treatments-xml/data/03/EB/87/03EB87FE1F60FD6CDEF7EAF7F8D5FE55.xml

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<mods:title id="DDD10875887A61891738467DD3A00D9B">A new Peruvian species of Megarthrus Curtis (Coleoptera: Staphylinidae Proteininae)</mods:title>
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<emphasis id="B936EAFA1F60FD68DEF7EAF7F8E9FCB2" bold="true" box="[151,421,189,216]" italics="true" pageId="2" pageNumber="583">Megarthrus wayqecha</emphasis>
Pérez, Rodríguez &amp; Asenjo
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, 
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(
<figureCitation id="13792A6D1F60FD68DEFFEAABF854FC91" box="[159,280,225,251]" captionStart-0="FIGURES 1–7" captionStart-1="FIGURES 8–16" captionStart-2="FIGURES 17–19" captionStart-3="FIGURES 20–21" captionStartId-0="2.[151,264,1549,1574]" captionStartId-1="3.[151,264,1763,1788]" captionStartId-2="4.[151,264,947,972]" captionStartId-3="4.[151,264,1676,1701]" captionTargetBox-0="[156,1432,348,1519]" captionTargetBox-1="[195,1399,436,1722]" captionTargetBox-2="[156,1430,393,916]" captionTargetBox-3="[151,1436,1066,1651]" captionTargetId-0="figure@2.[151,1436,343,1522]" captionTargetId-1="figure@3.[177,1411,417,1738]" captionTargetId-2="figure@4.[152,1436,381,922]" captionTargetId-3="figure@4.[151,1436,1066,1651]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionText-0="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." captionText-1="FIGURES 8–16. Megarthrus wayqecha sp. nov., holotype (8–14). 8, tergite VIII, dorsal view; 9, tergite VII, tergite VIII, sternite VIII, lateral view, 10, sternite VIII; 11, medial area of abdominal sternites II–III in lateral view; 12, hemitergites IX and sternite IX; 13, aedeagus, ventral view; 14, aedeagus, lateral view. Megarthrus wayqecha sp. nov., female paratype, from Wayqecha, Trocha Canopy 15, genital segments, dorsal view; 16, genital segments, lateral view." captionText-2="FIGURES 17–19. Megarthrus wayqecha sp. nov., male paratype. 17, aedeagus, ventral view (the side of the basal foramen); 18, aedeagus, lateral view; 19, aedeagus, dorsal view." captionText-3="FIGURES 20–21. Habitat of Megarthrus wayqecha sp. nov. 20, panoramic view of the cloud forests of the Kosñipata valley, where the biological station Wayqecha is located. 21, sifting leaf litter on Trocha Canopy from the biological station Wayqecha." figureDoi-0="http://doi.org/10.5281/zenodo.3661898" figureDoi-1="http://doi.org/10.5281/zenodo.3661900" figureDoi-2="http://doi.org/10.5281/zenodo.3661902" figureDoi-3="http://doi.org/10.5281/zenodo.3661904" httpUri-0="https://zenodo.org/record/3661898/files/figure.png" httpUri-1="https://zenodo.org/record/3661900/files/figure.png" httpUri-2="https://zenodo.org/record/3661902/files/figure.png" httpUri-3="https://zenodo.org/record/3661904/files/figure.png" pageId="2" pageNumber="583">Figs. 1–21</figureCitation>
)
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<emphasis id="B936EAFA1F60FD68DEF7EC47F878FA4C" bold="true" box="[151,308,1549,1574]" pageId="2" pageNumber="583">FIGURES 1–7</emphasis>
. 
<taxonomicName id="4C424D6B1F60FD68DF21EC44FB68FA4C" authority="Pérez &amp; Rodríguez &amp; Asenjo, 2020" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" box="[321,548,1550,1574]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="wayqecha" status="sp. nov.">
<emphasis id="B936EAFA1F60FD68DF21EC44FB68FA4C" box="[321,548,1550,1574]" italics="true" pageId="2" pageNumber="583">Megarthrus wayqecha</emphasis>
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<taxonomicNameLabel id="A20557811F60FD68DC4AEC44FB37FA4C" box="[554,635,1550,1574]" pageId="2" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
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, male holotype from Wayqecha, Trocha Canopy 
<emphasis id="B936EAFA1F60FD68DA0DEC47FDF1FA4C" bold="true" box="[1133,1213,1549,1574]" pageId="2" pageNumber="583">(2, 4–6)</emphasis>
. 
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antenna; 
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pronotum; 
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mesotrochanter and mesofemur; 
<emphasis id="B936EAFA1F60FD68DC58EC7BFB00FA20" bold="true" box="[568,588,1585,1610]" pageId="2" pageNumber="583">6,</emphasis>
mesotibia. 
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<emphasis id="B936EAFA1F60FD68DCA3EC78FAEBFA20" box="[707,935,1586,1610]" italics="true" pageId="2" pageNumber="583">Megarthrus wayqecha</emphasis>
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, male paratype from Wayqecha, Trocha Canopy 
<emphasis id="B936EAFA1F60FD68DE8CEC1FF810FA04" bold="true" box="[236,348,1621,1646]" pageId="2" pageNumber="583">(1, 3, 7). 1,</emphasis>
habitus, dorsal view; 
<emphasis id="B936EAFA1F60FD68DC5BEC1FFB04FA04" bold="true" box="[571,584,1621,1646]" pageId="2" pageNumber="583">3</emphasis>
, head, dorsal view; 
<emphasis id="B936EAFA1F60FD68DD74EC1CFA6AFA04" bold="true" box="[788,806,1622,1646]" pageId="2" pageNumber="583">7,</emphasis>
metaleg with peg-like setae in the apex of metatibia.
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:
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, cloud forest, / sifting of leaf litter, M. / 
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&amp; 
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.” “ 
<typeStatus id="54F9884A1F60FD68DC30ED42FBABFB48" box="[592,743,1800,1826]" pageId="2" pageNumber="583" type="holotype">HOLOTYPE</typeStatus>
[red label] / 
<emphasis id="B936EAFA1F60FD68DD18ED42FDCDFB48" box="[888,1153,1800,1826]" italics="true" pageId="2" pageNumber="583">
<taxonomicName id="4C424D6B1F60FD68DD18ED42FD3FFB48" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" box="[888,1139,1800,1826]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="wayqecha">Megarthrus wayqecha</taxonomicName>
/
</emphasis>
sp. nov. / Desig. Pérez 
<emphasis id="B936EAFA1F60FD68DBE8ED43F9F7FB2C" italics="true" pageId="2" pageNumber="583">et al.,</emphasis>
2019” (
<collectionCode id="ED53AE2D1F60FD68DF75ED66F83DFB2C" box="[277,369,1836,1862]" pageId="2" pageNumber="583">MUSM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2A3CB51F60FD68DFE3ED66F849FBE4" ID-GBIF-Occurrence="2572158487" collectingDate="2017-11-02" collectionCode="MUSM, MEKRB, CEMT, FMNH, MHNG, VMDC, MPEG, IADIZA" collectorName="Rodriguez &amp; L. Perez" country="Peru" county="Department" elevation="2931" latitude="-13.192445" location="Wayqecha" longLatPrecision="1" longitude="-71.58778" municipality="Kosnipata" pageId="2" pageNumber="583" specimenCount="130" specimenCount-female="66" specimenCount-male="64" stateProvince="Cusco" typeStatus="paratype">
<emphasis id="B936EAFA1F60FD68DFE3ED66FB75FB2C" bold="true" box="[387,569,1835,1862]" pageId="2" pageNumber="583">
<typeStatus id="54F9884A1F60FD68DFE3ED66F8B7FB2C" box="[387,507,1836,1862]" pageId="2" pageNumber="583" type="paratype">Paratypes</typeStatus>
(94):
</emphasis>
<specimenCount id="9D44FD611F60FD68DC20ED66FBC4FB2C" box="[576,648,1836,1862]" pageId="2" pageNumber="583" type="male">32♂♂</specimenCount>
, 
<specimenCount id="9D44FD611F60FD68DCF3ED66FB96FB2C" box="[659,730,1836,1862]" pageId="2" pageNumber="583" type="female">33♀♀</specimenCount>
labeled with the same data as holotype (
<specimenCount id="9D44FD611F60FD68DAFEED66FDABFB2C" box="[1182,1255,1836,1862]" pageId="2" pageNumber="583" type="male">21♂♂</specimenCount>
, 
<specimenCount id="9D44FD611F60FD68DA92ED66FC74FB2C" box="[1266,1336,1836,1862]" pageId="2" pageNumber="583" type="female">28♀♀</specimenCount>
<collectionCode id="ED53AE2D1F60FD68DB5FED66FCD4FB2C" box="[1343,1432,1836,1862]" pageId="2" pageNumber="583">MUSM</collectionCode>
; 
<specimenCount id="9D44FD611F60FD68DEF7ED1AF99EFB00" box="[151,210,1872,1898]" pageId="2" pageNumber="583" type="male">2♂♂</specimenCount>
, 
<specimenCount id="9D44FD611F60FD68DE80ED1AF84FFB00" box="[224,259,1872,1898]" pageId="2" pageNumber="583" type="female">1♀</specimenCount>
<collectionCode id="ED53AE2D1F60FD68DF6DED1AF838FB00" box="[269,372,1872,1898]" pageId="2" pageNumber="583">MEKRB</collectionCode>
; 
<specimenCount id="9D44FD611F60FD68DFE3ED1AF8EBFB00" box="[387,423,1872,1898]" pageId="2" pageNumber="583" type="male">1♂</specimenCount>
, 
<specimenCount id="9D44FD611F60FD68DFD7ED1AF896FB00" box="[439,474,1872,1898]" pageId="2" pageNumber="583" type="female">1♀</specimenCount>
<collectionCode id="ED53AE2D1F60FD68DF84ED1AFB78FB00" box="[484,564,1872,1898]" pageId="2" pageNumber="583">CEMT</collectionCode>
; 
<specimenCount id="9D44FD611F60FD68DC22ED1AFB31FB00" box="[578,637,1872,1898]" pageId="2" pageNumber="583" type="male">2♂♂</specimenCount>
