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<document id="423FE3449D91DE56CE4A5487857AC832" ID-DOI="10.11646/zootaxa.5486.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13209689" ID-ZooBank="46D2A955-0566-4711-A099-2C6947487E18" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1722787457646" checkinUser="plazi" docAuthor="Tovar-Hernandez, Maria Ana &amp; Jirkov, Igor A." docDate="2024" docId="03CFF948FF882B548B8376BCFD403946" docLanguage="en" docName="zootaxa.5486.1.2.pdf" docOrigin="Zootaxa 5486 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5486.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Jasmineira filatovae" docType="treatment" docVersion="4" lastPageNumber="61" masterDocId="FFF68130FF852B598B14762BFFC33C3F" masterDocTitle="Jasmineira filatovae Levenstein, 1961, the deepest known sabellid is a Potamethus Chamberlin, 1919: redescription, new combination and generic emendation" masterLastPageNumber="70" masterPageNumber="48" pageNumber="61" updateTime="1722949701387" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FEDFC7E58522647147FA54CAEAF99892">Jasmineira filatovae Levenstein, 1961, the deepest known sabellid is a Potamethus Chamberlin, 1919: redescription, new combination and generic emendation</mods:title>
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<mods:namePart id="FB3D7532885D5A6CE854225EF9FD844D">Tovar-Hernandez, Maria Ana</mods:namePart>
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<mods:affiliation id="40F26768EA8B30889BED54E36C665BF3">Facultad de Ciencias Biológicas, Laboratorio de Biosistemática, Universidad Autónoma de Nuevo León, Av. Pedro de Alba esq. Manuel L. Barragán, San Nicolás de los Garza, 66455, Nuevo León, México</mods:affiliation>
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<mods:namePart id="DC70703E8D57DAADF28F61459ACE0281">Jirkov, Igor A.</mods:namePart>
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<mods:affiliation id="8710FD44DCB97E48418E56567726DBDF">Department of General Ecology and Hydrobiology, Biological Faculty, Leninskiye Gory, 1, building 12, Moscow State University, Moscow, 119234, Russia</mods:affiliation>
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<treatment id="03CFF948FF882B548B8376BCFD403946" LSID="urn:lsid:plazi:treatment:03CFF948FF882B548B8376BCFD403946" httpUri="http://treatment.plazi.org/id/03CFF948FF882B548B8376BCFD403946" lastPageNumber="61" pageId="13" pageNumber="61">
<subSubSection id="C37C1BD5FF882B548B8376BCFCAF3C8D" box="[151,876,151,178]" pageId="13" pageNumber="61" type="nomenclature">
<paragraph id="8BD9485EFF882B548B8376BCFCAF3C8D" blockId="13.[151,876,151,178]" box="[151,876,151,178]" pageId="13" pageNumber="61">
<heading id="D091FF32FF882B548B8376BCFCAF3C8D" bold="true" box="[151,876,151,178]" fontSize="11" level="1" pageId="13" pageNumber="61" reason="1">
<emphasis id="B912944CFF882B548B8376BCFCAF3C8D" bold="true" box="[151,876,151,178]" italics="true" pageId="13" pageNumber="61">
Comparison of
<taxonomicName id="4C6633DDFF882B548A4776BCFE143C8D" ID-CoL="3QM9K" authority="(Levenstein, 1961)" box="[339,471,151,178]" class="Polychaeta" family="Sabellidae" genus="Jasmineira" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="filatovae">P. filatovae</taxonomicName>
with other species of
<taxonomicName id="4C6633DDFF882B5489C976BCFCAF3C8D" authorityName="Chamberlin" authorityYear="1919" box="[733,876,151,178]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="genus">Potamethus</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37C1BD5FF882B548B8376F4FD403946" pageId="13" pageNumber="61" type="description">
<paragraph id="8BD9485EFF882B548B8376F4FC713DEE" blockId="13.[151,1437,223,1401]" pageId="13" pageNumber="61">
<taxonomicName id="4C6633DDFF882B548B8376F4FED83CC6" ID-CoL="84SM9" authorityName="Chamberlin" authorityYear="1919" box="[151,283,223,249]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="genus">
<emphasis id="B912944CFF882B548B8376F4FED83CC6" box="[151,283,223,249]" italics="true" pageId="13" pageNumber="61">Potamethus</emphasis>
</taxonomicName>
is composed of 12 species including the new combination
