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<document id="3319A2EF8F827DA73C4BC4D8AAB0DC91" ID-ISSN="1464-5262" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629979807253" checkinUser="felipe" docAuthor="Rodney" docDate="2003" docId="03D687A1BE3A2B6528A0D6F7FE7874A6" docLanguage="en" docName="JNatHist.37.21.2615-2626.pdf" docOrigin="Journal of Natural History 37" docStyle="DocumentStyle:443144A73DAE3E092C51A378E4CD0138.2:JNatHist.2003-2004.journal_article.1cover" docStyleId="443144A73DAE3E092C51A378E4CD0138" docStyleName="JNatHist.2003-2004.journal_article.1cover" docStyleVersion="2" docTitle="Leptandria ashei Rodney 2003, sp. n." docType="treatment" docVersion="3" lastPageNumber="2621" masterDocId="FFEFFFD9BE3F2B622A32D528FFBB750A" masterDocTitle="Leptandria gen. n., a monophyletic group of Neotropical aleocharine Staphylinidae (Coleoptera)" masterLastPageNumber="2626" masterPageNumber="2615" pageNumber="2619" updateTime="1699132954488" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="A67A8126314584DEFB0467828B7567B3">
<mods:title id="47D8A3349D7D903CCA8B0B625DBC08FB">Leptandria gen. n., a monophyletic group of Neotropical aleocharine Staphylinidae (Coleoptera)</mods:title>
</mods:titleInfo>
<mods:name id="401D3BE12C1837A6F4A532DDAF5FDAAC" type="personal">
<mods:role id="AFE5A70D7FD4C2B00C889C50EF65F016">
<mods:roleTerm id="4A25D39A11EC1394860B7D54983EA8E4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5AE4A0EA30A2D554C7372BA2D406FC72">Rodney</mods:namePart>
</mods:name>
<mods:typeOfResource id="6D9DA1CBAA033C5C82903ACD4BF91D25">text</mods:typeOfResource>
<mods:relatedItem id="3A3760360A131BBE27BE5C3621C60504" type="host">
<mods:titleInfo id="9700A3425CA781648393D8D76EBCB0FE">
<mods:title id="ED212726549643B81B7E0ED56A154C8E">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="1451AD399312EBA06BE9F6E884C653B5">
<mods:date id="89E248418B4C2DE0F421580A5EA0BD74">2003</mods:date>
<mods:detail id="AE654F8927AFB72EE5BBFA0975BC9541" type="pubDate">
<mods:number id="14767187E75A668A6094D9DDD652061A">2003-10-10</mods:number>
</mods:detail>
<mods:detail id="F26DF50BA3AD31BA5FE31CE5DEA1BE43" type="volume">
<mods:number id="781756438EFED7E1DFFB42725F71BEBC">37</mods:number>
</mods:detail>
<mods:extent id="20A4131346E236A44FB4DD37FF571E07" unit="page">
<mods:start id="186CA87A05848E501395F08CCBCC01D2">2615</mods:start>
<mods:end id="75417918170C41ECBC3AE7A050B41788">2626</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4F6C9303B909A8349494FEF763F91FF6">journal article</mods:classification>
<mods:identifier id="C08F117B735A52E057EEE2E21034B675" type="ISSN">1464-5262</mods:identifier>
</mods:mods>
<treatment id="03D687A1BE3A2B6528A0D6F7FE7874A6" ID-DOI="http://doi.org/10.5281/zenodo.5271020" ID-Zenodo-Dep="5271020" LSID="urn:lsid:plazi:treatment:03D687A1BE3A2B6528A0D6F7FE7874A6" httpUri="http://treatment.plazi.org/id/03D687A1BE3A2B6528A0D6F7FE7874A6" lastPageId="7" lastPageNumber="2621" pageId="5" pageNumber="2619">
<subSubSection id="C365653CBE3A2B6728A0D6F7FCEA7112" pageId="5" pageNumber="2619" type="nomenclature">
<paragraph id="8BC036B7BE3A2B6728A0D6F7FC2076FD" blockId="5.[658,923,991,1048]" box="[658,923,991,1015]" pageId="5" pageNumber="2619">