, 
<specimenCount id="9D44FD611F60FD68DCEBED1AFBE2FB00" box="[651,686,1872,1898]" pageId="2" pageNumber="583" type="female">1♀</specimenCount>
<collectionCode id="ED53AE2D1F60FD68DCD8ED1AFA41FB00" box="[696,781,1872,1898]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="583" type="Museum">FMNH</collectionCode>
; 
<specimenCount id="9D44FD611F60FD68DD7BED1AFA1AFB00" box="[795,854,1872,1898]" pageId="2" pageNumber="583" type="male">2♂♂</specimenCount>
, 
<specimenCount id="9D44FD611F60FD68DD04ED1AFACBFB00" box="[868,903,1872,1898]" pageId="2" pageNumber="583" type="female">1♀</specimenCount>
<collectionCode id="ED53AE2D1F60FD68DDF1ED1AFAA6FB00" box="[913,1002,1872,1898]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="583" type="Museum">MHNG</collectionCode>
; 
<specimenCount id="9D44FD611F60FD68DD99ED1AFD51FB00" box="[1017,1053,1872,1898]" pageId="2" pageNumber="583" type="male">1♂</specimenCount>
, 
<specimenCount id="9D44FD611F60FD68DA4DED1AFD1CFB00" box="[1069,1104,1872,1898]" pageId="2" pageNumber="583" type="female">1♀</specimenCount>
<collectionCode id="ED53AE2D1F60FD68DA3AED1AFDFEFB00" box="[1114,1202,1872,1898]" pageId="2" pageNumber="583">VMDC</collectionCode>
; 
<specimenCount id="9D44FD611F60FD68DADFED1AFDB6FB00" box="[1215,1274,1872,1898]" pageId="2" pageNumber="583" type="male">2♂♂</specimenCount>
, 
<collectionCode id="ED53AE2D1F60FD68DB69ED1AFC16FB00" box="[1289,1370,1872,1898]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="583">MPEG</collectionCode>
; 
<specimenCount id="9D44FD611F60FD68DB08ED1AFCDBFB00" box="[1384,1431,1872,1898]" pageId="2" pageNumber="583" type="male">1 ♂</specimenCount>
, 
<collectionCode id="ED53AE2D1F60FD68DEF7ED3EF9B1FBE4" box="[151,253,1908,1934]" pageId="2" pageNumber="583">IADIZA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2A3CB51F60FD68DF6FED3EFBC7FBD8" ID-GBIF-Occurrence="2572158495" collectionCode="MUSM" country="Peru" pageId="2" pageNumber="583" specimenCount="3" specimenCount-male="3" typeStatus="holotype">
<specimenCount id="9D44FD611F60FD68DF6FED3EF803FBE4" box="[271,335,1908,1934]" pageId="2" pageNumber="583" type="male">3 ♂♂</specimenCount>
labeled as 
<typeStatus id="54F9884A1F60FD68DFAEED3EFB7CFBE4" box="[462,560,1908,1934]" pageId="2" pageNumber="583" type="holotype">holotype</typeStatus>
with additional green label “DNA BARCODING / VOUCHER / BOLD Sys- tems / CCDB-30362-B01 to B03” (
<collectionCode id="ED53AE2D1F60FD68DC47EDD2FBCFFBD8" box="[551,643,1944,1970]" pageId="2" pageNumber="583">MUSM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2A3CB51F60FD68DCF6EDD2FBE2FB90" ID-GBIF-Occurrence="2572158486" collectingDate="2017-10-23" collectionCode="MUSM" collectorName="Rodriguez &amp; L. Perez" country="Peru" county="Department" elevation="2950" latitude="-13.174742" location="Wayqecha Research" longLatPrecision="1" longitude="-71.58666" municipality="Kosnipata" pageId="2" pageNumber="583" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Cusco" typeStatus="paratype">
<specimenCount id="9D44FD611F60FD68DCF6EDD2FB9DFBD8" box="[662,721,1944,1970]" pageId="2" pageNumber="583" type="male">2♂♂</specimenCount>
, 
<specimenCount id="9D44FD611F60FD68DCBDEDD2FA59FBD8" box="[733,789,1944,1970]" pageId="2" pageNumber="583" type="female">6♀♀</specimenCount>
labeled “PERU: CU[
<collectingCounty id="629C4E641F60FD68DA66EDD2FDC2FBD8" box="[1030,1166,1944,1970]" pageId="2" pageNumber="583">Department</collectingCounty>
of 
<collectingRegion id="4986F80A1F60FD68DAD4EDDDFC4DFBD8" box="[1204,1281,1943,1970]" country="Peru" name="Cusco" pageId="2" pageNumber="583">Cuzco</collectingRegion>
], 
<collectingMunicipality id="6B99AC921F60FD68DB74EDD2FCC4FBD8" box="[1300,1416,1944,1970]" pageId="2" pageNumber="583">Kosñipata</collectingMunicipality>
, / 
<location id="8E9D60331F60FD68DEF7EDF6F836FBBC" LSID="urn:lsid:plazi:treatment:03EB87FE1F60FD6CDEF7EAF7F8D5FE55:8E9D60331F60FD68DEF7EDF6F836FBBC" box="[151,378,1980,2006]" country="Peru" county="Department" latitude="-13.174742" longLatPrecision="1" longitude="-71.58666" municipality="Kosnipata" name="Wayqecha Research" pageId="2" pageNumber="583" stateProvince="Cusco">Wayqecha Research</location>
/ 
<location id="8E9D60331F60FD68DFF0EDF6F890FBBC" LSID="urn:lsid:plazi:treatment:03EB87FE1F60FD6CDEF7EAF7F8D5FE55:8E9D60331F60FD68DFF0EDF6F890FBBC" box="[400,476,1980,2006]" country="Peru" county="Department" latitude="-13.174742" longLatPrecision="1" longitude="-71.58666" municipality="Kosnipata" name="Center" pageId="2" pageNumber="583" stateProvince="Cusco">Center</location>
, 
<geoCoordinate id="EE76502F1F60FD68DF89EDF6FBD4FBBC" box="[489,664,1980,2006]" degrees="13" direction="south" minutes="10" orientation="latitude" pageId="2" pageNumber="583" precision="1" seconds="29.07" value="-13.174742">13°10’29.07’’S</geoCoordinate>
,” “ 
<geoCoordinate id="EE76502F1F60FD68DCDAEDF1FA3DFBBC" box="[698,881,1979,2006]" degrees="71" direction="west" minutes="35" orientation="longitude" pageId="2" pageNumber="583" precision="1" seconds="11.98" value="-71.58666">71°35’11.98’’W</geoCoordinate>
, 
<quantity id="4CBA9B0D1F60FD68DD1DEDF1FA99FBBC" box="[893,981,1979,2006]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="2" pageNumber="583" unit="m" value="2950.0">
<elevation id="006FD1DB1F60FD68DD1DEDF1FA99FBBC" box="[893,981,1979,2006]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="2" pageNumber="583" unit="m" value="2950.0">2950 m</elevation>
</quantity>
, / 
<date id="FFFC10281F60FD68DD8BEDF6FD10FBBC" box="[1003,1116,1980,2006]" pageId="2" pageNumber="583" value="2017-10-23">
<collectingDate id="EFB8E9C01F60FD68DD8BEDF6FD10FBBC" box="[1003,1116,1980,2006]" pageId="2" pageNumber="583" value="2017-10-23">23.x.2017</collectingDate>
</date>
, cloud forest, /sifting of leaf litter, M. / 
<collectorName id="26B7533E1F60FD68DF69EDAAF831FB90" box="[265,381,2016,2042]" pageId="2" pageNumber="583">Rodríguez</collectorName>
&amp; 
<collectorName id="26B7533E1F60FD68DFC1EDAAF8B2FB90" box="[417,510,2016,2042]" pageId="2" pageNumber="583">L. Pérez</collectorName>
<emphasis id="B936EAFA1F60FD68DC65EDAAFB6BFB90" box="[517,551,2016,2042]" italics="true" pageId="2" pageNumber="583">leg</emphasis>
.” (
<collectionCode id="ED53AE2D1F60FD68DC2AEDAAFBEAFB90" box="[586,678,2016,2042]" pageId="2" pageNumber="583">MUSM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2A3CB51F60FD69DCD8EDAAFCD4FCD9" ID-GBIF-Occurrence="2572158492" collectingDate="1982-09-24" collectionCode="FMNH" collectorName="L. E. Watrous" country="Peru" county="Peru" lastPageId="3" lastPageNumber="584" location="Pillahuata. Manu" municipality="Dept." pageId="2" pageNumber="583" specimenCount="2" specimenCount-female="2" stateProvince="Cusco" typeStatus="paratype">
“ 
<collectingCounty id="629C4E641F60FD68DCA2EDAAFBB0FB90" box="[706,764,2016,2042]" pageId="2" pageNumber="583">Peru</collectingCounty>
: 
<collectingRegion id="4986F80A1F60FD68DD67EDAAFA03FB90" box="[775,847,2016,2042]" country="Peru" name="Cusco" pageId="2" pageNumber="583">Cuzco</collectingRegion>
<collectingMunicipality id="6B99AC921F60FD68DD35EDAAFAD9FB90" box="[853,917,2016,2042]" pageId="2" pageNumber="583">Dept.</collectingMunicipality>
/, 
<location id="8E9D60331F60FD68DDC9EDAAFD24FB90" LSID="urn:lsid:plazi:treatment:03EB87FE1F60FD6CDEF7EAF7F8D5FE55:8E9D60331F60FD68DDC9EDAAFD24FB90" box="[937,1128,2016,2042]" country="Peru" county="Peru" municipality="Dept." name="Pillahuata. Manu" pageId="2" pageNumber="583" stateProvince="Cusco">Pillahuata. Manu</location>
rd, / km 126, 
<date id="FFFC10281F60FD68DB67EDAAFCC5FB90" box="[1287,1417,2016,2042]" pageId="2" pageNumber="583" value="1982-09-24">
<collectingDate id="EFB8E9C01F60FD68DB67EDAAFCC5FB90" box="[1287,1417,2016,2042]" pageId="2" pageNumber="583" value="1982-09-24">24.IX.1982</collectingDate>
</date>
, / FMND #82-2861, ex” “vine litter &amp; fungus / nr. seepage area, / 
<collectorName id="26B7533E1F61FD69DD0CEAD3FD44FCD9" box="[876,1032,153,179]" pageId="3" pageNumber="584">L. E. Watrous</collectorName>
and G. / Mazurek ”(
<specimenCount id="9D44FD611F61FD69DA91EAD3FC7EFCD9" box="[1265,1330,153,179]" pageId="3" pageNumber="584" type="female">2 ♀♀</specimenCount>
<collectionCode id="ED53AE2D1F61FD69DB5AEAD3FCDCFCD9" box="[1338,1424,153,179]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="584" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2A3CB51F61FD69DEF7EAF7FDF1FCBD" ID-GBIF-Occurrence="2572158494" box="[151,1213,189,215]" collectingDate="1982-09-17" collectionCode="FMNH" collectorName="L. E. Watrous" country="Peru" county="Peru" location="Pillahuata. Manu" municipality="Dept." pageId="3" pageNumber="584" specimenCount="1" specimenCount-female="1" stateProvince="Cusco" typeStatus="paratype">
