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<document id="694375D9CB033E724D03D25982F12075" ID-DOI="10.5281/zenodo.188099" ID-GBIF-Dataset="ea3cfb7d-d5ca-4ea2-8f88-8e9e43df3b69" ID-ISSN="1175-5326" ID-Zenodo-Dep="188099" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461083685542" checkinUser="plazi" docAuthor="Hastings, Philip A. &amp; Springer, Victor G." docDate="2009" docId="03C43C38FFC63D4081A0172AFA43FA3D" docLanguage="en" docName="zt02120p014.pdf" docOrigin="Zootaxa 2120" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Blenniidae" docType="treatment" docVersion="8" lastPageNumber="8" masterDocId="FFFD4440FFC53D4581371155FFC4FFBA" masterDocTitle="Recognizing diversity in blennioid fish nomenclature (Teleostei: Blennioidei)" masterLastPageNumber="14" masterPageNumber="3" pageNumber="6" updateTime="1698587644791" updateUser="plazi">
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<mods:title id="E644927C9333449E41493A4001467E3A">Recognizing diversity in blennioid fish nomenclature (Teleostei: Blennioidei)</mods:title>
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<mods:namePart id="A57D93D5DD80C9C24E7321D655E1854A">Hastings, Philip A.</mods:namePart>
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<mods:namePart id="112350BCAA5ADCB4B81C3CE47D0C272D">Springer, Victor G.</mods:namePart>
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<mods:date id="4D94DB8C6472C72B9B00EE0FDAF89F56">2009</mods:date>
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<treatment id="03C43C38FFC63D4081A0172AFA43FA3D" ID-DOI="http://doi.org/10.5281/zenodo.5631531" ID-GBIF-Taxon="119618506" ID-Zenodo-Dep="5631531" LSID="urn:lsid:plazi:treatment:03C43C38FFC63D4081A0172AFA43FA3D" httpUri="http://treatment.plazi.org/id/03C43C38FFC63D4081A0172AFA43FA3D" lastPageId="5" lastPageNumber="8" pageId="3" pageNumber="6">
<subSubSection id="C377DEA5FFC63D4181A0172AFCF6FEF6" lastPageId="4" lastPageNumber="7" pageId="3" pageNumber="6" type="nomenclature">
<paragraph id="8BD28D2EFFC63D4681A0172AFED9F923" blockId="3.[151,285,1663,1689]" box="[151,285,1663,1689]" pageId="3" pageNumber="6">
<heading id="D09A3A42FFC63D4681A0172AFED9F923" bold="true" box="[151,285,1663,1689]" fontSize="11" level="1" pageId="3" pageNumber="6" reason="1">
<taxonomicName id="4C6DF6ADFFC63D4681A0172AFED9F923" ID-CoL="78Y" ID-ENA="56718" box="[151,285,1663,1689]" class="Actinopterygii" family="Blenniidae" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="6" phylum="Chordata" rank="family">
<emphasis id="B919513CFFC63D4681A0172AFED9F923" bold="true" box="[151,285,1663,1689]" pageId="3" pageNumber="6">Blenniidae</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD28D2EFFC63D4181F11798FCF6FEF6" blockId="3.[151,1437,1741,2038]" lastBlockId="4.[151,1437,152,1962]" lastPageId="4" lastPageNumber="7" pageId="3" pageNumber="6">
1)
<taxonomicName id="4C6DF6ADFFC63D4681D11798FEACF95C" box="[230,360,1741,1766]" class="Actinopterygii" family="Blenniidae" genus="Aspidontus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC63D4681D11798FEACF95C" box="[230,360,1741,1766]" italics="true" pageId="3" pageNumber="6">Aspidontus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C6DF6ADFFC63D46804F1798FC87F95D" authority="Quoy &amp; Gaimard" authorityName="Quoy &amp; Gaimard" box="[376,835,1741,1767]" class="Actinopterygii" family="Blenniidae" genus="Aspidontus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="6" phylum="Chordata" rank="species" species="taeniatus">
<emphasis id="B919513CFFC63D46804F1798FDA9F95C" box="[376,621,1741,1766]" italics="true" pageId="3" pageNumber="6">Aspidontus taeniatus</emphasis>
Quoy &amp; Gaimard
</taxonomicName>
(
<typeStatus id="54D6338CFFC63D4682631798FC4CF95D" box="[852,904,1741,1767]" pageId="3" pageNumber="6">type</typeStatus>
locality:
<collectingCountry id="F37ACDBEFFC63D4685371798FB8FF95D" box="[1024,1099,1741,1767]" name="Guam" pageId="3" pageNumber="6">Guam</collectingCountry>
,
<collectingCountry id="F37ACDBEFFC63D4685601798FAE5F95D" box="[1111,1313,1741,1767]" name="Northern Mariana Islands" pageId="3" pageNumber="6">Marianas Islands</collectingCountry>
) is widely distributed across the Indo-west and central Pacific where it mimics the cleaner wrasse
<taxonomicName id="4C6DF6ADFFC63D46859517A1FE8DF88E" authority="Valenciennes" authorityName="Valenciennes" class="Actinopterygii" family="Labridae" genus="Labroides" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="6" phylum="Chordata" rank="species" species="dimidiatus">
<emphasis id="B919513CFFC63D46859517A1FA58F8B7" box="[1186,1436,1780,1805]" italics="true" pageId="3" pageNumber="6">Labroides dimidiatus</emphasis>
(Valenciennes)
</taxonomicName>
(
<bibRefCitation id="EFFCF0DFFFC63D46806E164FFD0BF88E" author="Springer" box="[345,719,1818,1844]" pageId="3" pageNumber="14" refString="Springer, V. G. &amp; Smith-Vaniz, W. F. (1972) Mimetic relationships involving fishes of the family Blenniidae. Smithsonian Contributions to Zoology, 112, 1 - 36." type="journal article" year="1972">Springer &amp; Smith-Vaniz, 1972</bibRefCitation>
;
<bibRefCitation id="EFFCF0DFFFC63D4683EB164FFC05F88E" author="Smith-Vaniz" box="[732,961,1818,1844]" pageId="3" pageNumber="13" refString="Smith-Vaniz, W. F. (1976) The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae). Monographs of the Academy of Natural Sciences of Philadelphia, 19, 1 - 196." type="journal article" year="1976">Smith-Vaniz, 1976</bibRefCitation>
,
<bibRefCitation id="EFFCF0DFFFC63D4682FA164FFBCAF88E" author="Smith-Vaniz" box="[973,1038,1818,1844]" pageId="3" pageNumber="13" refString="Smith-Vaniz, W. F. (1987) The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae): an update. Proceedings Academy Natural Sciences Philadelphia, 139, 1 - 52." type="journal article" year="1987">1987</bibRefCitation>
).
<bibRefCitation id="EFFCF0DFFFC63D468514164FFAD5F88E" author="Smith-Vaniz" box="[1059,1297,1818,1844]" pageId="3" pageNumber="13" refString="Smith-Vaniz, W. F. (1976) The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae). Monographs of the Academy of Natural Sciences of Philadelphia, 19, 1 - 196." type="journal article" year="1976">Smith-Vaniz (1976)</bibRefCitation>
recognized two subspecies, the nominate occurring throughout the western and central Pacific, and
<emphasis id="B919513CFFC63D4685831614FAFBF8E0" box="[1204,1343,1857,1882]" italics="true" pageId="3" pageNumber="6">
A. t.
<taxonomicName id="4C6DF6ADFFC63D4685DA1614FAFBF8E0" authority="Fowler" authorityName="Fowler" box="[1261,1343,1857,1882]" class="Actinopterygii" family="Blenniidae" genus="Aspidontus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="6" phylum="Chordata" rank="species" species="tractus">tractus</taxonomicName>
</emphasis>
Fowler (
<typeStatus id="54D6338CFFC63D4681A8163DFF13F838" box="[159,215,1896,1922]" pageId="3" pageNumber="6">type</typeStatus>
locality:
<collectingCountry id="F37ACDBEFFC63D46806D163DFE08F838" box="[346,460,1896,1922]" name="Tanzania" pageId="3" pageNumber="6">Zanzibar</collectingCountry>
, Africa) from the Indian Ocean and Red Sea. These allopatric forms differ in coloration, with
<emphasis id="B919513CFFC63D46806016DAFE24F812" box="[343,480,1935,1960]" italics="true" pageId="3" pageNumber="6">
A. t.
<taxonomicName id="4C6DF6ADFFC63D4680B816DAFE24F812" box="[399,480,1935,1960]" class="Actinopterygii" family="Blenniidae" genus="Aspidontus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="6" phylum="Chordata" rank="species" species="tractus">tractus</taxonomicName>
</emphasis>
having a prominent black bar on the fleshy pectoral-fin base that is absent in
<emphasis id="B919513CFFC63D46845B16DAFEC7F874" italics="true" pageId="3" pageNumber="6">