<heading id="D08881DBBE3A2B6728A0D6F7FC2076FD" box="[658,923,991,1015]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="2619" reason="3">
<taxonomicName id="4C7F4D34BE3A2B6728A0D6F7FCF676FD" ID-CoL="6PDTY" authority="Rodney, 2003" authorityName="Rodney" authorityYear="2003" box="[658,845,991,1015]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="2619" phylum="Arthropoda" rank="species" species="ashei" status="sp. nov.">
<emphasis id="B90BEAA5BE3A2B6728A0D6F7FCF676FD" bold="true" box="[658,845,991,1015]" italics="true" pageId="5" pageNumber="2619">Leptandria ashei</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A23857DEBE3A2B67296ED6C8FC2076FD" box="[860,923,992,1015]" pageId="5" pageNumber="2619" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BC036B7BE3A2B6728EED128FCEA7112" blockId="5.[658,923,991,1048]" box="[732,849,1024,1048]" pageId="5" pageNumber="2619">(Figure 2)</paragraph>
</subSubSection>
<subSubSection id="C365653CBE3A2B652B53D11AFE7874A6" lastPageId="7" lastPageNumber="2621" pageId="5" pageNumber="2619" type="materials_examined">
<paragraph id="8BC036B7BE3A2B672B53D11AFB4F71A0" blockId="5.[313,1269,1073,1736]" pageId="5" pageNumber="2619">
<emphasis id="B90BEAA5BE3A2B672B53D11AFE717140" box="[353,458,1074,1098]" italics="true" pageId="5" pageNumber="2619">Material.</emphasis>
H
<collectionCode id="ED6EAE72BE3A2B672811D11AFD8A7140" box="[547,561,1074,1098]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="2619"></collectionCode>
<materialsCitation id="3B173CEABE3A2B672803D11AFEED71A0" collectingDate="1989-06-22" collectionCode="R, SEMC" collectorName="Ashe &amp; Leschen &amp; S. Hanley" country="Peru" location="Tambopata Prov." pageId="5" pageNumber="2619" specimenCount="1" typeStatus="holotype">
 male, [‘
<emphasis id="B90BEAA5BE3A2B6728E0D11AFCA67140" bold="true" box="[722,797,1074,1098]" pageId="5" pageNumber="2619">
<collectingCountry id="F3687627BE3A2B6728E0D11AFCA67140" box="[722,797,1074,1098]" name="Peru" pageId="5" pageNumber="2619">PERU</collectingCountry>
</emphasis>
:
<location id="8EA0606CBE3A2B672908D11AFBA87143" LSID="urn:lsid:plazi:treatment:03D687A1BE3A2B6528A0D6F7FE7874A6:8EA0606CBE3A2B672908D11AFBA87143" box="[826,1043,1074,1098]" country="Peru" name="Tambopata Prov." pageId="5" pageNumber="2619">Tambopata Prov.</location>
,
<locationDeviation id="9A722F95BE3A2B672E1CD119FE737160" location="Pto. Maldonaldo" pageId="5" pageNumber="2619">
<quantity id="4C879B52BE3A2B672E1CD119FBCA7140" box="[1070,1137,1073,1098]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="2619" unit="km" value="15.0">15 km</quantity>
NE Pto. Maldonaldo
</locationDeviation>
;
<date id="FFC11077BE3A2B672BE8D17AFDC07160" box="[474,635,1106,1130]" pageId="5" pageNumber="2619" value="1989-06-22">
<collectingDate id="EF85E99FBE3A2B672BE8D17AFDC07160" box="[474,635,1106,1130]" pageId="5" pageNumber="2619" value="1989-06-22">
22 June
<quantity id="4C879B52BE3A2B672871D17AFDC07160" box="[579,635,1106,1130]" metricMagnitude="6" metricUnit="m" metricValue="1.9892" pageId="5" pageNumber="2619" unit="m" value="1989200.0">1989</quantity>
</collectingDate>
</date>
, 200 m; J.