same data but “km 128, 
<date id="FFFC10281F61FD69DFCDEAF7FB62FCBD" box="[429,558,189,215]" pageId="3" pageNumber="584" value="1982-09-17">
<collectingDate id="EFB8E9C01F61FD69DFCDEAF7FB62FCBD" box="[429,558,189,215]" pageId="3" pageNumber="584" value="1982-09-17">17.IX.1982</collectingDate>
</date>
, FMND #82-249, ex litter &amp; seepage area” (
<specimenCount id="9D44FD611F61FD69DA4DEAF7FD1AFCBD" box="[1069,1110,189,215]" pageId="3" pageNumber="584" type="female">1 ♀</specimenCount>
<collectionCode id="ED53AE2D1F61FD69DA3EEAF7FDF8FCBD" box="[1118,1204,189,215]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="584" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2A3CB51F61FD69DAA9EAF7FA0BFC91" ID-GBIF-Occurrence="2572158493" collectingDate="1982-09-19" collectionCode="FMNH" collectorName="L. E. Watrous" country="Peru" county="Peru" location="Pillahuata. Manu" municipality="Dept." pageId="3" pageNumber="584" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cusco" typeStatus="paratype">
same data but “ 
<date id="FFFC10281F61FD69DB19EAF7F9BAFC91" pageId="3" pageNumber="584" value="1982-09-19">
<collectingDate id="EFB8E9C01F61FD69DB19EAF7F9BAFC91" pageId="3" pageNumber="584" value="1982-09-19">19. IX.1982</collectingDate>
</date>
, FMND #82-262, ex litter nr. falls” (
<specimenCount id="9D44FD611F61FD69DCECEAABFBF9FC91" box="[652,693,225,251]" pageId="3" pageNumber="584" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D44FD611F61FD69DCA0EAABFBAFFC91" box="[704,739,225,251]" pageId="3" pageNumber="584" type="female">1♀</specimenCount>
<collectionCode id="ED53AE2D1F61FD69DC89EAABFA73FC91" box="[745,831,225,251]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="584" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2A3CB51F61FD69DD30EAABFB3BFD75" ID-GBIF-Occurrence="2572158485" collectingDate="1982-09-23" collectionCode="FMNH" collectorName="L. E. Watrous" country="Peru" county="Peru" location="Pillahuata. Manu" municipality="Dept." pageId="3" pageNumber="584" specimenCount="2" specimenCount-female="2" stateProvince="Cusco" typeStatus="paratype">
same data but “ 
<date id="FFFC10281F61FD69DD98EAABFD30FC91" box="[1016,1148,225,251]" pageId="3" pageNumber="584" value="1982-09-23">
<collectingDate id="EFB8E9C01F61FD69DD98EAABFD30FC91" box="[1016,1148,225,251]" pageId="3" pageNumber="584" value="1982-09-23">23.IX.1982</collectingDate>
</date>
, FMND #82-281, ex litter &amp; fungus nr. seepage area” (
<specimenCount id="9D44FD611F61FD69DFB3EB4FFB5FFD75" box="[467,531,261,287]" pageId="3" pageNumber="584" type="female">2 ♀♀</specimenCount>
<collectionCode id="ED53AE2D1F61FD69DC79EB4FFB23FD75" box="[537,623,261,287]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="584" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2A3CB51F61FD69DCE2EB4FFB67FD29" ID-GBIF-Occurrence="2572158490" collectingDate="1982-09-25" collectionCode="FMNH, MHNG" collectorName="L. E. Watrous" country="Peru" county="Peru" location="Pillahuata. Manu" municipality="Dept." pageId="3" pageNumber="584" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Cusco" typeStatus="paratype">
same data but “ 
<date id="FFFC10281F61FD69DD4CEB4EFAFCFD75" box="[812,944,260,287]" pageId="3" pageNumber="584" value="1982-09-25">
<collectingDate id="EFB8E9C01F61FD69DD4CEB4EFAFCFD75" box="[812,944,260,287]" pageId="3" pageNumber="584" value="1982-09-25">25.IX.1982</collectingDate>
</date>
, FMND #82-293, ex litter, some nr. stream” (
<specimenCount id="9D44FD611F61FD69DEC1EB63F989FD29" box="[161,197,297,323]" pageId="3" pageNumber="584" type="male">1♂</specimenCount>
, 
<specimenCount id="9D44FD611F61FD69DEB3EB63F858FD29" box="[211,276,297,323]" pageId="3" pageNumber="584" type="female">2 ♀♀</specimenCount>
<collectionCode id="ED53AE2D1F61FD69DF7DEB63F83EFD29" box="[285,370,297,323]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="584" type="Museum">FMNH</collectionCode>
; 
<specimenCount id="9D44FD611F61FD69DF1FEB63F8F3FD29" box="[383,447,297,323]" pageId="3" pageNumber="584" type="male">2 ♂♂</specimenCount>
<collectionCode id="ED53AE2D1F61FD69DFA8EB63FB6FFD29" box="[456,547,297,323]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="3" pageNumber="584" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2A3CB51F61FD69DC58EB63F882FD0D" ID-GBIF-Occurrence="2572158496" collectingDate="1982-09-26" collectionCode="FMNH, MHNG" collectorName="L. E. Watrous" country="Peru" county="Peru" location="Pillahuata. Manu" municipality="Dept." pageId="3" pageNumber="584" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Cusco" typeStatus="paratype">
same data but “ 
<date id="FFFC10281F61FD69DC89EB63FA21FD29" box="[745,877,297,323]" pageId="3" pageNumber="584" value="1982-09-26">
<collectingDate id="EFB8E9C01F61FD69DC89EB63FA21FD29" box="[745,877,297,323]" pageId="3" pageNumber="584" value="1982-09-26">26.IX.1982</collectingDate>
</date>
, FMND #82-299, ex litter along stream” (
<specimenCount id="9D44FD611F61FD69DB34EB63FCCCFD29" box="[1364,1408,297,323]" pageId="3" pageNumber="584" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D44FD611F61FD69DBEEEB63F98DFD0D" pageId="3" pageNumber="584" type="female">2 ♀♀</specimenCount>
<collectionCode id="ED53AE2D1F61FD69DEA8EB07F850FD0D" box="[200,284,333,359]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="584" type="Museum">FMNH</collectionCode>
, 
<specimenCount id="9D44FD611F61FD69DF46EB07F829FD0D" box="[294,357,333,359]" pageId="3" pageNumber="584" type="female">2 ♀♀</specimenCount>
<collectionCode id="ED53AE2D1F61FD69DF0BEB07F88AFD0D" box="[363,454,333,359]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="3" pageNumber="584" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2A3CB51F61FD69DFB9EB07FDB2FD0D" ID-GBIF-Occurrence="2572158497" box="[473,1278,333,359]" collectingDate="1982-09-27" collectionCode="FMNH" collectorName="L. E. Watrous" country="Peru" county="Peru" location="Pillahuata. Manu" municipality="Dept." pageId="3" pageNumber="584" specimenCount="1" specimenCount-male="1" stateProvince="Cusco" typeStatus="paratype">
same data but “ 
<date id="FFFC10281F61FD69DCE3EB07FA4AFD0D" box="[643,774,333,359]" pageId="3" pageNumber="584" value="1982-09-27">
<collectingDate id="EFB8E9C01F61FD69DCE3EB07FA4AFD0D" box="[643,774,333,359]" pageId="3" pageNumber="584" value="1982-09-27">27.IX.1982</collectingDate>
</date>
, FMND #82-307, ex leaf litter”(
<specimenCount id="9D44FD611F61FD69DA15EB07FDD6FD0D" box="[1141,1178,333,359]" pageId="3" pageNumber="584" type="male">1 ♂</specimenCount>
<collectionCode id="ED53AE2D1F61FD69DAC0EB07FDBAFD0D" box="[1184,1270,333,359]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="584" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B2A3CB51F61FD69DB67EB07FD92FDE1" ID-GBIF-Occurrence="2572158488" country="Peru" location="All" pageId="3" pageNumber="584" specimenCount="2" typeStatus="paratype">
<location id="8E9D60331F61FD69DB67EB07FC67FD0D" LSID="urn:lsid:plazi:treatment:03EB87FE1F60FD6CDEF7EAF7F8D5FE55:8E9D60331F61FD69DB67EB07FC67FD0D" box="[1287,1323,333,359]" country="Peru" name="All" pageId="3" pageNumber="584">All</location>
<typeStatus id="54F9884A1F61FD69DB51EB07FCD0FD0D" box="[1329,1436,333,359]" pageId="3" pageNumber="584" type="paratype">paratypes</typeStatus>
labeled “ 
<typeStatus id="54F9884A1F61FD69DE98EB3BF8CBFDE1" box="[248,391,369,395]" pageId="3" pageNumber="584" type="paratype">PARATYPE</typeStatus>
[yellow label] / 
<emphasis id="B936EAFA1F61FD69DC20EB3BFA06FDE1" box="[576,842,369,395]" italics="true" pageId="3" pageNumber="584">
<taxonomicName id="4C424D6B1F61FD69DC20EB3BFA77FDE1" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" box="[576,827,369,395]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="species" species="wayqecha">Megarthrus wayqecha</taxonomicName>
/
</emphasis>
sp. nov. / Desig. Pérez 
<emphasis id="B936EAFA1F61FD69DA31EB38FDDCFDE1" box="[1105,1168,369,395]" italics="true" pageId="3" pageNumber="584">et al.,</emphasis>
2019”
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF3D66601F61FD69DEF7ECA9FD2AFB02" ID-DOI="http://doi.org/10.5281/zenodo.3661900" ID-Zenodo-Dep="3661900" httpUri="https://zenodo.org/record/3661900/files/figure.png" pageId="3" pageNumber="584" startId="3.[151,264,1763,1788]" targetBox="[195,1399,436,1722]" targetPageId="3">
<paragraph id="8BFD36E81F61FD69DEF7ECA9FD2AFB02" blockId="3.[151,1436,1763,1896]" pageId="3" pageNumber="584">
<emphasis id="B936EAFA1F61FD69DEF7ECA9F808FA96" bold="true" box="[151,324,1763,1788]" pageId="3" pageNumber="584">FIGURES 8–16</emphasis>
. 