A. t.
<taxonomicName id="4C6DF6ADFFC63D4681A016E0FEC7F874" box="[151,259,1973,1998]" class="Actinopterygii" family="Blenniidae" genus="Aspidontus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="6" phylum="Chordata" rank="species" species="taeniatus">taeniatus</taxonomicName>
</emphasis>
(although the only specimen available from the Cocos-Keeling Islands has only a faint indication of the bar;
<bibRefCitation id="EFFCF0DFFFC63D4681CC1689FE1BF84C" author="Smith-Vaniz" box="[251,479,2012,2038]" pageId="3" pageNumber="13" refString="Smith-Vaniz, W. F. (1976) The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae). Monographs of the Academy of Natural Sciences of Philadelphia, 19, 1 - 196." type="journal article" year="1976">Smith-Vaniz, 1976</bibRefCitation>
). The model,
<emphasis id="B919513CFFC63D4683BD1689FCF6F84F" box="[650,818,2012,2037]" italics="true" pageId="3" pageNumber="6">
<taxonomicName id="4C6DF6ADFFC63D4683BD1689FCE8F84F" box="[650,812,2012,2037]" class="Actinopterygii" family="Labridae" genus="Labroides" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="6" phylum="Chordata" rank="species" species="dimidiatus">L. dimidiatus</taxonomicName>
,
</emphasis>
also possesses a similar bar across the pectoral-fin base in the Indian Ocean and Red Sea that is absent from populations throughout the remainder of its range (
<bibRefCitation id="EFFCF0DFFFC13D4181A811EBFDC8FF62" author="Springer" box="[159,524,190,216]" pageId="4" pageNumber="14" refString="Springer, V. G. &amp; Smith-Vaniz, W. F. (1972) Mimetic relationships involving fishes of the family Blenniidae. Smithsonian Contributions to Zoology, 112, 1 - 36." type="journal article" year="1972">Springer &amp; Smith-Vaniz, 1972</bibRefCitation>
). Following the lead of
<bibRefCitation id="EFFCF0DFFFC13D41821411EBFBC8FF62" author="Smith-Vaniz" box="[803,1036,190,216]" pageId="4" pageNumber="13" refString="Smith-Vaniz, W. F. (1987) The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae): an update. Proceedings Academy Natural Sciences Philadelphia, 139, 1 - 52." type="journal article" year="1987">Smith-Vaniz (1987)</bibRefCitation>
who elevated two allopatric color morphs of
<taxonomicName id="4C6DF6ADFFC13D41802211B0FE68FF44" box="[277,428,229,254]" class="Actinopterygii" family="Blenniidae" genus="Meiacanthus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC13D41802211B0FE68FF44" box="[277,428,229,254]" italics="true" pageId="4" pageNumber="7">Meiacanthus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C6DF6ADFFC13D41808A11B0FDB5FF44" box="[445,625,229,254]" class="Actinopterygii" family="Blenniidae" genus="Meiacanthus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="oualanensis">
<emphasis id="B919513CFFC13D41808A11B0FDB5FF44" box="[445,625,229,254]" italics="true" pageId="4" pageNumber="7">M. oualanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DF6ADFFC13D41839E11B0FC9AFF44" box="[681,862,229,254]" class="Actinopterygii" family="Blenniidae" genus="Meiacanthus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="atrodorsalis">
<emphasis id="B919513CFFC13D41839E11B0FC9AFF44" box="[681,862,229,254]" italics="true" pageId="4" pageNumber="7">M. atrodorsalis</emphasis>
</taxonomicName>
) to full species, we recognize these two forms of
<taxonomicName id="4C6DF6ADFFC13D4181A01059FEDEFE9F" box="[151,282,268,293]" class="Actinopterygii" family="Blenniidae" genus="Aspidontus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC13D4181A01059FEDEFE9F" box="[151,282,268,293]" italics="true" pageId="4" pageNumber="7">Aspidontus</emphasis>
</taxonomicName>
as species. This is consistent with the treatment of
<bibRefCitation id="EFFCF0DFFFC13D41824D1059FB1CFE9C" author="Heemstra" box="[890,1240,268,294]" pageId="4" pageNumber="13" refString="Heemstra, P. &amp; Heemstra, E. (2004) Coastal Fishes of Southern Africa. National Inquiry Service Center and South African Institute for Aquatic Biodiversity, Grahamstown, South Africa, 488 p." type="book" year="2004">Heemstra &amp; Heemstra (2004)</bibRefCitation>
who recognized
<taxonomicName id="4C6DF6ADFFC13D4181A01067FECCFEF1" box="[151,264,306,331]" class="Actinopterygii" family="Blenniidae" genus="Aspidontus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="tractus">
<emphasis id="B919513CFFC13D4181A01067FECCFEF1" box="[151,264,306,331]" italics="true" pageId="4" pageNumber="7">A. tractus</emphasis>
</taxonomicName>
as the species present in South African waters.
</paragraph>
</subSubSection>
<subSubSection id="C377DEA5FFC13D4181F1100CFCD4F810" pageId="4" pageNumber="7" type="description">
<paragraph id="8BD28D2EFFC13D4181F1100CFB57FD12" blockId="4.[151,1437,152,1962]" pageId="4" pageNumber="7">
2)
<taxonomicName id="4C6DF6ADFFC13D4181D1100CFE47FEC8" box="[230,387,345,370]" class="Actinopterygii" family="Blenniidae" genus="Plagiotremus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC13D4181D1100CFE47FEC8" box="[230,387,345,370]" italics="true" pageId="4" pageNumber="7">Plagiotremus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C6DF6ADFFC13D4180A4100CFC69FEC9" authority="Whitley" authorityName="Whitley" box="[403,941,345,371]" class="Actinopterygii" family="Blenniidae" genus="Plagiotremus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="subSpecies" species="laudandus" subSpecies="laudandus">
<emphasis id="B919513CFFC13D4180A4100CFCF7FEC8" box="[403,819,345,370]" italics="true" pageId="4" pageNumber="7">Plagiotremus laudandus laudandus</emphasis>
(Whitley)
</taxonomicName>
(
<typeStatus id="54D6338CFFC13D41828A100CFC35FEC9" box="[957,1009,345,371]" pageId="4" pageNumber="7">type</typeStatus>
locality:
<collectingCountry id="F37ACDBEFFC13D41855E100CFAE5FEC9" box="[1129,1313,345,371]" name="New Caledonia" pageId="4" pageNumber="7">New Caledonia</collectingCountry>
) is widely distributed in the central and western Pacific Ocean with the exception of the
<collectingCountry id="F37ACDBEFFC13D41851010D5FB6AFE20" box="[1063,1198,384,410]" name="Fiji" pageId="4" pageNumber="7">Fiji Islands</collectingCountry>
. At that locality it is replaced by
<taxonomicName id="4C6DF6ADFFC13D41801D10F2FC44FE7A" authority="Smith-Vaniz. The" authorityName="Smith-Vaniz. The" box="[298,896,422,448]" class="Actinopterygii" family="Blenniidae" genus="Plagiotremus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="subSpecies" species="laudandus" subSpecies="flavus">
<emphasis id="B919513CFFC13D41801D10F2FD59FE7A" box="[298,669,423,448]" italics="true" pageId="4" pageNumber="7">Plagiotremus laudandus flavus</emphasis>
Smith-Vaniz. The
</taxonomicName>
two forms differ in coloration (
<bibRefCitation id="EFFCF0DFFFC13D4185C910F3FF1CFE5D" author="Smith-Vaniz" pageId="4" pageNumber="13" refString="Smith-Vaniz, W. F. (1976) The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae). Monographs of the Academy of Natural Sciences of Philadelphia, 19, 1 - 196." type="journal article" year="1976">Smith-Vaniz, 1976</bibRefCitation>
,
<bibRefCitation id="EFFCF0DFFFC13D4181D21098FEE2FE5D" author="Smith-Vaniz" box="[229,294,461,487]" pageId="4" pageNumber="13" refString="Smith-Vaniz, W. F. (1987) The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae): an update. Proceedings Academy Natural Sciences Philadelphia, 139, 1 - 52." type="journal article" year="1987">1987</bibRefCitation>
;
