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<mods:title id="3593B0EDFD67CB5E544F91C7208264DB">Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="8B26931C86C9C661B4050394434B6624">Khaustov, Alexander A.</mods:namePart>
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<mods:namePart id="623C16C48050DCB1F553C3E5B8A2B305">Frolov, Andrey V.</mods:namePart>
<mods:affiliation id="00874C678379F3B1D86AE9D0F433D7EA">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<mods:namePart id="B98194A5F8E78295B7566141E4F17249">Akhmetova, Lilia A.</mods:namePart>
<mods:affiliation id="7E7579FE746193EA8F476D4DA2D7A93E">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
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<treatment id="03EB878FFFBA744E81B43862FE22FB5F" ID-DOI="http://doi.org/10.5281/zenodo.13630075" ID-GBIF-Taxon="240057119" ID-Zenodo-Dep="13630075" LSID="urn:lsid:plazi:treatment:03EB878FFFBA744E81B43862FE22FB5F" httpUri="http://treatment.plazi.org/id/03EB878FFFBA744E81B43862FE22FB5F" lastPageId="29" lastPageNumber="366" pageId="28" pageNumber="365">
<subSubSection id="C3586512FFBA744F81B43862FBEBFABB" box="[151,1077,1337,1364]" pageId="28" pageNumber="365" type="nomenclature">
<paragraph id="8BFD3699FFBA744F81B43862FBEBFABB" blockId="28.[151,1077,1337,1364]" box="[151,1077,1337,1364]" pageId="28" pageNumber="365">
<heading id="D0B581F5FFBA744F81B43862FBEBFABB" bold="true" box="[151,1077,1337,1364]" fontSize="11" level="1" pageId="28" pageNumber="365" reason="1">
<emphasis id="B936EA8BFFBA744F81B43862FBEBFABB" bold="true" box="[151,1077,1337,1364]" pageId="28" pageNumber="365">
Key to world species of
<taxonomicName id="4C424D1AFFBA744F809C3861FDAFFABB" authorityName="Rack" authorityYear="1993" box="[447,625,1337,1364]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="B936EA8BFFBA744F809C3861FDAFFABB" bold="true" box="[447,625,1337,1364]" italics="true" pageId="28" pageNumber="365">Spatulaphorus</emphasis>
</taxonomicName>
(based on
<bibRefCitation id="EFD34B68FFBA744F83D43862FBF3FABB" author="Khaustov, A. A. &amp; Frolov, A. V." box="[759,1069,1338,1364]" pageId="28" pageNumber="365" pagination="277 - 292" refId="ref16197" refString="Khaustov, A. A. &amp; Frolov, A. V. (2018) New taxa of pygmephoroid mites (Acari: Pygmephoroidea: Pygmephoridae, Neopygmephoridae) phoretic on Enoplotrupes sharpi (Coleoptera: Geotrupidae) from Thailand. Zootaxa, 4442 (2), 277 - 292. https: // doi. org / 10.11646 / zootaxa. 4442.2.5" type="journal article" year="2018">Khaustov &amp; Frolov 2018</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3586512FFBA744E81B43827FE22FB5F" lastPageId="29" lastPageNumber="366" pageId="28" pageNumber="365" type="key">
<keyStep id="30B62E3CFFBA744F81B43827FA43FA5E" pageId="28" pageNumber="365">
<paragraph id="8BFD3699FFBA744F81B43827FA42FA7B" blockId="28.[151,1437,1407,1986]" box="[151,1436,1407,1429]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43827FA42FA7B" box="[151,1436,1407,1429]" pageId="28" pageNumber="365">1. Genu I with two setae................................................................................. 2</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBA744F81B438C5FA43FA5E" blockId="28.[151,1437,1407,1986]" box="[151,1437,1436,1458]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B438C5FA43FA5E" box="[151,1437,1436,1458]" pageId="28" pageNumber="365">
- Genu I with one seta................................................
<taxonomicName id="4C424D1AFFBA744F82DC38C5FAD3FA5E" authority="Khaustov, 2007 a" authorityName="Khaustov" authorityYear="2007" box="[1023,1293,1436,1458]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="365" phylum="Arthropoda" rank="species" species="copridis">
<emphasis id="B936EA8BFFBA744F82DC38C5FBB8FA5E" box="[1023,1126,1436,1458]" italics="true" pageId="28" pageNumber="365">S. copridis</emphasis>
<bibRefCitation id="EFD34B68FFBA744F854F38C4FAD3FA5E" author="Khaustov, A. A." box="[1132,1293,1436,1458]" pageId="28" pageNumber="365" pagination="155 - 157" refId="ref15930" refString="Khaustov, A. A. (2007 a) A new species of the genus Spatulaphorus (Acari: Heterostigmata: Pygmephoridae) associated with Copris lunaris (Coleoptera: Scarabaeidae) from Crimea. Acarina, 15, 155 - 157." type="journal article" year="2007">Khaustov, 2007a</bibRefCitation>
</taxonomicName>
(Crimea,
<collectingCountry id="F3557609FFBA744F844D38C4FA48FA5E" box="[1390,1430,1436,1457]" name="Iran" pageId="28" pageNumber="365">Iran</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBA744F81B438E2FA42FA02" pageId="28" pageNumber="365">
<paragraph id="8BFD3699FFBA744F81B438E2FA42FA20" blockId="28.[151,1437,1407,1986]" box="[151,1436,1466,1488]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B438E2FA42FA20" box="[151,1436,1466,1488]" pageId="28" pageNumber="365">
2. Alveoli of setae
<emphasis id="B936EA8BFFBA744F80A338E2FE7CFA20" box="[384,418,1466,1487]" italics="true" pageId="28" pageNumber="365">ps1</emphasis>
and
<emphasis id="B936EA8BFFBA744F80F338E2FE2FFA20" box="[464,497,1466,1487]" italics="true" pageId="28" pageNumber="365">ps2</emphasis>
contiguous; tarsus IV with six setae (
<emphasis id="B936EA8BFFBA744F827838E2FCA5FA20" box="[859,891,1466,1487]" italics="true" pageId="28" pageNumber="365">pl”</emphasis>
present).................................... 3
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBA744F81B4388FFA42FA02" blockId="28.[151,1437,1407,1986]" box="[151,1436,1495,1517]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B4388FFA42FA02" box="[151,1436,1495,1517]" pageId="28" pageNumber="365">
- Alveoli of setae
<emphasis id="B936EA8BFFBA744F80A3388FFE7CFA03" box="[384,418,1495,1516]" italics="true" pageId="28" pageNumber="365">ps1</emphasis>
and
<emphasis id="B936EA8BFFBA744F80F3388FFE2FFA03" box="[464,497,1495,1516]" italics="true" pageId="28" pageNumber="365">ps2</emphasis>
clearly separated; tarsus IV with five setae (
<emphasis id="B936EA8BFFBA744F82BE388FFC62FA03" box="[925,956,1495,1516]" italics="true" pageId="28" pageNumber="365">pl”</emphasis>
absent).......
