120 lines
14 KiB
XML
120 lines
14 KiB
XML
<document id="6FA53969674C06D5903C8F4D4ADE3694" ID-CLB-Dataset="45917" ID-DOI="10.5281/zenodo.279260" ID-GBIF-Dataset="ffbcaa56-6a77-4df4-b944-508fb074171f" ID-ISSN="1175-5326" ID-Zenodo-Dep="279260" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460413266680" checkinUser="plazi" docAuthor="Shear, William A." docDate="2011" docId="03C47A60FFCA725D60C05C9F7BEA7977" docLanguage="en" docName="zt03109p038.pdf" docOrigin="Zootaxa 3109" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pseudotremia culveri Shear, 2011, n. sp." docType="treatment" docVersion="9" lastPageNumber="32" masterDocId="FFFD0218FFD5724260575C087F167D21" masterDocTitle="Cave millipeds of the United States. X. New species and records of the genus Pseudotremia Cope. 2. Species from Virginia, USA (Diplopoda, Chordeumatida, Cleidogonidae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="32" updateTime="1698301493022" updateUser="ExternalLinkService">
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<mods:titleInfo id="F140B28E1770C91E7E3FE5DE7B61EEA3">
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<mods:title id="293CCE84814286DDA6895680E691E59D">Cave millipeds of the United States. X. New species and records of the genus Pseudotremia Cope. 2. Species from Virginia, USA (Diplopoda, Chordeumatida, Cleidogonidae)</mods:title>
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<mods:name id="6AEC0F7F862945DA3CB313EBE50ACC22" type="personal">
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<mods:roleTerm id="3AB389B2DF0F68DE44B32524E5E1D667">Author</mods:roleTerm>
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<mods:namePart id="B52A11118FC28B570D122BC8AD3E8703">Shear, William A.</mods:namePart>
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<mods:typeOfResource id="0E6E86337E4C1557FCD5DD2D17ED6FC2">text</mods:typeOfResource>
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<mods:titleInfo id="4109ABB0F6CB701AF20D5D738CBA19C9">
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<mods:title id="32C0011D37DD455A0DA06EADC272B786">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="571CC74339D6A05B512A42174DA63257">
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<mods:date id="E882610A5380C31932E34005AE72958E">2011</mods:date>
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<mods:detail id="17DEB8FC252667C4EDDFFED4B5D61A1F" type="volume">
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<mods:number id="6506F3C6470674FE3E38F89CB6F14AEF">3109</mods:number>
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<mods:start id="96046DE73966CEB6EE4F27B435986B73">1</mods:start>
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<mods:classification id="51E51FE829C45309AE91750C8C86177D">journal article</mods:classification>
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<mods:identifier id="025CF5732D03D39265FA9050991B7B48" type="CLB-Dataset">45917</mods:identifier>
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<mods:identifier id="18379AC96F7A35AF781068B5DBA04A42" type="DOI">10.5281/zenodo.279260</mods:identifier>
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<mods:identifier id="09483A4555AA56DC7AC7B33861A0170D" type="GBIF-Dataset">ffbcaa56-6a77-4df4-b944-508fb074171f</mods:identifier>
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<mods:identifier id="18FBF4EC8F196111128F136FE6136625" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B3C5D7403C493560CA1335990DF1D84B" type="Zenodo-Dep">279260</mods:identifier>
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<treatment id="03C47A60FFCA725D60C05C9F7BEA7977" ID-DOI="http://doi.org/10.5281/zenodo.6186463" ID-GBIF-Taxon="119424130" ID-Zenodo-Dep="6186463" LSID="urn:lsid:plazi:treatment:03C47A60FFCA725D60C05C9F7BEA7977" httpUri="http://treatment.plazi.org/id/03C47A60FFCA725D60C05C9F7BEA7977" lastPageNumber="32" pageId="31" pageNumber="32">
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<subSubSection id="C37798FDFFCA725D60C05C9F7CA17C1F" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="8BD2CB76FFCA725D60C05C9F7EFE7D90" blockId="31.[151,488,151,210]" box="[151,488,151,177]" pageId="31" pageNumber="32">
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<heading id="D09A7C1AFFCA725D60C05C9F7EFE7D90" bold="true" box="[151,488,151,177]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
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<emphasis id="B9191764FFCA725D60C05C9F7EFE7D90" bold="true" box="[151,488,151,177]" pageId="31" pageNumber="32">
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<emphasis id="B9191764FFCA725D60C05C9F7E887D90" bold="true" box="[151,414,151,177]" italics="true" pageId="31" pageNumber="32">
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<taxonomicName id="4C6DB0F5FFCA725D60C05C9F7E8E7D90" box="[151,408,151,177]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="culveri" status="sp. nov.">Pseudotremia culveri</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A22AAA1FFFCA725D61F15C9F7EFE7D90" box="[422,488,151,177]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD2CB76FFCA725D60C05CB27E3F7DF3" blockId="31.[151,488,151,210]" box="[151,297,186,210]" pageId="31" pageNumber="32">