<bibRefCitation id="EFFCF0DFFFC13D4180031098FDACFE5D" author="Smith-Vaniz" box="[308,616,461,487]" pageId="4" pageNumber="13" refString="Smith-Vaniz, W. F., Satapoomin, &amp; Allen, G. R. (2001) Meiacanthus urostigma, a new fangblenny from the northeastern Indian Ocean, with discussion and examples of mimicry in species of Meiacanthus (Teleostei: Blenniidae: Nemophini). Aqua, Journal of Ichthyology and Aquatic Biology, 5, 25 - 43." type="journal article" year="2001">Smith-Vaniz et al., 2001</bibRefCitation>
):
<taxonomicName id="4C6DF6ADFFC13D4183BF1098FCCFFE5C" box="[648,779,461,486]" class="Actinopterygii" family="Blenniidae" genus="Plagiotremus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="subSpecies" species="laudandus" subSpecies="flavus">
<emphasis id="B919513CFFC13D4183BF1098FCCFFE5C" box="[648,779,461,486]" italics="true" pageId="4" pageNumber="7">P. l. flavus</emphasis>
</taxonomicName>
is entirely yellow, while
<taxonomicName id="4C6DF6ADFFC13D41857E1098FAC7FE5C" box="[1097,1283,461,486]" class="Actinopterygii" family="Blenniidae" genus="Plagiotremus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="subSpecies" species="laudandus" subSpecies="laudandus">
<emphasis id="B919513CFFC13D41857E1098FAC7FE5C" box="[1097,1283,461,486]" italics="true" pageId="4" pageNumber="7">P. l. laudandus</emphasis>
</taxonomicName>
is variously pigmented with dark stripes. Both of these forms mimic species of the blenniid genus
<taxonomicName id="4C6DF6ADFFC13D4185A310A1FAE8FDB7" box="[1172,1324,500,525]" class="Actinopterygii" family="Blenniidae" genus="Meiacanthus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC13D4185A310A1FAE8FDB7" box="[1172,1324,500,525]" italics="true" pageId="4" pageNumber="7">Meiacanthus</emphasis>
</taxonomicName>
and their distributions closely parallel those of their model species, with the yellow form,
<emphasis id="B919513CFFC13D418574134FFB00FD89" box="[1091,1220,538,563]" italics="true" pageId="4" pageNumber="7">
<taxonomicName id="4C6DF6ADFFC13D418574134FFB7BFD89" box="[1091,1215,538,563]" class="Actinopterygii" family="Blenniidae" genus="Plagiotremus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="subSpecies" species="laudandus" subSpecies="flavus">P. l. flavus</taxonomicName>
,
</emphasis>
co-occurring with the yellow species
<taxonomicName id="4C6DF6ADFFC13D4180431314FCDDFDE1" authority="Gunther" authorityName="Gunther" box="[372,793,577,603]" class="Actinopterygii" family="Blenniidae" genus="Meiacanthus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="oualanensis">
<emphasis id="B919513CFFC13D4180431314FD5AFDE0" box="[372,670,577,602]" italics="true" pageId="4" pageNumber="7">Meiacanthus oualanensis</emphasis>
(Günther)
</taxonomicName>
in the
<collectingCountry id="F37ACDBEFFC13D41825F1314FC2EFDE1" box="[872,1002,577,603]" name="Fiji" pageId="4" pageNumber="7">Fiji Islands</collectingCountry>
(
<bibRefCitation id="EFFCF0DFFFC13D4182CE1314FB1DFDE1" author="Smith-Vaniz" box="[1017,1241,577,603]" pageId="4" pageNumber="13" refString="Smith-Vaniz, W. F. (1976) The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae). Monographs of the Academy of Natural Sciences of Philadelphia, 19, 1 - 196." type="journal article" year="1976">Smith-Vaniz, 1976</bibRefCitation>
,
<bibRefCitation id="EFFCF0DFFFC13D4185D21314FAE0FDE1" author="Smith-Vaniz" box="[1253,1316,577,603]" pageId="4" pageNumber="13" refString="Smith-Vaniz, W. F. (1987) The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae): an update. Proceedings Academy Natural Sciences Philadelphia, 139, 1 - 52." type="journal article" year="1987">1987</bibRefCitation>
). We elect to elevate them to species status following a similar conclusion made for the two allopatric color morphs of their model
<taxonomicName id="4C6DF6ADFFC13D41801313DAFE7FFD12" box="[292,443,655,680]" class="Actinopterygii" family="Blenniidae" genus="Meiacanthus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC13D41801313DAFE7FFD12" box="[292,443,655,680]" italics="true" pageId="4" pageNumber="7">Meiacanthus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C6DF6ADFFC13D4180FA13DAFD44FD12" box="[461,640,655,680]" class="Actinopterygii" family="Blenniidae" genus="Meiacanthus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="oualanensis">
<emphasis id="B919513CFFC13D4180FA13DAFD44FD12" box="[461,640,655,680]" italics="true" pageId="4" pageNumber="7">M. oualanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DF6ADFFC13D41838E13DAFCAAFD12" box="[697,878,655,680]" class="Actinopterygii" family="Blenniidae" genus="Meiacanthus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="atrodorsalis">
<emphasis id="B919513CFFC13D41838E13DAFCAAFD12" box="[697,878,655,680]" italics="true" pageId="4" pageNumber="7">M. atrodorsalis</emphasis>
</taxonomicName>
) by
<bibRefCitation id="EFFCF0DFFFC13D41829313DBFB4BFD12" author="Smith-Vaniz" box="[932,1167,654,680]" pageId="4" pageNumber="13" refString="Smith-Vaniz, W. F. (1987) The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae): an update. Proceedings Academy Natural Sciences Philadelphia, 139, 1 - 52." type="journal article" year="1987">Smith-Vaniz (1987)</bibRefCitation>
.
</paragraph>
<paragraph id="8BD28D2EFFC13D4181F113E0FC5EFB76" blockId="4.[151,1437,152,1962]" pageId="4" pageNumber="7">
3)
<emphasis id="B919513CFFC13D4181D013E0FE5FFD74" box="[231,411,693,718]" italics="true" pageId="4" pageNumber="7">
<taxonomicName id="4C6DF6ADFFC13D4181D013E0FE53FD74" box="[231,407,693,718]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="genus">Ophioblennius</taxonomicName>
.
</emphasis>
<bibRefCitation id="EFFCF0DFFFC13D41809413E0FD99FD75" author="Springer" box="[419,605,693,719]" pageId="4" pageNumber="14" refString="Springer, V. G. (1962) A review of the blenniid fishes of the genus Ophioblennius Gill. Copeia, 1962, 426 - 433." type="journal article" year="1962">Springer (1962)</bibRefCitation>
reviewed the salariine genus
<emphasis id="B919513CFFC13D41828A13E0FBB5FD74" box="[957,1137,693,718]" italics="true" pageId="4" pageNumber="7">
<taxonomicName id="4C6DF6ADFFC13D41828A13E0FBA8FD74" box="[957,1132,693,718]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="genus">Ophioblennius</taxonomicName>
,
</emphasis>
recognizing two species, each with two subspecies. In the Pacific,
<taxonomicName id="4C6DF6ADFFC13D4183A61389FB5BFD4F" box="[657,1183,732,757]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="subSpecies" species="steindachneri" subSpecies="steindachneri">
<emphasis id="B919513CFFC13D4183A61389FB5BFD4F" box="[657,1183,732,757]" italics="true" pageId="4" pageNumber="7">Ophioblennius steindachneri steindachneri</emphasis>
</taxonomicName>
<collectingCountry id="F37ACDBEFFC13D4185901389FB3CFD4C" box="[1191,1272,732,758]" name="Jordan" pageId="4" pageNumber="7">Jordan</collectingCountry>
&amp; Evermann (
<typeStatus id="54D6338CFFC13D4181A81257FF17FCA6" box="[159,211,770,796]" pageId="4" pageNumber="7">type</typeStatus>
locality: near Mazatlán,
<collectingCountry id="F37ACDBEFFC13D4183371257FD98FCA6" box="[512,604,770,796]" name="Mexico" pageId="4" pageNumber="7">Mexico</collectingCountry>
) is found throughout the tropical eastern Pacific Ocean mainland from
<collectingCountry id="F37ACDBEFFC13D4181A0127CFF35FCF9" box="[151,241,809,835]" name="Mexico" pageId="4" pageNumber="7">Mexico</collectingCountry>
to
<collectingCountry id="F37ACDBEFFC13D41802F127CFE8AFCF9" box="[280,334,809,835]" name="Peru" pageId="4" pageNumber="7">Peru</collectingCountry>
and at the oceanic islands of Galápagos, Cocos, Malpelo and Revillagegido.