<taxonomicName id="4C424D1AFFBA744F8549388FFB0AFA03" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[1130,1236,1495,1516]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="365" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov.">
<emphasis id="B936EA8BFFBA744F8549388FFB0AFA03" box="[1130,1236,1495,1516]" italics="true" pageId="28" pageNumber="365">S. reductus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBA744F85F9388FFAF8FA03" bold="true" box="[1242,1318,1495,1516]" pageId="28" pageNumber="365">
<taxonomicNameLabel id="A20557F0FFBA744F85F9388FFAF8FA03" box="[1242,1318,1495,1516]" pageId="28" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F3557609FFBA744F8410388FFA48FA02" box="[1331,1430,1495,1517]" name="Argentina" pageId="28" pageNumber="365">Argentina</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBA744F81B438ACFA42F9C8" pageId="28" pageNumber="365">
<paragraph id="8BFD3699FFBA744F81B438ACFA42F9E6" blockId="28.[151,1437,1407,1986]" box="[151,1436,1524,1546]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B438ACFA42F9E6" box="[151,1436,1524,1546]" pageId="28" pageNumber="365">
3. Seta
<emphasis id="B936EA8BFFBA744F803238ACFEEFF9E6" box="[273,305,1524,1545]" italics="true" pageId="28" pageNumber="365">pl”</emphasis>
of tarsus III spiniform.......................................................................... 4
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBA744F81B43B4AFA42F9C8" blockId="28.[151,1437,1407,1986]" box="[151,1436,1554,1576]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43B4AFA42F9C8" box="[151,1436,1554,1576]" pageId="28" pageNumber="365">
- setae
<emphasis id="B936EA8BFFBA744F803B3B4AFEE6F9C8" box="[280,312,1554,1575]" italics="true" pageId="28" pageNumber="365">pl”</emphasis>
of tarsus III not modified...................................................................... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBA744F81B43B77FA42F98E" pageId="28" pageNumber="365">
<paragraph id="8BFD3699FFBA744F81B43B77FA42F9AA" blockId="28.[151,1437,1407,1986]" box="[151,1436,1583,1605]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43B77FA42F9AA" box="[151,1436,1583,1605]" pageId="28" pageNumber="365">
4. Seta
<emphasis id="B936EA8BFFBA744F80323B77FEF4F9AB" box="[273,298,1583,1604]" italics="true" pageId="28" pageNumber="365">tc</emphasis>
of tarsus III spiniform........................................................................... 5
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBA744F81B43B15FA42F98E" blockId="28.[151,1437,1407,1986]" box="[151,1436,1612,1634]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43B15FA42F98E" box="[151,1436,1612,1634]" pageId="28" pageNumber="365">
- Seta
<emphasis id="B936EA8BFFBA744F80323B15FEF4F98D" box="[273,298,1613,1634]" italics="true" pageId="28" pageNumber="365">tc</emphasis>
of tarsus III not modified........................................................................ 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBA744F81B43B31FA42F973" pageId="28" pageNumber="365">
<paragraph id="8BFD3699FFBA744F81B43B31FA42F96F" blockId="28.[151,1437,1407,1986]" box="[151,1436,1641,1664]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43B31FA42F96F" box="[151,1436,1641,1664]" pageId="28" pageNumber="365">5. Claws on tarsi II and III distinctly thickened basally; tibiotarsus I thick, distinctly wider than genu I................... 6</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBA744F81B43BDFFA42F973" blockId="28.[151,1437,1407,1986]" box="[151,1436,1671,1693]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43BDFFA42F973" box="[151,1436,1671,1693]" pageId="28" pageNumber="365">- Claws on tarsi II and III not thickened basally; tibiotarsus I cylindrical, as wide as genu I............................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBA744F81B43BFCFA43F91A" pageId="28" pageNumber="365">
<paragraph id="8BFD3699FFBA744F81B43BFCFA42F937" blockId="28.[151,1437,1407,1986]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43BFCFA42F937" pageId="28" pageNumber="365">
6. Setae
<emphasis id="B936EA8BFFBA744F803F3BFCFEEDF956" box="[284,307,1700,1721]" italics="true" pageId="28" pageNumber="365">c1</emphasis>
and
<emphasis id="B936EA8BFFBA744F80423BFCFEA6F956" box="[353,376,1700,1721]" italics="true" pageId="28" pageNumber="365">c2</emphasis>
subequal in length; setae
<emphasis id="B936EA8BFFBA744F83533BFCFD57F956" box="[624,649,1700,1721]" italics="true" pageId="28" pageNumber="365">h1</emphasis>
longer than distance between their bases......................................................................................
<taxonomicName id="4C424D1AFFBA744F82583B9AFB6DF938" authority="(Mahunka, 1980)" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" baseAuthorityName="Mahunka" baseAuthorityYear="1980" box="[891,1203,1730,1752]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="365" phylum="Arthropoda" rank="species" species="inbricatipes" status="comb. nov.">
<emphasis id="B936EA8BFFBA744F82583B9AFBD8F938" box="[891,1030,1730,1751]" italics="true" pageId="28" pageNumber="365">S. inbricatipes</emphasis>
(
<bibRefCitation id="EFD34B68FFBA744F85303B9AFB72F938" author="Mahunka, S." box="[1043,1196,1730,1752]" pageId="28" pageNumber="365" pagination="377 - 399" refId="ref16707" refString="Mahunka, S. (1980) Data to the knowledge of mites preserved in the &quot; Berlese collection &quot; (Acari: Tarsonemida, Oribatida). I. Acta Zoologica Academiae Scientiarum Hungaricae, 26, 377 - 399." type="journal article" year="1980">Mahunka, 1980</bibRefCitation>
)
</taxonomicName>
<emphasis id="B936EA8BFFBA744F859A3B9AFAF8F938" bold="true" box="[1209,1318,1730,1751]" pageId="28" pageNumber="365">
<taxonomicNameLabel id="A20557F0FFBA744F859A3B9AFAF8F938" box="[1209,1318,1730,1751]" pageId="28" pageNumber="365" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F3557609FFBA744F84103B9AFA48F937" box="[1331,1430,1730,1752]" name="Argentina" pageId="28" pageNumber="365">Argentina</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBA744F81B43B87FA43F91A" blockId="28.[151,1437,1407,1986]" box="[151,1437,1759,1781]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43B87FA43F91A" box="[151,1437,1759,1781]" pageId="28" pageNumber="365">
- Setae
<emphasis id="B936EA8BFFBA744F803F3B87FEEDF91B" box="[284,307,1759,1780]" italics="true" pageId="28" pageNumber="365">c1</emphasis>
distinctly shorter than
<emphasis id="B936EA8BFFBA744F83333B87FDF8F91B" box="[528,550,1759,1780]" italics="true" pageId="28" pageNumber="365">c2</emphasis>
; setae
<emphasis id="B936EA8BFFBA744F834A3B87FD5FF91B" box="[617,641,1759,1780]" italics="true" pageId="28" pageNumber="365">h1</emphasis>
distinctly shorter than distance between their bases
<taxonomicName id="4C424D1AFFBA744F85A13B87FB27F91A" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[1154,1273,1759,1781]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="365" phylum="Arthropoda" rank="species" species="spinisetus" status="sp. nov.">
<emphasis id="B936EA8BFFBA744F85A13B87FB27F91A" box="[1154,1273,1759,1781]" italics="true" pageId="28" pageNumber="365">S. spinisetus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBA744F85DC3B87FA95F91B" bold="true" box="[1279,1355,1759,1780]" pageId="28" pageNumber="365">
<taxonomicNameLabel id="A20557F0FFBA744F85DC3B87FA95F91B" box="[1279,1355,1759,1780]" pageId="28" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F3557609FFBA744F847B3B87FA4BF91A" box="[1368,1429,1759,1781]" name="Brazil" pageId="28" pageNumber="365">Brazil</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBA744F81B43BA5FA43F8C0" pageId="28" pageNumber="365">
<paragraph id="8BFD3699FFBA744F81B43BA5FA42F8FE" blockId="28.[151,1437,1407,1986]" box="[151,1436,1788,1810]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43BA5FA42F8FE" box="[151,1436,1788,1810]" pageId="28" pageNumber="365">
7. Setae
<emphasis id="B936EA8BFFBA744F803F3BA4FEE2F8FE" box="[284,316,1788,1809]" italics="true" pageId="28" pageNumber="365">sc2</emphasis>
not modified, setiform......................................................................... 8
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBA744F81B43A42FA43F8C0" blockId="28.[151,1437,1407,1986]" box="[151,1437,1817,1840]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43A42FA43F8C0" box="[151,1437,1817,1840]" pageId="28" pageNumber="365">
- Setae
<emphasis id="B936EA8BFFBA744F803F3A42FEE2F8C0" box="[284,316,1818,1839]" italics="true" pageId="28" pageNumber="365">sc2</emphasis>
foliate.............................................
