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<mods:title id="30477790514E87179517F4540B1D9EB9">Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera</mods:title>
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<mods:namePart id="569B0D0B7223CBA89935BB052D754088">Winterton, Shaun L.</mods:namePart>
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<mods:namePart id="F71DBDC66D6034388CC5B5769B1015F8">Martins, Caleb Califre</mods:namePart>
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<treatment id="03C47176FFA98D1F7AD201D4FCF69025" ID-DOI="http://doi.org/10.5281/zenodo.5631465" ID-GBIF-Taxon="156146937" ID-Zenodo-Dep="5631465" LSID="urn:lsid:plazi:treatment:03C47176FFA98D1F7AD201D4FCF69025" httpUri="http://treatment.plazi.org/id/03C47176FFA98D1F7AD201D4FCF69025" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<subSubSection id="C37793EBFFA98D1E7AD201D4FE8794F2" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="8BD2C060FFA98D1E7AD201D4FE499493" blockId="42.[151,489,1684,1743]" box="[151,489,1684,1710]" pageId="42" pageNumber="43">
<heading id="D09A770CFFA98D1E7AD201D4FE499493" bold="true" box="[151,489,1684,1710]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<taxonomicName id="4C6DBBE3FFA98D1E7AD201D4FE499493" ID-CoL="63Q8H" ID-ENA="560941" authority="McLachlan, 1867" authorityName="McLachlan" authorityYear="1867" box="[151,489,1684,1710]" class="Insecta" family="Osmylidae" genus="Porismus" kingdom="Animalia" order="Neuroptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FFA98D1E7AD201D4FE499493" bold="true" box="[151,489,1684,1710]" pageId="42" pageNumber="43">
<emphasis id="B9191C72FFA98D1E7AD201D4FEA99493" bold="true" box="[151,265,1684,1710]" italics="true" pageId="42" pageNumber="43">Porismus</emphasis>
<bibRefCitation id="EFFCBD91FFA98D1E7B5501D4FE499493" author="McLachlan" box="[272,489,1684,1710]" firstAuthor="McLachlan" pageId="42" pageNumber="43" pagination="230 - 281" refId="ref51479" refString="McLachlan, R. (1867) [1868] New genera and species, &amp; c., of neuropterous insects; and a revision of Mr. F. Walker's British Museum Catalogue of Neuroptera, part ii. (1853), as far as the end of the genus Myrmeleon. Journal of the Linnean Society of London, Zoology, 9, 230 - 281. https: // doi. org / 10.1111 / j. 1096 - 3642.1867. tb 00195. x" type="journal article" year="1867">McLachlan, 1867</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8BD2C060FFA98D1E7AD201F6FE8794F2" blockId="42.[151,489,1684,1743]" box="[151,295,1718,1743]" pageId="42" pageNumber="43">(Figs 2426)</paragraph>
</subSubSection>
<subSubSection id="C37793EBFFA98D1E7AD201BEFCCC9529" box="[151,876,1790,1813]" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8BD2C060FFA98D1E7AD201BEFCCC9529" blockId="42.[151,876,1790,1813]" box="[151,876,1790,1813]" pageId="42" pageNumber="43">
<emphasis id="B9191C72FFA98D1E7AD201BEFE829529" bold="true" box="[151,290,1790,1812]" pageId="42" pageNumber="43">
<typeStatus id="54D67EC2FFA98D1E7AD201BEFF6C9529" box="[151,204,1790,1812]" pageId="42" pageNumber="43">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C6DBBE3FFA98D1E7B6C01BEFD689529" authority="Burmeister, 1839: 184" authorityName="Burmeister" authorityPageNumber="184" authorityYear="1839" box="[297,712,1790,1813]" class="Insecta" family="Osmylidae" genus="Osmylus" kingdom="Animalia" order="Neuroptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="strigatus">
<emphasis id="B9191C72FFA98D1E7B6C01BEFE409528" box="[297,480,1790,1813]" italics="true" pageId="42" pageNumber="43">Osmylus strigatus</emphasis>
<bibRefCitation id="EFFCBD91FFA98D1E7BA301BEFD389529" author="Burmeister" box="[486,664,1790,1812]" firstAuthor="Burmeister" pageId="42" pageNumber="43" pagination="757 - 1050" refId="ref47828" refString="Burmeister, H. C. C. (1839) Handbuch der Entomologie. Zweiter Band. Besondere Entomologie. Zweite Abtheilung. Kaukerfe. Gymnognatha. Theod. Chr. Friedr. Enslin, Berlin, xii + 294 pp. [pp. i-xii + 757 - 1050]" type="book chapter" year="1839">Burmeister, 1839</bibRefCitation>
: 184
</taxonomicName>
(by monotypy).