<taxonomicName id="4C6633DDFF882B5488A876F4FB693CC6" authorityName="Tovar-Hernandez &amp; Jirkov" authorityYear="2024" baseAuthorityName="Levenstein" baseAuthorityYear="1961" box="[956,1194,223,249]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="filatovae">
<emphasis id="B912944CFF882B5488A876F4FB693CC6" box="[956,1194,223,249]" italics="true" pageId="13" pageNumber="61">Potamethus filatovae</emphasis>
</taxonomicName>
proposed here (
<bibRefCitation id="EFF735AFFF882B548E7576F4FEDA3D22" author="Capa, M. &amp; Giangrande, A. &amp; Nogueira, J. M. M. &amp; Tovar-Hernandez, M. A." pageId="13" pageNumber="61" refId="ref9752" refString="Capa, M., Giangrande, A., Nogueira, J. M. M. &amp; Tovar-Hernandez, M. A. (2019) Sabellidae Latreille, 1825. In: Purschke, G., Boggemann, M. &amp; Westheide, W. (Eds.), Handbook of Zoology, Annelida. Vol. 2. Pleistoannelida, Sedentaria II. De Gruyter, Berlin, pp. 164 - 212." type="book" year="2019">
Capa
<emphasis id="B912944CFF882B548B83772FFF173D22" box="[151,212,259,285]" italics="true" pageId="13" pageNumber="61">et al.</emphasis>
2019
</bibRefCitation>
,
<bibRefCitation id="EFF735AFFF882B548A3C7728FEA03D22" author="Capa, M. &amp; Kupriyanova, E. &amp; Nogueira, J. M. M. &amp; Bick, A. &amp; Tovar-Hernandez, M. A." box="[296,355,259,285]" pageId="13" pageNumber="61" refId="ref9832" refString="Capa, M., Kupriyanova, E., Nogueira, J. M. M., Bick, A. &amp; Tovar-Hernandez, M. A. (2021) Fanworms: Yesterday, Today and Tomorrow. Diversity Special Issue Systematics and Biodiversity of Annelids, 13 (3), 130. https: // doi. org / 10.3390 / d 13030130" type="journal volume" year="2021">2021</bibRefCitation>
,
<bibRefCitation id="EFF735AFFF882B548A667728FD5B3D22" author="Read, G. &amp; Fauchald, K." box="[370,664,259,285]" pageId="13" pageNumber="61" refId="ref11461" refString="Read, G. &amp; Fauchald, K. (2024 b) World Polychaeta Database. Potamethus Chamberlin, 1919. World Register of Marine Species. Available from: https: // www. marinespecies. org / aphia. php? p = taxdetails &amp; id = 129543 (accessed 23 May 2024)" type="journal volume" year="2024">Read &amp; Fauchald, 2024b</bibRefCitation>
) (
<tableCitation id="C6E47DE5FF882B5489A67728FCCD3D22" box="[690,782,259,285]" captionStart="TABLE 2" captionStartId="9.[151,239,575,601]" captionTargetPageId="9" captionText="TABLE 2. Species of Potamethus currently considered valid, distinctive features and presence of uncinal humps." httpUri="http://table.plazi.org/id/DF1918D6FF8C2B508B837414FA923E66" pageId="13" pageNumber="61" tableUuid="DF1918D6FF8C2B508B837414FA923E66">Table 2</tableCitation>
). Based on the revision of original descriptions and the contribution provided by
<bibRefCitation id="EFF735AFFF882B548AD2770CFD763D7E" author="Knight-Jones, P." box="[454,693,295,321]" pageId="13" pageNumber="61" pagination="245 - 295" refId="ref10699" refString="Knight-Jones, P. (1983) Contributions to the taxonomy of Sabellidae (Polychaeta). Zoological Journal of the Linnean Society, 79 (3), 245 - 295. https: // doi. org / 10.1111 / j. 1096 - 3642.1983. tb 01167. x" type="journal article" year="1983">Knight-Jones (1983)</bibRefCitation>
, we recognize three features potentially useful to comparative purposes at the interspecific level: the shape of dorsal collar margin and ventral collar lappets, and the presence or absence of ventral shields (
<tableCitation id="C6E47DE5FF882B548AD67744FDDA3DB6" box="[450,537,367,393]" captionStart="TABLE 2" captionStartId="9.[151,239,575,601]" captionTargetPageId="9" captionText="TABLE 2. Species of Potamethus currently considered valid, distinctive features and presence of uncinal humps." httpUri="http://table.plazi.org/id/DF1918D6FF8C2B508B837414FA923E66" pageId="13" pageNumber="61" tableUuid="DF1918D6FF8C2B508B837414FA923E66">Table 2</tableCitation>
). However, a revision based on
<typeStatus id="54DDF6FCFF882B54886E775BFC763DB5" box="[890,949,368,394]" pageId="13" pageNumber="61">types</typeStatus>
is out the scope of this paper.At the generic level, the presence of oval to circular plates or moldures on each side of the anterior peristomial ring and humps in thoracic and abdominal uncini might be diagnostic as explained below.