<collectorName id="268A5361BE3A2B672930D17AFC857160" box="[770,830,1106,1130]" pageId="5" pageNumber="2619">Ashe</collectorName>
,
<collectionCode id="ED6EAE72BE3A2B67297DD17AFCDC7163" box="[847,871,1106,1129]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="2619">R</collectionCode>
.
<collectorName id="268A5361BE3A2B672945D17AFC6D7160" box="[887,982,1106,1130]" pageId="5" pageNumber="2619">Leschen</collectorName>
, #223; ex. flower beds’/ ‘
<typeStatus id="54C48815BE3A2B672B0FD15AFE607180" box="[317,475,1138,1162]" pageId="5" pageNumber="2619" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="4C7F4D34BE3A2B672BDAD15AFCBB7180" authority="Hanley" authorityName="Hanley" authorityYear="2003" box="[488,768,1138,1162]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="2619" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="B90BEAA5BE3A2B672BDAD15AFD187180" box="[488,675,1138,1162]" italics="true" pageId="5" pageNumber="2619">Leptandria ashei</emphasis>
Hanley
</taxonomicName>
, desig.
<collectionCode id="ED6EAE72BE3A2B672964D15AFCD57183" box="[854,878,1138,1161]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="2619">R</collectionCode>
.
<collectorName id="268A5361BE3A2B672948D15AFC557180" box="[890,1006,1138,1162]" pageId="5" pageNumber="2619">S. Hanley</collectorName>
, 2001’ (red)] (
<collectionCode id="ED6EAE72BE3A2B672EA2D15AFB5B7180" box="[1168,1248,1138,1162]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="5" pageNumber="2619" type="Museum">SEMC</collectionCode>
). P
</materialsCitation>
<collectionCode id="ED6EAE72BE3A2B672B64D1BAFED871A0" box="[342,355,1170,1194]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="2619"></collectionCode>
<collectionCode id="ED6EAE72BE3A2B672B42D1BAFEC671A0" box="[368,381,1170,1194]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="2619"></collectionCode>
<materialsCitation id="3B173CEABE3A2B672B4FD1BAFB5471A0" box="[381,1263,1170,1194]" collectingDate="1989-06-22" collectionCode="SEMC" collectorName="Ashe &amp; Leschen &amp; S. Hanley" country="Peru" location="Tambopata Prov." pageId="5" pageNumber="2619" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
,
<specimenCount id="9D79FD3EBE3A2B672B8CD1BAFD8D71A0" box="[446,566,1170,1194]" pageId="5" pageNumber="2619" type="male">nine males</specimenCount>
,
<specimenCount id="9D79FD3EBE3A2B672872D1BAFD7971A0" box="[576,706,1170,1194]" pageId="5" pageNumber="2619" type="female">five females</specimenCount>
with the same label data as the holotype (
<collectionCode id="ED6EAE72BE3A2B672EA2D1BAFB5B71A0" box="[1168,1248,1170,1194]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="5" pageNumber="2619" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC036B7BE3A2B672B53D19AFC7570E3" blockId="5.[313,1269,1073,1736]" pageId="5" pageNumber="2619">
<emphasis id="B90BEAA5BE3A2B672B53D19AFE6E71C3" box="[353,469,1202,1225]" italics="true" pageId="5" pageNumber="2619">Diagnosis.</emphasis>
This species differs from
<taxonomicName id="4C7F4D34BE3A2B672921D19AFC1771C3" authorityName="Rodney" authorityYear="2003" box="[787,940,1201,1225]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="2619" phylum="Arthropoda" rank="species" species="tishechkini">
<emphasis id="B90BEAA5BE3A2B672921D19AFC1771C3" box="[787,940,1201,1225]" italics="true" pageId="5" pageNumber="2619">L. tishechkini</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D34BE3A2B6729D8D19AFBC371C3" authorityName="Rodney" authorityYear="2003" box="[1002,1144,1201,1225]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="2619" phylum="Arthropoda" rank="species" species="ecitophila">
<emphasis id="B90BEAA5BE3A2B6729D8D19AFBC371C3" box="[1002,1144,1201,1225]" italics="true" pageId="5" pageNumber="2619">L. ecitophila</emphasis>
</taxonomicName>
by having small eyes, overall length about 0.3–0.4 times the length of the temples; a more or less oval pronotum with the area along the midline slightly extended posteriorly; and the abdominal sternum VIII without a modified duct opening.