<taxonomicName id="4C6DF6ADFFC13D4185DA127CFDB3FCD0" authority="Springer" authorityName="Springer" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="subSpecies" species="steindachneri" subSpecies="clippertonensis">
<emphasis id="B919513CFFC13D4185DA127CFDC1FCD3" italics="true" pageId="4" pageNumber="7">Ophioblennius steindachneri clippertonensis</emphasis>
Springer
</taxonomicName>
is endemic to
<collectingCountry id="F37ACDBEFFC13D4182181205FC77FCD0" box="[815,947,848,874]" name="Clipperton Island" pageId="4" pageNumber="7">Clipperton</collectingCountry>
Atoll. These forms differ in number of dorsal-fin rays (33-35, modally
<quantity id="4C9520CBFFC13D4183251223FD8AFC2A" box="[530,590,886,912]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="4" pageNumber="7" unit="in" value="34.0">34 in</quantity>
<taxonomicName id="4C6DF6ADFFC13D4183611222FD33FC2A" box="[598,759,887,912]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="B919513CFFC13D4183611222FD33FC2A" box="[598,759,887,912]" italics="true" pageId="4" pageNumber="7">steindachneri</emphasis>
</taxonomicName>
versus 34-36, modally
<quantity id="4C9520CBFFC13D4185381223FB88FC2A" box="[1039,1100,886,912]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="4" pageNumber="7" unit="in" value="35.0">35 in</quantity>
<taxonomicName id="4C6DF6ADFFC13D4185631222FACCFC2A" box="[1108,1288,887,912]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="clippertonensis">
<emphasis id="B919513CFFC13D4185631222FACCFC2A" box="[1108,1288,887,912]" italics="true" pageId="4" pageNumber="7">clippertonensis</emphasis>
</taxonomicName>
), number of anal-fin rays (24-26, modally
<quantity id="4C9520CBFFC13D41833012C8FD82FC0D" box="[519,582,925,951]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" pageId="4" pageNumber="7" unit="in" value="25.0">25 in</quantity>
<taxonomicName id="4C6DF6ADFFC13D41837912C8FD30FC0C" box="[590,756,925,950]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="B919513CFFC13D41837912C8FD30FC0C" box="[590,756,925,950]" italics="true" pageId="4" pageNumber="7">steindachneri</emphasis>
</taxonomicName>
versus 25-27, modally
<quantity id="4C9520CBFFC13D41852212C8FB90FC0D" box="[1045,1108,925,951]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="4" pageNumber="7" unit="in" value="26.0">26 in</quantity>
<taxonomicName id="4C6DF6ADFFC13D41856A12C8FAD3FC0C" box="[1117,1303,925,950]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="clippertonensis">
<emphasis id="B919513CFFC13D41856A12C8FAD3FC0C" box="[1117,1303,925,950]" italics="true" pageId="4" pageNumber="7">clippertonensis</emphasis>
</taxonomicName>
), and total number of nuchal cirri (8-19, average 11.2-13.7 1 depending on locality, in
<taxonomicName id="4C6DF6ADFFC13D4185351292FB66FC5A" box="[1026,1186,967,992]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="B919513CFFC13D4185351292FB66FC5A" box="[1026,1186,967,992]" italics="true" pageId="4" pageNumber="7">steindachneri</emphasis>
</taxonomicName>
versus 6-14, average 9.7 1, in
<taxonomicName id="4C6DF6ADFFC13D4181C612A4FE60FBB0" box="[241,420,1009,1034]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="clippertonensis">
<emphasis id="B919513CFFC13D4181C612A4FE60FBB0" box="[241,420,1009,1034]" italics="true" pageId="4" pageNumber="7">clippertonensis</emphasis>
</taxonomicName>
). In addition, significant population differentiation in
<emphasis id="B919513CFFC13D41851412A4FB04FBB0" box="[1059,1216,1009,1034]" italics="true" pageId="4" pageNumber="7">cytochrome b</emphasis>
was demonstrated for the
<collectingCountry id="F37ACDBEFFC13D4181D81542FEB4FB8B" box="[239,368,1047,1073]" name="Clipperton Island" pageId="4" pageNumber="7">Clipperton</collectingCountry>
population relative to other eastern Pacific populations (
<bibRefCitation id="EFFCF0DFFFC13D4185171542FB3CFB8B" author="Muss" box="[1056,1272,1047,1073]" pageId="4" pageNumber="13" refString="Muss, A., Robertson, D. R., Stepien, C. A., Wirtz, P. &amp; Bowen, B. W. (2001) Phylogeography of Ophioblennius: the role of ocean currents and geography in reef fish evolution. Evolution, 55, 561 - 572." type="journal article" year="2001">Muss et al., 2001</bibRefCitation>
). While these authors indicated that the level of genetic differentiation was not at the level of most species, other authors have recognized similar levels of genetic divergence as indicative of species level differences (
<bibRefCitation id="EFFCF0DFFFC13D4184331531FF13FB1F" author="Rocha" pageId="4" pageNumber="13" refString="Rocha, L. A., Craig, M. T. &amp; Bowen, B. W. (2007) Phylogeography and the conservation of coral reef fishes. Coral Reefs, 26, 501 - 512." type="journal article" year="2007">Rocha et al., 2007</bibRefCitation>
). This, together with the well-documented differences in morphology, which
<bibRefCitation id="EFFCF0DFFFC13D41856815DEFAF7FB1F" box="[1119,1331,1163,1189]" pageId="4" pageNumber="13" refString="Muss, A., Robertson, D. R., Stepien, C. A., Wirtz, P. &amp; Bowen, B. W. (2001) Phylogeography of Ophioblennius: the role of ocean currents and geography in reef fish evolution. Evolution, 55, 561 - 572." type="journal article">Muss et al. (2001)</bibRefCitation>
ignored, clearly justifies recognition of these two forms as distinct species.
</paragraph>
<paragraph id="8BD28D2EFFC13D4181F1158DFCD4F810" blockId="4.[151,1437,152,1962]" pageId="4" pageNumber="7">
4)
<taxonomicName id="4C6DF6ADFFC13D4181C6158CFE59FB48" box="[241,413,1241,1266]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC13D4181C6158CFE59FB48" box="[241,413,1241,1266]" italics="true" pageId="4" pageNumber="7">Ophioblennius</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFFCF0DFFFC13D41808F158DFDB9FB48" author="Springer" box="[440,637,1240,1266]" pageId="4" pageNumber="14" refString="Springer, V. G. (1962) A review of the blenniid fishes of the genus Ophioblennius Gill. Copeia, 1962, 426 - 433." type="journal article" year="1962">Springer (1962)</bibRefCitation>
recognized two Atlantic subspecies,
<taxonomicName id="4C6DF6ADFFC13D41855E158CFE04FAA3" authority="Valenciennes" authorityName="Valenciennes" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="subSpecies" species="atlanticus" subSpecies="atlanticus">
<emphasis id="B919513CFFC13D41855E158CFECFFAA2" italics="true" pageId="4" pageNumber="7">Ophioblennius atlanticus atlanticus</emphasis>
(Valenciennes)
</taxonomicName>
from the west coast of Africa and adjacent islands and the east coast of
<collectingCountry id="F37ACDBEFFC13D41842715AAFA93FAA3" box="[1296,1367,1279,1305]" name="Brazil" pageId="4" pageNumber="7">Brazil</collectingCountry>
(
<typeStatus id="54D6338CFFC13D41845015AAFA5FFAA3" box="[1383,1435,1279,1305]" pageId="4" pageNumber="7">type</typeStatus>
locality: Madeira) and
<taxonomicName id="4C6DF6ADFFC13D4180871473FC19FAFA" authority="Silvester" authorityName="Silvester" box="[432,989,1318,1344]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="subSpecies" species="atlanticus" subSpecies="macclurei">
<emphasis id="B919513CFFC13D4180871473FC9DFA85" box="[432,857,1318,1343]" italics="true" pageId="4" pageNumber="7">Ophioblennius atlanticus macclurei</emphasis>
(Silvester)
</taxonomicName>
from throughout the Caribbean (
<typeStatus id="54D6338CFFC13D4184501473FA58FAFA" box="[1383,1436,1318,1344]" pageId="4" pageNumber="7">type</typeStatus>
locality:
<collectingCountry id="F37ACDBEFFC13D4180331419FE57FADC" box="[260,403,1356,1382]" name="Puerto Rico" pageId="4" pageNumber="7">Puerto Rico</collectingCountry>
). These forms differ in number of dorsal-fin rays (33-36, modally
<quantity id="4C9520CBFFC13D4185A21419FB15FADC" box="[1173,1233,1356,1382]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="4" pageNumber="7" unit="in" value="34.0">34 in</quantity>
<taxonomicName id="4C6DF6ADFFC13D4185E01419FA8FFADF" box="[1239,1355,1356,1381]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="atlanticus">
