042DC7CFCC7AA0AEDD0C3A67D942068">
<mods:date id="94E383DC67C913B88C5FF28BF92FC465">2012</mods:date>
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<mods:number id="F32C31C5CCED7A5209730439DC3DC678">2012-04-06</mods:number>
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<mods:classification id="2E168148021F20F3699FEB8BFD4680B2">journal article</mods:classification>
<mods:identifier id="8B05E913C7D3B9B87E565FDF12A0B264" type="DOI">10.5281/zenodo.5174200</mods:identifier>
<mods:identifier id="2D8BD7668C004B690F95C0F18255E29F" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="E07143E1F9D9549F0C0178FC6F89DDA8" type="Zenodo-Dep">5174200</mods:identifier>
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<treatment id="03FA8783FF98FF945CF0F9592ED7F145" LSID="urn:lsid:plazi:treatment:03FA8783FF98FF945CF0F9592ED7F145" httpUri="http://treatment.plazi.org/id/03FA8783FF98FF945CF0F9592ED7F145" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4">
<subSubSection id="C349651EFF98FF965CF0F9592D56F285" box="[192,553,1768,1787]" pageId="5" pageNumber="4" type="nomenclature">
<paragraph id="8BEC3695FF98FF965CF0F9592D56F285" blockId="5.[192,553,1768,1819]" box="[192,553,1768,1787]" pageId="5" pageNumber="4">
<heading id="D0A481F9FF98FF965CF0F9592D56F285" bold="true" box="[192,553,1768,1787]" fontSize="10" level="2" pageId="5" pageNumber="4" reason="3">
<emphasis id="B927EA87FF98FF965CF0F9592D56F285" bold="true" box="[192,553,1768,1787]" pageId="5" pageNumber="4">
<taxonomicName id="4C534D16FF98FF965CF0F9592EAAF285" authority="Blahnik &amp; Arefina-Armitage &amp; J, 2012" authorityName="Blahnik &amp; Arefina-Armitage &amp; J" authorityYear="2012" box="[192,469,1768,1787]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="aculeata" status="sp. nov.">
<emphasis id="B927EA87FF98FF965CF0F9592EAAF285" bold="true" box="[192,469,1768,1787]" italics="true" pageId="5" pageNumber="4">Chimarra aculeata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21457FCFF98FF965DEEF9592D56F285" box="[478,553,1768,1787]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C349651EFF98FF965CF0F8B92E43F365" box="[192,316,1800,1819]" pageId="5" pageNumber="4" type="description">
<paragraph id="8BEC3695FF98FF965CF0F8B92E43F365" blockId="5.[192,553,1768,1819]" box="[192,316,1800,1819]" pageId="5" pageNumber="4">
(
<figureCitation id="13682A10FF98FF965CF9F8B92E4EF365" box="[201,305,1800,1819]" captionStart="Figure 1" captionStartId="6.[192,278,1870,1887]" captionTargetBox="[219,1435,619,1833]" captionTargetId="figure-185@6.[192,1440,586,1851]" captionTargetPageId="6" captionText="Figure 1. Chimarra aculeata sp. n., male genitalia. A) Lateral; B) Dorsal; C) Ventral; D) Distal portion of left inferior appendage, dorsal; E) Phallic apparatus, lateral; F) Phallic apparatus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5174202" httpUri="https://zenodo.org/record/5174202/files/figure.png" pageId="5" pageNumber="4">Figure 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C349651EFF98FF955CF0F8F92E19F525" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4" type="diagnosis">
<paragraph id="8BEC3695FF98FF955CF0F8F92E19F525" blockId="5.[192,1440,1864,1915]" lastBlockId="6.[192,1440,200,347]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
<emphasis id="B927EA87FF98FF965CF0F8F92E2AF325" bold="true" box="[192,341,1864,1883]" pageId="5" pageNumber="4">Diagnosis.</emphasis>
This species is possibly most similar to
<taxonomicName id="4C534D16FF98FF965F7EF8F92B32F325" authority="Mey. Both" authorityName="Mey. Both" box="[846,1101,1864,1883]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="uncula">
<emphasis id="B927EA87FF98FF965F7EF8F92CB7F325" box="[846,968,1864,1883]" italics="true" pageId="5" pageNumber="4">C. uncula</emphasis>
Mey. Both
</taxonomicName>
species have paired spinelike projections from dorsolateral margins of segment IX and also a modified tergum VIII, with a mesal invagination and accompanying spines. The differences, however, are considerable. The armature of tergum VIII is especially distinctive in
<taxonomicName id="4C534D16FF9BFF955EF5FF592C2EF485" authorityName="Blahnik &amp; Arefina-Armitage &amp; J" authorityYear="2012" box="[709,849,232,251]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B927EA87FF9BFF955EF5FF592DA7F485" box="[709,728,232,251]" italics="true" pageId="6" pageNumber="5">C</emphasis>
.