<taxonomicName id="4C424D6B1F61FD69DF34ECAEFB76FA96" authority="Pérez &amp; Rodríguez &amp; Asenjo, 2020" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" box="[340,570,1764,1788]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="species" species="wayqecha" status="sp. nov.">
<emphasis id="B936EAFA1F61FD69DF34ECAEFB76FA96" box="[340,570,1764,1788]" italics="true" pageId="3" pageNumber="584">Megarthrus wayqecha</emphasis>
</taxonomicName>
<emphasis id="B936EAFA1F61FD69DC24ECAEFBD4FA96" bold="true" box="[580,664,1764,1788]" pageId="3" pageNumber="584">
<taxonomicNameLabel id="A20557811F61FD69DC24ECAEFBD4FA96" box="[580,664,1764,1788]" pageId="3" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (8 
<emphasis id="B936EAFA1F61FD69DD7FECAEFA60FA96" bold="true" box="[799,812,1764,1788]" pageId="3" pageNumber="584">–</emphasis>
14). 
<emphasis id="B936EAFA1F61FD69DD3DECA9FA3DFA96" bold="true" box="[861,881,1763,1788]" pageId="3" pageNumber="584">8,</emphasis>
tergite VIII, dorsal view; 
<emphasis id="B936EAFA1F61FD69DAE8ECA9FDD9FA96" bold="true" box="[1160,1173,1763,1788]" pageId="3" pageNumber="584">9</emphasis>
, tergite VII, tergite VIII, sternite VIII, lateral view, 
<emphasis id="B936EAFA1F61FD69DFC9ED4DF88EFB4A" bold="true" box="[425,450,1799,1824]" pageId="3" pageNumber="584">10</emphasis>
, sternite VIII; 
<emphasis id="B936EAFA1F61FD69DC3EED4DFB3AFB4A" bold="true" box="[606,630,1799,1824]" pageId="3" pageNumber="584">11</emphasis>
, medial area of abdominal sternites II 
<emphasis id="B936EAFA1F61FD69DA62ED42FD43FB4A" bold="true" box="[1026,1039,1800,1824]" pageId="3" pageNumber="584">–</emphasis>
III in lateral view; 
<emphasis id="B936EAFA1F61FD69DAB5ED4DFDB8FB4A" bold="true" box="[1237,1268,1799,1824]" pageId="3" pageNumber="584">12,</emphasis>
hemitergites IX and sternite IX; 
<emphasis id="B936EAFA1F61FD69DF5DED61F81AFB2E" bold="true" box="[317,342,1835,1860]" pageId="3" pageNumber="584">13</emphasis>
, aedeagus, ventral view; 
<emphasis id="B936EAFA1F61FD69DC3BED61FB38FB2E" bold="true" box="[603,628,1835,1860]" pageId="3" pageNumber="584">14</emphasis>
, aedeagus, lateral view. 
<taxonomicName id="4C424D6B1F61FD69DD10ED66FD18FB2E" authority="Pérez &amp; Rodríguez &amp; Asenjo, 2020" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" box="[880,1108,1836,1860]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="species" species="wayqecha" status="sp. nov.">
<emphasis id="B936EAFA1F61FD69DD10ED66FD18FB2E" box="[880,1108,1836,1860]" italics="true" pageId="3" pageNumber="584">Megarthrus wayqecha</emphasis>
</taxonomicName>
<emphasis id="B936EAFA1F61FD69DA3BED66FDE0FB2E" bold="true" box="[1115,1196,1836,1860]" pageId="3" pageNumber="584">
<taxonomicNameLabel id="A20557811F61FD69DA3BED66FDE0FB2E" box="[1115,1196,1836,1860]" pageId="3" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female paratype, from Wayqecha, Trocha Canopy 
<emphasis id="B936EAFA1F61FD69DFCFED05F882FB02" bold="true" box="[431,462,1871,1896]" pageId="3" pageNumber="584">15,</emphasis>
genital segments, dorsal view; 
<emphasis id="B936EAFA1F61FD69DD6EED05FA61FB02" bold="true" box="[782,813,1871,1896]" pageId="3" pageNumber="584">16,</emphasis>
genital segments, lateral view.
</paragraph>
</caption>
<subSubSection id="C35865631F61FD6EDEA7EDDFF819FD0D" lastPageId="4" lastPageNumber="585" pageId="3" pageNumber="584" type="diagnosis">
<paragraph id="8BFD36E81F61FD6EDEA7EDDFF819FD0D" blockId="3.[151,1436,1941,2041]" lastBlockId="4.[151,1436,153,359]" lastPageId="4" lastPageNumber="585" pageId="3" pageNumber="584">
<emphasis id="B936EAFA1F61FD69DEA7EDDFF80EFBDA" bold="true" box="[199,322,1941,1968]" pageId="3" pageNumber="584">Diagnosis.</emphasis>
Among the members of the genus possessing the eleventh antennomeres piriform, 
<taxonomicName id="4C424D6B1F61FD69DB78EDDCF84BFBBE" authority="Pérez &amp; Rodríguez &amp; Asenjo, 2020" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="species" species="wayqecha" status="sp. nov.">
<emphasis id="B936EAFA1F61FD69DB78EDDCF84BFBBE" italics="true" pageId="3" pageNumber="584">Megarthrus wayqecha</emphasis>
</taxonomicName>
<emphasis id="B936EAFA1F61FD69DF6EEDF0F824FBBE" bold="true" box="[270,360,1978,2004]" pageId="3" pageNumber="584">
<taxonomicNameLabel id="A20557811F61FD69DF6EEDF0F824FBBE" box="[270,360,1978,2004]" pageId="3" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
uniquely shares with the Argentinian 
<taxonomicName id="4C424D6B1F61FD69DD72EDF1FD62FBBE" authority="Bruch, 1940" authorityName="Bruch" authorityYear="1940" box="[786,1070,1978,2004]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="species" species="ogloblini">
<emphasis id="B936EAFA1F61FD69DD72EDF1FAD0FBBE" box="[786,924,1978,2004]" italics="true" pageId="3" pageNumber="584">M. ogloblini</emphasis>
Bruch, 1940
</taxonomicName>
(Fig. 147: Cuccodoro 2011) and the Colombian 
<taxonomicName id="4C424D6B1F61FD69DF24ED95FBABFB92" authority="Lopez-Garcia et al., 2011" authorityName="Lopez-Garcia" authorityYear="2011" box="[324,743,2014,2041]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B936EAFA1F61FD69DF24ED95F88EFB92" box="[324,450,2014,2040]" italics="true" pageId="3" pageNumber="584">M. andinus</emphasis>
<bibRefCitation id="EFD34B191F61FD69DFA9ED94FBABFB92" author="Lopez-Garcia, M. M. &amp; Mendez-Rojas, D. M. &amp; Navarrete-Heredia, J. L." box="[457,743,2014,2041]" pageId="3" pageNumber="584" pagination="62 - 64" refId="ref4619" refString="Lopez-Garcia, M. M., Mendez-Rojas, D. M. &amp; Navarrete-Heredia, J. L. (2011) First record and a new species of Megarthrus Curtis (Coleoptera, Staphylinidae, Proteininae) from the Colombian Central Andes. Zootaxa, 2916 (1), 62 - 64. https: // doi. org / 10.11646 / zootaxa. 2916.1.5" type="journal article" year="2011">
López-García 
<emphasis id="B936EAFA1F61FD69DC0AED95FBD7FB92" box="[618,667,2014,2040]" italics="true" pageId="3" pageNumber="584">et al</emphasis>
., 2011
</bibRefCitation>
</taxonomicName>
the pronotum shallowly notched laterobasally with the lateral outline forming 4 distinct angles. However 
<taxonomicName id="4C424D6B1F66FD6EDCE1EAD0FA4CFCD9" box="[641,768,153,179]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B936EAFA1F66FD6EDCE1EAD0FA4CFCD9" box="[641,768,153,179]" italics="true" pageId="4" pageNumber="585">M. andinus</emphasis>
</taxonomicName>
is readily distinguished from these species by its conspicuous anterior adsutural elytral humps, and 
<taxonomicName id="4C424D6B1F66FD6EDC14EAF4FA4DFCBD" authorityName="Bruch" authorityYear="1940" box="[628,769,189,215]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="ogloblini">
<emphasis id="B936EAFA1F66FD6EDC14EAF4FA4DFCBD" box="[628,769,189,215]" italics="true" pageId="4" pageNumber="585">M. ogloblini</emphasis>
</taxonomicName>
has the male metatibiae cylindrical bearing peg-like setae on apical half, while males of 
<taxonomicName id="4C424D6B1F66FD6EDF8DEAA8FBCFFC91" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" box="[493,643,225,251]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="wayqecha">
<emphasis id="B936EAFA1F66FD6EDF8DEAA8FBCFFC91" box="[493,643,225,251]" italics="true" pageId="4" pageNumber="585">M. wayqecha</emphasis>
</taxonomicName>
have the metatibiae enlarged and notched with peg-like setae grouped subapically. The shape of the aedeagus of 
<taxonomicName id="4C424D6B1F66FD6EDC16EB4CFA40FD75" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" box="[630,780,261,287]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="wayqecha">
<emphasis id="B936EAFA1F66FD6EDC16EB4CFA40FD75" box="[630,780,261,287]" italics="true" pageId="4" pageNumber="585">M. wayqecha</emphasis>
</taxonomicName>
is also diagnostic: the median lobe in lateral view has the ventral face subapically straight with the tip of apex acute (
<figureCitation id="13792A6D1F66FD6EDD4EEB63FACCFD29" box="[814,896,297,323]" captionStart="FIGURES 17–19" captionStartId="4.[151,264,947,972]" captionTargetBox="[156,1430,393,916]" captionTargetId="figure@4.[152,1436,381,922]" captionTargetPageId="4" captionText="FIGURES 17–19. Megarthrus wayqecha sp. nov., male paratype. 17, aedeagus, ventral view (the side of the basal foramen); 18, aedeagus, lateral view; 19, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3661902" httpUri="https://zenodo.org/record/3661902/files/figure.png" pageId="4" pageNumber="585">Fig. 18</figureCitation>
) and the dorsal valve (seen in dorsal view) is Vshaped (
<figureCitation id="13792A6D1F66FD6EDE94EB07F804FD0D" box="[244,328,333,359]" captionStart="FIGURES 17–19" captionStartId="4.[151,264,947,972]" captionTargetBox="[156,1430,393,916]" captionTargetId="figure@4.[152,1436,381,922]" captionTargetPageId="4" captionText="FIGURES 17–19. Megarthrus wayqecha sp. nov., male paratype. 17, aedeagus, ventral view (the side of the basal foramen); 18, aedeagus, lateral view; 19, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3661902" httpUri="https://zenodo.org/record/3661902/files/figure.png" pageId="4" pageNumber="585">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3D66601F66FD6EDEF7E9F9FBFAFF9A" ID-DOI="http://doi.org/10.5281/zenodo.3661902" ID-Zenodo-Dep="3661902" httpUri="https://zenodo.org/record/3661902/files/figure.png" pageId="4" pageNumber="585" startId="4.[151,264,947,972]" targetBox="[156,1430,393,916]" targetPageId="4">
<paragraph id="8BFD36E81F66FD6EDEF7E9F9FBFAFF9A" blockId="4.[151,1436,947,1008]" pageId="4" pageNumber="585">
<emphasis id="B936EAFA1F66FD6EDEF7E9F9F819FFA6" bold="true" box="[151,341,947,972]" pageId="4" pageNumber="585">FIGURES 17–19.</emphasis>
<taxonomicName id="4C424D6B1F66FD6EDF3CE9FEFB0CFFA6" authority="Pérez &amp; Rodríguez &amp; Asenjo, 2020" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" box="[348,576,948,972]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="wayqecha" status="sp. nov.">
<emphasis id="B936EAFA1F66FD6EDF3CE9FEFB0CFFA6" box="[348,576,948,972]" italics="true" pageId="4" pageNumber="585">Megarthrus wayqecha</emphasis>
</taxonomicName>
<emphasis id="B936EAFA1F66FD6EDC27E9FEFBD5FFA6" bold="true" box="[583,665,948,972]" pageId="4" pageNumber="585">
<taxonomicNameLabel id="A20557811F66FD6EDC27E9FEFBD5FFA6" box="[583,665,948,972]" pageId="4" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype. 