<taxonomicName id="4C424D1AFFBA744F82843A42FB6AF8C0" authority="Khaustov, 2005" authorityName="Khaustov" authorityYear="2005" box="[935,1204,1817,1840]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="365" phylum="Arthropoda" rank="species" species="vladimiri">
<emphasis id="B936EA8BFFBA744F82843A42FBC9F8C0" box="[935,1047,1818,1839]" italics="true" pageId="28" pageNumber="365">S. vladimiri</emphasis>
<bibRefCitation id="EFD34B68FFBA744F853E3A42FB6AF8C0" author="Khaustov, A. A." box="[1053,1204,1817,1840]" pageId="28" pageNumber="365" pagination="35 - 42" refId="ref15889" refString="Khaustov, A. A. (2005) Three new species of mite family Pygmephoridae (Acari: Heterostigmata) associated with scarab beetles (Coleoptera: Scarabaeidae) from Ukraine. Acarina, 13, 35 - 42." type="journal article" year="2005">Khaustov, 2005</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBA744F85E33A42FACFF8DF" box="[1216,1297,1818,1840]" name="Ukraine" pageId="28" pageNumber="365">Ukraine</collectingCountry>
,
<collectingRegion id="4986F87BFFBA744F84383A42FABAF8DF" box="[1307,1380,1818,1840]" country="Ukraine" name="Crimea" pageId="28" pageNumber="365">Crimea</collectingRegion>
,
<collectingCountry id="F3557609FFBA744F844D3A42FA48F8C0" box="[1390,1430,1818,1839]" name="Iran" pageId="28" pageNumber="365">Iran</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBA744F81B43A6FFA42F885" pageId="28" pageNumber="365">
<paragraph id="8BFD3699FFBA744F81B43A6FFA42F8A2" blockId="28.[151,1437,1407,1986]" box="[151,1436,1847,1869]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43A6FFA42F8A2" box="[151,1436,1847,1869]" pageId="28" pageNumber="365">
8. Setae
<emphasis id="B936EA8BFFBA744F803F3A6FFEF6F8A3" box="[284,296,1847,1868]" italics="true" pageId="28" pageNumber="365">d</emphasis>
,
<emphasis id="B936EA8BFFBA744F80173A6FFEE5F8A3" box="[308,315,1847,1868]" italics="true" pageId="28" pageNumber="365">f</emphasis>
, and
<emphasis id="B936EA8BFFBA744F804C3A6FFE56F8A3" box="[367,392,1847,1868]" italics="true" pageId="28" pageNumber="365">h1</emphasis>
foliate, setae
<emphasis id="B936EA8BFFBA744F832F3A6FFDF0F8A3" box="[524,558,1847,1868]" italics="true" pageId="28" pageNumber="365">ps3</emphasis>
slightly longer than
<emphasis id="B936EA8BFFBA744F83D73A6FFCF4F8A3" box="[756,810,1847,1868]" italics="true" pageId="28" pageNumber="365">ps1-2</emphasis>
............
<taxonomicName id="4C424D1AFFBA744F82E83A6FFAE7F8A3" authority="Khaustov and Trach, 2012" authorityName="Khaustov and Trach" authorityYear="2012" box="[971,1337,1847,1869]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="365" phylum="Arthropoda" rank="species" species="venustus">
<emphasis id="B936EA8BFFBA744F82E83A6FFC09F8A3" box="[971,983,1847,1868]" italics="true" pageId="28" pageNumber="365">S</emphasis>
.
<emphasis id="B936EA8BFFBA744F82C03A60FBE8F8A2" box="[995,1078,1848,1869]" italics="true" pageId="28" pageNumber="365">venustus</emphasis>
<bibRefCitation id="EFD34B68FFBA744F851F3A6FFAE7F8A3" author="Khaustov, A. A. &amp; Trach, V. A." box="[1084,1337,1847,1869]" pageId="28" pageNumber="365" pagination="480 - 485" refId="ref16479" refString="Khaustov, A. A. &amp; Trach, V. A. (2012) A new species and new records of mites of the genus Spatulaphorus Rack (Acari: Heterostigmata: Pygmephoridae) from Ukraine. International Journal of Acarology, 38, 480 - 485. https: // doi. org / 10.1080 / 01647954.2012.677475" type="journal article" year="2012">Khaustov and Trach, 2012</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBA744F84663A6FFA4BF8A2" box="[1349,1429,1847,1869]" name="Ukraine" pageId="28" pageNumber="365">Ukraine</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBA744F81B43A0DFA42F885" blockId="28.[151,1437,1407,1986]" box="[151,1436,1876,1898]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43A0DFA42F885" box="[151,1436,1876,1898]" pageId="28" pageNumber="365">
- Setae
<emphasis id="B936EA8BFFBA744F80383A0CFEF9F886" box="[283,295,1876,1897]" italics="true" pageId="28" pageNumber="365">d</emphasis>
,
<emphasis id="B936EA8BFFBA744F80103A0CFEE4F886" box="[307,314,1876,1897]" italics="true" pageId="28" pageNumber="365">f</emphasis>
, and
<emphasis id="B936EA8BFFBA744F804D3A0CFE5BF886" box="[366,389,1876,1897]" italics="true" pageId="28" pageNumber="365">h1</emphasis>
not foliate, setae
<emphasis id="B936EA8BFFBA744F830D3A0CFD91F886" box="[558,591,1876,1897]" italics="true" pageId="28" pageNumber="365">ps3</emphasis>
at least twice longer than
<emphasis id="B936EA8BFFBA744F826B3A0CFCA3F886" box="[840,893,1876,1897]" italics="true" pageId="28" pageNumber="365">ps1-2</emphasis>
...
<taxonomicName id="4C424D1AFFBA744F828A3A0DFAD3F886" authority="Khaustov and Trach, 2018" authorityName="Khaustov and Trach" authorityYear="2018" box="[937,1293,1876,1898]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="365" phylum="Arthropoda" rank="species" species="altaicus">
<emphasis id="B936EA8BFFBA744F828A3A0DFBD5F886" box="[937,1035,1876,1898]" italics="true" pageId="28" pageNumber="365">S. altaicus</emphasis>
<bibRefCitation id="EFD34B68FFBA744F85313A0CFAD3F886" author="Khaustov, A. A. &amp; Trach, V. A." box="[1042,1293,1876,1898]" pageId="28" pageNumber="365" pagination="123 - 131" refId="ref16539" refString="Khaustov, A. A. &amp; Trach, V. A. (2018) New species of Spatulaphorus (Acari: Heterostigmata: Pygmephoridae) associated with Geotrupes baicalicus (Coleoptera: Geotrupidae) from Altai Republic, Russia. Systematic &amp; Applied Acarology, 23, 123 - 131. https: // doi. org / 10.11158 / saa. 23.1.10" type="journal article" year="2018">Khaustov and Trach, 2018</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBA744F843A3A0CFA83F885" box="[1305,1373,1876,1898]" name="Russia" pageId="28" pageNumber="365">Russia</collectingCountry>
:
<collectingRegion id="4986F87BFFBA744F84463A0CFA48F885" box="[1381,1430,1876,1898]" country="Russia" name="Altay" pageId="28" pageNumber="365">Altai</collectingRegion>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBA744F81B43A2AFA42F82D" pageId="28" pageNumber="365">
<paragraph id="8BFD3699FFBA744F81B43A2AFA42F868" blockId="28.[151,1437,1407,1986]" box="[151,1436,1906,1928]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43A2AFA42F868" box="[151,1436,1906,1928]" pageId="28" pageNumber="365">9. Claws on tarsi II and III distinctly thickened basally; tibiotarsus I thick, distinctly wider than genu I.................. 10</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBA744F81B43AD7FA42F82D" blockId="28.[151,1437,1407,1986]" pageId="28" pageNumber="365">
<keyLead id="30B395ACFFBA744F81B43AD7FA42F82D" pageId="28" pageNumber="365">
- Claws on tarsi II and III not thickened basally; tibiotarsus I cylindrical, as wide as genu I..............................................................................