<emphasis id="B919513CFFC13D4185E01419FA8FFADF" box="[1239,1355,1356,1381]" italics="true" pageId="4" pageNumber="7">atlanticus</emphasis>
</taxonomicName>
versus 31-33, modally
<quantity id="4C9520CBFFC13D4180631426FE55FA37" box="[340,401,1395,1421]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="4" pageNumber="7" unit="in" value="32.0">32 in</quantity>
<taxonomicName id="4C6DF6ADFFC13D4180AE1426FDCAFA36" box="[409,526,1395,1420]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="macclurei">
<emphasis id="B919513CFFC13D4180AE1426FDCAFA36" box="[409,526,1395,1420]" italics="true" pageId="4" pageNumber="7">macclurei</emphasis>
</taxonomicName>
) and number of anal-fin rays (24-25, modally
<quantity id="4C9520CBFFC13D41850D1426FBB3FA37" box="[1082,1143,1395,1421]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" pageId="4" pageNumber="7" unit="in" value="25.0">25 in</quantity>
<taxonomicName id="4C6DF6ADFFC13D4185481426FB37FA36" box="[1151,1267,1395,1420]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="atlanticus">
<emphasis id="B919513CFFC13D4185481426FB37FA36" box="[1151,1267,1395,1420]" italics="true" pageId="4" pageNumber="7">atlanticus</emphasis>
</taxonomicName>
versus 22-24, modally
<quantity id="4C9520CBFFC13D41803214CFFE80FA0E" box="[261,324,1434,1460]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="4" pageNumber="7" unit="in" value="23.0">23 in</quantity>
<taxonomicName id="4C6DF6ADFFC13D41807A14CFFE0CFA09" box="[333,456,1434,1459]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="macclurei">
<emphasis id="B919513CFFC13D41807A14CFFE0CFA09" box="[333,456,1434,1459]" italics="true" pageId="4" pageNumber="7">macclurei</emphasis>
</taxonomicName>
). Based on relatively few specimens,
<bibRefCitation id="EFFCF0DFFFC13D41829114CFFBA8FA0E" author="Springer" box="[934,1132,1434,1460]" pageId="4" pageNumber="14" refString="Springer, V. G. (1962) A review of the blenniid fishes of the genus Ophioblennius Gill. Copeia, 1962, 426 - 433." type="journal article" year="1962">Springer (1962)</bibRefCitation>
reluctantly included the Brazilian population with the eastern Atlantic population under
<taxonomicName id="4C6DF6ADFFC13D4182AE1494FB97FA60" box="[921,1107,1473,1498]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="subSpecies" species="atlanticus" subSpecies="atlanticus">
<emphasis id="B919513CFFC13D4182AE1494FB97FA60" box="[921,1107,1473,1498]" italics="true" pageId="4" pageNumber="7">
<collectingCountry id="F37ACDBEFFC13D4182AE1494FC76FA60" box="[921,946,1473,1498]" name="Iceland" pageId="4" pageNumber="7">O</collectingCountry>
. a. atlanticus
</emphasis>
</taxonomicName>
but noted that the name
<taxonomicName id="4C6DF6ADFFC13D4184481494FE9AF9BB" authority="Ribeiro" authorityName="Ribeiro" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="trinitatis">
<emphasis id="B919513CFFC13D4184481494FF39F9BA" italics="true" pageId="4" pageNumber="7">
<collectingCountry id="F37ACDBEFFC13D4184481494FA5CFA60" box="[1407,1432,1473,1498]" name="Iceland" pageId="4" pageNumber="7">O</collectingCountry>
. trinitatis
</emphasis>
Ribeiro
</taxonomicName>
(
<typeStatus id="54D6338CFFC13D41805814B2FE60F9BB" box="[367,420,1511,1537]" pageId="4" pageNumber="7">type</typeStatus>
locality: Trindade
<collectingCountry id="F37ACDBEFFC13D4183A714B2FD1EF9BB" box="[656,730,1511,1537]" name="Iceland" pageId="4" pageNumber="7">Island</collectingCountry>
) was available should additional material and data become available that showed them to be distinct.
<bibRefCitation id="EFFCF0DFFFC13D4183B7175BFC97F992" box="[640,851,1550,1576]" pageId="4" pageNumber="13" refString="Muss, A., Robertson, D. R., Stepien, C. A., Wirtz, P. &amp; Bowen, B. W. (2001) Phylogeography of Ophioblennius: the role of ocean currents and geography in reef fish evolution. Evolution, 55, 561 - 572." type="journal article">Muss et al. (2001)</bibRefCitation>
surveyed variation in the
<emphasis id="B919513CFFC13D4185B6175BFAD9F99D" box="[1153,1309,1550,1575]" italics="true" pageId="4" pageNumber="7">cytochrome b</emphasis>
gene from
<taxonomicName id="4C6DF6ADFFC13D4181A01761FE8DF9F7" box="[151,329,1588,1613]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC13D4181A01761FE8DF9F7" box="[151,329,1588,1613]" italics="true" pageId="4" pageNumber="7">Ophioblennius</emphasis>
</taxonomicName>
populations throughout the Atlantic basin, reporting significant structure that parallels the biogeographic regions designated by earlier workers and coincides with the subspecies designated by
<bibRefCitation id="EFFCF0DFFFC13D418400170EFF2DF926" author="Springer" pageId="4" pageNumber="14" refString="Springer, V. G. (1962) A review of the blenniid fishes of the genus Ophioblennius Gill. Copeia, 1962, 426 - 433." type="journal article" year="1962">Springer (1962)</bibRefCitation>
. In addition they found significant differences between the eastern Atlantic and Brazilian populations of this genus. Consequently, we recognize three species of
<taxonomicName id="4C6DF6ADFFC13D41827917FCFC3FF978" box="[846,1019,1705,1730]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC13D41827917FCFC3FF978" box="[846,1019,1705,1730]" italics="true" pageId="4" pageNumber="7">Ophioblennius</emphasis>
</taxonomicName>
in the Atlantic:
<taxonomicName id="4C6DF6ADFFC13D4185F417FCFA98F978" box="[1219,1372,1705,1730]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="atlanticus">
<emphasis id="B919513CFFC13D4185F417FCFA98F978" box="[1219,1372,1705,1730]" italics="true" pageId="4" pageNumber="7">
<collectingCountry id="F37ACDBEFFC13D4185F417FCFB1FF978" box="[1219,1243,1705,1730]" name="Iceland" pageId="4" pageNumber="7">O</collectingCountry>
. atlanticus
</emphasis>
</taxonomicName>
from the eastern Atlantic,
<taxonomicName id="4C6DF6ADFFC13D4180A3179AFDF6F952" box="[404,562,1743,1768]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="macclurei">
<emphasis id="B919513CFFC13D4180A3179AFDF6F952" box="[404,562,1743,1768]" italics="true" pageId="4" pageNumber="7">
<collectingCountry id="F37ACDBEFFC13D4180A3179AFE69F952" box="[404,429,1743,1768]" name="Iceland" pageId="4" pageNumber="7">O</collectingCountry>
. macclurei
</emphasis>
</taxonomicName>
from the northwestern Atlantic, and
<taxonomicName id="4C6DF6ADFFC13D4182C1179AFB41F952" box="[1014,1157,1743,1768]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="species" species="trinitatis">
<emphasis id="B919513CFFC13D4182C1179AFB41F952" box="[1014,1157,1743,1768]" italics="true" pageId="4" pageNumber="7">
<collectingCountry id="F37ACDBEFFC13D4182C1179AFBCBF952" box="[1014,1039,1743,1768]" name="Iceland" pageId="4" pageNumber="7">O</collectingCountry>
. trinitatis
</emphasis>
</taxonomicName>
from the southwestern Atlantic. These taxa were also recognized as distinct species in a recent paper on biogeography of Atlantic reef fishes (Floeter et al., 2007). This does not, however, fully resolve the status of all Atlantic
<taxonomicName id="4C6DF6ADFFC13D4185DF1649FA50F88F" box="[1256,1428,1820,1845]" class="Actinopterygii" family="Blenniidae" genus="Ophioblennius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC13D4185DF1649FA50F88F" box="[1256,1428,1820,1845]" italics="true" pageId="4" pageNumber="7">Ophioblennius</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFFCF0DFFFC13D4181A01616FEABF8E7" box="[151,367,1859,1885]" pageId="4" pageNumber="13" refString="Muss, A., Robertson, D. R., Stepien, C. A., Wirtz, P. &amp; Bowen, B. W. (2001) Phylogeography of Ophioblennius: the role of ocean currents and geography in reef fish evolution. Evolution, 55, 561 - 572." type="journal article">Muss et al. (2001)</bibRefCitation>
demonstrated significant genetic differences among populations found on oceanic islands of the Atlantic and C. Baldwin (personal communication) reports morphological evidence supporting the recognition of additional species in the Atlantic basin.