<emphasis id="B936EAFA1F66FD6EDD24E9F9FA11FFA6" bold="true" box="[836,861,947,972]" pageId="4" pageNumber="585">17</emphasis>
, aedeagus, ventral view (the side of the basal foramen); 
<emphasis id="B936EAFA1F66FD6EDEF7E99DF9FCFF9A" bold="true" box="[151,176,983,1008]" pageId="4" pageNumber="585">18</emphasis>
, aedeagus, lateral view; 
<emphasis id="B936EAFA1F66FD6EDFCBE99DF888FF9A" bold="true" box="[427,452,983,1008]" pageId="4" pageNumber="585">19</emphasis>
, aedeagus, dorsal view.
</paragraph>
</caption>
<caption id="DF3D66601F66FD6EDEF7ECC6F98EFA87" ID-DOI="http://doi.org/10.5281/zenodo.3661904" ID-Zenodo-Dep="3661904" httpUri="https://zenodo.org/record/3661904/files/figure.png" pageId="4" pageNumber="585" startId="4.[151,264,1676,1701]" targetBox="[151,1436,1066,1651]" targetPageId="4">
<paragraph id="8BFD36E81F66FD6EDEF7ECC6F98EFA87" blockId="4.[151,1436,1676,1773]" pageId="4" pageNumber="585">
<emphasis id="B936EAFA1F66FD6EDEF7ECC6F819FACF" bold="true" box="[151,341,1676,1701]" pageId="4" pageNumber="585">FIGURES 20–21.</emphasis>
Habitat of 
<taxonomicName id="4C424D6B1F66FD6EDFA9ECC7FBE1FACF" authority="Pérez &amp; Rodríguez &amp; Asenjo, 2020" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" box="[457,685,1677,1701]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="wayqecha" status="sp. nov.">
<emphasis id="B936EAFA1F66FD6EDFA9ECC7FBE1FACF" box="[457,685,1677,1701]" italics="true" pageId="4" pageNumber="585">Megarthrus wayqecha</emphasis>
</taxonomicName>
<emphasis id="B936EAFA1F66FD6EDCD4ECC7FA60FACF" bold="true" box="[692,812,1676,1701]" pageId="4" pageNumber="585">
<taxonomicNameLabel id="A20557811F66FD6EDCD4ECC7FA49FACF" box="[692,773,1677,1701]" pageId="4" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
20,
</emphasis>
panoramic view of the cloud forests of the Kosñipata valley, where the biological station Wayqecha is located. 
<emphasis id="B936EAFA1F66FD6EDCFCECFAFBF7FAA3" bold="true" box="[668,699,1712,1737]" pageId="4" pageNumber="585">21,</emphasis>
sifting leaf litter on Trocha Canopy from the biological station Wayqecha.
</paragraph>
</caption>
<subSubSection id="C35865631F66FD6FDEA7ED50FB4DFA1D" lastPageId="5" lastPageNumber="586" pageId="4" pageNumber="585" type="description">
<paragraph id="8BFD36E81F66FD6FDEA7ED50FD65FCBD" blockId="4.[151,1437,1818,2026]" lastBlockId="5.[151,1437,153,2015]" lastPageId="5" lastPageNumber="586" pageId="4" pageNumber="585">
<emphasis id="B936EAFA1F66FD6EDEA7ED50F815FB5F" bold="true" box="[199,345,1818,1845]" pageId="4" pageNumber="585">Description.</emphasis>
<typeStatus id="54F9884A1F66FD6EDF00ED51F884FB5F" box="[352,456,1819,1845]" pageId="4" pageNumber="585" type="holotype">Holotype</typeStatus>
male (BL=3.11, BW=1.18, PL+EL=1.82). Body dark brown with appendages and legs somewhat paler, antennomeres 9–11 paler than antennomeres 1–8 (
<figureCitation id="13792A6D1F66FD6EDDF2ED75FA98FB33" box="[914,980,1855,1881]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1549,1574]" captionTargetBox="[156,1432,348,1519]" captionTargetId="figure@2.[151,1436,343,1522]" captionTargetPageId="2" captionText="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3661898" httpUri="https://zenodo.org/record/3661898/files/figure.png" pageId="4" pageNumber="585">Fig. 1</figureCitation>
). Dorsal pubescence of body fairly uniform, frontal setae directed forward, median area of frons less dense than on vertex; pronotum, elytra and abdomen with setae directed backward, denser on pronotum than on elytra, pronotal setae curved and recumbent, denser on pronotal disc; elytral setae curved and recumbent, becoming denser near humeral area with interspaces as wide as puncture diameters; metaventral pubescence uniform, with small protuberance near posterior margin; pubescence on abdominal tergites parallel, uniform. Frons with granulation conspicuous, with granulae about as high as their diameter or higher; pronotum and elytra granulofossulate, with granulation small.
</paragraph>
<paragraph id="8BFD36E81F67FD6FDEA7EAABFD7EFD0D" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="586">
Head (
<figureCitation id="13792A6D1F67FD6FDF72EAABF818FC91" box="[274,340,225,251]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1549,1574]" captionTargetBox="[156,1432,348,1519]" captionTargetId="figure@2.[151,1436,343,1522]" captionTargetPageId="2" captionText="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3661898" httpUri="https://zenodo.org/record/3661898/files/figure.png" pageId="5" pageNumber="586">Fig. 3</figureCitation>
) wider than long (HW=0.71, HL=0.53), mesal portion of disc strongly convex in lateral view; Ushaped frontal impression shallow; eyes medium size on posterior half; clypeus carinate, nearly rounded; temples convex in dorsal view. Antenna (
<figureCitation id="13792A6D1F67FD6FDC6DEB63FB1EFD29" box="[525,594,297,323]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1549,1574]" captionTargetBox="[156,1432,348,1519]" captionTargetId="figure@2.[151,1436,343,1522]" captionTargetPageId="2" captionText="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3661898" httpUri="https://zenodo.org/record/3661898/files/figure.png" pageId="5" pageNumber="586">Fig. 2</figureCitation>
) symmetrical and filiform, 2.1 times longer than pronotum, antennomeres 1–4 without pubescence, antennomeres 5–11 pubescent; antennomere 11 piriform.
</paragraph>
<paragraph id="8BFD36E81F67FD6FDEA7EB3BFC2CFE55" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="586">
Pronotum wider than long (PW=1.27, PL=0.65), 1.2 times longer than head, anterior margin slightly concave and posterior margin slightly convex and bilobed in the middle (
<figureCitation id="13792A6D1F67FD6FDD14EBDFFAF7FDC5" box="[884,955,405,431]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1549,1574]" captionTargetBox="[156,1432,348,1519]" captionTargetId="figure@2.[151,1436,343,1522]" captionTargetPageId="2" captionText="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3661898" httpUri="https://zenodo.org/record/3661898/files/figure.png" pageId="5" pageNumber="586">Fig. 4</figureCitation>
). Pronotum with disc strongly convex and deeply depressed near middle of lateral edges, lateral and posterior edges shallowly depressed along disc. Anterior angles projected forward, lateral contours forming four denticles, posterior angles weakly projected backwards. Medial groove punctate and flat, slightly narrower in middle (
<figureCitation id="13792A6D1F67FD6FDD48E84BFA21FE71" box="[808,877,513,539]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1549,1574]" captionTargetBox="[156,1432,348,1519]" captionTargetId="figure@2.[151,1436,343,1522]" captionTargetPageId="2" captionText="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3661898" httpUri="https://zenodo.org/record/3661898/files/figure.png" pageId="5" pageNumber="586">Fig. 4</figureCitation>
). Hypomeron with inner edge wider in its median portion, ridged from anterior margin to laterobasal angle, without a discal pit. Proventral medial ridge absent.