<taxonomicName id="4C424D1AFFBA744F821C3AF5FAD3F82E" authority="Rahiminejad and Hajiqanbar, 2011" authorityName="Rahiminejad and Hajiqanbar" authorityYear="2011" box="[831,1293,1964,1986]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="28" pageNumber="365" phylum="Arthropoda" rank="species" species="gorganica">
<emphasis id="B936EA8BFFBA744F821C3AF5FC67F82D" box="[831,953,1965,1986]" italics="true" pageId="28" pageNumber="365">S. gorganica</emphasis>
<bibRefCitation id="EFD34B68FFBA744F829C3AF4FAD3F82E" author="Rahiminejad, V. &amp; Hajiqanbar, H. &amp; Fathipour, Y." box="[959,1293,1964,1986]" pageId="28" pageNumber="365" pagination="547 - 551" refId="ref16749" refString="Rahiminejad, V., Hajiqanbar, H. &amp; Fathipour, Y. (2011) A new species of the genus Spatulaphorus (Acari: Heterostigmatina: Pygmephoridae) phoretic on Geotrupes spiniger (Coleoptera: Geotrupidae) from Iran. Annales Zoologici, 61, 547 - 551. https: // doi. org / 10.3161 / 000345411 X 603391" type="journal article" year="2011">Rahiminejad and Hajiqanbar, 2011</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBA744F84393AF4FA9AF82E" box="[1306,1348,1964,1985]" name="Iran" pageId="28" pageNumber="365">Iran</collectingCountry>
, Crimea)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43DC0FA43FEEA" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43DC0FA42FF25" blockId="29.[151,1437,151,1200]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43DC0FA42FF25" pageId="29" pageNumber="366">
10. Alveoli of setae
<emphasis id="B936EA8BFFBB744E805D3DC0FE57FF42" box="[382,393,152,173]" italics="true" pageId="29" pageNumber="366">e</emphasis>
and
<emphasis id="B936EA8BFFBB744E80953DC0FE63FF42" box="[438,445,152,173]" italics="true" pageId="29" pageNumber="366">f</emphasis>
as well as
<emphasis id="B936EA8BFFBB744E83073DCFFDE2FF43" box="[548,572,151,172]" italics="true" pageId="29" pageNumber="366">h1</emphasis>
and
<emphasis id="B936EA8BFFBB744E834A3DCFFD5FFF43" box="[617,641,151,172]" italics="true" pageId="29" pageNumber="366">h2</emphasis>
clearly separated; setae
<emphasis id="B936EA8BFFBB744E824A3DC0FC41FF42" box="[873,927,152,173]" italics="true" pageId="29" pageNumber="366">ps1-2</emphasis>
smooth; seta
<emphasis id="B936EA8BFFBB744E85013DCFFB9FFF43" box="[1058,1089,151,172]" italics="true" pageId="29" pageNumber="366">pl”</emphasis>
of tarsus IV about twice longer than maximum width of tarsus IV........................................................................... 11
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43D8BFA43FEEA" blockId="29.[151,1437,151,1200]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43D8BFA43FEEA" pageId="29" pageNumber="366">
- Alveoli of setae
<emphasis id="B936EA8BFFBB744E80A13D8BFE53FF07" box="[386,397,211,232]" italics="true" pageId="29" pageNumber="366">e</emphasis>
and
<emphasis id="B936EA8BFFBB744E809F3D8AFE1DFF08" box="[444,451,210,231]" italics="true" pageId="29" pageNumber="366">f</emphasis>
as well as
<emphasis id="B936EA8BFFBB744E830C3D8AFD99FF08" box="[559,583,210,231]" italics="true" pageId="29" pageNumber="366">h1</emphasis>
and
<emphasis id="B936EA8BFFBB744E83543D8AFD51FF08" box="[631,655,210,231]" italics="true" pageId="29" pageNumber="366">h2</emphasis>
contiguous; setae
<emphasis id="B936EA8BFFBB744E82613D8AFCA6FF08" box="[834,888,210,231]" italics="true" pageId="29" pageNumber="366">ps1-2</emphasis>
barbed; seta
<emphasis id="B936EA8BFFBB744E82DA3D8AFBC7FF08" box="[1017,1049,210,231]" italics="true" pageId="29" pageNumber="366">pl”</emphasis>
of tarsus IV about as long as maximum width of tarsus IV.................................................
<taxonomicName id="4C424D1AFFBB744E82D03DA8FAE3FEEA" authority="Trejo-Palacios, 2023" authorityName="Trejo-Palacios" authorityYear="2023" box="[1011,1341,239,261]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B936EA8BFFBB744E82D03DA8FBAEFEEA" box="[1011,1136,240,261]" italics="true" pageId="29" pageNumber="366">S. mexicanus</emphasis>
<bibRefCitation id="EFD34B68FFBB744E85553DB7FAE3FEEA" author="Trejo-Palacios, S. J. &amp; Toledo-Hernandez, V. H. &amp; Paredes-Leon, R. &amp; Corona-Lopez, A. M." box="[1142,1341,239,261]" pageId="29" pageNumber="366" pagination="77 - 87" refId="ref16938" refString="Trejo-Palacios, S. J., Toledo-Hernandez, V. H., Paredes-Leon, R. &amp; Corona-Lopez, A. M. (2023) A new species and new records of Pygmephoridae (Acari: Heterostigmata) phoretic on Coleoptera from Mexico. Systematic &amp; Applied Acarology, 28 (1), 77 - 87. https: // doi. org / 10.11158 / saa. 28.1.8" type="journal article" year="2023">Trejo-Palacios, 2023</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBB744E846A3DB7FA4BFEEA" box="[1353,1429,239,261]" name="Mexico" pageId="29" pageNumber="366">Mexico</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43C55FA43FED0" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43C55FA43FECD" blockId="29.[151,1437,151,1200]" box="[151,1437,269,291]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43C55FA43FECD" box="[151,1437,269,291]" pageId="29" pageNumber="366">
11. Dorsal idiosomal setae short; setae
<emphasis id="B936EA8BFFBB744E83163C55FD92FECD" box="[565,588,269,290]" italics="true" pageId="29" pageNumber="366">c1</emphasis>
,
<emphasis id="B936EA8BFFBB744E837B3C55FD81FECD" box="[600,607,269,290]" italics="true" pageId="29" pageNumber="366">f</emphasis>
, and
<emphasis id="B936EA8BFFBB744E83B03C55FD75FECD" box="[659,683,269,290]" italics="true" pageId="29" pageNumber="366">h1</emphasis>
shorter than distances between their bases...