</paragraph>
</subSubSection>
<subSubSection id="C377DEA5FFC13D4081A01682FF37FE7A" lastPageId="5" lastPageNumber="8" pageId="4" pageNumber="7" type="key">
<paragraph id="8BD28D2EFFC13D4181A01682FC3AF854" blockId="4.[151,1022,2007,2030]" box="[151,1022,2007,2030]" pageId="4" pageNumber="7">
<keyLead id="309C2E1BFFC13D4181A01682FC3AF854" box="[151,1022,2007,2030]" pageId="4" pageNumber="7">
<keyStep id="3099958BFFC13D4181A01682FC3AF854" box="[151,1022,2007,2030]" pageId="4" pageNumber="7">
<key id="FFFCDA1FFFC13D4181A01682FC3AF854" box="[151,1022,2007,2030]" pageId="4" pageNumber="7">
1.
<bibRefCitation id="EFFCF0DFFFC13D4181FA1682FEB3F854" author="Springer" box="[205,375,2007,2030]" pageId="4" pageNumber="14" refString="Springer, V. G. (1962) A review of the blenniid fishes of the genus Ophioblennius Gill. Copeia, 1962, 426 - 433." type="journal article" year="1962">Springer (1962)</bibRefCitation>
gave only frequency distributions, which we have averaged.
</key>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8BD28D2EFFC03D4081F111CDFF37FE7A" blockId="5.[151,1437,152,1415]" pageId="5" pageNumber="8">
5)
<emphasis id="B919513CFFC03D4081C711CDFE4DFF0B" box="[240,393,152,177]" italics="true" pageId="5" pageNumber="8">
<taxonomicName id="4C6DF6ADFFC03D4081C711CDFE41FF0B" ID-CoL="62DW9" box="[240,389,152,177]" class="Actinopterygii" family="Blenniidae" genus="Atrosalarias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="genus">Atrosalarias</taxonomicName>
.
</emphasis>
<bibRefCitation id="EFFCF0DFFFC03D4080AD11CDFCEEFF08" author="Springer" box="[410,810,152,178]" pageId="5" pageNumber="14" refString="Springer, V. G. &amp; Smith-Vaniz, W. F. (1968) Systematics and distribution of the monotypic Indo-Pacific blenniid fish genus Atrosalarias. Proceedings of the United States National Museum, 124, 1 - 12." type="journal article" year="1968">Springer &amp; Smith-Vaniz (1968)</bibRefCitation>
reviewed the systematics of the salariine genus
<taxonomicName id="4C6DF6ADFFC03D4081A011EAFEEAFF62" ID-CoL="62DW9" box="[151,302,191,216]" class="Actinopterygii" family="Blenniidae" genus="Atrosalarias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC03D4081A011EAFEEAFF62" box="[151,302,191,216]" italics="true" pageId="5" pageNumber="8">Atrosalarias</emphasis>
</taxonomicName>
, recognizing one species with two subspecies; another species,
<taxonomicName id="4C6DF6ADFFC03D40850B11EAFA99FF62" ID-CoL="JM8K" box="[1084,1373,191,216]" class="Actinopterygii" family="Blenniidae" genus="Atrosalarias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="hosokawai">
<emphasis id="B919513CFFC03D40850B11EAFA99FF62" box="[1084,1373,191,216]" italics="true" pageId="5" pageNumber="8">Atrosalarias hosokawai</emphasis>
</taxonomicName>
, was recently described (
<bibRefCitation id="EFFCF0DFFFC03D40804811B0FD4BFF45" author="Suzuki" box="[383,655,229,255]" pageId="5" pageNumber="14" refString="Suzuki, T., &amp; Senou, H. (1999). Atrosalarias hosokawai, a new species of blenny (Perciformes: Blenniidae) from the western Pacific. Ichthyological Research, 46, 225 - 330." type="journal article" year="1999">Suzuki &amp; Senou, 1999</bibRefCitation>
) from the western Pacific.
<taxonomicName id="4C6DF6ADFFC03D4082E511B0FA44FF45" ID-CoL="JM8H" authority="Ruppell" authorityName="Ruppell" box="[978,1408,229,255]" class="Actinopterygii" family="Blenniidae" genus="Atrosalarias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="subSpecies" species="fuscus" subSpecies="fuscus">
<emphasis id="B919513CFFC03D4082E511B0FAC3FF44" box="[978,1287,229,254]" italics="true" pageId="5" pageNumber="8">Atrosalarias fuscus fuscus</emphasis>
(Rüppell)
</taxonomicName>
is found in the Indian Ocean including the Red Sea (
<typeStatus id="54D6338CFFC03D4083C41059FCECFE9C" box="[755,808,268,294]" pageId="5" pageNumber="8">type</typeStatus>
locality:
<collectingCountry id="F37ACDBEFFC03D4082941059FC3CFE9C" box="[931,1016,268,294]" name="Eritrea" pageId="5" pageNumber="8">Eritrea</collectingCountry>
, Red Sea), and
<taxonomicName id="4C6DF6ADFFC03D4085821059FE4EFEF6" ID-CoL="JM8J" authority="Gunther" authorityName="Gunther" class="Actinopterygii" family="Blenniidae" genus="Atrosalarias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="subSpecies" species="fuscus" subSpecies="holomelas">
<emphasis id="B919513CFFC03D4085821059FED4FEF1" italics="true" pageId="5" pageNumber="8">Atrosalarias fuscus holomelas</emphasis>
(Günther)
</taxonomicName>
is found primarily in the southern portion of the central and western Pacific Ocean (
<typeStatus id="54D6338CFFC03D4084501067FA5FFEF6" box="[1383,1435,306,332]" pageId="5" pageNumber="8">type</typeStatus>
locality: Cebu,
<collectingCountry id="F37ACDBEFFC03D40805F100CFD92FEC9" box="[360,598,345,371]" name="Philippines" pageId="5" pageNumber="8">Philippine Islands</collectingCountry>
). These forms differ primarily in the number of dorsal-fin spines (consistently
<quantity id="4C9520CBFFC03D40800310D5FEB4FE20" box="[308,368,384,410]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="5" pageNumber="8" unit="in" value="11.0">11 in</quantity>
<taxonomicName id="4C6DF6ADFFC03D40804010D5FE04FE23" ID-CoL="JM8H" box="[375,448,384,409]" class="Actinopterygii" family="Blenniidae" genus="Atrosalarias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="fuscus">
<emphasis id="B919513CFFC03D40804010D5FE04FE23" box="[375,448,384,409]" italics="true" pageId="5" pageNumber="8">fuscus</emphasis>
</taxonomicName>
versus 9-11, strongly modally 10, in
<taxonomicName id="4C6DF6ADFFC03D40824D10D5FC37FE23" ID-CoL="JM8J" box="[890,1011,384,409]" class="Actinopterygii" family="Blenniidae" genus="Atrosalarias" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="holomelas">
<emphasis id="B919513CFFC03D40824D10D5FC37FE23" box="[890,1011,384,409]" italics="true" pageId="5" pageNumber="8">holomelas</emphasis>
</taxonomicName>
) and are here recognized as distinct species.
</paragraph>
</subSubSection>
<subSubSection id="C377DEA5FFC03D4081F11098FA43FA3D" pageId="5" pageNumber="8" type="description">
<paragraph id="8BD28D2EFFC03D4081F11098FCD3FD75" blockId="5.[151,1437,152,1415]" pageId="5" pageNumber="8">
6)
<taxonomicName id="4C6DF6ADFFC03D4081DF1098FE56FE5C" box="[232,402,461,486]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC03D4081DF1098FE56FE5C" box="[232,402,461,486]" italics="true" pageId="5" pageNumber="8">Entomacrodus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C6DF6ADFFC03D4080941098FD1FFE5C" box="[419,731,461,486]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="thalassinus">
<emphasis id="B919513CFFC03D4080941098FD1FFE5C" box="[419,731,461,486]" italics="true" pageId="5" pageNumber="8">Entomacrodus thalassinus</emphasis>
</taxonomicName>
(
<collectingCountry id="F37ACDBEFFC03D4083DA1098FCF9FE5D" box="[749,829,461,487]" name="Jordan" pageId="5" pageNumber="8">Jordan</collectingCountry>
&amp; Seale) was described based on specimens from
<collectingCountry id="F37ACDBEFFC03D4081A010A1FE95FDB4" box="[151,337,500,526]" name="Samoa" pageId="5" pageNumber="8">Western Samoa</collectingCountry>
and later recorded from the central Pacific to the Indian Ocean.