</paragraph>
<paragraph id="8BD9485EFF882B548BD377F0FB863E5E" blockId="13.[151,1437,223,1401]" pageId="13" pageNumber="61">
<taxonomicName id="4C6633DDFF882B548BD377F0FE6E3DCA" box="[199,429,475,501]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="filitovae">
<emphasis id="B912944CFF882B548BD377F0FE6E3DCA" box="[199,429,475,501]" italics="true" pageId="13" pageNumber="61">Potamethus filitovae</emphasis>
</taxonomicName>
, as most species in
<taxonomicName id="4C6633DDFF882B54899677F0FCC53DCA" ID-CoL="84SM9" authorityName="Chamberlin" authorityYear="1919" box="[642,774,475,501]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="genus">
<emphasis id="B912944CFF882B54899677F0FCC53DCA" box="[642,774,475,501]" italics="true" pageId="13" pageNumber="61">Potamethus</emphasis>
</taxonomicName>
, have pronounced V-shaped dorsal collar margins, exposing the anterior peristomial ring and forming well developed pockets, whereas in
<taxonomicName id="4C6633DDFF882B548F10742BFF113E02" ID-CoL="4M39W" authority="Hartmann-Schroder, 1977" authorityName="Hartmann-Schroder" authorityYear="1977" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="breviuncatus">
<emphasis id="B912944CFF882B548F10742BFB723E26" box="[1028,1201,511,537]" italics="true" pageId="13" pageNumber="61">P. breviuncatus</emphasis>
<bibRefCitation id="EFF735AFFF882B548FAC77D4FF113E02" author="Hartmann-Schroder, G." pageId="13" pageNumber="61" pagination="65 - 99" refId="ref10551" refString="Hartmann-Schroder, G. (1977) Polychaeten aus dem Sublitoral und Bathyal vor der portugiesischen und marokkanischen Kuste. Auswertung der Fahrt 8 (1967) von F. S. ' Meteor'. Meteor Forschungergebnisse, Series D, 26, 65 - 99." type="journal article" year="1977">Hartmann-Schröder, 1977</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B548BC9740FFE363E01" ID-CoL="4M39X" authority="(Eliason, 1951)" authorityName="Nares Deep W" baseAuthorityName="Eliason" baseAuthorityYear="1951" box="[221,501,547,574]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="dubius">
<emphasis id="B912944CFF882B548BC9740FFE803E02" box="[221,323,547,573]" italics="true" pageId="13" pageNumber="61">P. dubius</emphasis>
(Eliason, 1951)
</taxonomicName>
and
<taxonomicName id="4C6633DDFF882B54893E740FFC473E02" ID-CoL="6W7KG" authority="(Johansson, 1922)" authorityName="Sagami" baseAuthorityName="Johansson" baseAuthorityYear="1922" box="[554,900,547,573]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="japonicus">
<emphasis id="B912944CFF882B54893E740FFD713E02" box="[554,690,548,573]" italics="true" pageId="13" pageNumber="61">P. japonicus</emphasis>
(
<bibRefCitation id="EFF735AFFF882B5489D47408FCB83E02" author="Johansson, K. E." box="[704,891,547,573]" pageId="13" pageNumber="61" pagination="1 - 11" refId="ref10598" refString="Johansson, K. E. (1922) On some new tubicolous annelids from Japan, the Bonin Islands and the Antarctic. Arkiv for Zoologi, 15 (2), 1 - 11." type="journal article" year="1922">Johansson, 1922</bibRefCitation>
)
</taxonomicName>
dorsal collar margins form two shallow, convex semi-circles and in
<taxonomicName id="4C6633DDFF882B548A667463FD6F3E5D" ID-CoL="787KJ" authority="Hartman, 1965" authorityName="Hartman" authorityYear="1965" box="[370,684,583,610]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="singularis">
<emphasis id="B912944CFF882B548A667463FE3E3E5E" box="[370,509,583,609]" italics="true" pageId="13" pageNumber="61">P. singularis</emphasis>
Hartman, 1965
</taxonomicName>
these margins are oblique (
<tableCitation id="C6E47DE5FF882B5488F4746CFBFB3E5E" box="[992,1080,583,609]" captionStart="TABLE 2" captionStartId="9.[151,239,575,601]" captionTargetPageId="9" captionText="TABLE 2. Species of Potamethus currently considered valid, distinctive features and presence of uncinal humps." httpUri="http://table.plazi.org/id/DF1918D6FF8C2B508B837414FA923E66" pageId="13" pageNumber="61" tableUuid="DF1918D6FF8C2B508B837414FA923E66">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8BD9485EFF882B548BD37440FC303F06" blockId="13.[151,1437,223,1401]" pageId="13" pageNumber="61">
The ventral lappets of the dorsal collar in the species of
<taxonomicName id="4C6633DDFF882B5488767440FC253EBA" ID-CoL="84SM9" authorityName="Chamberlin" authorityYear="1919" box="[866,998,619,645]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="genus">
<emphasis id="B912944CFF882B5488767440FC253EBA" box="[866,998,619,645]" italics="true" pageId="13" pageNumber="61">Potamethus</emphasis>
</taxonomicName>
can be triangular (
<taxonomicName id="4C6633DDFF882B548FDC7447FAB43EBA" ID-CoL="4M39W" authorityName="Hartmann-Schroder" authorityYear="1977" box="[1224,1399,619,645]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="breviuncatus">