<taxonomicName id="4C424D1AFFBB744E85553C55FADAFECD" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[1142,1284,269,290]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="brevisetosus" status="sp. nov.">
<emphasis id="B936EA8BFFBB744E85553C55FADAFECD" box="[1142,1284,269,290]" italics="true" pageId="29" pageNumber="366">S. brevisetosus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBB744E84293C55FA88FECD" bold="true" box="[1290,1366,269,290]" pageId="29" pageNumber="366">
<taxonomicNameLabel id="A20557F0FFBB744E84293C55FA88FECD" box="[1290,1366,269,290]" pageId="29" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F3557609FFBB744E84403C55FA48FECD" box="[1379,1430,269,290]" name="United States of America" pageId="29" pageNumber="366">USA</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43C73FA43FED0" blockId="29.[151,1437,151,1200]" box="[151,1437,298,320]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43C73FA43FED0" box="[151,1437,298,320]" pageId="29" pageNumber="366">
- Dorsal idiosomal setae long; setae
<emphasis id="B936EA8BFFBB744E83133C72FD99FED0" box="[560,583,298,319]" italics="true" pageId="29" pageNumber="366">c1</emphasis>
,
<emphasis id="B936EA8BFFBB744E83713C72FD87FED0" box="[594,601,298,319]" italics="true" pageId="29" pageNumber="366">f</emphasis>
, and
<emphasis id="B936EA8BFFBB744E83AD3C72FD78FED0" box="[654,678,298,319]" italics="true" pageId="29" pageNumber="366">h1</emphasis>
longer than distances between their bases....
<taxonomicName id="4C424D1AFFBB744E85563C73FADAFED0" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[1141,1284,298,320]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="longisetosus" status="sp. nov.">
<emphasis id="B936EA8BFFBB744E85563C73FADAFED0" box="[1141,1284,298,320]" italics="true" pageId="29" pageNumber="366">S. longisetosus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBB744E84293C72FA88FED0" bold="true" box="[1290,1366,298,319]" pageId="29" pageNumber="366">
<taxonomicNameLabel id="A20557F0FFBB744E84293C72FA88FED0" box="[1290,1366,298,319]" pageId="29" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F3557609FFBB744E84403C72FA48FED0" box="[1379,1430,298,319]" name="United States of America" pageId="29" pageNumber="366">USA</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43C10FA42FE95" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43C10FA42FEB2" blockId="29.[151,1437,151,1200]" box="[151,1436,327,349]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43C10FA42FEB2" box="[151,1436,327,349]" pageId="29" pageNumber="366">
12. Seta
<emphasis id="B936EA8BFFBB744E80323C10FEF4FEB2" box="[273,298,328,349]" italics="true" pageId="29" pageNumber="366">tc</emphasis>
of tarsus III spiniform.......................................................................... 13
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43C3DFA42FE95" blockId="29.[151,1437,151,1200]" box="[151,1436,357,379]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43C3DFA42FE95" box="[151,1436,357,379]" pageId="29" pageNumber="366">
- Seta
<emphasis id="B936EA8BFFBB744E80323C3DFEF4FE95" box="[273,298,357,378]" italics="true" pageId="29" pageNumber="366">tc</emphasis>
of tarsus III not modified....................................................................... 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43CDAFA42FE3C" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43CDAFA43FE78" blockId="29.[151,1437,151,1200]" box="[151,1437,386,408]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43CDAFA43FE78" box="[151,1437,386,408]" pageId="29" pageNumber="366">
13. Tibiotarsus I with a small, almost straight claw, and no counterpart........
<taxonomicName id="4C424D1AFFBB744E82F43CDBFAF6FE78" authority="Dastych and Rack, 1993" authorityName="Dastych and Rack" authorityYear="1993" box="[983,1320,386,408]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B936EA8BFFBB744E82F43CDBFBE9FE78" box="[983,1079,386,408]" italics="true" pageId="29" pageNumber="366">S. foliatus</emphasis>
<bibRefCitation id="EFD34B68FFBB744E851E3CDAFAF6FE78" author="Dastych, H. &amp; Rack, G." box="[1085,1320,386,408]" pageId="29" pageNumber="366" pagination="265 - 84" refId="ref15517" refString="Dastych, H. &amp; Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article" year="1993">Dastych and Rack, 1993</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBB744E84173CDAFA48FE78" box="[1332,1430,386,407]" name="Botswana" pageId="29" pageNumber="366">Botswana</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43CF8FA42FE3C" blockId="29.[151,1437,151,1200]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43CF8FA42FE3C" pageId="29" pageNumber="366">
- Tibiotarsus I with large, hook-like claw and a distinct counterpart opposite to the claw base............................................................................................
<taxonomicName id="4C424D1AFFBB744E82D83CE5FAEBFE3D" authority="Dastych and Rack, 1993" authorityName="Dastych and Rack" authorityYear="1993" box="[1019,1333,445,467]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="B936EA8BFFBB744E82D83CE5FB9AFE3D" box="[1019,1092,445,466]" italics="true" pageId="29" pageNumber="366">S. langi</emphasis>
<bibRefCitation id="EFD34B68FFBB744E85693CE5FAEBFE3D" author="Dastych, H. &amp; Rack, G." box="[1098,1333,445,467]" pageId="29" pageNumber="366" pagination="265 - 84" refId="ref15517" refString="Dastych, H. &amp; Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article" year="1993">Dastych and Rack, 1993</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBB744E84623CE5FA4BFE3C" box="[1345,1429,445,467]" name="Vietnam" pageId="29" pageNumber="366">Vietnam</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43C82FA42FDE2" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43C82FA42FE1F" blockId="29.[151,1437,151,1200]" box="[151,1436,474,496]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43C82FA42FE1F" box="[151,1436,474,496]" pageId="29" pageNumber="366">
14. Seta
<emphasis id="B936EA8BFFBB744E80323C83FEF4FE1F" box="[273,298,475,496]" italics="true" pageId="29" pageNumber="366">tc</emphasis>
of tarsus II spiniform and curved................................................................. 15
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43CA0FA42FDE2" blockId="29.[151,1437,151,1200]" box="[151,1436,503,525]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43CA0FA42FDE2" box="[151,1436,503,525]" pageId="29" pageNumber="366">
- Seta
<emphasis id="B936EA8BFFBB744E80323CA0FEF4FDE2" box="[273,298,504,525]" italics="true" pageId="29" pageNumber="366">tc</emphasis>
of tarsus II not modified........................................................................ 17
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43F4DFA42FDA7" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43F4DFA42FDC4" blockId="29.[151,1437,151,1200]" box="[151,1436,533,555]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43F4DFA42FDC4" box="[151,1436,533,555]" pageId="29" pageNumber="366">
15. Setae
<emphasis id="B936EA8BFFBB744E803F3F4EFEF9FDC4" box="[284,295,534,555]" italics="true" pageId="29" pageNumber="366">e</emphasis>
situated distinctly posteriad setae
<emphasis id="B936EA8BFFBB744E83533F4DFDA9FDC5" box="[624,631,533,554]" italics="true" pageId="29" pageNumber="366">f</emphasis>
................................................................ 16
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43F6BFA42FDA7" blockId="29.[151,1437,151,1200]" box="[151,1436,562,584]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43F6BFA42FDA7" box="[151,1436,562,584]" pageId="29" pageNumber="366">
- Setae
<emphasis id="B936EA8BFFBB744E803F3F6BFEF9FDA7" box="[284,295,563,584]" italics="true" pageId="29" pageNumber="366">e</emphasis>
situated slightly anteriad setae
<emphasis id="B936EA8BFFBB744E83773F6AFD85FDA8" box="[596,603,562,583]" italics="true" pageId="29" pageNumber="366">f</emphasis>
.....................................