<taxonomicName id="4C7F4D34BE3A2B672E09D039FB4E7023" authorityName="Hanley" authorityYear="2003" box="[1083,1269,1297,1321]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="2619" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="B90BEAA5BE3A2B672E09D039FB4E7023" box="[1083,1269,1297,1321]" italics="true" pageId="5" pageNumber="2619">Leptandria ashei</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C7F4D34BE3A2B672BFAD01AFDEF7043" authorityName="Rodney" authorityYear="2003" box="[456,596,1329,1353]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="2619" phylum="Arthropoda" rank="species" species="ecitophila">
<emphasis id="B90BEAA5BE3A2B672BFAD01AFDEF7043" box="[456,596,1329,1353]" italics="true" pageId="5" pageNumber="2619">L. ecitophila</emphasis>
</taxonomicName>
by having relatively short antennae, typically reaching the apex of the pronotum when extended posteriorly (in
<taxonomicName id="4C7F4D34BE3A2B672988D07AFBF07063" authorityName="Rodney" authorityYear="2003" box="[954,1099,1361,1385]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="2619" phylum="Arthropoda" rank="species" species="ecitophila">
<emphasis id="B90BEAA5BE3A2B672988D07AFBF07063" box="[954,1099,1361,1385]" italics="true" pageId="5" pageNumber="2619">L. ecitophila</emphasis>
</taxonomicName>
, the antennae reach to the middle of the elytra when extended posteriorly), no distinctive secondary sexual features in the males (in
<taxonomicName id="4C7F4D34BE3A2B6728ADD0B9FC9170A3" authorityName="Rodney" authorityYear="2003" box="[671,810,1425,1449]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="2619" phylum="Arthropoda" rank="species" species="ecitophila">
<emphasis id="B90BEAA5BE3A2B6728ADD0B9FC9170A3" box="[671,810,1425,1449]" italics="true" pageId="5" pageNumber="2619">L. ecitophila</emphasis>
</taxonomicName>
, males possess various secondary sexual features), and no lobe-like structures in the female genitalic tract (in
<taxonomicName id="4C7F4D34BE3A2B672E6DD099FB5670C2" authorityName="Rodney" authorityYear="2003" box="[1119,1261,1456,1480]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="2619" phylum="Arthropoda" rank="species" species="ecitophila">
<emphasis id="B90BEAA5BE3A2B672E6DD099FB5670C2" box="[1119,1261,1456,1480]" italics="true" pageId="5" pageNumber="2619">L. ecitophila</emphasis>
</taxonomicName>
, females possess lobe-like structures in the genitalic tract).