</paragraph>
<paragraph id="8BFD36E81F67FD6FDEA7E803FBC0F831" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="586">
Scutellum triangular with anterior border rounded and posterior border slightly acute, margins slightly raised and with pubescence directed backward. Elytra wider than long (EL=0.96, EW=1.30), 1.8 times longer than pronotum, gradually widened toward apex (
<figureCitation id="13792A6D1F67FD6FDC27E8DBFBC0FEC1" box="[583,652,657,683]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1549,1574]" captionTargetBox="[156,1432,348,1519]" captionTargetId="figure@2.[151,1436,343,1522]" captionTargetPageId="2" captionText="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3661898" httpUri="https://zenodo.org/record/3661898/files/figure.png" pageId="5" pageNumber="586">Fig. 2</figureCitation>
), humeral callus raised, moderately convex; disc with low swellings, moderately depressed posteriorly and along lateral external edge, the latter finely carinate, slightly arcuate in dorsal view. Protrochanter without transverse ridge, small, subtriangular with one macroseta near the apex; profemur oblong with base flattened; protibia thin and shorter than profemur, inner face with many setae in the apical two thirds; protarsomere 1–4 dorsoventrally setose, protarsomere 1 as long as protarsomere 2–3 combined, protarsomere 5 with scarce setae, longer than protarsomere 1. Mesotrochanter subtriangular with peg-like setae arranged in two rows, one row with 7–8 peg-like setae and the second with 4–5 peg-like setae, and bearing one subapical macroseta (
<figureCitation id="13792A6D1F67FD6FDEC0E9C7F9A9FFCD" box="[160,229,908,935]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1549,1574]" captionTargetBox="[156,1432,348,1519]" captionTargetId="figure@2.[151,1436,343,1522]" captionTargetPageId="2" captionText="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3661898" httpUri="https://zenodo.org/record/3661898/files/figure.png" pageId="5" pageNumber="586">Fig. 5</figureCitation>
); mesofemur oblong; mesotibia sinuate, shorter than mesofemur, with peg-like setae arranged in two rows, a row along 4/5 apical length and a short row from base to middle, and some apical peg-like setae not aligned (
<figureCitation id="13792A6D1F67FD6FDB0FE9FBF9EBFF9A" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1549,1574]" captionTargetBox="[156,1432,348,1519]" captionTargetId="figure@2.[151,1436,343,1522]" captionTargetPageId="2" captionText="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3661898" httpUri="https://zenodo.org/record/3661898/files/figure.png" pageId="5" pageNumber="586">Fig. 6</figureCitation>
); mesotarsomeres similar to protarsomeres. Metatrochanter subtriangular, incrassate (
<figureCitation id="13792A6D1F67FD6FDA06E99FFDE6FF85" box="[1126,1194,981,1007]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1549,1574]" captionTargetBox="[156,1432,348,1519]" captionTargetId="figure@2.[151,1436,343,1522]" captionTargetPageId="2" captionText="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3661898" httpUri="https://zenodo.org/record/3661898/files/figure.png" pageId="5" pageNumber="586">Fig. 7</figureCitation>
); metafemur strongly very robust about twice as long as broad (0.71/0.30) (
<figureCitation id="13792A6D1F67FD6FDC86E9B3FA65F879" box="[742,809,1017,1043]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1549,1574]" captionTargetBox="[156,1432,348,1519]" captionTargetId="figure@2.[151,1436,343,1522]" captionTargetPageId="2" captionText="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3661898" httpUri="https://zenodo.org/record/3661898/files/figure.png" pageId="5" pageNumber="586">Fig. 7</figureCitation>
); metatibia gradually enlarged toward apex, shorter than metafemur, with notched at subapical quarter constriction, and with peg-like setae grouped on apical quarter (
<figureCitation id="13792A6D1F67FD6FDB0FEE57F9EBF831" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1549,1574]" captionTargetBox="[156,1432,348,1519]" captionTargetId="figure@2.[151,1436,343,1522]" captionTargetPageId="2" captionText="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3661898" httpUri="https://zenodo.org/record/3661898/files/figure.png" pageId="5" pageNumber="586">Fig. 7</figureCitation>
); metatarsomeres similar to protarsomeres.
</paragraph>
<paragraph id="8BFD36E81F67FD6FDEA7EE2FF8AAF9F5" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="586">
Tergites III–VII pubescent with one pair of paratergites; tergites II–III covered by elytra, tergite IV partially covered. Medial process of sternite II shorter and smaller than medial process of sternite III (
<figureCitation id="13792A6D1F67FD6FDACCEEC3FDB2F8C9" box="[1196,1278,1161,1187]" captionStart="FIGURES 8–16" captionStartId="3.[151,264,1763,1788]" captionTargetBox="[195,1399,436,1722]" captionTargetId="figure@3.[177,1411,417,1738]" captionTargetPageId="3" captionText="FIGURES 8–16. Megarthrus wayqecha sp. nov., holotype (8–14). 8, tergite VIII, dorsal view; 9, tergite VII, tergite VIII, sternite VIII, lateral view, 10, sternite VIII; 11, medial area of abdominal sternites II–III in lateral view; 12, hemitergites IX and sternite IX; 13, aedeagus, ventral view; 14, aedeagus, lateral view. Megarthrus wayqecha sp. nov., female paratype, from Wayqecha, Trocha Canopy 15, genital segments, dorsal view; 16, genital segments, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3661900" httpUri="https://zenodo.org/record/3661900/files/figure.png" pageId="5" pageNumber="586">Fig. 11</figureCitation>
). Sternite III– VII rectangular with pubescence. Tergite VIII triangular with anterior angles slightly projected forward, posterior margin forming an obtuse angle (
<figureCitation id="13792A6D1F67FD6FDC6AEE9BFB01F881" box="[522,589,1233,1259]" captionStart="FIGURES 8–16" captionStartId="3.[151,264,1763,1788]" captionTargetBox="[195,1399,436,1722]" captionTargetId="figure@3.[177,1411,417,1738]" captionTargetPageId="3" captionText="FIGURES 8–16. Megarthrus wayqecha sp. nov., holotype (8–14). 8, tergite VIII, dorsal view; 9, tergite VII, tergite VIII, sternite VIII, lateral view, 10, sternite VIII; 11, medial area of abdominal sternites II–III in lateral view; 12, hemitergites IX and sternite IX; 13, aedeagus, ventral view; 14, aedeagus, lateral view. Megarthrus wayqecha sp. nov., female paratype, from Wayqecha, Trocha Canopy 15, genital segments, dorsal view; 16, genital segments, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3661900" httpUri="https://zenodo.org/record/3661900/files/figure.png" pageId="5" pageNumber="586">Fig. 8</figureCitation>
), with small and broad medial apical projection convex in lateral view (
<figureCitation id="13792A6D1F67FD6FDB0FEE9BF9EBF965" captionStart="FIGURES 8–16" captionStartId="3.[151,264,1763,1788]" captionTargetBox="[195,1399,436,1722]" captionTargetId="figure@3.[177,1411,417,1738]" captionTargetPageId="3" captionText="FIGURES 8–16. Megarthrus wayqecha sp. nov., holotype (8–14). 8, tergite VIII, dorsal view; 9, tergite VII, tergite VIII, sternite VIII, lateral view, 10, sternite VIII; 11, medial area of abdominal sternites II–III in lateral view; 12, hemitergites IX and sternite IX; 13, aedeagus, ventral view; 14, aedeagus, lateral view. Megarthrus wayqecha sp. nov., female paratype, from Wayqecha, Trocha Canopy 15, genital segments, dorsal view; 16, genital segments, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3661900" httpUri="https://zenodo.org/record/3661900/files/figure.png" pageId="5" pageNumber="586">Fig. 9</figureCitation>
). Sternite VIII trapezoidal, with lateral margin slightly convex, posterior margin strongly concave and membranous with four short macrosetae, anterior margin characteristic, inner plate asymmetrical with apex acute exceeding a little the anterior margin of sternite VIII (
<figureCitation id="13792A6D1F67FD6FDC16EF77FB85F93D" box="[630,713,1341,1367]" captionStart="FIGURES 8–16" captionStartId="3.[151,264,1763,1788]" captionTargetBox="[195,1399,436,1722]" captionTargetId="figure@3.[177,1411,417,1738]" captionTargetPageId="3" captionText="FIGURES 8–16. Megarthrus wayqecha sp. nov., holotype (8–14). 8, tergite VIII, dorsal view; 9, tergite VII, tergite VIII, sternite VIII, lateral view, 10, sternite VIII; 11, medial area of abdominal sternites II–III in lateral view; 12, hemitergites IX and sternite IX; 13, aedeagus, ventral view; 14, aedeagus, lateral view. Megarthrus wayqecha sp. nov., female paratype, from Wayqecha, Trocha Canopy 15, genital segments, dorsal view; 16, genital segments, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3661900" httpUri="https://zenodo.org/record/3661900/files/figure.png" pageId="5" pageNumber="586">Fig. 10</figureCitation>
). Hemitergites IX slightly asymmetrical, left hemitergite slightly bigger in ventral view, anterior margin of right hemitergite more concave than left (
<figureCitation id="13792A6D1F67FD6FDA55EF2BFDC4F911" box="[1077,1160,1377,1403]" captionStart="FIGURES 8–16" captionStartId="3.[151,264,1763,1788]" captionTargetBox="[195,1399,436,1722]" captionTargetId="figure@3.[177,1411,417,1738]" captionTargetPageId="3" captionText="FIGURES 8–16. Megarthrus wayqecha sp. nov., holotype (8–14). 8, tergite VIII, dorsal view; 9, tergite VII, tergite VIII, sternite VIII, lateral view, 10, sternite VIII; 11, medial area of abdominal sternites II–III in lateral view; 12, hemitergites IX and sternite IX; 13, aedeagus, ventral view; 14, aedeagus, lateral view. Megarthrus wayqecha sp. nov., female paratype, from Wayqecha, Trocha Canopy 15, genital segments, dorsal view; 16, genital segments, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3661900" httpUri="https://zenodo.org/record/3661900/files/figure.png" pageId="5" pageNumber="586">Fig. 12</figureCitation>
). Sternite IX asymmetrical, oblong and thin (
<figureCitation id="13792A6D1F67FD6FDFE5EFCFF894F9F5" box="[389,472,1413,1439]" captionStart="FIGURES 8–16" captionStartId="3.[151,264,1763,1788]" captionTargetBox="[195,1399,436,1722]" captionTargetId="figure@3.[177,1411,417,1738]" captionTargetPageId="3" captionText="FIGURES 8–16. Megarthrus wayqecha sp. nov., holotype (8–14). 8, tergite VIII, dorsal view; 9, tergite VII, tergite VIII, sternite VIII, lateral view, 10, sternite VIII; 11, medial area of abdominal sternites II–III in lateral view; 12, hemitergites IX and sternite IX; 13, aedeagus, ventral view; 14, aedeagus, lateral view. Megarthrus wayqecha sp. nov., female paratype, from Wayqecha, Trocha Canopy 15, genital segments, dorsal view; 16, genital segments, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3661900" httpUri="https://zenodo.org/record/3661900/files/figure.png" pageId="5" pageNumber="586">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8BFD36E81F67FD6FDEA7EFE3FC39F98D" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="586">
Aedeagus as in 