<taxonomicName id="4C424D1AFFBB744E85093F6BFB0FFDA8" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[1066,1233,562,584]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="venezuelaensis" status="sp. nov.">
<emphasis id="B936EA8BFFBB744E85093F6BFB0FFDA8" box="[1066,1233,562,584]" italics="true" pageId="29" pageNumber="366">S. venezuelaensis</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBB744E85F43F6AFAFDFDA8" bold="true" box="[1239,1315,562,583]" pageId="29" pageNumber="366">
<taxonomicNameLabel id="A20557F0FFBB744E85F43F6AFAFDFDA8" box="[1239,1315,562,583]" pageId="29" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F3557609FFBB744E84133F6AFA48FDA7" box="[1328,1430,562,584]" name="Venezuela" pageId="29" pageNumber="366">Venezuela</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43F17FA42FD6C" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43F17FA42FD8A" blockId="29.[151,1437,151,1200]" box="[151,1436,591,613]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43F17FA42FD8A" box="[151,1436,591,613]" pageId="29" pageNumber="366">
16. Idiosomal tergites with big puncta (
<figureCitation id="13792A1CFFBB744E83153F17FD5CFD8A" box="[566,642,591,613]" captionStart="FIGURE 7" captionStartId="7.[151,250,1947,1971]" captionTargetBox="[151,1435,352,1924]" captionTargetId="figure-97@7.[151,1435,345,1924]" captionTargetPageId="7" captionText="FIGURE 7. Spatulaphorus longisetosus sp. nov., female: A—left leg III, dorsal aspect; B—left leg IV, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618625" httpUri="https://zenodo.org/record/13618625/files/figure.png" pageId="29" pageNumber="366">Fig. 7A</figureCitation>
)....................
<taxonomicName id="4C424D1AFFBB744E82A93F08FB26FD8A" authority="Khaustov and Frolov, 2021" authorityName="Khaustov and Frolov" authorityYear="2021" box="[906,1272,591,613]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="porosus">
<emphasis id="B936EA8BFFBB744E82A93F08FC33FD8A" box="[906,1005,592,613]" italics="true" pageId="29" pageNumber="366">S. porosus</emphasis>
<bibRefCitation id="EFD34B68FFBB744E82D03F17FB26FD8A" author="Khaustov, A. A. &amp; Frolov, A. V." box="[1011,1272,591,613]" pageId="29" pageNumber="366" pagination="333 - 345" refId="ref16337" refString="Khaustov, A. A. &amp; Frolov, A. V. (2021) Two new species and a new record of Spatulaphorus (Acari &quot; Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae) from French Guiana. Acarina, 29 (2), 333 - 345. https: // doi. org / 10.21684 / 0132 - 8077 - 2021 - 29 - 2 - 233 - 245" type="journal article" year="2021">Khaustov and Frolov, 2021</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBB744E84263F17FA48FD8A" box="[1285,1430,591,613]" name="French Guiana" pageId="29" pageNumber="366">French Guiana</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43F35FA42FD6C" blockId="29.[151,1437,151,1200]" box="[151,1436,621,643]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43F35FA42FD6C" box="[151,1436,621,643]" pageId="29" pageNumber="366">
- Idiosomal tergites only with tiny puncta (
<figureCitation id="13792A1CFFBB744E834D3F35FD66FD6D" box="[622,696,621,643]" captionStart="FIGURE 1" captionStartId="1.[151,250,1956,1980]" captionTargetBox="[151,1435,639,1933]" captionTargetId="figure-207@1.[151,1435,633,1933]" captionTargetPageId="1" captionText="FIGURE 1. Spatulaphorus brevisetosus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." figureDoi="http://doi.org/10.5281/zenodo.13618613" httpUri="https://zenodo.org/record/13618613/files/figure.png" pageId="29" pageNumber="366">Fig. 1A</figureCitation>
)................
<taxonomicName id="4C424D1AFFBB744E82B63F35FB26FD6D" authority="Khaustov and Frolov, 2021" authorityName="Khaustov and Frolov" authorityYear="2021" box="[917,1272,621,643]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="dignus">
<emphasis id="B936EA8BFFBB744E82B63F35FC33FD6D" box="[917,1005,621,642]" italics="true" pageId="29" pageNumber="366">S. dignus</emphasis>
<bibRefCitation id="EFD34B68FFBB744E82D03F35FB26FD6D" author="Khaustov, A. A. &amp; Frolov, A. V." box="[1011,1272,621,643]" pageId="29" pageNumber="366" pagination="333 - 345" refId="ref16337" refString="Khaustov, A. A. &amp; Frolov, A. V. (2021) Two new species and a new record of Spatulaphorus (Acari &quot; Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae) from French Guiana. Acarina, 29 (2), 333 - 345. https: // doi. org / 10.21684 / 0132 - 8077 - 2021 - 29 - 2 - 233 - 245" type="journal article" year="2021">Khaustov and Frolov, 2021</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBB744E84263F35FA48FD6C" box="[1285,1430,621,643]" name="French Guiana" pageId="29" pageNumber="366">French Guiana</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43FD2FA42FD52" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43FD2FA42FD70" blockId="29.[151,1437,151,1200]" box="[151,1436,650,672]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43FD2FA42FD70" box="[151,1436,650,672]" pageId="29" pageNumber="366">
17. Distances between setae
<emphasis id="B936EA8BFFBB744E80F23FD2FDD7FD70" box="[465,521,650,672]" italics="true" pageId="29" pageNumber="366">h1-h1</emphasis>
and
<emphasis id="B936EA8BFFBB744E831B3FD2FDAEFD70" box="[568,624,650,672]" italics="true" pageId="29" pageNumber="366">h1-h2</emphasis>
subequal......................................................... 18
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43FF0FA42FD52" blockId="29.[151,1437,151,1200]" box="[151,1436,679,701]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43FF0FA42FD52" box="[151,1436,679,701]" pageId="29" pageNumber="366">
- Distances between setae
<emphasis id="B936EA8BFFBB744E80F23FFFFDD7FD53" box="[465,521,679,701]" italics="true" pageId="29" pageNumber="366">h1-h1</emphasis>
much greater than
<emphasis id="B936EA8BFFBB744E83E13FFFFD25FD53" box="[706,763,679,701]" italics="true" pageId="29" pageNumber="366">h1-h2</emphasis>
..................................................... 20
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43F9DFA42FD17" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43F9DFA42FD35" blockId="29.[151,1437,151,1200]" box="[151,1436,709,731]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43F9DFA42FD35" box="[151,1436,709,731]" pageId="29" pageNumber="366">
18. Setae
<emphasis id="B936EA8BFFBB744E803F3F9DFEE3FD35" box="[284,317,709,730]" italics="true" pageId="29" pageNumber="366">ps3</emphasis>
subequal to or shorter than
<emphasis id="B936EA8BFFBB744E836A3F9DFDA0FD35" box="[585,638,709,730]" italics="true" pageId="29" pageNumber="366">ps1-2</emphasis>
............................................................... 19
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43FBBFA42FD17" blockId="29.[151,1437,151,1200]" box="[151,1436,738,760]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43FBBFA42FD17" box="[151,1436,738,760]" pageId="29" pageNumber="366">
- Setae
<emphasis id="B936EA8BFFBB744E803F3FBAFEE3FD18" box="[284,317,738,759]" italics="true" pageId="29" pageNumber="366">ps3</emphasis>
distinctly longer than
<emphasis id="B936EA8BFFBB744E83363FBAFD94FD18" box="[533,586,738,759]" italics="true" pageId="29" pageNumber="366">ps1-2</emphasis>
.......................
<taxonomicName id="4C424D1AFFBB744E82523FBBFAD2FD18" authority="Dastych, Rack and Camerik, 1997" authorityName="Dastych, Rack and Camerik" authorityYear="1997" box="[881,1292,738,760]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="luriei">
<emphasis id="B936EA8BFFBB744E82523FBBFC63FD18" box="[881,957,738,760]" italics="true" pageId="29" pageNumber="366">S. luriei</emphasis>
<bibRefCitation id="EFD34B68FFBB744E82E03FBAFAD2FD18" author="Dastych, H. &amp; Rack, G. &amp; Camerik, A. M." box="[963,1292,738,760]" pageId="29" pageNumber="366" pagination="145 - 151" refId="ref15568" refString="Dastych, H., Rack, G. &amp; Camerik, A. M. (1997) A new species of the genus Spatulaphorus Rack (Acari: Heterostigmata, Pygmephoridae) associated with scarab beetles in South Africa. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 94, 145 - 151." type="journal article" year="1997">Dastych, Rack and Camerik, 1997</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBB744E843A3FBAFA48FD17" box="[1305,1430,738,760]" name="South Africa" pageId="29" pageNumber="366">South Africa</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43E58FA43FCDD" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43E58FA42FCFA" blockId="29.[151,1437,151,1200]" box="[151,1436,767,789]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43E58FA42FCFA" box="[151,1436,767,789]" pageId="29" pageNumber="366">
19. Setae (
<emphasis id="B936EA8BFFBB744E80073E58FEE5FCFA" box="[292,315,768,789]" italics="true" pageId="29" pageNumber="366">pv</emphasis>
) on tarsus III foliate, setae
<emphasis id="B936EA8BFFBB744E83613E58FDA9FCFA" box="[578,631,768,789]" italics="true" pageId="29" pageNumber="366">ps1-2</emphasis>
distinctly barbed.....