<emphasis id="B912944CFF882B548FDC7447FAB43EBA" box="[1224,1399,619,645]" italics="true" pageId="13" pageNumber="61">P. breviuncatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B548E9C7447FDEB3E96" ID-CoL="4M39Z" authority="Hartmann-Schroder, 1977" authorityName="Hartmann-Schroder" authorityYear="1977" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="filiformis">
<emphasis id="B912944CFF882B548E9C7447FF3D3E96" italics="true" pageId="13" pageNumber="61">P. filiformis</emphasis>
<bibRefCitation id="EFF735AFFF882B548A1074A4FDEB3E96" author="Hartmann-Schroder, G." box="[260,552,655,681]" pageId="13" pageNumber="61" pagination="65 - 99" refId="ref10551" refString="Hartmann-Schroder, G. (1977) Polychaeten aus dem Sublitoral und Bathyal vor der portugiesischen und marokkanischen Kuste. Auswertung der Fahrt 8 (1967) von F. S. ' Meteor'. Meteor Forschungergebnisse, Series D, 26, 65 - 99." type="journal article" year="1977">Hartmann-Schröder, 1977</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B54892674BBFD7F3E96" ID-CoL="787KJ" authorityName="Hartman" authorityYear="1965" box="[562,700,655,681]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="singularis">
<emphasis id="B912944CFF882B54892674BBFD7F3E96" box="[562,700,655,681]" italics="true" pageId="13" pageNumber="61">P. singularis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6633DDFF882B5489E474BBFBCC3E96" authority="Ehlers, 1887" authorityName="Ehlers" authorityYear="1887" box="[752,1039,655,681]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="spatiferus">
<emphasis id="B912944CFF882B5489E474BBFCBB3E96" box="[752,888,655,681]" italics="true" pageId="13" pageNumber="61">P. spatiferus</emphasis>
<bibRefCitation id="EFF735AFFF882B54886B74A4FBCC3E96" author="Ehlers, E." box="[895,1039,655,681]" pageId="13" pageNumber="61" pagination="1 - 335" refId="ref10023" refString="Ehlers, E. (1887) Reports on the results of dredging, under the direction of L. F. Pourtales, during the years 1868 - 1870, and of Alexander Agassiz, in the Gulf of Mexico (1877 - 78), and in the Caribbean Sea (1878 - 79), in the U. S. Coast Survey steamer &quot; Blake &quot;, Lieut-Com. C. D. Sigsbee, U. S. N. and Commander J. R. Bartlett, U. S. N., commanding. XXXI. Report on the Annelids. Memoirs of the Museum of Comparative Zoology at Harvard College, 15, 1 - 335. https: // doi. org / 10.5962 / bhl. title. 65639" type="journal article" year="1887">Ehlers, 1887</bibRefCitation>
</taxonomicName>
), rounded (
<taxonomicName id="4C6633DDFF882B548F9874BBFAC13E96" box="[1164,1282,655,681]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="filitovae">
<emphasis id="B912944CFF882B548F9874BBFAC13E96" box="[1164,1282,655,681]" italics="true" pageId="13" pageNumber="61">P. filitovae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B548E1A74BBFA563E96" ID-CoL="6W7KG" authorityName="Sagami" baseAuthorityName="Johansson" baseAuthorityYear="1922" box="[1294,1429,656,681]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="japonicus">
<emphasis id="B912944CFF882B548E1A74BBFA563E96" box="[1294,1429,656,681]" italics="true" pageId="13" pageNumber="61">P. japonicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B548B83749FFE213EF2" ID-CoL="787KG" authority="(Hansen, 1878)" baseAuthorityName="Hansen" baseAuthorityYear="1878" box="[151,482,691,717]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="malmgreni">
<emphasis id="B912944CFF882B548B83749FFEE83EF2" box="[151,299,691,717]" italics="true" pageId="13" pageNumber="61">P. malmgreni</emphasis>
(Hansen, 1878)
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B548AFB749FFCEA3EF2" ID-CoL="6VVKJ" authority="Moore, 1923" authorityName="Moore" authorityYear="1923" box="[495,809,691,718]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="mucronatus">
<emphasis id="B912944CFF882B548AFB749FFD4D3EF2" box="[495,654,692,717]" italics="true" pageId="13" pageNumber="61">P. mucronatus</emphasis>
<bibRefCitation id="EFF735AFFF882B548981749FFCEA3EF2" author="Moore, J. P." box="[661,809,691,718]" pageId="13" pageNumber="61" pagination="179 - 259" refId="ref11287" refString="Moore, J. P. (1923) The polychaetous annelids dredged by the U. S. S. ' ' Albatross' ' off the coast of southern California in 1904. IV. Spionidae to Sabellariidae. Proceedings of the Academy of Natural Sciences of Philadelphia, 75, 179 - 259." type="journal article" year="1923">Moore, 1923</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B548821749FFBBA3EF2" ID-CoL="6W7KJ" authority="(McIntosh, 1916)" baseAuthorityName="McIntosh" baseAuthorityYear="1916" box="[821,1145,691,717]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="murrayi">