<figureCitation id="13792A6D1F67FD6FDF19EFE3F8B2F9A9" box="[377,510,1449,1475]" captionStart="FIGURES 8–16" captionStartId="3.[151,264,1763,1788]" captionTargetBox="[195,1399,436,1722]" captionTargetId="figure@3.[177,1411,417,1738]" captionTargetPageId="3" captionText="FIGURES 8–16. Megarthrus wayqecha sp. nov., holotype (8–14). 8, tergite VIII, dorsal view; 9, tergite VII, tergite VIII, sternite VIII, lateral view, 10, sternite VIII; 11, medial area of abdominal sternites II–III in lateral view; 12, hemitergites IX and sternite IX; 13, aedeagus, ventral view; 14, aedeagus, lateral view. Megarthrus wayqecha sp. nov., female paratype, from Wayqecha, Trocha Canopy 15, genital segments, dorsal view; 16, genital segments, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3661900" httpUri="https://zenodo.org/record/3661900/files/figure.png" pageId="5" pageNumber="586">Figs. 13–14</figureCitation>
(internal sac evaginated), with median lobe oval-shaped in ventral view (
<figureCitation id="13792A6D1F67FD6FDB5BEFE3FCC2F9A9" box="[1339,1422,1449,1475]" captionStart="FIGURES 8–16" captionStartId="3.[151,264,1763,1788]" captionTargetBox="[195,1399,436,1722]" captionTargetId="figure@3.[177,1411,417,1738]" captionTargetPageId="3" captionText="FIGURES 8–16. Megarthrus wayqecha sp. nov., holotype (8–14). 8, tergite VIII, dorsal view; 9, tergite VII, tergite VIII, sternite VIII, lateral view, 10, sternite VIII; 11, medial area of abdominal sternites II–III in lateral view; 12, hemitergites IX and sternite IX; 13, aedeagus, ventral view; 14, aedeagus, lateral view. Megarthrus wayqecha sp. nov., female paratype, from Wayqecha, Trocha Canopy 15, genital segments, dorsal view; 16, genital segments, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3661900" httpUri="https://zenodo.org/record/3661900/files/figure.png" pageId="5" pageNumber="586">Fig. 13</figureCitation>
), in lateral view with ventral face subapically straight with the tip of apex acute (
<figureCitation id="13792A6D1F67FD6FDA6EEF87FD2DF98D" box="[1038,1121,1485,1511]" captionStart="FIGURES 8–16" captionStartId="3.[151,264,1763,1788]" captionTargetBox="[195,1399,436,1722]" captionTargetId="figure@3.[177,1411,417,1738]" captionTargetPageId="3" captionText="FIGURES 8–16. Megarthrus wayqecha sp. nov., holotype (8–14). 8, tergite VIII, dorsal view; 9, tergite VII, tergite VIII, sternite VIII, lateral view, 10, sternite VIII; 11, medial area of abdominal sternites II–III in lateral view; 12, hemitergites IX and sternite IX; 13, aedeagus, ventral view; 14, aedeagus, lateral view. Megarthrus wayqecha sp. nov., female paratype, from Wayqecha, Trocha Canopy 15, genital segments, dorsal view; 16, genital segments, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3661900" httpUri="https://zenodo.org/record/3661900/files/figure.png" pageId="5" pageNumber="586">Fig. 14</figureCitation>
), dorsal valve V-shaped.
</paragraph>
<paragraph id="8BFD36E81F67FD6FDEF7EFBBFB4DFA1D" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="586">
<emphasis id="B936EAFA1F67FD6FDEF7EFBBF9A7FA61" box="[151,235,1521,1547]" italics="true" pageId="5" pageNumber="586">Female</emphasis>
. Similar to male, except mesotrochanter without peg-like setae; mesotibia without peg-like; metafemur thin; metatibia without peg-like setae; tergite VIII in lateral view with little and thin medioapical projection convex; sternite VIII without inner plate and posterior margin almost straight; genital segments as in 
<figureCitation id="13792A6D1F67FD6FDAF6EC73FC51FA39" box="[1174,1309,1592,1619]" captionStart="FIGURES 8–16" captionStartId="3.[151,264,1763,1788]" captionTargetBox="[195,1399,436,1722]" captionTargetId="figure@3.[177,1411,417,1738]" captionTargetPageId="3" captionText="FIGURES 8–16. Megarthrus wayqecha sp. nov., holotype (8–14). 8, tergite VIII, dorsal view; 9, tergite VII, tergite VIII, sternite VIII, lateral view, 10, sternite VIII; 11, medial area of abdominal sternites II–III in lateral view; 12, hemitergites IX and sternite IX; 13, aedeagus, ventral view; 14, aedeagus, lateral view. Megarthrus wayqecha sp. nov., female paratype, from Wayqecha, Trocha Canopy 15, genital segments, dorsal view; 16, genital segments, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3661900" httpUri="https://zenodo.org/record/3661900/files/figure.png" pageId="5" pageNumber="586">Figs. 15–16</figureCitation>
; gonocoxal plate lacking mediodorsal ridge.
</paragraph>
</subSubSection>
<subSubSection id="C35865631F67FD6CDEA7ECCBF986FCBD" lastPageId="6" lastPageNumber="587" pageId="5" pageNumber="586" type="discussion">
<paragraph id="8BFD36E81F67FD6FDEA7ECCBFC14FBD1" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="586">
<emphasis id="B936EAFA1F67FD6FDEA7ECCBF877FAF1" bold="true" box="[199,315,1665,1691]" pageId="5" pageNumber="586">Remarks.</emphasis>
The new species 
<taxonomicName id="4C424D6B1F67FD6FDC72ECC8FBE7FAF1" authority="Pérez &amp; Rodríguez &amp; Asenjo, 2020" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" box="[530,683,1665,1691]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="wayqecha" status="sp. nov.">
<emphasis id="B936EAFA1F67FD6FDC72ECC8FBE7FAF1" box="[530,683,1665,1691]" italics="true" pageId="5" pageNumber="586">M. wayqecha</emphasis>
</taxonomicName>
<emphasis id="B936EAFA1F67FD6FDCD6ECCBFA5FFAF1" bold="true" box="[694,787,1665,1691]" pageId="5" pageNumber="586">
<taxonomicNameLabel id="A20557811F67FD6FDCD6ECCBFA5FFAF1" box="[694,787,1665,1691]" pageId="5" pageNumber="586" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is included in the definition of 
<taxonomicName id="4C424D6B1F67FD6FDAEFECCBFCDFFAF1" authorityName="Bierig" authorityYear="1940" box="[1167,1427,1665,1691]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B936EAFA1F67FD6FDAEFECCBFCDFFAF1" box="[1167,1427,1665,1691]" italics="true" pageId="5" pageNumber="586">Megarthrus inaequalis</emphasis>
</taxonomicName>
- supergroup proposed by 
<bibRefCitation id="EFD34B191F67FD6FDFD2ECEFFBC0FAD5" author="Cuccodoro, G." box="[434,652,1701,1727]" pageId="5" pageNumber="586" pagination="107 - 147" refId="ref4167" refString="Cuccodoro, G. (2011 a) Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae). Revue Suisse de Zoologie, 118 (1), 107 - 147. https: // doi. org / 10.5962 / bhl. part. 117802" type="journal article" year="2011">Cuccodoro (2011a)</bibRefCitation>
based on the following characters: presence of a long prohypomeral ridge extended posteriorly to the laterobasal angle, frontal pubescence directed forward (
<figureCitation id="13792A6D1F67FD6FDAE7EC83FDB8FA89" box="[1159,1268,1737,1763]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1549,1574]" captionTargetBox="[156,1432,348,1519]" captionTargetId="figure@2.[151,1436,343,1522]" captionTargetPageId="2" captionText="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3661898" httpUri="https://zenodo.org/record/3661898/files/figure.png" pageId="5" pageNumber="586">Figs. 1, 3</figureCitation>
), antennomere 11 piriform (
<figureCitation id="13792A6D1F67FD6FDF45ECA7F8DCFB6D" box="[293,400,1773,1799]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1549,1574]" captionTargetBox="[156,1432,348,1519]" captionTargetId="figure@2.[151,1436,343,1522]" captionTargetPageId="2" captionText="FIGURES 1–7. Megarthrus wayqecha sp. nov., male holotype from Wayqecha, Trocha Canopy (2, 4–6). 2, antenna; 4, pronotum; 5, mesotrochanter and mesofemur; 6, mesotibia. Megarthrus wayqecha sp. nov., male paratype from Wayqecha, Trocha Canopy (1, 3, 7). 1, habitus, dorsal view; 3, head, dorsal view; 7, metaleg with peg-like setae in the apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3661898" httpUri="https://zenodo.org/record/3661898/files/figure.png" pageId="5" pageNumber="586">Figs. 1, 2</figureCitation>
), and lack of adventral adhesive setae on the first male protarsomere. Within this supergroup only 
<taxonomicName id="4C424D6B1F67FD6FDEB2ED58F825FB41" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" box="[210,361,1809,1835]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="wayqecha">
<emphasis id="B936EAFA1F67FD6FDEB2ED58F825FB41" box="[210,361,1809,1835]" italics="true" pageId="5" pageNumber="586">M. wayqecha</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C424D6B1F67FD6FDF18ED58F8B4FB41" box="[376,504,1809,1835]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B936EAFA1F67FD6FDF18ED58F8B4FB41" box="[376,504,1809,1835]" italics="true" pageId="5" pageNumber="586">M. andinus</emphasis>
</taxonomicName>
and the Costa Rican 
<taxonomicName id="4C424D6B1F67FD6FDC8FED58FD6BFB41" authority="Bierig, 1940" authorityName="Bierig" authorityYear="1940" box="[751,1063,1809,1835]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="mammiger">
<emphasis id="B936EAFA1F67FD6FDC8FED58FAC2FB41" box="[751,910,1810,1835]" italics="true" pageId="5" pageNumber="586">M. mammiger</emphasis>
Bierig, 1940
</taxonomicName>
, lack a mediodorsal ridge on the female gonocoxal plate. However, 
<taxonomicName id="4C424D6B1F67FD6FDC7EED7CFBF7FB25" authorityName="Bierig" authorityYear="1940" box="[542,699,1846,1871]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="mammiger">
<emphasis id="B936EAFA1F67FD6FDC7EED7CFBF7FB25" box="[542,699,1846,1871]" italics="true" pageId="5" pageNumber="586">M. mammiger</emphasis>
</taxonomicName>
has the pronotum conspicuously notched laterobasally, while it is only slightly notched laterobasally in 
<taxonomicName id="4C424D6B1F67FD6FDC5DED10FB9EFB19" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" box="[573,722,1881,1907]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="wayqecha">
<emphasis id="B936EAFA1F67FD6FDC5DED10FB9EFB19" box="[573,722,1881,1907]" italics="true" pageId="5" pageNumber="586">M. wayqecha</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C424D6B1F67FD6FDD68ED10FACBFB19" box="[776,903,1881,1907]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B936EAFA1F67FD6FDD68ED10FACBFB19" box="[776,903,1881,1907]" italics="true" pageId="5" pageNumber="586">M. andinus</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C424D6B1F67FD6FDDF4ED13FDC3FB19" authorityName="Pérez &amp; Rodríguez &amp; Asenjo" authorityYear="2020" box="[916,1167,1881,1907]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="wayqecha">
<emphasis id="B936EAFA1F67FD6FDDF4ED13FDC3FB19" box="[916,1167,1881,1907]" italics="true" pageId="5" pageNumber="586">Megarthrus wayqecha</emphasis>
</taxonomicName>
differs notably from 
<taxonomicName id="4C424D6B1F67FD6FDB1EED10F9BCFBFD" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B936EAFA1F67FD6FDB1EED10F9BCFBFD" italics="true" pageId="5" pageNumber="586">M. andinus</emphasis>
</taxonomicName>
by its elytra lacking conspicuous antebasal adsutural humps. The sexually dimorphic features on the male legs, the shape of the aedeagal ventral wall, and the conformation of the female valvifers are also diagnostic.