<taxonomicName id="4C6DF6ADFFC03D40856810A1FE4BFD8E" authority="Springer" authorityName="Springer" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="subSpecies" species="thalassinus" subSpecies="longicirrus">
<emphasis id="B919513CFFC03D40856810A1FEDAFD89" italics="true" pageId="5" pageNumber="8">Entomacrodus thalassinus longicirrus</emphasis>
Springer
</taxonomicName>
was described from the South
<collectingCountry id="F37ACDBEFFC03D40823B134FFC90FD8E" box="[780,852,538,564]" name="China" pageId="5" pageNumber="8">China</collectingCountry>
Sea and Gulf of
<collectingCountry id="F37ACDBEFFC03D40851F134FFB50FD8E" box="[1064,1172,538,564]" name="Thailand" pageId="5" pageNumber="8">Thailand</collectingCountry>
(
<typeStatus id="54D6338CFFC03D408593134FFB1EFD8E" box="[1188,1242,538,564]" pageId="5" pageNumber="8">type</typeStatus>
locality: Kram Islands, Gulf of
<collectingCountry id="F37ACDBEFFC03D40806F1314FE06FDE1" box="[344,450,577,603]" name="Thailand" pageId="5" pageNumber="8">Thailand</collectingCountry>
) and subsequently recorded from
<collectingCountry id="F37ACDBEFFC03D4082611314FC68FDE1" box="[854,940,577,603]" name="Taiwan" pageId="5" pageNumber="8">Taiwan</collectingCountry>
(Springer, 1972). The two forms differ in length of the supraorbital cirrus (significantly longer in
<taxonomicName id="4C6DF6ADFFC03D408211133DFC6CFD3B" box="[806,936,616,641]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="longicirrus">
<emphasis id="B919513CFFC03D408211133DFC6CFD3B" box="[806,936,616,641]" italics="true" pageId="5" pageNumber="8">longicirrus</emphasis>
</taxonomicName>
), pigmentation (darker in
<taxonomicName id="4C6DF6ADFFC03D4085E9133DFAA4FD3B" box="[1246,1376,616,641]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="longicirrus">
<emphasis id="B919513CFFC03D4085E9133DFAA4FD3B" box="[1246,1376,616,641]" italics="true" pageId="5" pageNumber="8">longicirrus</emphasis>
</taxonomicName>
) and body size (larger in
<taxonomicName id="4C6DF6ADFFC03D4080BA13DAFDD0FD12" box="[397,532,655,680]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="longicirrus">
<emphasis id="B919513CFFC03D4080BA13DAFDD0FD12" box="[397,532,655,680]" italics="true" pageId="5" pageNumber="8">longicirrus</emphasis>
</taxonomicName>
). Although designated as a subspecies (
<bibRefCitation id="EFFCF0DFFFC03D4082C913DBFB7DFD12" author="Springer" box="[1022,1209,654,680]" pageId="5" pageNumber="14" refString="Springer, V. G. (1967) Revision of the cirumtropical shorefish genus Entomacrodus (Blenniidae: Salariinae). Proceedings of the United States National Museum, 122, 1 - 150." type="journal article" year="1967">Springer, 1967</bibRefCitation>
), these differences warrant recognition of these forms as separate species.
</paragraph>
<paragraph id="8BD28D2EFFC03D4081F11389FDA1FBBE" blockId="5.[151,1437,152,1415]" pageId="5" pageNumber="8">
7)
<taxonomicName id="4C6DF6ADFFC03D4081D21389FE4BFD4F" box="[229,399,732,757]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC03D4081D21389FE4BFD4F" box="[229,399,732,757]" italics="true" pageId="5" pageNumber="8">Entomacrodus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C6DF6ADFFC03D4080A91389FD6CFD4F" box="[414,680,732,757]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="stellifer">
<emphasis id="B919513CFFC03D4080A91389FD6CFD4F" box="[414,680,732,757]" italics="true" pageId="5" pageNumber="8">Entomacrodus stellifer</emphasis>
</taxonomicName>
(
<collectingCountry id="F37ACDBEFFC03D4083801389FCC3FD4C" box="[695,775,732,758]" name="Jordan" pageId="5" pageNumber="8">Jordan</collectingCountry>
&amp; Snyder) was originally described based on specimens from Wakanoura,
<collectingCountry id="F37ACDBEFFC03D4080411257FE04FCA6" box="[374,448,770,796]" name="Japan" pageId="5" pageNumber="8">Japan</collectingCountry>
.
<taxonomicName id="4C6DF6ADFFC03D4080FB1257FCEBFCA6" authority="Herre" authorityName="Herre" box="[460,815,770,796]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="lighti">
<emphasis id="B919513CFFC03D4080FB1257FD0CFCA1" box="[460,712,770,795]" italics="true" pageId="5" pageNumber="8">Entomacrodus lighti</emphasis>
(Herre)
</taxonomicName>
, described based on specimens from Dodd
<collectingCountry id="F37ACDBEFFC03D40847E1257FA5CFCA6" box="[1353,1432,770,796]" name="Iceland" pageId="5" pageNumber="8">Island</collectingCountry>
, Amoy,
<collectingCountry id="F37ACDBEFFC03D4081C5127CFEFBFCF9" box="[242,319,809,835]" name="China" pageId="5" pageNumber="8">China</collectingCountry>
, was considered a subspecies of
<taxonomicName id="4C6DF6ADFFC03D4083EF127CFCFCFCF8" box="[728,824,809,834]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="stellifer">
<emphasis id="B919513CFFC03D4083EF127CFCFCFCF8" box="[728,824,809,834]" italics="true" pageId="5" pageNumber="8">stellifer</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFFCF0DFFFC03D40825F127CFBF5FCF9" author="Springer" box="[872,1073,809,835]" pageId="5" pageNumber="14" refString="Springer, V. G. (1967) Revision of the cirumtropical shorefish genus Entomacrodus (Blenniidae: Salariinae). Proceedings of the United States National Museum, 122, 1 - 150." type="journal article" year="1967">Springer (1967)</bibRefCitation>
. They differ in coloration (in
<taxonomicName id="4C6DF6ADFFC03D4081A01205FF35FCD3" box="[151,241,848,873]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="stellifer">
<emphasis id="B919513CFFC03D4081A01205FF35FCD3" box="[151,241,848,873]" italics="true" pageId="5" pageNumber="8">stellifer</emphasis>
</taxonomicName>
the side of the body has dark bands either broken up by reticulations, or with inclusions of fine pale spots or dashes, while in
<taxonomicName id="4C6DF6ADFFC03D4080F21222FDC0FC2A" box="[453,516,887,912]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="lighti">
<emphasis id="B919513CFFC03D4080F21222FDC0FC2A" box="[453,516,887,912]" italics="true" pageId="5" pageNumber="8">lighti</emphasis>
</taxonomicName>
the side of the body is more-or-less uniformly dark) and in pattern of sexual dimorphism (selected meristic characters are dimorphic in
<taxonomicName id="4C6DF6ADFFC03D40824312C8FC72FC0C" box="[884,950,925,950]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="lighti">
<emphasis id="B919513CFFC03D40824312C8FC72FC0C" box="[884,950,925,950]" italics="true" pageId="5" pageNumber="8">lighti</emphasis>
</taxonomicName>
, but not in
<taxonomicName id="4C6DF6ADFFC03D40857012C8FB62FC0C" box="[1095,1190,925,950]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="stellifer">
<emphasis id="B919513CFFC03D40857012C8FB62FC0C" box="[1095,1190,925,950]" italics="true" pageId="5" pageNumber="8">stellifer</emphasis>
</taxonomicName>
). These differences warrant recognition of these as separate species whose distributions partially parallel those of
<taxonomicName id="4C6DF6ADFFC03D4085C21291FA58FC67" box="[1269,1436,964,989]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="thalassinus">
<emphasis id="B919513CFFC03D4085C21291FA58FC67" box="[1269,1436,964,989]" italics="true" pageId="5" pageNumber="8">E. thalassinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DF6ADFFC03D4081FF12BFFEAFFBB9" box="[200,363,1002,1027]" class="Actinopterygii" family="Blenniidae" genus="Entomacrodus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="longicirrus">
<emphasis id="B919513CFFC03D4081FF12BFFEAFFBB9" box="[200,363,1002,1027]" italics="true" pageId="5" pageNumber="8">E. longicirrus</emphasis>
</taxonomicName>
(see above;
<tableCitation id="C6EFB895FFC03D4080C912BFFD90FBBE" box="[510,596,1002,1028]" captionStart="TABLE 1" captionStartId="2.[151,239,151,175]" captionTargetBox="[151,1432,202,1969]" captionTargetPageId="2" captionText="TABLE 1. Distributions of blennioid species discussed in this paper. Case numbers refer to species accounts in the text." httpUri="http://table.plazi.org/id/DF12DDA6FFC73D4781A011C2FA54FF15" pageId="5" pageNumber="8" tableUuid="DF12DDA6FFC73D4781A011C2FA54FF15">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BD28D2EFFC03D4081F11544FA43FA3D" blockId="5.[151,1437,152,1415]" pageId="5" pageNumber="8">