<taxonomicName id="4C424D1AFFBB744E82513E58FAEFFCFA" authority="Khaustov and Frolov, 2018" authorityName="Khaustov and Frolov" authorityYear="2018" box="[882,1329,767,789]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="enoplotruporum">
<emphasis id="B936EA8BFFBB744E82513E58FBF8FCFB" box="[882,1062,767,789]" italics="true" pageId="29" pageNumber="366">S. enoplotruporum</emphasis>
<bibRefCitation id="EFD34B68FFBB744E850F3FA7FAEFFCFA" author="Khaustov, A. A. &amp; Frolov, A. V." box="[1068,1329,767,789]" pageId="29" pageNumber="366" pagination="277 - 292" refId="ref16197" refString="Khaustov, A. A. &amp; Frolov, A. V. (2018) New taxa of pygmephoroid mites (Acari: Pygmephoroidea: Pygmephoridae, Neopygmephoridae) phoretic on Enoplotrupes sharpi (Coleoptera: Geotrupidae) from Thailand. Zootaxa, 4442 (2), 277 - 292. https: // doi. org / 10.11646 / zootaxa. 4442.2.5" type="journal article" year="2018">Khaustov and Frolov, 2018</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBB744E841E3FA7FA4BFCFA" box="[1341,1429,767,789]" name="Thailand" pageId="29" pageNumber="366">Thailand</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43E45FA43FCDD" blockId="29.[151,1437,151,1200]" box="[151,1437,797,819]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43E45FA43FCDD" box="[151,1437,797,819]" pageId="29" pageNumber="366">
- Setae (
<emphasis id="B936EA8BFFBB744E80073E45FE9DFCDD" box="[292,323,797,818]" italics="true" pageId="29" pageNumber="366">pv)</emphasis>
on tarsus III not foliate, setae
<emphasis id="B936EA8BFFBB744E83453E45FD42FCDD" box="[614,668,797,818]" italics="true" pageId="29" pageNumber="366">ps1-2</emphasis>
smooth.................
<taxonomicName id="4C424D1AFFBB744E82983E45FAF6FCDD" authority="Dastych and Rack, 1993" authorityName="Dastych and Rack" authorityYear="1993" box="[955,1320,797,819]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="camerikae">
<emphasis id="B936EA8BFFBB744E82983E45FBE9FCDD" box="[955,1079,797,818]" italics="true" pageId="29" pageNumber="366">S. camerikae</emphasis>
<bibRefCitation id="EFD34B68FFBB744E851E3E45FAF6FCDD" author="Dastych, H. &amp; Rack, G." box="[1085,1320,797,819]" pageId="29" pageNumber="366" pagination="265 - 84" refId="ref15517" refString="Dastych, H. &amp; Rack, G. (1993) Spatulaphorus gen. n. and three new species of pygmephorid mites associated with scarab beetles in Botswana and Vietnam (Acari: Heterostigmata). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 90, 265 - 84." type="journal article" year="1993">Dastych and Rack, 1993</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBB744E84173E45FA48FCDD" box="[1332,1430,797,818]" name="Botswana" pageId="29" pageNumber="366">Botswana</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43E62FA42FC64" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43E62FA42FCA0" blockId="29.[151,1437,151,1200]" box="[151,1436,826,848]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43E62FA42FCA0" box="[151,1436,826,848]" pageId="29" pageNumber="366">
20. Apodemes 4 long, extending beyond setae
<emphasis id="B936EA8BFFBB744E835D3E62FD48FCA0" box="[638,662,826,847]" italics="true" pageId="29" pageNumber="366">3b</emphasis>
; apodemes 3 weakly developed.................................... 21
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43E00FA42FC64" blockId="29.[151,1437,151,1200]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43E00FA42FC64" pageId="29" pageNumber="366">
- Apodemes 4 vestigial; apodemes 3 well developed between bases of setae
<emphasis id="B936EA8BFFBB744E828D3E00FC18FC82" box="[942,966,856,877]" italics="true" pageId="29" pageNumber="366">3a</emphasis>
and trochanter III..........................................................................................
<taxonomicName id="4C424D1AFFBB744E85243E2DFAE7FC64" authority="Khaustov, 2005" authorityName="Khaustov" authorityYear="2005" box="[1031,1337,885,907]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="geotruporum">
<emphasis id="B936EA8BFFBB744E85243E2DFB42FC64" box="[1031,1180,885,907]" italics="true" pageId="29" pageNumber="366">S. geotruporum</emphasis>
<bibRefCitation id="EFD34B68FFBB744E85803E2DFAE7FC64" author="Khaustov, A. A." box="[1187,1337,885,907]" pageId="29" pageNumber="366" pagination="35 - 42" refId="ref15889" refString="Khaustov, A. A. (2005) Three new species of mite family Pygmephoridae (Acari: Heterostigmata) associated with scarab beetles (Coleoptera: Scarabaeidae) from Ukraine. Acarina, 13, 35 - 42." type="journal article" year="2005">Khaustov, 2005</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBB744E84663E2DFA4BFC64" box="[1349,1429,885,907]" name="Ukraine" pageId="29" pageNumber="366">Ukraine</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43ECAFA43FC2A" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43ECAFA42FC48" blockId="29.[151,1437,151,1200]" box="[151,1436,914,936]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43ECAFA42FC48" box="[151,1436,914,936]" pageId="29" pageNumber="366">21. Tibiotarsus I thickened, almost ovoid in shape and distinctly wider than genu I................................... 22</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43EE8FA43FC2A" blockId="29.[151,1437,151,1200]" box="[151,1437,943,965]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43EE8FA43FC2A" box="[151,1437,943,965]" pageId="29" pageNumber="366">
- Tibiotarsus I cylindrical, subequal in width with genu I........................
<taxonomicName id="4C424D1AFFBB744E853F3EE8FAEDFC2A" authority="(Mahunka, 1970)" baseAuthorityName="Mahunka" baseAuthorityYear="1970" box="[1052,1331,943,965]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="geotrupi">
<emphasis id="B936EA8BFFBB744E853F3EE8FB5BFC2A" box="[1052,1157,944,965]" italics="true" pageId="29" pageNumber="366">S. geotrupi</emphasis>
(
<bibRefCitation id="EFD34B68FFBB744E85B13EF7FAF5FC2A" author="Mahunka, S." box="[1170,1323,943,965]" pageId="29" pageNumber="366" pagination="137 - 174" refId="ref16668" refString="Mahunka, S. (1970) Considerations on the systematics of the Tarsonemina and the description of new European taxa (Acari: Trombidiformes). Acta Zoologica Academiae Scientiarum Hungaricae, 16, 137 - 174." type="journal article" year="1970">Mahunka, 1970</bibRefCitation>
)
</taxonomicName>
(
<collectingCountry id="F3557609FFBB744E841C3EE8FA48FC2A" box="[1343,1430,944,965]" name="Hungary" pageId="29" pageNumber="366">Hungary</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43E95FA42FBF2" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43E95FA42FC0D" blockId="29.[151,1437,151,1200]" box="[151,1436,973,995]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43E95FA42FC0D" box="[151,1436,973,995]" pageId="29" pageNumber="366">22. Apodemes 5 short, weakly sclerotized................................................................... 23</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43EB3FA42FBF2" blockId="29.[151,1437,151,1200]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43EB3FA42FBF2" pageId="29" pageNumber="366">
- Apodemes 5 long, thick and well sclerotized, extending beyond bases of trochanter IV................................................................................................
