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<mods:title id="88204381816828B67A0BEE7E9C2D2906">Revision of the cranial anatomy and phylogenetic relationships of the Eocene minute boas Messelophis variatus and Messelophis ermannorum (Serpentes, Booidea)</mods:title>
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<mods:namePart id="9F44D9508DFB3545BB4BAA5A6ADE1855">Scanferla, Agustín</mods:namePart>
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<mods:namePart id="9E62EF1F27BACBF0279D6D0AA434937D">Smith, Krister T.</mods:namePart>
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<mods:namePart id="F76F11D2D20A34E3970526749D6623C4">Schaal, Stephan F. K.</mods:namePart>
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<taxonomicName id="4C114D1BFFCDFFFF920824FFFC3A4D89" authorityName="Scanferla &amp; Smith &amp; Schaal" authorityYear="2016" box="[821,994,197,218]" class="Reptilia" family="Boidae" genus="Rieppelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="203" phylum="Chordata" rank="genus">RIEPPELOPHIS</taxonomicName>
ERMANNORUM
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(
<bibRefCitation id="EF804B69FFCDFFFF959E24FFFBF14DAB" author="Schaal S &amp; Baszio S" pageId="21" pageNumber="203" pagination="67 - 77" refId="ref14919" refString="Schaal S, Baszio S. 2004. Messelophis ermannorum n. sp., eine neue Zwergboa (Serpentes: Boidae: Tropidopheinae) aus dem Mittel-Eozan von Messel. Courier Forschungsinstitut Senckenberg 252: 67 - 77." type="journal article" year="2004">SCHAAL &amp; BASZIO, 2004</bibRefCitation>
)
<emphasis id="B965EA8AFFCDFFFF950424D2FB664DAA" bold="true" box="[1081,1214,229,250]" pageId="21" pageNumber="203">COMB. NOV.</emphasis>
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</paragraph>
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<paragraph id="8BAE3698FFCDFFFF921E2531FC5E4C70" blockId="21.[803,1226,267,320]" box="[803,902,267,288]" pageId="21" pageNumber="203">
<emphasis id="B965EA8AFFCDFFFF921E2531FC5E4C70" box="[803,902,267,288]" italics="true" pageId="21" pageNumber="203">
<typeStatus id="54AA883AFFCDFFFF921E2531FC5E4C70" box="[803,902,267,288]" pageId="21" pageNumber="203" type="holotype">Holotype</typeStatus>
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</paragraph>
<paragraph id="8BAE3698FFCDFFFF921E2510FB124C10" blockId="21.[803,1226,267,320]" box="[803,1226,298,320]" pageId="21" pageNumber="203">
<materialsCitation id="3B793CC5FFCDFFFF921E2510FB124C10" box="[803,1226,298,320]" collectionCode="SMF" pageId="21" pageNumber="203" specimenCode="ME 1812" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED00AE5DFFCDFFFF921E2510FC824C6F" box="[803,858,298,319]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="21" pageNumber="203">SMF</collectionCode>
<specimenCode id="DBB79EE3FFCDFFFF925C2510FC124C6F" box="[865,970,298,319]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="21" pageNumber="203">ME 1812</specimenCode>
, five trunk vertebrae.
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<emphasis id="B965EA8AFFCDFFFF921E2556FBC84CD1" box="[803,1040,364,385]" italics="true" pageId="21" pageNumber="203">Locality and horizon</emphasis>
</paragraph>
<paragraph id="8BAE3698FFCDFFFF921E25B0FADD4CEF" blockId="21.[803,1418,364,447]" pageId="21" pageNumber="203">
Middle Messel Formation, Messel Pit near Darmstadt (
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), middle Eocene (
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11).
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<paragraph id="8BAE3698FFCDFFFF921E25D1FC4B4F50" blockId="21.[803,1419,491,1216]" box="[803,915,491,512]" pageId="21" pageNumber="203">
<emphasis id="B965EA8AFFCDFFFF921E25D1FC4B4F50" box="[803,915,491,512]" italics="true" pageId="21" pageNumber="203">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BAE3698FFCDFFFF921E2633FB394990" blockId="21.[803,1419,491,1216]" pageId="21" pageNumber="203">
A very small booid snake that can be distinguished from all other members of
<taxonomicName id="4C114D1BFFCDFFFF95642612FB124F6D" authorityName="LINNAEUS" authorityYear="1758" box="[1113,1226,552,573]" pageId="21" pageNumber="204" rank="subOrder" subOrder="Serpentes">Serpentes</taxonomicName>
by the following combination of characters: edentulous premaxilla with ascending process; nasal articulates with the medial frontal pillar; lacrimal duct opens ventrally; large fingerlike medial foot process; postorbital applied to parietal and frontal; parietal with parietal table and a pointed posterior process in sagittal crest; maxilla with two maxillary foramina, and 22 tooth positions and teeth diminishing in size posteriorly; ectopterygoid with forked maxillary process; contact with the pterygoid via a concave surface in the lateral surface of the pterygoid; parietal table present; supraorbital process of parietal well developed; supratemporal short with a conspicuous lip in its contact region with the quadrate; 183 precloacal and 36 cloacalpostcloacal vertebrae; neural spine of precloacal vertebrae robust, occupying almost half the length of the neural arch; paracotylar foramina absent; prezygapophyseal process very weakly developed; haemal keel well developed; and paired haemapophyses in postcloacal vertebrae absent and replaced by barely defined paired protuberances located below of the condyle.
</paragraph>
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