</paragraph>
<paragraph id="8BFD36E81F67FD6FDEA7ED8FFD1CFBB5" blockId="5.[151,1437,153,2015]" box="[199,1104,1989,2015]" pageId="5" pageNumber="586">
Aedeagus with the internal sac not evaginated from a 
<typeStatus id="54F9884A1F67FD6FDD42ED8FFACEFBB5" box="[802,898,1989,2015]" pageId="5" pageNumber="586" type="paratype">paratype</typeStatus>
as in 
<figureCitation id="13792A6D1F67FD6FDDA4ED8FFD07FBB5" box="[964,1099,1989,2015]" captionStart="FIGURES 17–19" captionStartId="4.[151,264,947,972]" captionTargetBox="[156,1430,393,916]" captionTargetId="figure@4.[152,1436,381,922]" captionTargetPageId="4" captionText="FIGURES 17–19. Megarthrus wayqecha sp. nov., male paratype. 17, aedeagus, ventral view (the side of the basal foramen); 18, aedeagus, lateral view; 19, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3661902" httpUri="https://zenodo.org/record/3661902/files/figure.png" pageId="5" pageNumber="586">Figs. 17–19</figureCitation>
.
</paragraph>
<paragraph id="8BFD36E81F64FD6CDEA7EAD3F986FCBD" blockId="6.[151,1437,153,575]" pageId="6" pageNumber="587">
Three male 
<typeStatus id="54F9884A1F64FD6CDF2BEAD3F8FAFCD9" box="[331,438,153,179]" pageId="6" pageNumber="587" type="paratype">paratypes</typeStatus>
and 
<specimenCount id="9D44FD611F64FD6CDF8CEAD3FB16FCD9" box="[492,602,153,179]" pageId="6" pageNumber="587" type="female">16 female</specimenCount>
<typeStatus id="54F9884A1F64FD6CDC00EAD3FB87FCD9" box="[608,715,153,179]" pageId="6" pageNumber="587" type="paratype">paratypes</typeStatus>
have the antennomere 9 yellow in apical half and brown in basal half.
</paragraph>
</subSubSection>
<subSubSection id="C35865631F64FD6CDEA7EAAAFB98FDC5" pageId="6" pageNumber="587" type="distribution">
<paragraph id="8BFD36E81F64FD6CDEA7EAAAFB98FDC5" blockId="6.[151,1437,153,575]" pageId="6" pageNumber="587">
<emphasis id="B936EAFA1F64FD6CDEA7EAAAFB02FC91" bold="true" box="[199,590,224,251]" pageId="6" pageNumber="587">Distribution and natural history</emphasis>
. The specimens fron wayquecha were collected in a cloud forest between 
<quantity id="4CBA9B0D1F64FD6CDEF7EB4FF9A7FD75" box="[151,235,261,287]" metricMagnitude="3" metricUnit="m" metricValue="2.931" pageId="6" pageNumber="587" unit="m" value="2931.0">2931 m</quantity>
and 
<quantity id="4CBA9B0D1F64FD6CDF40EB4EF83AFD75" box="[288,374,260,287]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="6" pageNumber="587" unit="m" value="2950.0">2950 m</quantity>
, from the biological station “Wayqecha”, a private area recently protected by Peruvian law with the aim to preserve the biodiversity of Peruvian Yungas (MINAM, 2016) (
<figureCitation id="13792A6D1F64FD6CDDB5EB63FD11FD29" box="[981,1117,297,323]" captionStart="FIGURES 20–21" captionStartId="4.[151,264,1676,1701]" captionTargetBox="[151,1436,1066,1651]" captionTargetId="figure@4.[151,1436,1066,1651]" captionTargetPageId="4" captionText="FIGURES 20–21. Habitat of Megarthrus wayqecha sp. nov. 20, panoramic view of the cloud forests of the Kosñipata valley, where the biological station Wayqecha is located. 21, sifting leaf litter on Trocha Canopy from the biological station Wayqecha." figureDoi="http://doi.org/10.5281/zenodo.3661904" httpUri="https://zenodo.org/record/3661904/files/figure.png" pageId="6" pageNumber="587">Figs. 20, 21</figureCitation>
). Wayqecha is placed 
<quantity id="4CBA9B0D1F64FD6CDB35EB63FCD0FD29" box="[1365,1436,297,323]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="587" unit="km" value="19.0">19 km</quantity>
SW of Pillahuata (
<geoCoordinate id="EE76502F1F64FD6CDF08EB07F88AFD0D" box="[360,454,333,359]" degrees="13" direction="south" minutes="08" orientation="latitude" pageId="6" pageNumber="587" precision="925" value="-13.133333">13°08’S</geoCoordinate>
, 
<geoCoordinate id="EE76502F1F64FD6CDFB3EB06FB71FD0D" box="[467,573,332,359]" degrees="71" direction="west" minutes="25" orientation="longitude" pageId="6" pageNumber="587" precision="925" value="-71.416664">71°25’W</geoCoordinate>
), the 
<typeStatus id="54F9884A1F64FD6CDC1CEB07FBE0FD0D" box="[636,684,333,359]" pageId="6" pageNumber="587">type</typeStatus>
locality of 
<taxonomicName id="4C424D6B1F64FD6CDD4EEB04FD27FD0D" authority="Cuccodoro, 2001" authorityName="Cuccodoro" authorityYear="2001" box="[814,1131,333,359]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="machu">
<emphasis id="B936EAFA1F64FD6CDD4EEB04FAD3FD0D" box="[814,927,333,359]" italics="true" pageId="6" pageNumber="587">M. machu</emphasis>
Cuccodoro, 2001
</taxonomicName>
. According to the information provided by the collector, the specimens were found in leaf litter with moss, ferns, and bamboo leaves alongside the roots and trunks of trees as observed in 
<figureCitation id="13792A6D1F64FD6CDC1DEBDFFB9CFDC5" box="[637,720,405,431]" captionStart="FIGURES 20–21" captionStartId="4.[151,264,1676,1701]" captionTargetBox="[151,1436,1066,1651]" captionTargetId="figure@4.[151,1436,1066,1651]" captionTargetPageId="4" captionText="FIGURES 20–21. Habitat of Megarthrus wayqecha sp. nov. 20, panoramic view of the cloud forests of the Kosñipata valley, where the biological station Wayqecha is located. 21, sifting leaf litter on Trocha Canopy from the biological station Wayqecha." figureDoi="http://doi.org/10.5281/zenodo.3661904" httpUri="https://zenodo.org/record/3661904/files/figure.png" pageId="6" pageNumber="587">Fig. 21</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35865631F64FD6CDEA7EBF3F8D5FE55" pageId="6" pageNumber="587" type="etymology">
<paragraph id="8BFD36E81F64FD6CDEA7EBF3FA8DFD9D" blockId="6.[151,1437,153,575]" pageId="6" pageNumber="587">
<emphasis id="B936EAFA1F64FD6CDEA7EBF3F802FDB9" bold="true" box="[199,334,441,467]" pageId="6" pageNumber="587">Etymology.</emphasis>
The specific name “Wayqecha” refers to the locality of collection of this new species. Wayqecha means ‘little brother’ in Quechua language. This is a noun in apposition.
</paragraph>
<paragraph id="8BFD36E81F64FD6CDEA7E84AF8D5FE55" blockId="6.[151,1437,153,575]" pageId="6" pageNumber="587">
<emphasis id="B936EAFA1F64FD6CDEA7E84AF80FFE71" bold="true" box="[199,323,512,539]" pageId="6" pageNumber="587">Barcoding</emphasis>
. The mitochondrial DNA sequences (COI) of three 
<typeStatus id="54F9884A1F64FD6CDDFDE84BFD44FE71" box="[925,1032,513,539]" pageId="6" pageNumber="587" type="paratype">paratypes</typeStatus>
were identical (
<tableCitation id="C6C003531F64FD6CDAA3E84BFC55FE71" box="[1219,1305,513,539]" captionStart="TABLE 1" captionStartId="6.[151,244,620,647]" captionTargetBox="[163,1334,698,843]" captionText="TABLE 1. Accession number in GenBank and barcoding voucher codes in BOLD Systems for COI sequences from three paratypes of Megarthrus wayqecha sp. nov." httpUri="http://table.plazi.org/id/DF3D66601F64FD6CDEF7E826FB8EFEC1" pageId="6" pageNumber="587" tableUuid="DF3D66601F64FD6CDEF7E826FB8EFEC1">Table 1</tableCitation>
). One COI sequence is as follows:
</paragraph>
</subSubSection>
</treatment>
</document>