</paragraph>
<paragraph id="8BC036B7BE3A2B652B53D0D9FC037447" blockId="5.[313,1269,1073,1736]" lastBlockId="7.[313,1268,181,428]" lastPageId="7" lastPageNumber="2621" pageId="5" pageNumber="2619">
<emphasis id="B90BEAA5BE3A2B672B53D0D9FE537302" box="[353,488,1521,1544]" italics="true" pageId="5" pageNumber="2619">Description.</emphasis>
Body length
<quantity id="4C879B52BE3A2B6728A0D0D8FCA37302" box="[658,792,1520,1544]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="4.2" metricValueMin="2.4" pageId="5" pageNumber="2619" unit="mm" value="3.3" valueMax="4.2" valueMin="2.4">2.4–4.2 mm</quantity>
. Head piceous to black, pronotum, elytra, and abdomen brown to dark brown. Head, pronotum, elytra and abdomen covered throughout with short pile of yellow microsetae; integument heavily punctured, moderately glossy. Head with eyes small, length about 0.3–0.4 times length of temples, distinct carina present below each eye, pubescence concentrated at vertex, primarily directed slightly anteriorly towards midline. Antennae (figure 2A) short in length, just reaching to apex of pronotum when extended posteriorly, widest part at segments 8–11, apical segment acute to rounded. Pronotum (figure 2B) more or less oval with outer apical angles slightly sinuate to straight, area along midline slightly extended posteriorly, pubescence evenly distributed, primarily directed posteriorly, each lateral margin of disc with four small macrosetae. Elytra (figure 2C) with each elytron about 1.2 times longer than wide, outer apical margin distinctly sinuate, surface with setiferous punctures, evenly pubescent, with pubescence directed toward apico-lateral angles. Tarsi (figure 2D) with single, long empodial bristle, length slightly shorter than tarsal claws. Mesosternum (figure 2E) without medial carina. Males often with secondary sexual features of small, medioapical denticle on each elytron (figure 2C), U-shaped, longitudinal raised area on abdominal tergum IV, narrow, longitudinal carina on abdominal tergum VII (figure 2G). Tergum IX as depicted (figure 2H). Female genitalic tract with two unsclerotized lobes with numerous, distinct denticles and spines (figure 2F). Spermatheca as depicted (figure 2I). Aedeagus (figure 2J), paramere (figure 2K) as depicted.
</paragraph>
<caption id="DF00663FBE392B642B0BD03AFC52707D" pageId="6" pageNumber="2620" startId="6.[313,329,1298,1319]" targetBox="[313,1268,179,1272]" targetPageId="6">
<paragraph id="8BC036B7BE392B642B0BD03AFC52707D" blockId="6.[313,1268,1298,1399]" pageId="6" pageNumber="2620">
F. 2.
<taxonomicName id="4C7F4D34BE392B642BA7D03AFDF8702D" authority="Rodney, 2003" authorityName="Rodney" authorityYear="2003" box="[405,579,1298,1319]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="2620" phylum="Arthropoda" rank="species" species="ashei" status="sp. nov.">
<emphasis id="B90BEAA5BE392B642BA7D03AFDF8702D" box="[405,579,1298,1319]" italics="true" pageId="6" pageNumber="2620">Leptandria ashei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857DEBE392B642860D03AFD2E702D" box="[594,661,1298,1319]" pageId="6" pageNumber="2620" rank="species">sp. n.</taxonomicNameLabel>
: (A) antenna; (B) pronotum; (C) left elytron (male); (D) tarsus of hind leg; (E) meso- and metathoracic region; (F) genitalic tract (female); (G) abdominal tergum VII (male); (H) abdominal tergum IX; (I) spermatheca, lateral view; (J) aedeagus, lateral view; (K) paramere, lateral view.
</paragraph>
</caption>
<paragraph id="8BC036B7BE382B652B53D47CFE7874A6" blockId="7.[313,1268,181,428]" pageId="7" pageNumber="2621">
<emphasis id="B90BEAA5BE382B652B53D47CFE777466" box="[353,460,340,364]" italics="true" pageId="7" pageNumber="2621">Remarks.</emphasis>
This species was referred to as ‘undescribed species 1’ in the phylogeny portion in Hanley (2002). The adults of
<taxonomicName id="4C7F4D34BE382B652928D45DFCCE7486" authorityName="Rodney" authorityYear="2003" box="[794,885,372,396]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2621" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="B90BEAA5BE382B652928D45DFCCE7486" box="[794,885,372,396]" italics="true" pageId="7" pageNumber="2621">L. ashei</emphasis>
</taxonomicName>
were collected from unidentified flower beds.
</paragraph>
</subSubSection>
</treatment>
</document>
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