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<figureCitation id="1356D7F3FFCA725D60C05CB27E3F7DF3" box="[151,297,186,210]" captionStart-0="FIGURES 92 – 98" captionStart-1="FIGURES 99, 100" captionStart-2="FIGURES 101 – 109" captionStartId-0="30.[151,264,1924,1947]" captionStartId-1="31.[151,264,1973,1996]" captionStartId-2="32.[151,264,1919,1942]" captionTargetBox-0="[169,1417,155,1900]" captionTargetBox-1="[167,1424,1183,1941]" captionTargetBox-2="[151,1436,195,1892]" captionTargetId-0="figure@30.[169,1417,155,1901]" captionTargetId-1="figure@31.[151,1435,1166,1949]" captionTargetId-2="figure@32.[151,1436,195,1893]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionTargetPageId-2="32" captionText-0="FIGURES 92 – 98. New species of Pseudotremia. Figs. 92 – 97, Pseudotremia piscator, n. sp. Fig. 92, ocelli. Fig. 93, metazonite, lateral view. Fig. 94, gonopods, lateral view. Fig. 95, gonopods, anterior view. Fig. 96, gonopods, ventral view. Fig. 97, right ninth leg, anterior view. Fig. 98, P. culveri, n. sp., eleventh coxae, posterior view." captionText-1="FIGURES 99, 100. Pseudotremia culveri, n, sp. Fig. 99, gonopods, anterior view. Fig. 100, gonopods, lateral view." captionText-2="FIGURES 101 – 109. New species of Pseudotremia. Figs. 101 – 104, Pseudotremia culveri, n. sp. Fig. 101, ocelli. Fig. 102, metazonite, dorsal view. Fig. 103, left ninth leg, anterior view. Fig. 104, metazonite, lateral view. Figs. 105 – 109, Pseudotremia salfodina, n. sp. Fig. 105, left ninth leg, anterior view. Fig. 106, gonopods, lateral view. Fig. 107, gonopods, anterior view. Fig. 108, gonopods, ventral view. Fig. 109, ocelli." httpUri-0="https://zenodo.org/record/279275/files/figure.png" httpUri-1="https://zenodo.org/record/279276/files/figure.png" httpUri-2="https://zenodo.org/record/279277/files/figure.png" pageId="31" pageNumber="32">Figs. 98–104</figureCitation>
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</paragraph>
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<paragraph id="8BD2CB76FFCA725D60C05D0B7CA17C1F" blockId="31.[151,1436,258,318]" pageId="31" pageNumber="32">
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<emphasis id="B9191764FFCA725D60C05D0B7FF57C3A" bold="true" box="[151,227,259,283]" pageId="31" pageNumber="32">
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<typeStatus id="54D675D4FFCA725D60C05D0B7FCC7C3A" box="[151,218,259,283]" pageId="31" pageNumber="32">Types</typeStatus>
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:
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</emphasis>
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Male
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<typeStatus id="54D675D4FFCA725D61795D0A7E867C3B" box="[302,400,258,282]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
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and female
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<typeStatus id="54D675D4FFCA725D624B5D0B7D917C3A" box="[540,647,259,283]" pageId="31" pageNumber="32" type="paratype">paratypes</typeStatus>
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from Fisher Cave,
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<quantity id="4C956693FFCA725D63355D0A7CB87C3B" box="[866,942,258,282]" metricMagnitude="3" metricUnit="m" metricValue="5.632704" pageId="31" pageNumber="32" unit="mi" value="3.5">3.5 mi</quantity>
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. NNW of Kyles Ford,
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<emphasis id="B9191764FFCA725D64F85D0C7A187C3B" box="[1199,1294,259,283]" italics="true" pageId="31" pageNumber="32">Lee Co.,</emphasis>
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VIRGINIA, collected by J. R. Holsinger, T. C. Barr, Jr., T. C. Barr III,
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<date id="FFD3EDB6FFCA725D63485D2E7CA57C1F" box="[799,947,294,318]" pageId="31" pageNumber="32" value="1979-07-17">17 July 1979</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37798FDFFCA725D60905D667C027CD3" pageId="31" pageNumber="32" type="diagnosis">
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<paragraph id="8BD2CB76FFCA725D60905D667C027CD3" blockId="31.[151,1437,366,1111]" pageId="31" pageNumber="32">
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<emphasis id="B9191764FFCA725D60905D667E557CA6" bold="true" box="[199,323,366,391]" pageId="31" pageNumber="32">Diagnosis:</emphasis>
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Like
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<taxonomicName id="4C6DB0F5FFCA725D61D15D787EE57CA7" box="[390,499,366,391]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nodosa">
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<emphasis id="B9191764FFCA725D61D15D787EE57CA7" box="[390,499,366,391]" italics="true" pageId="31" pageNumber="32">P. nodosa</emphasis>
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</taxonomicName>
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from its
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<typeStatus id="54D675D4FFCA725D620C5D677D9D7CA6" box="[603,651,367,391]" pageId="31" pageNumber="32">type</typeStatus>
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locality, specimens of this species have nodules across the back margins of the metazonites, but often only four to six (fig. 102); the gonopods are distinguished from those of
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<taxonomicName id="4C6DB0F5FFCA725D651C5D9B7A8A7C8A" box="[1355,1436,403,427]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nodosa">
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<emphasis id="B9191764FFCA725D651C5D9B7A8A7C8A" box="[1355,1436,403,427]" italics="true" pageId="31" pageNumber="32">nodosa</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C6DB0F5FFCA725D60915DBF7E517CEF" box="[198,327,439,462]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="inexpectata">