8)
<taxonomicName id="4C6DF6ADFFC03D4081CE1544FE6AFB90" box="[249,430,1041,1066]" class="Actinopterygii" family="Blenniidae" genus="Microlipophrys" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC03D4081CE1544FE6AFB90" box="[249,430,1041,1066]" italics="true" pageId="5" pageNumber="8">Microlipophrys</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFFCF0DFFFC03D4080E71544FD5CFB91" author="Zander" box="[464,664,1041,1067]" pageId="5" pageNumber="14" refString="Zander, C. D. (1986 a) Blenniidae. In: Whitehead, P. J. P., Bauchot, M. L., Hureau, J. C., Nielsen, J. &amp; Tortonese, E. (Eds.), Fishes of the North-eastern Atlantic and Mediterranean (Volume 3). UNESCO, Paris, pp. 1096 - 1112." type="journal article" year="1986" yearSuffix="a">Zander (1986a)</bibRefCitation>
recognized two subspecies of
<taxonomicName id="4C6DF6ADFFC03D4085681544FEF8FBE8" authority="Vinciguerra" authorityName="Vinciguerra" class="Actinopterygii" family="Blenniidae" genus="Microlipophrys" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="nigriceps">
<emphasis id="B919513CFFC03D4085681544FA58FB90" box="[1119,1436,1041,1066]" italics="true" pageId="5" pageNumber="8">Microlipophrys nigriceps</emphasis>
(Vinciguerra)
</taxonomicName>
(formerly in the genus
<taxonomicName id="4C6DF6ADFFC03D40836D156DFD12FBEB" box="[602,726,1080,1105]" class="Actinopterygii" family="Blenniidae" genus="Lipophrys" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="genus">
<emphasis id="B919513CFFC03D40836D156DFD12FBEB" box="[602,726,1080,1105]" italics="true" pageId="5" pageNumber="8">Lipophrys</emphasis>
</taxonomicName>
), the nominate form from coastal regions of the northern Mediterranean (
<typeStatus id="54D6338CFFC03D408054150BFE58FBC2" box="[355,412,1118,1144]" pageId="5" pageNumber="8">type</typeStatus>
locality: Brazza
<collectingCountry id="F37ACDBEFFC03D4083B3150BFD12FBC2" box="[644,726,1118,1144]" name="Iceland" pageId="5" pageNumber="8">Island</collectingCountry>
, Dalmatia), and
<taxonomicName id="4C6DF6ADFFC03D40829B150AFED3FB25" authority="Castanos" authorityName="Castanos" class="Actinopterygii" family="Blenniidae" genus="Microlipophrys" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="subSpecies" species="nigriceps" subSpecies="portmahonis">
<emphasis id="B919513CFFC03D40829B150AFA59FBC2" box="[940,1437,1119,1144]" italics="true" pageId="5" pageNumber="8">Microlipophrys nigriceps portmahonis</emphasis>
(Castaños)
</taxonomicName>
, from the southern Mediterranean and off the Balearics (
<typeStatus id="54D6338CFFC03D40829915D0FC26FB25" box="[942,994,1157,1183]" pageId="5" pageNumber="8">type</typeStatus>
locality:
<collectingCountry id="F37ACDBEFFC03D40856F15D0FB07FB25" box="[1112,1219,1157,1183]" name="Spain" pageId="5" pageNumber="8">Menorca</collectingCountry>
, Balearic Islands).
<bibRefCitation id="EFFCF0DFFFC03D4081A015F9FEEDFB7C" author="Bath" box="[151,297,1196,1222]" pageId="5" pageNumber="12" refString="Bath, H. (1996) Beitrag zur Osteologie der Arteen der Tribus Parablenniini Die Beziehungen der Knochen des Schadeldaches zum Seitenorgan-System und zu den Weichteilbildungen der Kopfoberseite sowie die systematische Bedeutung der Befund nebst Bemerkungen zu Lupinoblennius dispar Herre 1942 (Pisces: Blenniidae). Senckenbergiana Biologica, 76, 65 - 92." type="journal article" year="1996">Bath (1996)</bibRefCitation>
also considered
<taxonomicName id="4C6DF6ADFFC03D4080C115F9FC79FB7C" authority="Bath" authorityName="Bath" box="[502,957,1196,1222]" class="Actinopterygii" family="Blenniidae" genus="Lipophrys" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="subSpecies" species="nigriceps" subSpecies="cypriacus">
<emphasis id="B919513CFFC03D4080C115F9FCACFB7F" box="[502,872,1196,1221]" italics="true" pageId="5" pageNumber="8">Lipophrys nigriceps cypriacus</emphasis>
(Bath)
</taxonomicName>
as a valid subspecies, although
<bibRefCitation id="EFFCF0DFFFC03D40847015F9FF36FB56" author="Zander" pageId="5" pageNumber="14" refString="Zander, C. D. (1986 a) Blenniidae. In: Whitehead, P. J. P., Bauchot, M. L., Hureau, J. C., Nielsen, J. &amp; Tortonese, E. (Eds.), Fishes of the North-eastern Atlantic and Mediterranean (Volume 3). UNESCO, Paris, pp. 1096 - 1112." type="journal article" year="1986" yearSuffix="a">Zander (1986a)</bibRefCitation>
had previously synonymized it with
<taxonomicName id="4C6DF6ADFFC03D40839E1586FCBFFB56" box="[681,891,1235,1260]" class="Actinopterygii" family="Blenniidae" genus="Lipophrys" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="subSpecies" species="nigriceps" subSpecies="portmahonis">
<emphasis id="B919513CFFC03D40839E1586FCBFFB56" box="[681,891,1235,1260]" italics="true" pageId="5" pageNumber="8">L. n. portmahonis</emphasis>
</taxonomicName>
. These forms differ in the presence of a black caudal peduncle spot in
<emphasis id="B919513CFFC03D40809815ACFD87FAA8" box="[431,579,1273,1298]" italics="true" pageId="5" pageNumber="8">portmahonis</emphasis>
and
<emphasis id="B919513CFFC03D40834C15ACFD29FAA8" box="[635,749,1273,1298]" italics="true" pageId="5" pageNumber="8">cypriacus</emphasis>
that is absent in
<taxonomicName id="4C6DF6ADFFC03D40829815ACFB25FAA9" authority="(Zander, 1986a)" baseAuthorityName="Zander" baseAuthorityYear="1986" box="[943,1249,1273,1299]" class="Actinopterygii" family="Blenniidae" genus="Lipophrys" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="nigriceps">
<emphasis id="B919513CFFC03D40829815ACFBD8FAA8" box="[943,1052,1273,1298]" italics="true" pageId="5" pageNumber="8">nigriceps</emphasis>
(
<bibRefCitation id="EFFCF0DFFFC03D40851D15ACFB1CFAA9" author="Zander" box="[1066,1240,1273,1299]" pageId="5" pageNumber="14" refString="Zander, C. D. (1986 a) Blenniidae. In: Whitehead, P. J. P., Bauchot, M. L., Hureau, J. C., Nielsen, J. &amp; Tortonese, E. (Eds.), Fishes of the North-eastern Atlantic and Mediterranean (Volume 3). UNESCO, Paris, pp. 1096 - 1112." type="journal article" year="1986" yearSuffix="a">Zander, 1986a</bibRefCitation>
)
</taxonomicName>
. In addition, the northern form (
<taxonomicName id="4C6DF6ADFFC03D4080781475FE79FA83" box="[335,445,1312,1337]" class="Actinopterygii" family="Blenniidae" genus="Lipophrys" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="8" phylum="Chordata" rank="species" species="nigriceps">
<emphasis id="B919513CFFC03D4080781475FE79FA83" box="[335,445,1312,1337]" italics="true" pageId="5" pageNumber="8">nigriceps</emphasis>
</taxonomicName>
) exhibits overall reduced pigmentation associated with its cave-dwelling habits, as well as considerable color variation within a single population (
<bibRefCitation id="EFFCF0DFFFC03D4082A61413FBF1FADA" author="Zander" box="[913,1077,1350,1376]" pageId="5" pageNumber="14" refString="Zander, C. D. (1980) Morphological and ecological investigations on sympatric Lipophrys species (Blenniidae, Pisces). Helgolander Meeresunters, 34, 91 - 110." type="journal article" year="1980">Zander, 1980</bibRefCitation>
). Although warranting further study, we follow recent workers on these fishes (e.g.,
<bibRefCitation id="EFFCF0DFFFC03D40823C1438FC3EFA3D" author="Almada" box="[779,1018,1389,1415]" pageId="5" pageNumber="12" refString="Almada, V. C., Oliveira, R. F., Goncalves, E. J., Almeida, A. J., Santos, R. S. &amp; Wirtz, P. (2001) Patterns of diversity of the north-eastern Atlantic blenniid fish fauna (Pisces: Blenniidae). Global Ecology &amp; Biogeography, 10, 411 - 422." type="journal article" year="2001">Almada, et al., 2001</bibRefCitation>
) in considering them conspecific.
</paragraph>
</subSubSection>
</treatment>
</document>
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