</paragraph>
</subSubSection>
<subSubSection id="C37793EBFFA98D1E7AD2007CFAC295D8" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="8BD2C060FFA98D1E7AD2007CFAC295D8" blockId="42.[151,1436,1852,2021]" pageId="42" pageNumber="43">
<emphasis id="B9191C72FFA98D1E7AD2007CFEB19568" bold="true" box="[151,273,1852,1877]" pageId="42" pageNumber="43">Diagnosis.</emphasis>
Wings with highly distinctive contrasting colour pattern; wings ovoid; body glossy black with head bright orange; RP with numerous crossveins irregularly arranged; CuA and CuP strongly sinuous in both wings, branching near wing margin in FW; HW CuA with few dichotomous branches; female forecoxa with 1115 short pedicellate setae and a prominent preapical process on dorsal surface; female tergite 9 strongly produced posteroventrally; sternite 8 with posterior end forked and with two small setose lobes; spermathecae rounded.
</paragraph>
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<subSubSection id="C37793EBFFA88D1F7A8207D7FCF69025" pageId="43" pageNumber="44" type="discussion">
<paragraph id="8BD2C060FFA88D1F7A8207D7FCF69025" blockId="43.[151,1437,151,536]" pageId="43" pageNumber="44">
<emphasis id="B9191C72FFA88D1F7A8207D7FEE8928D" bold="true" box="[199,328,151,176]" pageId="43" pageNumber="44">Comments</emphasis>
. Commonly called the pied lacewing this highly distinctive species is placed in a separate subfamily based on a series of autapomorphies features in the wings such as the branching pattern of medial and cubital veins, lack of gradate series crossveins and multiple
<emphasis id="B9191C72FFA88D1F790407A1FCCD92C5" box="[833,877,225,248]" italics="true" pageId="43" pageNumber="44">sc-r</emphasis>
crossveins.
<taxonomicName id="4C6DBBE3FFA88D1F79BD07A1FB6C92C5" box="[1016,1228,225,248]" class="Insecta" family="Osmylidae" genus="Porismus" kingdom="Animalia" order="Neuroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="strigatus">
<emphasis id="B9191C72FFA88D1F79BD07A1FB6C92C5" box="[1016,1228,225,248]" italics="true" pageId="43" pageNumber="44">Porismus strigatus</emphasis>
</taxonomicName>
is associated with
<emphasis id="B9191C72FFA88D1F7AD20644FEB49320" box="[151,276,260,285]" italics="true" pageId="43" pageNumber="44">Eucalyotus</emphasis>
trees during the autumn months in eastern
<collectingCountry id="F37A80F0FFA88D1F78BD0644FCFE9320" box="[760,862,260,285]" name="Australia" pageId="43" pageNumber="44">Australia</collectingCountry>
when adults have been observed ovipositing under tree bark; larvae are sub-corticolous on these trees and not associated with riparian habitats (
<bibRefCitation id="EFFCBD91FFA88D1F7E9E0667FF749359" author="Winterton" firstAuthor="Winterton" pageId="43" pageNumber="44" pagination="555 - 574" refId="ref55173" refString="Winterton, S. L., Zhao, J., Garzon-Orduna, I., Wang, Y. &amp; Liu, Z. (2017) Phylogeny of lance lacewings (Neuroptera: Osmylidae). Systematic Entomology, 42, 555 - 574. https: // doi. org / 10.1111 / syen. 12231" type="journal article" year="2017">
Winterton
<emphasis id="B9191C72FFA88D1F7F1C0669FA35937D" box="[1369,1429,295,320]" italics="true" pageId="43" pageNumber="44">et al.</emphasis>
, 2017
</bibRefCitation>
).