<emphasis id="B927EA87FF9BFF955ED6FF592C2EF485" box="[742,849,232,251]" italics="true" pageId="6" pageNumber="5">aculeata</emphasis>
</taxonomicName>
in that the dorsal margin has a distinctive pair of brushes composed of many spines bordering the mesal invagination; also, the apices of the resulting lobes each have a small tassel of spines. The shape of the inferior appendages is also quite different in the 2 species.
</paragraph>
</subSubSection>
<subSubSection id="C349651EFF9BFF945CF0FE392A71F005" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5" type="description">
<paragraph id="8BEC3695FF9BFF945CF0FE392C22F565" blockId="6.[192,1441,392,571]" lastBlockId="7.[192,1441,200,1147]" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5">
<emphasis id="B927EA87FF9BFF955CF0FE392E1CF5E5" bold="true" box="[192,355,392,411]" pageId="6" pageNumber="5">Description.</emphasis>
Forewing length (male)
<quantity id="4CAB9B70FF9BFF955E4BFE392DAAF5E5" box="[635,725,392,411]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="6" pageNumber="5" unit="mm" value="6.1">6.1 mm</quantity>
. Color (in alcohol) yellowish-brown. Head elongate (postocular parietal sclerite extended behind eye). Mesoscutellar setal wart pear-shaped, with usual anterior suture. Maxillary palps with 1
<superScript id="7C269BDDFF9BFF955DE4FE742E9EF5AE" attach="left" box="[468,481,453,464]" fontSize="6" pageId="6" pageNumber="5">st</superScript>
segment very short, 2
<superScript id="7C269BDDFF9BFF955EDDFE742D80F5AE" attach="left" box="[749,767,453,464]" fontSize="6" pageId="6" pageNumber="5">nd</superScript>
segment relatively elongate (nearly as long as segment 3), 3
<superScript id="7C269BDDFF9BFF955CC3FE542E7CF58E" attach="left" box="[243,259,485,496]" fontSize="6" pageId="6" pageNumber="5">rd</superScript>
segment elongate, subequal to segment 5, 4
<superScript id="7C269BDDFF9BFF955F21FE542C5FF58E" attach="left" box="[785,800,485,496]" fontSize="6" pageId="6" pageNumber="5">th</superScript>
segment short. Tarsi of males highly modified, tarsal claws enlarged and sinuous, asymmetric in length and curvature. Forewing with Rs deflected (curved toward anal margin) and weakly, sinuously inflected before base of discal cell; veins at base of discal cell not or hardly swollen, length of discal cell about 3 times width;
<emphasis id="B927EA87FF9AFF945FBDFF792CC7F4A5" box="[909,952,200,219]" italics="true" pageId="7" pageNumber="6">r-m</emphasis>
and
<emphasis id="B927EA87FF9AFF945FDEFF792B79F4A5" box="[1006,1030,200,219]" italics="true" pageId="7" pageNumber="6">m</emphasis>
cross-veins hyaline (unpigmented),
<emphasis id="B927EA87FF9AFF945CF0FF592FB3F485" box="[192,204,232,251]" italics="true" pageId="7" pageNumber="6">s</emphasis>
not hyaline,
<emphasis id="B927EA87FF9AFF945D55FF592E0EF485" box="[357,369,232,251]" italics="true" pageId="7" pageNumber="6">s</emphasis>
and
<emphasis id="B927EA87FF9AFF945D99FF592EAAF485" box="[425,469,232,251]" italics="true" pageId="7" pageNumber="6">r-m</emphasis>
linearly aligned,
<emphasis id="B927EA87FF9AFF945E95FF592DC2F485" box="[677,701,232,251]" italics="true" pageId="7" pageNumber="6">m</emphasis>
distinctly proximal; anal veins without apical fork (2A and 3A both apparently looped to 1A). Hind wing with Sc and R
</paragraph>
<caption id="DF2C661DFF9BFF955CF0F8FF2BCFF305" ID-DOI="http://doi.org/10.5281/zenodo.5174202" ID-Zenodo-Dep="5174202" httpUri="https://zenodo.org/record/5174202/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,278,1870,1887]" targetBox="[219,1435,619,1833]" targetPageId="6">
<paragraph id="8BEC3695FF9BFF955CF0F8FF2BCFF305" blockId="6.[192,1440,1870,1915]" pageId="6" pageNumber="5">
<emphasis id="B927EA87FF9BFF955CF0F8FF2E4EF321" bold="true" box="[192,305,1870,1887]" pageId="6" pageNumber="5">Figure 1.</emphasis>
<taxonomicName id="4C534D16FF9BFF955D08F8FF2D6CF321" authority="Blahnik &amp; Arefina-Armitage &amp; J, 2012" authorityName="Blahnik &amp; Arefina-Armitage &amp; J" authorityYear="2012" box="[312,531,1870,1887]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="aculeata" status="sp. nov.">
<emphasis id="B927EA87FF9BFF955D08F8FF2D6CF321" box="[312,531,1870,1887]" italics="true" pageId="6" pageNumber="5">Chimarra aculeata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21457FCFF9BFF955E29F8FF2D27F321" box="[537,600,1870,1887]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
, male genitalia. A) Lateral; B) Dorsal; C) Ventral; D) Distal portion of left inferior appendage, dorsal; E) Phallic apparatus, lateral; F) Phallic apparatus, dorsal.
</paragraph>
</caption>
<paragraph id="8BEC3695FF9AFF945F6DFEA62C19F55C" blockId="7.[192,1441,200,1147]" box="[861,870,279,290]" pageId="7" pageNumber="6">1</paragraph>
<paragraph id="8BEC3695FF9AFF945F5CFEB92BA1F565" blockId="7.[192,1441,200,1147]" box="[876,1246,264,283]" pageId="7" pageNumber="6">narrowly separated, not fused.</paragraph>
<paragraph id="8BEC3695FF9AFF945CC0FE992A71F005" blockId="7.[192,1441,200,1147]" pageId="7" pageNumber="6">
<emphasis id="B927EA87FF9AFF945CC0FE992EDDF545" box="[240,418,296,315]" italics="true" pageId="7" pageNumber="6">Male genitalia</emphasis>
: Segment VIII with sternum very short, tergum highly modified; tergum, as viewed laterally, with ventral margin about 2 times length of sternum, dorsum of tergum about 2 times length at base, narrowing and acute posteriorly, apex with projecting tract of spinose setae, setae somewhat mesally directed, as viewed dorsally; tergum, as viewed dorsally, with deep, more or less Vshaped mesal emargination, emargination extending almost to anterior margin, lateral margins of emargination with dense, leaf-shaped tract of spinose setae, tract narrow at base, tapering, acuminate apically. Segment IX unsclerotized dorsally, with distinct, very short dorsolateral apodemes on anterior margin; as viewed laterally, with anterior margin extending nearly linearly from apodemes, anteroventral margin not produced (truncate as viewed dorsally or ventrally); dorsolateral margins of tergum, on either side, with posteriorly projecting, spine-like process, extending about as far as lobe of tergum VIII, processes slightly mesally curved as viewed dorsally; width of segment below processes relatively short (anterior and posterior margins narrowly separated), posterior margin lengthened below preanal appendages, angulate at about level of inferior appendages; ventral process very short, subtriangular, broadest basally, projecting ventrally. Tergum X with lateral lobes moderately elongate, simple in structure; in lateral view, linear with rounded apex, in dorsal view with slight bulge on lateral margin at about middle, apices rounded; lobes with numerous sensilla, almost uniformly distributed. Mesal lobes of tergum X short, digitate, weakly sclerotized, posteriorly directed, forming about 45 degree angle with lateral lobe, each lobe (apparently) with 3 minute sensilla, 2 apically and 1 at midlength on lateral margin. Preanal appendages very small and rounded, somewhat flattened laterally, fused basally to posterior margin of segment IX. Inferior appendages, as viewed laterally, moderately elongate, linear, uniform in width, subtruncate apically; as viewed ventrally, with lateral margins rounded, mesal margins subparallel in basal half, narrowing apically, apices rather broadly rounded; appendages, as viewed caudally, with pair of minute sclerotized spine-like projections or cusps on mesal margin. Phallotheca relatively short, tubular, with usual basal expansion. Endotheca apparently short, with paired tracts of short spines. Phallotremal sclerite complex prominent, extending over half length of phallotheca, composed of ring structure with very short, paired, spinose ventral “rods”; ring open dorsally, on each side with projecting, elongate, curved, spine-like dorsolateral sclerites.
</paragraph>
</subSubSection>
<subSubSection id="C349651EFF9AFF945CF0FB192BCFF0A5" pageId="7" pageNumber="6" type="materials_examined">
<paragraph id="8BEC3695FF9AFF945CF0FB192BCFF0A5" blockId="7.[192,1440,1189,1243]" pageId="7" pageNumber="6">
<emphasis id="B927EA87FF9AFF945CF0FB192E9EF0C5" bold="true" box="[192,481,1192,1211]" pageId="7" pageNumber="6">Material examined.</emphasis>
<materialsCitation id="3B3B3CC8FF9AFF945DDEFB192BCFF0A5" collectingDate="2000-09-11" collectingDateMax="2000-09-16" collectingDateMin="2000-09-11" collectionCode="AMNH" collectorName="C. Johnson &amp; K. Long" country="Vietnam" elevation="1170" latitude="22.767834" location="Ha Giang Province" longLatPrecision="12" longitude="104.825165" pageId="7" pageNumber="6" specimenCount="1" stateProvince="Ha Giang" typeStatus="holotype">
<emphasis id="B927EA87FF9AFF945DDEFB192D0CF0C5" bold="true" box="[494,627,1192,1211]" pageId="7" pageNumber="6">
<typeStatus id="54E88837FF9AFF945DDEFB192D0CF0C5" box="[494,627,1192,1211]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
</emphasis>
male:
<emphasis id="B927EA87FF9AFF945EE5FB192C18F0C5" bold="true" box="[725,871,1192,1211]" pageId="7" pageNumber="6">
<collectingCountry id="F3447605FF9AFF945EE5FB192C18F0C5" box="[725,871,1192,1211]" name="Vietnam" pageId="7" pageNumber="6">VIETNAM</collectingCountry>
</emphasis>
:
<collectingRegion id="4997F877FF9AFF945F4DFB192BFDF0C5" box="[893,1154,1192,1211]" country="Vietnam" name="Ha Giang" pageId="7" pageNumber="6">Ha Giang Province</collectingRegion>
,
<quantity id="4CAB9B70FF9AFF9458A4FB192B86F0C5" box="[1172,1273,1192,1211]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="7" pageNumber="6" unit="m" value="1170.0">
<elevation id="007ED1A6FF9AFF9458A4FB192B86F0C5" box="[1172,1273,1192,1211]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="7" pageNumber="6" unit="m" value="1170.0">1170 m</elevation>
</quantity>
,
<geoCoordinate id="EE675052FF9AFF94593CFB192AE4F0C5" box="[1292,1435,1189,1211]" degrees="22" direction="north" minutes="46.07" orientation="latitude" pageId="7" pageNumber="6" precision="9" value="22.767834">
22
<superScript id="7C269BDDFF9AFF94591BFB142A4CF0CE" attach="none" box="[1323,1331,1189,1200]" fontSize="6" pageId="7" pageNumber="6">o</superScript>
46.07N
</geoCoordinate>
,
<geoCoordinate id="EE675052FF9AFF945CF0FB792E28F0A5" box="[192,343,1221,1243]" degrees="104" direction="east" minutes="49.51" orientation="longitude" pageId="7" pageNumber="6" precision="9" value="104.825165">
104
<superScript id="7C269BDDFF9AFF945CDEFB742F89F0AE" attach="right" box="[238,246,1221,1232]" fontSize="6" pageId="7" pageNumber="6">o</superScript>
49.51E
</geoCoordinate>
,
<date id="FFED1055FF9AFF945D52FB792D03F0A5" box="[354,636,1224,1243]" pageId="7" pageNumber="6" value="2000-09-11" valueMax="2000-09-16" valueMin="2000-09-11">
<collectingDate id="EFA9E9BDFF9AFF945D52FB792D03F0A5" box="[354,636,1224,1243]" pageId="7" pageNumber="6" value="2000-09-11" valueMax="2000-09-16" valueMin="2000-09-11">11-16 September 2000</collectingDate>
</date>
,
<collectingMethod id="52124E82FF9AFF945EB8FB792C55F0A5" box="[648,810,1224,1243]" pageId="7" pageNumber="6">Malaise trap</collectingMethod>
,
<collectorName id="26A65343FF9AFF945F06FB792CBAF0A5" box="[822,965,1224,1243]" pageId="7" pageNumber="6">C. Johnson</collectorName>
,
<collectorName id="26A65343FF9AFF945FE0FB792B4DF0A5" box="[976,1074,1224,1243]" pageId="7" pageNumber="6">K. Long</collectorName>
(
<emphasis id="B927EA87FF9AFF945872FB792BE0F0A5" bold="true" box="[1090,1183,1224,1243]" pageId="7" pageNumber="6">
<collectionCode id="ED42AE50FF9AFF945872FB792BE0F0A5" box="[1090,1183,1224,1243]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="7" pageNumber="6">AMNH</collectionCode>
</emphasis>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C349651EFF9AFF945CF0FAB92ED7F145" pageId="7" pageNumber="6" type="etymology">
<paragraph id="8BEC3695FF9AFF945CF0FAB92ED7F145" blockId="7.[192,1440,1288,1339]" pageId="7" pageNumber="6">
<emphasis id="B927EA87FF9AFF945CF0FAB92E20F165" bold="true" box="[192,351,1288,1307]" pageId="7" pageNumber="6">Etymology.</emphasis>
This species is named for the spinose setae on tergum VIII (
<taxonomicName id="4C534D16FF9AFF945879FAB92BCBF165" authorityName="Blahnik &amp; Arefina-Armitage &amp; J" authorityYear="2012" box="[1097,1204,1288,1307]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B927EA87FF9AFF945879FAB92BCBF165" box="[1097,1204,1288,1307]" italics="true" pageId="7" pageNumber="6">aculeata</emphasis>
</taxonomicName>
is Latin for having or bearing spines).
</paragraph>
</subSubSection>
</treatment>
</document>