<emphasis id="B912944CFF882B548821749FFC683EF2" box="[821,939,692,717]" italics="true" pageId="13" pageNumber="61">P. murrayi</emphasis>
(
<bibRefCitation id="EFF735AFFF882B5488AE7498FBB13EF2" author="McIntosh, W. C." box="[954,1138,691,717]" pageId="13" pageNumber="61" pagination="1 - 68" refId="ref11148" refString="McIntosh, W. C. (1916) Notes from the Gatty Marine Laboratory, St. Andrews. No. XXXVIII. 1. On the British Sabellidae. 2. On the Sabellidae dredged by H. M. S. ' Porcupine' in 1869 and 1870, and by H. M. S. ' Knight Errant' in 1882. 3. On the Terebellidae and Sabellidae dredged in the Gulf of St. Lawrence, Canada by Dr. Whiteaves in 1871 - 73. 4. On the Sabellidae dredged by Canon A. M. Norman off Norway and Finmark. The Annals and Magazine of Natural History, Series 8, 17 (97), 1 - 68. https: // doi. org / 10.1080 / 00222931608693750" type="journal article" year="1916">McIntosh, 1916</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B548F92749FFA5F3EF2" ID-CoL="77VKJ" authority="(Pixell, 1913))" authorityName="Weddell" baseAuthorityName="Pixell" baseAuthorityYear="1913" box="[1158,1436,691,717]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="scotiae">
<emphasis id="B912944CFF882B548F92749FFB323EF2" box="[1158,1265,692,717]" italics="true" pageId="13" pageNumber="61">P. scotiae</emphasis>
(
<bibRefCitation id="EFF735AFFF882B548E147498FA493EF2" author="Pixell, H. L. M." box="[1280,1418,691,717]" pageId="13" pageNumber="61" pagination="347 - 358" refId="ref11343" refString="Pixell, H. L. M. (1913) Polychaeta of the families Serpulidae and Sabellidae, collected by the Scottish National Antarctic Expedition. Transactions of the Royal Society of Edinburgh, 49 (2), 347 - 358. https: // doi. org / 10.1017 / S 0080456800003987" type="journal article" year="1913">Pixell, 1913</bibRefCitation>
))
</taxonomicName>
or bilobed (
<taxonomicName id="4C6633DDFF882B548A0874F3FE473ECE" ID-CoL="4M39X" authorityName="Nares Deep W" baseAuthorityName="Eliason" baseAuthorityYear="1951" box="[284,388,727,753]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="dubius">
<emphasis id="B912944CFF882B548A0874F3FE473ECE" box="[284,388,727,753]" italics="true" pageId="13" pageNumber="61">P. dubius</emphasis>
</taxonomicName>
). Within each category, the length of ventral lappets might be diagnostic. For example, in
<taxonomicName id="4C6633DDFF882B548E9C74F3FED03F2A" ID-CoL="6VVKL" authorityName="Ehlers" authorityYear="1887" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="spathiferus">
<emphasis id="B912944CFF882B548E9C74F3FED03F2A" italics="true" pageId="13" pageNumber="61">P. spathiferus</emphasis>
</taxonomicName>
, the triangular ventral lappets are 2.5 times longer than the chaetiger 1 length, while in the rest of species with triangular lappets, they are short (not longer than chaetiger 1) (
<tableCitation id="C6E47DE5FF882B54889A7534FC263F06" box="[910,997,799,825]" captionStart="TABLE 2" captionStartId="9.[151,239,575,601]" captionTargetPageId="9" captionText="TABLE 2. Species of Potamethus currently considered valid, distinctive features and presence of uncinal humps." httpUri="http://table.plazi.org/id/DF1918D6FF8C2B508B837414FA923E66" pageId="13" pageNumber="61" tableUuid="DF1918D6FF8C2B508B837414FA923E66">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8BD9485EFF882B548BD37568FF353FF6" blockId="13.[151,1437,223,1401]" pageId="13" pageNumber="61">
The presence of large, brown, oval, semi-circular to circular moldures on each side of anterior peristomial ring is reported for
<taxonomicName id="4C6633DDFF882B548A547543FE753FBE" baseAuthorityName="McIntosh" baseAuthorityYear="1916" box="[320,438,872,897]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="murrayi">
<emphasis id="B912944CFF882B548A547543FE753FBE" box="[320,438,872,897]" italics="true" pageId="13" pageNumber="61">P. murrayi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B548AD17543FDEF3FBE" authorityName="Nares Deep W" baseAuthorityName="Eliason" baseAuthorityYear="1951" box="[453,556,871,897]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="dubius">
<emphasis id="B912944CFF882B548AD17543FDEF3FBE" box="[453,556,871,897]" italics="true" pageId="13" pageNumber="61">P. dubius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B54892F7543FD7A3FBE" authorityName="Tovar-Hernandez &amp; Jirkov" authorityYear="2024" baseAuthorityName="Levenstein" baseAuthorityYear="1961" box="[571,697,871,897]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="filatovae">
<emphasis id="B912944CFF882B54892F7543FD7A3FBE" box="[571,697,871,897]" italics="true" pageId="13" pageNumber="61">P. filatovae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B5489DC7543FCA23FBE" authorityName="Ehlers" authorityYear="1887" box="[712,865,871,897]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="spathiferus">
<emphasis id="B912944CFF882B5489DC7543FCA23FBE" box="[712,865,871,897]" italics="true" pageId="13" pageNumber="61">P. spathiferus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6633DDFF882B54888D7543FBED3FBE" baseAuthorityName="Hansen" baseAuthorityYear="1878" box="[921,1070,871,897]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="malmgreni">
<emphasis id="B912944CFF882B54888D7543FBED3FBE" box="[921,1070,871,897]" italics="true" pageId="13" pageNumber="61">P. malmgreni</emphasis>
</taxonomicName>
(
<tableCitation id="C6E47DE5FF882B548F29754CFB553FBE" box="[1085,1174,871,897]" captionStart="TABLE 2" captionStartId="9.[151,239,575,601]" captionTargetPageId="9" captionText="TABLE 2. Species of Potamethus currently considered valid, distinctive features and presence of uncinal humps." httpUri="http://table.plazi.org/id/DF1918D6FF8C2B508B837414FA923E66" pageId="13" pageNumber="61" tableUuid="DF1918D6FF8C2B508B837414FA923E66">Table 2</tableCitation>
). The function of these organs is unknown, but
<bibRefCitation id="EFF735AFFF882B548AA575A0FDB23F9A" author="McIntosh, W. C." box="[433,625,907,933]" pageId="13" pageNumber="61" pagination="1 - 68" refId="ref11148" refString="McIntosh, W. C. (1916) Notes from the Gatty Marine Laboratory, St. Andrews. No. XXXVIII. 1. On the British Sabellidae. 2. On the Sabellidae dredged by H. M. S. ' Porcupine' in 1869 and 1870, and by H. M. S. ' Knight Errant' in 1882. 3. On the Terebellidae and Sabellidae dredged in the Gulf of St. Lawrence, Canada by Dr. Whiteaves in 1871 - 73. 4. On the Sabellidae dredged by Canon A. M. Norman off Norway and Finmark. The Annals and Magazine of Natural History, Series 8, 17 (97), 1 - 68. https: // doi. org / 10.1080 / 00222931608693750" type="journal article" year="1916">McIntosh (1916)</bibRefCitation>
suggested that this structure is probably related to feeding in abyssal habitats.
</paragraph>
<paragraph id="8BD9485EFF882B548BD375F8FECA38D5" blockId="13.[151,1437,223,1401]" pageId="13" pageNumber="61">
The presence or absence of ventral glandular shields in species of
<taxonomicName id="4C6633DDFF882B5488B475F8FBE73FD2" authorityName="Chamberlin" authorityYear="1919" box="[928,1060,979,1005]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="genus">
<emphasis id="B912944CFF882B5488B475F8FBE73FD2" box="[928,1060,979,1005]" italics="true" pageId="13" pageNumber="61">Potamethus</emphasis>
</taxonomicName>
was already remarked by
<bibRefCitation id="EFF735AFFF882B548E5175F8FEEA382E" author="Knight-Jones, P." pageId="13" pageNumber="61" pagination="245 - 295" refId="ref10699" refString="Knight-Jones, P. (1983) Contributions to the taxonomy of Sabellidae (Polychaeta). Zoological Journal of the Linnean Society, 79 (3), 245 - 295. https: // doi. org / 10.1111 / j. 1096 - 3642.1983. tb 01167. x" type="journal article" year="1983">Knight-Jones (1983)</bibRefCitation>
. Whereas some species, such as
<taxonomicName id="4C6633DDFF882B54899F75D3FD3E382E" baseAuthorityName="Eliason" baseAuthorityYear="1951" box="[651,765,1015,1041]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="dubious">
<emphasis id="B912944CFF882B54899F75D3FD3E382E" box="[651,765,1015,1041]" italics="true" pageId="13" pageNumber="61">P. dubious</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B54881D75D3FCBC382E" box="[777,895,1015,1041]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="filitovae">
<emphasis id="B912944CFF882B54881D75D3FCBC382E" box="[777,895,1015,1041]" italics="true" pageId="13" pageNumber="61">P. filitovae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B54889F75D3FBD1382E" authorityName="Sagami" baseAuthorityName="Johansson" baseAuthorityYear="1922" box="[907,1042,1016,1041]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="japonicus">
<emphasis id="B912944CFF882B54889F75D3FBD1382E" box="[907,1042,1016,1041]" italics="true" pageId="13" pageNumber="61">P. japonicus</emphasis>
</taxonomicName>
and
<emphasis id="B912944CFF882B548F5075D3FB77382E" box="[1092,1204,1016,1041]" italics="true" pageId="13" pageNumber="61">
<taxonomicName id="4C6633DDFF882B548F5075D3FB73382E" authorityName="Weddell" baseAuthorityName="Pixell" baseAuthorityYear="1913" box="[1092,1200,1016,1041]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="scotiae">P. scotiae</taxonomicName>
,
</emphasis>
have well developed shields at least in the thorax, in others (
<taxonomicName id="4C6633DDFF882B5489487237FD25380A" authorityName="Molokai" baseAuthorityName="Treadwell" baseAuthorityYear="1906" box="[604,742,1051,1077]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="elongatus">
<emphasis id="B912944CFF882B5489487237FD25380A" box="[604,742,1051,1077]" italics="true" pageId="13" pageNumber="61">P. elongatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B5489E27237FCB9380A" authorityName="Hartmann-Schroder" authorityYear="1977" box="[758,890,1051,1077]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="filiformis">
<emphasis id="B912944CFF882B5489E27237FCB9380A" box="[758,890,1051,1077]" italics="true" pageId="13" pageNumber="61">P. filiformis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B54889D7237FBE9380A" authorityName="Moore" authorityYear="1923" box="[905,1066,1052,1077]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="mucronatus">
<emphasis id="B912944CFF882B54889D7237FBE9380A" box="[905,1066,1052,1077]" italics="true" pageId="13" pageNumber="61">P. mucronatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B548F2E7237FB72380A" baseAuthorityName="McIntosh" baseAuthorityYear="1916" box="[1082,1201,1052,1077]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="murrayi">
<emphasis id="B912944CFF882B548F2E7237FB72380A" box="[1082,1201,1052,1077]" italics="true" pageId="13" pageNumber="61">P. murrayi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6633DDFF882B548FD57237FA8D380A" authorityName="Hartman" authorityYear="1965" box="[1217,1358,1051,1077]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="singularis">
<emphasis id="B912944CFF882B548FD57237FA8D380A" box="[1217,1358,1051,1077]" italics="true" pageId="13" pageNumber="61">P. singularis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6633DDFF882B548E9C7237FED03866" authorityName="Ehlers" authorityYear="1887" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="spathiferus">
<emphasis id="B912944CFF882B548E9C7237FED03866" italics="true" pageId="13" pageNumber="61">P. spathiferus</emphasis>
</taxonomicName>
) the glandular epithelium of the entire body lacks shields. However,
<taxonomicName id="4C6633DDFF882B548F07726BFB653866" baseAuthorityName="Hansen" baseAuthorityYear="1878" box="[1043,1190,1087,1113]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="malmgreni">
<emphasis id="B912944CFF882B548F07726BFB653866" box="[1043,1190,1087,1113]" italics="true" pageId="13" pageNumber="61">P. malmgreni</emphasis>
</taxonomicName>
from the
<typeStatus id="54DDF6FCFF882B548E06726BFA813865" box="[1298,1346,1088,1114]" pageId="13" pageNumber="61">type</typeStatus>
locality (Norwegian Sea) lacks shields as reported by
<bibRefCitation id="EFF735AFFF882B5489877248FCBF3842" author="Knight-Jones, P." box="[659,892,1123,1149]" pageId="13" pageNumber="61" pagination="245 - 295" refId="ref10699" refString="Knight-Jones, P. (1983) Contributions to the taxonomy of Sabellidae (Polychaeta). Zoological Journal of the Linnean Society, 79 (3), 245 - 295. https: // doi. org / 10.1111 / j. 1096 - 3642.1983. tb 01167. x" type="journal article" year="1983">Knight-Jones (1983)</bibRefCitation>
, whereas specimens examined in this study from the Norwegian Sea and the Arctic Ocean have well-developed shields in both thorax and abdomen (
<figureCitation id="135D54DBFF882B548FE472ACFA89389E" box="[1264,1354,1159,1185]" captionStart="FIGURE 6" captionStartId="15.[151,250,1785,1809]" captionTargetBox="[172,1421,181,1761]" captionTargetId="figure-32@15.[159,1428,181,1761]" captionTargetPageId="15" captionText="FIGURE 6. Potamethus malmgreni (Hansen, 1878) from the Norwegian Sea stained with methylene blue, light microscopy images. A) Collar and base of radiolar crown, ventral view, B) same, dorsal view, C) same, ventral view, D) ventral shields and transition of thorax and abdomen, E) ventral shields from posterior abdomen, F) thoracic uncini, arrow points to uncinal hump. Abbreviations: 1, 2) number of chaetigers, apr) anterior peristomial ring, pl) parallel lamella, vsa) ventral shields of abdominal segments, vsc) ventral shield of collar, vst) ventral shield of thoracic segments. Scale bars: AC, E) 500 µm, D) 1 mm, F) not scaled." figureDoi="http://doi.org/10.5281/zenodo.13209703" httpUri="https://zenodo.org/record/13209703/files/figure.png" pageId="13" pageNumber="61">Fig. 6D</figureCitation>
). Thus, infraspecific variation of ventral shields needs to be accessed not only for
<taxonomicName id="4C6633DDFF882B5488F07287FBBA38FA" baseAuthorityName="Hansen" baseAuthorityYear="1878" box="[996,1145,1195,1221]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="malmgreni">
<emphasis id="B912944CFF882B5488F07287FBBA38FA" box="[996,1145,1195,1221]" italics="true" pageId="13" pageNumber="61">P. malmgreni</emphasis>
</taxonomicName>
, but also in all species of the genus.
</paragraph>
<paragraph id="8BD9485EFF882B548BD372D8FD403946" blockId="13.[151,1437,223,1401]" pageId="13" pageNumber="61">
Finally, even though the number of thoracic segments cannot be used to confidently distinguish species within
<taxonomicName id="4C6633DDFF882B548B83733CFECF390E" authorityName="Latreille" authorityYear="1825" box="[151,268,1303,1329]" class="Polychaeta" family="Sabellidae" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="family">Sabellidae</taxonomicName>
(
<bibRefCitation id="EFF735AFFF882B548A09733CFE5E390E" author="Capa, M." box="[285,413,1303,1329]" pageId="13" pageNumber="61" pagination="537 - 567" refId="ref9709" refString="Capa, M. (2007) Taxonomic revision and phylogenetic relationships of apomorphic sabellids (Sabellidae: Polychaeta) from Australia. Invertebrate Systematics, 21, 537 - 567. https: // doi. org / 10.1071 / IS 07002" type="journal article" year="2007">Capa 2007</bibRefCitation>
,
<bibRefCitation id="EFF735AFFF882B548ABE733CFCF3390E" author="Tovar-Hernandez, M. A. &amp; Dean, H." box="[426,816,1303,1330]" pageId="13" pageNumber="61" refId="ref11732" refString="Tovar-Hernandez, M. A. &amp; Dean, H. (2014) A new gregarious sabellid worm from the Gulf of California reproduced by spontaneous fission (Polychaeta, Sabellidae). Journal of the Marine Biological Association the United Kingdom, 94 (5)," type="journal volume" year="2014">Tovar-Hernández and Dean, 2014</bibRefCitation>
,
<bibRefCitation id="EFF735AFFF882B54882A733CFBD4390D" author="Keppel, E. &amp; Tovar-Hernandez, M. A. &amp; Ruiz, G." box="[830,1047,1303,1330]" pageId="13" pageNumber="61" pagination="499 - 518" refId="ref10635" refString="Keppel, E., Tovar-Hernandez, M. A. &amp; Ruiz, G. (2015) First record of Branchiomma coheni (Polychaeta: Sabellidae) along the US East coast and update of non indigenous species. Zootaxa, 4058 (4), 499 - 518. https: // doi. org / 10.11646 / zootaxa. 4058.4.3" type="journal article" year="2015">
Keppel
<emphasis id="B912944CFF882B54888C7333FC10390E" box="[920,979,1303,1329]" italics="true" pageId="13" pageNumber="61">et al.</emphasis>
2015
</bibRefCitation>
,
<bibRefCitation id="EFF735AFFF882B548F31733CFAB1390E" author="Tovar-Hernandez, M. A. &amp; Garcia-Garza, M. E. &amp; de Leon-Gonzalez, J. A." box="[1061,1394,1303,1329]" pageId="13" pageNumber="61" refId="ref11799" refString="Tovar-Hernandez, M. A., Garcia-Garza, M. E. &amp; de Leon-Gonzalez, J. A. (2020) Sclerozoan and fouling sabellid worms (Annelida: Sabellidae) from Mexico with the establishment of two new species. Biodiversity Data Journal, 8, e 57471. https: // doi. org / 10.3897 / BDJ. 8. e 57471" type="journal volume" year="2020">
Tovar-Hernández
<emphasis id="B912944CFF882B548FE57333FAEF390E" box="[1265,1324,1303,1329]" italics="true" pageId="13" pageNumber="61">et al.</emphasis>
2020
</bibRefCitation>
),
<taxonomicName id="4C6633DDFF882B548E9C7333FEC7396A" authorityName="Sagami" baseAuthorityName="Johansson" baseAuthorityYear="1922" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="japonicus">
<emphasis id="B912944CFF882B548E9C7333FEC7396A" italics="true" pageId="13" pageNumber="61">P. japonicus</emphasis>
</taxonomicName>
was described with 17 thoracic segments,
<taxonomicName id="4C6633DDFF882B5489FE7317FC55396A" authorityName="Hartmann-Schroder" authorityYear="1977" box="[746,918,1339,1365]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="breviuncatus">
<emphasis id="B912944CFF882B5489FE7317FC55396A" box="[746,918,1339,1365]" italics="true" pageId="13" pageNumber="61">P. breviuncatus</emphasis>
</taxonomicName>
with 10,
<taxonomicName id="4C6633DDFF882B548F177317FB61396A" authorityName="Moore" authorityYear="1923" box="[1027,1186,1340,1365]" class="Polychaeta" family="Sabellidae" genus="Potamethus" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="61" phylum="Annelida" rank="species" species="mucronatus">
<emphasis id="B912944CFF882B548F177317FB61396A" box="[1027,1186,1340,1365]" italics="true" pageId="13" pageNumber="61">P. mucronatus</emphasis>
</taxonomicName>
with 9 and the rest of species with 8 thoracic chaetigers (
<tableCitation id="C6E47DE5FF882B54890A7374FDB53946" box="[542,630,1375,1401]" captionStart="TABLE 2" captionStartId="9.[151,239,575,601]" captionTargetPageId="9" captionText="TABLE 2. Species of Potamethus currently considered valid, distinctive features and presence of uncinal humps." httpUri="http://table.plazi.org/id/DF1918D6FF8C2B508B837414FA923E66" pageId="13" pageNumber="61" tableUuid="DF1918D6FF8C2B508B837414FA923E66">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>