<taxonomicName id="4C424D1AFFBB744E82D93950FB61FBF3" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[1018,1215,1031,1053]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="madagascariensis" status="sp. nov.">
<emphasis id="B936EA8BFFBB744E82D93950FB61FBF3" box="[1018,1215,1031,1053]" italics="true" pageId="29" pageNumber="366">S. madagascariensis</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBB744E85E53950FACCFBF2" bold="true" box="[1222,1298,1032,1053]" pageId="29" pageNumber="366">
<taxonomicNameLabel id="A20557F0FFBB744E85E53950FACCFBF2" box="[1222,1298,1032,1053]" pageId="29" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F3557609FFBB744E843C395FFA48FBF2" box="[1311,1430,1031,1053]" name="Madagascar" pageId="29" pageNumber="366">Madagascar</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B4397DFA43FBB8" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B4397DFA42FBD5" blockId="29.[151,1437,151,1200]" box="[151,1436,1061,1083]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B4397DFA42FBD5" box="[151,1436,1061,1083]" pageId="29" pageNumber="366">
23. Setae
<emphasis id="B936EA8BFFBB744E803F397DFEE3FBD5" box="[284,317,1061,1082]" italics="true" pageId="29" pageNumber="366">ps3</emphasis>
subequal with
<emphasis id="B936EA8BFFBB744E80EC397DFDDAFBD5" box="[463,516,1061,1082]" italics="true" pageId="29" pageNumber="366">ps1-2</emphasis>
.......................................................................... 24
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B4391BFA43FBB8" blockId="29.[151,1437,151,1200]" box="[151,1437,1090,1112]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B4391BFA43FBB8" box="[151,1437,1090,1112]" pageId="29" pageNumber="366">
- Setae
<emphasis id="B936EA8BFFBB744E803F391AFEE3FBB8" box="[284,317,1090,1111]" italics="true" pageId="29" pageNumber="366">ps3</emphasis>
more than twice longer than
<emphasis id="B936EA8BFFBB744E8375391AFD52FBB8" box="[598,652,1090,1111]" italics="true" pageId="29" pageNumber="366">ps1-2</emphasis>
........................
<taxonomicName id="4C424D1AFFBB744E829F391BFABEFBB8" authority="Sobhi and Hajiqanbar, 2017" authorityName="Sobhi and Hajiqanbar" authorityYear="2017" box="[956,1376,1090,1112]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="gymnopleuri">
<emphasis id="B936EA8BFFBB744E829F391BFB93FBB8" box="[956,1101,1090,1112]" italics="true" pageId="29" pageNumber="366">S. gymnopleuri</emphasis>
<bibRefCitation id="EFD34B68FFBB744E8570391AFABEFBB8" author="Sobhi, M. &amp; Hajiqanbar, H. &amp; Mortazavi, A." box="[1107,1376,1090,1112]" pageId="29" pageNumber="366" pagination="217 - 223" refId="ref16872" refString="Sobhi, M., Hajiqanbar, H. &amp; Mortazavi, A. (2017) A new species of the genus Spatulaphorus (Acari: Heterostigmata: Pygmephoridae) associated with Gymnopleurus flagellatus (Coleoptera: Scarabaeidae) from Iran. Systematic &amp; Applied Acarology, 22, 217 - 223. https: // doi. org / 10.11158 / saa. 22.2.6" type="journal article" year="2017">Sobhi and Hajiqanbar, 2017</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBB744E844F391AFA4BFBB8" box="[1388,1429,1090,1111]" name="Iran" pageId="29" pageNumber="366">Iran</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B62E3CFFBB744E81B43938FE22FB5F" pageId="29" pageNumber="366">
<paragraph id="8BFD3699FFBB744E81B43938FA43FB9A" blockId="29.[151,1437,151,1200]" box="[151,1437,1119,1141]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43938FA43FB9A" box="[151,1437,1119,1141]" pageId="29" pageNumber="366">
24. Posterior margin of posterior sternal plate deeply incised; setae
<emphasis id="B936EA8BFFBB744E82663938FCA4FB9A" box="[837,890,1120,1141]" italics="true" pageId="29" pageNumber="366">ps1-2</emphasis>
weakly barbed..........
<taxonomicName id="4C424D1AFFBB744E85BD3938FB27FB9A" authorityName="Khaustov &amp; Frolov &amp; Akhmetova" authorityYear="2024" box="[1182,1273,1120,1141]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="incisus" status="sp. nov.">
<emphasis id="B936EA8BFFBB744E85BD3938FB27FB9A" box="[1182,1273,1120,1141]" italics="true" pageId="29" pageNumber="366">S. incisus</emphasis>
</taxonomicName>
<emphasis id="B936EA8BFFBB744E85DC3938FA95FB9A" bold="true" box="[1279,1355,1120,1141]" pageId="29" pageNumber="366">
<taxonomicNameLabel id="A20557F0FFBB744E85DC3938FA95FB9A" box="[1279,1355,1120,1141]" pageId="29" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F3557609FFBB744E847B3907FA4BFB9A" box="[1368,1429,1119,1141]" name="Brazil" pageId="29" pageNumber="366">Brazil</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="8BFD3699FFBB744E81B43925FE22FB5F" blockId="29.[151,1437,151,1200]" pageId="29" pageNumber="366">
<keyLead id="30B395ACFFBB744E81B43925FE22FB5F" pageId="29" pageNumber="366">
- Posterior margin of posterior sternal plate only slightly concave; setae
<emphasis id="B936EA8BFFBB744E82AB3925FC63FB7D" box="[904,957,1149,1170]" italics="true" pageId="29" pageNumber="366">ps1-2</emphasis>
smooth...
<taxonomicName id="4C424D1AFFBB744E85633925FECCFB40" authority="Khaustov and Frolov, 2017" authorityName="Khaustov and Frolov" authorityYear="2017" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="29" pageNumber="366" phylum="Arthropoda" rank="species" species="braziliensis">
<emphasis id="B936EA8BFFBB744E85633925FB92FB7D" box="[1088,1100,1149,1170]" italics="true" pageId="29" pageNumber="366">S</emphasis>
.
<emphasis id="B936EA8BFFBB744E857B3925FB19FB7D" box="[1112,1223,1149,1170]" italics="true" pageId="29" pageNumber="366">braziliensis</emphasis>
<bibRefCitation id="EFD34B68FFBB744E85EE3925FECCFB40" author="Khaustov, A. A. &amp; Frolov, A. V." pageId="29" pageNumber="366" pagination="501 - 521" refId="ref16128" refString="Khaustov, A. A. &amp; Frolov, A. V. (2017) New species of heterostigmatic mites (Acari: Heterostigmata: Athyreacaridae, Dolichocybidae, Pygmephoridae) associated with scarab beetles (Coleoptera: Geotrupidae, Scarabaeidae) from Brazil. Zootaxa, 4294 (5), 501 - 521. https: // doi. org / 10.11646 / zootaxa. 4294.5.1" type="journal article" year="2017">Khaustov and Frolov, 2017</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3557609FFBB744E803C39C2FE83FB5F" box="[287,349,1178,1200]" name="Brazil" pageId="29" pageNumber="366">Brazil</collectingCountry>
,
<collectingCountry id="F3557609FFBB744E804539C2FE28FB5F" box="[358,502,1178,1200]" name="French Guiana" pageId="29" pageNumber="366">French Guiana</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>