<bibRefCitation id="EFFCBD91FFA88D1F7AAD060DFE2A9359" author="Kruger" box="[232,394,332,357]" firstAuthor="Kruger" pageId="43" pageNumber="44" pagination="3 - 123" refId="ref49715" refString="Kruger, L. (1913 a) Osmylidae. Beitrage zu einer Monographie der Neuropteren-Familie der Osmyliden. II. Charakteristik der Familie, Unterfamilien und Gattungen auf Grund des Geaders. Stettiner Entomologische Zeitung, 74, 3 - 123." type="journal article" year="1913">Krüger (1913)</bibRefCitation>
erected Porisminae along with Eidoporisminae as separate subfamilies based on distinctive venational characters, and although the male and female abdominal sclerites and genitalic morphology reflect a close relationship with
<taxonomicName id="4C6DBBE3FFA88D1F7BDC06D4FDE19390" authorityName="Kruger" authorityYear="1913" box="[409,577,404,429]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="subFamily" subFamily="Stenosmylinae">Stenosmylinae</taxonomicName>
, their status as separate subfamilies is maintained (discussed in
<bibRefCitation id="EFFCBD91FFA88D1F7F4306D5FA309390" author="New" box="[1286,1424,404,429]" firstAuthor="New" pageId="43" pageNumber="44" pagination="763 - 770" refId="ref52676" refString="New, T. R. (1983 b) Revision of the osmylid subfamilies Porisminae and Eidoporisminae (Insecta: Neuroptera). Australian Journal of Zoology, 31, 763 - 770. https: // doi. org / 10.1071 / ZO 9830763" type="journal article" year="1983 b">New, 1983b</bibRefCitation>
).
<bibRefCitation id="EFFCBD91FFA88D1F7AD206F7FE1693ED" author="Winterton" box="[151,438,439,464]" firstAuthor="Winterton" pageId="43" pageNumber="44" pagination="555 - 574" refId="ref55173" refString="Winterton, S. L., Zhao, J., Garzon-Orduna, I., Wang, Y. &amp; Liu, Z. (2017) Phylogeny of lance lacewings (Neuroptera: Osmylidae). Systematic Entomology, 42, 555 - 574. https: // doi. org / 10.1111 / syen. 12231" type="journal article" year="2017">
Winterton
<emphasis id="B9191C72FFA88D1F7B5F06F9FEFF93ED" box="[282,351,439,464]" italics="true" pageId="43" pageNumber="44">et al.</emphasis>
(2017
</bibRefCitation>
a,b) also concluded that Porisminae was closely related to Eidoporisminae and
<taxonomicName id="4C6DBBE3FFA88D1F7AD2069CFE9E93C8" authorityName="Kruger" authorityYear="1913" box="[151,318,476,501]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="subFamily" subFamily="Stenosmylinae">Stenosmylinae</taxonomicName>
, consistent with previous proposals (
<bibRefCitation id="EFFCBD91FFA88D1F78BD069CFC5593C9" author="Esben-Petersen" box="[760,1013,476,501]" firstAuthor="Esben-Petersen" pageId="43" pageNumber="44" pagination="1 - 5" refId="ref48410" refString="Esben-Petersen, P. (1917) New and little-known Australian Osmylidae. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, Kobenhaven, 68, 1 - 5." type="journal article" year="1917">Esben-Petersen, 1917</bibRefCitation>
,
<bibRefCitation id="EFFCBD91FFA88D1F7E43069CFB1D93C9" author="Kimmins" box="[1030,1213,476,501]" firstAuthor="Kimmins" pageId="43" pageNumber="44" pagination="165 - 201" refId="ref49518" refString="Kimmins, D. E. (1940) A revision of the osmylid subfamilies Stenosmylinae and Kalosmylinae (Neuroptera). Novitates Zoologicae, 42, 165 - 201." type="journal article" year="1940">Kimmins, 1940</bibRefCitation>
,
<bibRefCitation id="EFFCBD91FFA88D1F7E8B069DFAFF93C8" author="New" box="[1230,1375,476,501]" firstAuthor="New" pageId="43" pageNumber="44" pagination="763 - 770" refId="ref52676" refString="New, T. R. (1983 b) Revision of the osmylid subfamilies Porisminae and Eidoporisminae (Insecta: Neuroptera). Australian Journal of Zoology, 31, 763 - 770. https: // doi. org / 10.1071 / ZO 9830763" type="journal article" year="1983 b">New, 1983b</bibRefCitation>
), but relationships amongst the three subfamilies were inconclusive.
</paragraph>
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