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<emphasis id="B9191764FFCA725D60915DBF7E517CEF" box="[198,327,439,462]" italics="true" pageId="31" pageNumber="32">inexpectata</emphasis>
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</taxonomicName>
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by the double-pronged, but short and massive, VCP (fig. 100), and the ninth legs (fig. 103) are also quite different from those found in the other two species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37798FDFFCA725D60905DF77EB07F7F" pageId="31" pageNumber="32" type="etymology">
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<paragraph id="8BD2CB76FFCA725D60905DF77EB07F7F" blockId="31.[151,1437,366,1111]" pageId="31" pageNumber="32">
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<emphasis id="B9191764FFCA725D60905DF77E477F36" bold="true" box="[199,337,511,535]" pageId="31" pageNumber="32">Etymology:</emphasis>
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The species epithet honors David Culver, dean of American speleobiologists, in gratitude for all he has done to make possible the publication of several of the papers in this series. Suggested vernacular name: Culver’s Cave Milliped.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37798FDFFCA725D60905E627BEA7977" pageId="31" pageNumber="32" type="description">
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<paragraph id="8BD2CB76FFCA725D60905E627A5B7FCF" blockId="31.[151,1437,366,1111]" pageId="31" pageNumber="32">
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<emphasis id="B9191764FFCA725D60905E627E4D7FA2" bold="true" box="[199,347,618,643]" pageId="31" pageNumber="32">Description:</emphasis>
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Male
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<typeStatus id="54D675D4FFCA725D61F55E627D127FA3" box="[418,516,618,642]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
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<quantity id="4C956693FFCA725D625C5E627D4C7FA3" box="[523,602,618,642]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="31" pageNumber="32" unit="mm" value="17.0">17 mm</quantity>
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long,
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<quantity id="4C956693FFCA725D62C85E627DE37FA3" box="[671,757,618,642]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="31" pageNumber="32" unit="mm" value="1.7">1.7 mm</quantity>
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wide. 10 poorly defined unpigmented ocelli on each side of head (fig. 101), may appear eyeless under dissecting light microscope. Metazonites with well-developed shoulders anteriorly, shoulders reduced, segments cylindrical posteriorly, dorsum smooth, with 4–6 round nodules on posterior margins of midbody segments (figs. 102, 104); 6–8 lateral striae, mostly incomplete. Without pigment.
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</paragraph>
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<paragraph id="8BD2CB76FFCA725D60905EF27CC67ECB" blockId="31.[151,1437,366,1111]" pageId="31" pageNumber="32">Gonopods (figs. 99, 100) somewhat reduced; angiocoxites widely separated at base, sharply converging distally, MAPs with short but stout preapical spine, apical spine absent; LAPs short, evenly curved, distally narrowing, not divided (fig. 100). Colpocoxites large, erect, distinctly mitten-shaped, “hand” of mitten narrow, diverging (figs. 99). VCP large, two-branched, branches semicircularly curved (fig. 99). Ninth legs (fig.103) with distal segment slightly shorter than basal, traces of articulations of postfemoral segments present; coxoprefemur sharply constricted at point of fusion of coxa and prefemur, some degree of articulation possible, coxal portion with squarish median knob, prefemur with peglike knob just distal to midway in length.</paragraph>
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<paragraph id="8BD2CB76FFCA725D60905FFE7EC6792F" blockId="31.[151,1437,366,1111]" box="[199,464,1014,1038]" pageId="31" pageNumber="32">Female similar to male.</paragraph>
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<paragraph id="8BD2CB76FFCA725D609058137BEA7977" blockId="31.[151,1437,366,1111]" pageId="31" pageNumber="32">
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<emphasis id="B9191764FFCA725D609058137E057912" bold="true" box="[199,275,1051,1075]" pageId="31" pageNumber="32">Notes:</emphasis>
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At the
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<typeStatus id="54D675D4FFCA725D613358137E827912" box="[356,404,1051,1075]" pageId="31" pageNumber="32">type</typeStatus>
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locality, this species is syntopic with
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<taxonomicName id="4C6DB0F5FFCA725D636F58147CB97912" box="[824,943,1052,1075]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="piscator" status="sp. nov.">
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<emphasis id="B9191764FFCA725D636F58147CB97912" box="[824,943,1052,1075]" italics="true" pageId="31" pageNumber="32">P. piscator</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22AAA1FFFCA725D63E258137CE47913" box="[949,1010,1051,1074]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
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, but cannot be confused with it as that species is a larger, pigmented, troglophilic form with distinctive gonopods and a very rough dorsum.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |