treatments-xml/data/03/A8/D7/03A8D74FFF9FFFFCC9C2F9874588F850.xml

<document id="387BEA9B62A76CE34BEC1CD14C87D9B8" ID-CLB-Dataset="29235" ID-DOI="10.11646/zootaxa.4497.1.1" ID-GBIF-Dataset="d5b94fdb-db02-4530-85aa-12d9757e3313" ID-ISSN="1175-5326" ID-Zenodo-Dep="1451056" ID-ZooBank="890E26F1-1B03-4F13-9299-A5E8BED8439D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538973324849" checkinUser="plazi" docAuthor="Grehan, John R. &amp; Mielke, Os. G. C." docDate="2018" docId="03A8D74FFF9FFFFCC9C2F9874588F850" docLanguage="en" docName="zootaxa.4497.1.1.pdf" docOrigin="Zootaxa 4497 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pfitzneriella antonkozlovi ♂, Grehan &amp; C. Mielke 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="9" masterDocId="FF91AF37FF98FFF4C955FF924474FFBB" masterDocTitle="New species of Dugdaleiella, gen. nov., Kozloviella, gen. nov., and Pfitzneriella Viette from upper elevation Andes of Ecuador and Peru (Lepidoptera: Hepialidae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="8" updateTime="1698567105610" updateUser="ExternalLinkService">
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<mods:title id="0939B666DA8FCF0A33DD430BF162F6B6">New species of Dugdaleiella, gen. nov., Kozloviella, gen. nov., and Pfitzneriella Viette from upper elevation Andes of Ecuador and Peru (Lepidoptera: Hepialidae)</mods:title>
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<mods:namePart id="35658F85F3ACCB68C4ACAD0393F8A810">Grehan, John R.</mods:namePart>
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<mods:namePart id="105AA5AE744DF79FDE8231B37BC82D73">Mielke, Os. G. C.</mods:namePart>
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<mods:date id="99EC3E95F11DD6B9B821B0FBC1A66FE3">2018</mods:date>
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<treatment id="03A8D74FFF9FFFFCC9C2F9874588F850" ID-DOI="http://doi.org/10.5281/zenodo.5957836" ID-GBIF-Taxon="148651433" ID-Zenodo-Dep="5957836" LSID="urn:lsid:plazi:treatment:03A8D74FFF9FFFFCC9C2F9874588F850" httpUri="http://treatment.plazi.org/id/03A8D74FFF9FFFFCC9C2F9874588F850" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection id="C31B35D2FF9FFFF3C9C2F98746F9F9EB" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BBE6659FF9FFFF3C9C2F9874662F994" blockId="7.[151,653,1557,1616]" box="[151,534,1557,1583]" pageId="7" pageNumber="8">
<heading id="D0F6D135FF9FFFF3C9C2F9874662F994" bold="true" box="[151,534,1557,1583]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B975BA4BFF9FFFF3C9C2F9874662F994" bold="true" box="[151,534,1557,1583]" pageId="7" pageNumber="8">
<taxonomicName id="4C011DDAFF9FFFF3C9C2F98745B9F994" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[151,461,1557,1583]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="antonkozlovi" status="sp. nov.">
<emphasis id="B975BA4BFF9FFFF3C9C2F98745B9F994" bold="true" box="[151,461,1557,1583]" italics="true" pageId="7" pageNumber="8">Pfitzneriella antonkozlovi</emphasis>
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<taxonomicNameLabel id="A2460730FF9FFFF3C881F9874662F994" box="[468,534,1557,1583]" pageId="7" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="8BBE6659FF9FFFF3C9C2F9A546F9F9EB" blockId="7.[151,653,1557,1616]" box="[151,653,1591,1616]" pageId="7" pageNumber="8">
(
<figureCitation id="133A7ADCFF9FFFF3C9CAF9A5456BF9EB" box="[159,287,1591,1616]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1707,1730]" captionTargetBox="[151,1436,195,1660]" captionTargetId="figure@4.[151,1436,195,1660]" captionTargetPageId="4" captionText="FIGURES 1–7. Voucher specimens. 1, Kozloviella viazmenskyi gen. n., sp. n. male HT (CMNH). 1a, dorsal; 1b, ventral. 2, Pfitzneriella antonkozlovi sp. n. male HT (CMNH), 2a, dorsal; 2b, ventral. 3, P. olafi sp. n. male HT (MNHN), 3a, dorsal; 3b, ventral. 4, P. rawlinsi sp. n. male, 4a, HT dorsal (CMNH); 4b, PT dorsal; 4c, PT ventral (CGCM). 5, P. remota male HT dorsal (MNHN). 6, P. titarenkoi sp. n. male HT (CMNH), 6a, dorsal; 6b, ventral. 7, P. yuliyakovalevae sp. n. male HT (CMNH), 7a, dorsal; 7b, ventral. Female PT (CMNH); 7c, dorsal; 7d, ventral. Photos: Ernst Brockmann (Fig. 5), Jane Hyland (Fig. 4a), Anton Kozlov (Figs 1, 2, 6, 7), and Carlos Mielke (Figs 3, 4b, 4c)." httpUri="https://zenodo.org/record/1451058/files/figure.png" pageId="7" pageNumber="8">Figs 2a, 2b</figureCitation>
, 
<figureCitation id="133A7ADCFF9FFFF3C87FF9A5453CF9EB" box="[298,328,1591,1616]" captionStart="FIGURES 8–9" captionStartId="5.[151,264,1506,1529]" captionTargetBox="[151,1435,193,1486]" captionTargetId="figure@5.[151,1435,193,1486]" captionTargetPageId="5" captionText="FIGURES 8–9. Voucher specimens. 8, Dugdaleiella monticola male. 8a, LT (ZMHB); 8b, Cotopaxi, Ecuador (CMNH); 8c, Volcán Chiles, Ecuador (CMNH); 8d, male, Volcán Chiles, Ecuador (CMNH). 9, Labial palpi (male unless otherwise specified), 9a, Kozloviella viazmenskyi sp. n.; 9b, Pfitzneriella antonkozlovi sp. n.; 9c, P. rawlinsi sp. n.; 9d, P. titarenkoi sp. n.; 9e, P. yuliyakovalevae sp. n.; 9f, P. yuliyakovalevae sp. n. (female); 9g, Dugdaleiella monticola. Photos: Jane Hyland (Fig. 8), John Grehan (Figs 9a–b, d–h), and Carlos Mielke (Fig. 9c). Scale 1 mm (when present)." httpUri="https://zenodo.org/record/1451060/files/figure.png" pageId="7" pageNumber="8">9b</figureCitation>
, 
<figureCitation id="133A7ADCFF9FFFF3C801F9A545F5F9EB" box="[340,385,1591,1616]" captionStart="FIGURE 10" captionStartId="6.[151,250,1649,1671]" captionTargetBox="[152,1437,195,1610]" captionTargetId="figure@6.[152,1437,195,1610]" captionTargetPageId="6" captionText="FIGURE 10. Wing venation (male unless otherwise specified). 10a, Kozloviella viazmenskyi sp. n.; 10b, P. antonkozlovi sp. n.; 10c, P. olafi sp. n.; 10d, P. rawlinsi sp. n.; 10e, P. remota (from photo only, venation incomplete due to damaged condition of HT); 10f, P. yuliyakovalevae sp. n.; 10g, P. yuliyakovalevae sp. n. (female); 10h, Dugdaleiella monticola." httpUri="https://zenodo.org/record/1451062/files/figure.png" pageId="7" pageNumber="8">10b</figureCitation>
, 
<figureCitation id="133A7ADCFF9FFFF3C8D9F9A545C3F9EB" box="[396,439,1591,1616]" captionStart="FIGURE 11" captionStartId="7.[151,250,1429,1451]" captionTargetBox="[150,1436,810,1407]" captionTargetId="figure@7.[150,1436,810,1407]" captionTargetPageId="7" captionText="FIGURE 11. Wing markings. 11a; Kozloviella viazmenskyi sp. n., 11b; Pfitzneriella antonkozlovi sp. n.; 11c, P. olafi sp. n.; 11d, P. rawlinsi sp. n.; 11e, P. remota; 11f, P. yuliyakovalevae sp. n. Blue line—edge of white banding." httpUri="https://zenodo.org/record/1451064/files/figure.png" pageId="7" pageNumber="8">11b</figureCitation>
, 
<figureCitation id="133A7ADCFF9FFFF3C896F9AA4651F9EB" box="[451,549,1592,1616]" captionStart="FIGURES 12–14. 12" captionStartId="9.[151,264,1172,1195]" captionTargetBox="[150,1436,193,1151]" captionTargetId="figure@9.[150,1436,193,1151]" captionTargetPageId="9" captionText="FIGURES 12–14. 12, Antenna. Pfitzneriella rawlinsi sp. n. (unscaled). 13, Legs. 13a, Kozloviella viazmenskyi sp. n.; 13b, P. olafi sp. n.; 13c, P. rawlinsi sp. n.; 13d, P. titarenkoi sp. n.; 13e, P. yuliyakovalevae sp. n. 14, Metatibia. 14a, P. antonkozlovi sp. n.; 14b, P. rawlinsi sp. n.; 14c, P. titarenkoi; 14d, P. yuliyakovalevae sp. n.; 14e, P. antonkozlovi sp. n.; 14f, Dugdaleiella monticola. Photos: John Grehan (Figs 12, 14e, 14f), Carlos Mielke (Figs 13,14a–d). Scale 1 mm." httpUri="https://zenodo.org/record/1451066/files/figure.png" pageId="7" pageNumber="8">14a, 14e</figureCitation>
, 
<figureCitation id="133A7ADCFF9FFFF3CB65F9A54629F9EB" box="[560,605,1591,1616]" captionStart="FIGURES 15–16" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[197,1386,199,1851]" captionTargetId="figure@10.[197,1386,199,1851]" captionTargetPageId="10" captionText="FIGURES 15–16. Tergosternal connection. 15a, Kozloviella viazmenskyi sp. n.; 15b, Pfitzneriella yuliyakovalevae sp. n.; 15c, P. rawlinsi sp. n.; 15d, Dugdaleiella monticola. 16, Abdomen (males unless otherwise specified). 16a, K. viazmenskyi sp. n.; 16b, P. antonkozlovi sp. n.; 16c, P. olafi sp. n.; 16d, P. rawlinsi sp. n.; 16e, P. titarenkoi sp. n.; 16f, P. yuliyakovalevae sp. n.; 16g, P. yuliyakovalevae sp. n. (female); 16h–16j, Dugdaleiella monticola. 16h, tergum II; 16i, sternum II; 16j, tergum and sternum VIII. Photos: John Grehan (Figs 15d, 16h–16j), Carlos Mielke (Figs 15a–15c, 16a–16g)." httpUri="https://zenodo.org/record/1451068/files/figure.png" pageId="7" pageNumber="8">16b</figureCitation>
, 
<figureCitation id="133A7ADCFF9FFFF3CB3DF9AA46F0F9EB" box="[616,644,1592,1616]" captionStart="FIGURES 17–25" captionStartId="11.[151,264,1644,1667]" captionTargetBox="[151,1435,193,1629]" captionTargetId="figure@11.[151,1435,193,1629]" captionTargetPageId="11" captionText="FIGURES 17–25. Genitalia (male unless otherwise specified). 17, Kozloviella viazmenskyi sp. n. HT. 17a, ventral; 17b, lateral. 18, Pfitzneriella antonkozlovi sp. n. HT, ventral. 19, P. olafi sp. n. HT, ventral. 20, P. rawlinsi sp. n. HT, ventral. 21, P. remota HT (redrawn from Viette (1951a)). 22, P. titarenkoi sp. n. HT, ventral. 23, P. yuliyakovalevae sp. n. 23a, ventral; 23b, lateral. 24, P. yuliyakovalevae sp. n. PT female. 24a, posterior; 24b, lateral. 25, Dugdaleiella monticola. 25a, LT (ZMHB); 25b, Cotopaxi, Parque Nacional Cotopaxi. Dissection JRG 136 (CMNH). sa = saccus, fs = fultura superior, fi = fultura inferior, as = apodemal suture. Scale 1 mm (when present). Photos: Carlos Mielke (Figs 17–20, 22–25b)." httpUri="https://zenodo.org/record/1451070/files/figure.png" pageId="7" pageNumber="8">18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF9FFFF3C9C2F9ED463BF821" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BBE6659FF9FFFF3C9C2F9ED463BF821" blockId="7.[151,1437,1663,2018]" pageId="7" pageNumber="8">
<emphasis id="B975BA4BFF9FFFF3C9C2F9ED457BF923" bold="true" box="[151,271,1663,1688]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
The broadly triangular base and sharply narrowed apical region of the valvae distinguishes this species from all other 
<taxonomicName id="4C011DDAFF9FFFF3C861F93645DFF906" authorityName="Stephens" authorityYear="1829" box="[308,427,1700,1725]" class="Insecta" family="Hepialidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
we have examined. In 
<taxonomicName id="4C011DDAFF9FFFF3CBFFF9374682F906" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[682,758,1700,1725]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="olafi" status="sp. nov.">
<emphasis id="B975BA4BFF9FFFF3CBFFF9374682F906" box="[682,758,1700,1725]" italics="true" pageId="7" pageNumber="8">P. olafi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9FFFF3CBA8F9364748F906" box="[765,828,1700,1725]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B975BA4BFF9FFFF3CBA8F9364748F906" bold="true" box="[765,828,1700,1725]" pageId="7" pageNumber="8">sp. n.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C011DDAFF9FFFF3CA25F9374048F906" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[880,1084,1700,1725]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="yuliyakovalevae" status="sp. nov.">
<emphasis id="B975BA4BFF9FFFF3CA25F9374048F906" box="[880,1084,1700,1725]" italics="true" pageId="7" pageNumber="8">P. yuliyakovalevae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9FFFF3CD17F936400BF906" box="[1090,1151,1700,1725]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B975BA4BFF9FFFF3CD17F936400BF906" bold="true" box="[1090,1151,1700,1725]" pageId="7" pageNumber="8">sp. n.</emphasis>
</taxonomicNameLabel>
the apex is rounded. In 
<taxonomicName id="4C011DDAFF9FFFF3CCDDF9374485F95B" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="rawlinsi" status="sp. nov.">
<emphasis id="B975BA4BFF9FFFF3CCDDF9374485F95B" italics="true" pageId="7" pageNumber="8">P. rawlinsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9FFFF3C9ACF9554543F95B" box="[249,311,1735,1760]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B975BA4BFF9FFFF3C9ACF9554543F95B" bold="true" box="[249,311,1735,1760]" pageId="7" pageNumber="8">sp. n.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C011DDAFF9FFFF3C825F95B4582F95B" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[368,502,1735,1760]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="titarenkoi" status="sp. nov.">
<emphasis id="B975BA4BFF9FFFF3C825F95B4582F95B" box="[368,502,1735,1760]" italics="true" pageId="7" pageNumber="8">P. titarenkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9FFFF3C8A8F9554648F95B" box="[509,572,1735,1760]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B975BA4BFF9FFFF3C8A8F9554648F95B" bold="true" box="[509,572,1735,1760]" pageId="7" pageNumber="8">sp. n.</emphasis>
</taxonomicNameLabel>
the apex is pointed, but without forming a distinct narrowed region. The FW color tone and pattern is most similar to 
<taxonomicName id="4C011DDAFF9FFFF3CB09F97F475AF8BE" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[604,814,1772,1797]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="yuliyakovalevae" status="sp. nov.">
<emphasis id="B975BA4BFF9FFFF3CB09F97F475AF8BE" box="[604,814,1772,1797]" italics="true" pageId="7" pageNumber="8">P. yuliyakovalevae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9FFFF3CA63F97E4701F8BF" box="[822,885,1772,1797]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B975BA4BFF9FFFF3CA63F97E4724F8BE" bold="true" box="[822,848,1772,1797]" pageId="7" pageNumber="8">sp</emphasis>
. 
<emphasis id="B975BA4BFF9FFFF3CA35F97E471BF8BE" bold="true" box="[864,879,1772,1797]" pageId="7" pageNumber="8">n</emphasis>
.
</taxonomicNameLabel>
, but central region between CuA2 and CuP with broad yellowish brown band curving posterior to dark grayish brown patch along central CuA2. The FW color tone differs from the yellowish brown of 
<taxonomicName id="4C011DDAFF9FFFF3CB6DF8A846FCF8E9" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[568,648,1849,1874]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="olafi" status="sp. nov.">
<emphasis id="B975BA4BFF9FFFF3CB6DF8A846FCF8E9" box="[568,648,1849,1874]" italics="true" pageId="7" pageNumber="8">P. olafi</emphasis>
</taxonomicName>
, 
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<emphasis id="B975BA4BFF9FFFF3CBCCF8AB46ACF8E9" bold="true" box="[665,728,1849,1874]" pageId="7" pageNumber="8">sp. n.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="4C011DDAFF9FFFF3CBB5F8A8472EF8E9" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[736,858,1849,1874]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="rawlinsi" status="sp. nov.">
<emphasis id="B975BA4BFF9FFFF3CBB5F8A8472EF8E9" box="[736,858,1849,1874]" italics="true" pageId="7" pageNumber="8">P. rawlinsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9FFFF3CA37F8AB47D5F8E9" box="[866,929,1849,1874]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B975BA4BFF9FFFF3CA37F8AB47D5F8E9" bold="true" box="[866,929,1849,1874]" pageId="7" pageNumber="8">sp. n.</emphasis>
</taxonomicNameLabel>
, and 
<taxonomicName id="4C011DDAFF9FFFF3CAB6F8A8403FF8EA" authorityName="Pfitzner" authorityYear="1906" box="[995,1099,1850,1873]" class="Insecta" family="Hepialidae" genus="Triodia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="remota">
<emphasis id="B975BA4BFF9FFFF3CAB6F8A8403FF8EA" box="[995,1099,1850,1873]" italics="true" pageId="7" pageNumber="8">P. remota</emphasis>
</taxonomicName>
, the indistinct markings on a dark grayish brown background in 
<taxonomicName id="4C011DDAFF9FFFF3CB72F8CC46C4F8CD" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[551,688,1885,1910]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="titarenkoi" status="sp. nov.">
<emphasis id="B975BA4BFF9FFFF3CB72F8CC46C4F8CD" box="[551,688,1885,1910]" italics="true" pageId="7" pageNumber="8">P. titarenkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9FFFF3CBEDF8CF4682F8CD" box="[696,758,1885,1910]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B975BA4BFF9FFFF3CBEDF8CF4682F8CD" bold="true" box="[696,758,1885,1910]" pageId="7" pageNumber="8">sp. n.</emphasis>
</taxonomicNameLabel>
, and the light grayish brown with pale to indistinct grayish white markings in 
<taxonomicName id="4C011DDAFF9FFFF3C83DF813458EF821" authority="Grehan &amp; Mielke, 2018" authorityName="Grehan &amp; Mielke" authorityYear="2018" box="[360,506,1921,1946]" class="Insecta" family="Hepialidae" genus="Dugdaleiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B975BA4BFF9FFFF3C83DF813458EF821" box="[360,506,1921,1946]" italics="true" pageId="7" pageNumber="8">Dugdaleiella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF9FFFF3CB54F813463BF821" box="[513,591,1921,1946]" pageId="7" pageNumber="8" rank="genus">
<emphasis id="B975BA4BFF9FFFF3CB54F813463BF821" bold="true" box="[513,591,1921,1946]" pageId="7" pageNumber="8">gen. n.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF9FFFFCC992F83745E4FA48" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="8BBE6659FF9FFFF3C992F8374666F859" blockId="7.[151,1437,1663,2018]" pageId="7" pageNumber="8">
<emphasis id="B975BA4BFF9FFFF3C992F837452DF805" bold="true" box="[199,345,1957,1982]" pageId="7" pageNumber="8">Description.</emphasis>
Male (
<figureCitation id="133A7ADCFF9FFFF3C8FDF837465DF805" box="[424,553,1957,1982]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1707,1730]" captionTargetBox="[151,1436,195,1660]" captionTargetId="figure@4.[151,1436,195,1660]" captionTargetPageId="4" captionText="FIGURES 1–7. Voucher specimens. 1, Kozloviella viazmenskyi gen. n., sp. n. male HT (CMNH). 1a, dorsal; 1b, ventral. 2, Pfitzneriella antonkozlovi sp. n. male HT (CMNH), 2a, dorsal; 2b, ventral. 3, P. olafi sp. n. male HT (MNHN), 3a, dorsal; 3b, ventral. 4, P. rawlinsi sp. n. male, 4a, HT dorsal (CMNH); 4b, PT dorsal; 4c, PT ventral (CGCM). 5, P. remota male HT dorsal (MNHN). 6, P. titarenkoi sp. n. male HT (CMNH), 6a, dorsal; 6b, ventral. 7, P. yuliyakovalevae sp. n. male HT (CMNH), 7a, dorsal; 7b, ventral. Female PT (CMNH); 7c, dorsal; 7d, ventral. Photos: Ernst Brockmann (Fig. 5), Jane Hyland (Fig. 4a), Anton Kozlov (Figs 1, 2, 6, 7), and Carlos Mielke (Figs 3, 4b, 4c)." httpUri="https://zenodo.org/record/1451058/files/figure.png" pageId="7" pageNumber="8">Figs 2a, 2b</figureCitation>
): Wingspan 
<quantity id="4CF9CBBCFF9FFFF3CBE3F837477CF806" box="[694,776,1957,1982]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="7" pageNumber="8" unit="mm" value="46.0">46 mm</quantity>
; FW length: 
<quantity id="4CF9CBBCFF9FFFF3CACFF834479FF806" box="[922,1003,1958,1982]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="8" unit="mm" value="22.0">22 mm</quantity>
, width: 
<quantity id="4CF9CBBCFF9FFFF3CD11F83440F3F806" box="[1092,1159,1958,1982]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="8" unit="mm" value="9.0">9 mm</quantity>
, ratio 2.4: 1; HW length: 
<quantity id="4CF9CBBCFF9FFFF3C9C2F858449CF85A" box="[151,232,1994,2018]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="8" unit="mm" value="22.0">22 mm</quantity>
, width: 
<quantity id="4CF9CBBCFF9FFFF3C817F85845F1F85A" box="[322,389,1994,2018]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="8" unit="mm" value="8.0">8 mm</quantity>
, ratio 2.8: 1.
</paragraph>
<paragraph id="8BBE6659FF90FFFCC992FF0540A1FEDE" blockId="8.[151,1437,151,2027]" pageId="8" pageNumber="9">
<emphasis id="B975BA4BFF90FFFCC992FF054570FF0B" box="[199,260,151,176]" italics="true" pageId="8" pageNumber="9">Head</emphasis>
. Frons and vertex covered with yellowish brown piliform scales tipped with dark brown. Interocularantennal scales piliform, overlapping eye. Antenna filiform, blackish-brown, 44 flagellomeres, annuli laterally compressed, with numerous sensillae chaetica, dorsoventrally wide over much of basal and central flagellum, tapering to tip, terminal segment in the form of a narrowed, elongate triangle; scape rounded, covered with lamellar scales. Clypeus unscaled, projecting anteriorly as a transverse ridge. Labium subsquare. Labial palpus long (extend beyond length of labial scales), two segmented, distal palpomere twice as long as basal (
<figureCitation id="133A7ADCFF90FFFCCD20FEDE40BCFEDE" box="[1141,1224,332,357]" captionStart="FIGURES 8–9" captionStartId="5.[151,264,1506,1529]" captionTargetBox="[151,1435,193,1486]" captionTargetId="figure@5.[151,1435,193,1486]" captionTargetPageId="5" captionText="FIGURES 8–9. Voucher specimens. 8, Dugdaleiella monticola male. 8a, LT (ZMHB); 8b, Cotopaxi, Ecuador (CMNH); 8c, Volcán Chiles, Ecuador (CMNH); 8d, male, Volcán Chiles, Ecuador (CMNH). 9, Labial palpi (male unless otherwise specified), 9a, Kozloviella viazmenskyi sp. n.; 9b, Pfitzneriella antonkozlovi sp. n.; 9c, P. rawlinsi sp. n.; 9d, P. titarenkoi sp. n.; 9e, P. yuliyakovalevae sp. n.; 9f, P. yuliyakovalevae sp. n. (female); 9g, Dugdaleiella monticola. Photos: Jane Hyland (Fig. 8), John Grehan (Figs 9a–b, d–h), and Carlos Mielke (Fig. 9c). Scale 1 mm (when present)." httpUri="https://zenodo.org/record/1451060/files/figure.png" pageId="8" pageNumber="9">Fig. 9b</figureCitation>
).
</paragraph>
<paragraph id="8BBE6659FF90FFFCC992FEFD41E9FC35" blockId="8.[151,1437,151,2027]" pageId="8" pageNumber="9">
<emphasis id="B975BA4BFF90FFFCC992FEFD456CFE33" box="[199,280,367,392]" italics="true" pageId="8" pageNumber="9">Thorax</emphasis>
. Dorsally covered with yellowish brown scales, scutum III pale greyish brown, free of scales other than posterior and medial regions, obscured from direct view by overlapping mesothoracic scales; ventral thorax yellowish brown. Pro- and meso- legs missing; metaleg pale yellowish brown with basal tibial gland (
<figureCitation id="133A7ADCFF90FFFCCC61FE2541E0FE6B" box="[1332,1428,439,464]" captionStart="FIGURES 12–14. 12" captionStartId="9.[151,264,1172,1195]" captionTargetBox="[150,1436,193,1151]" captionTargetId="figure@9.[150,1436,193,1151]" captionTargetPageId="9" captionText="FIGURES 12–14. 12, Antenna. Pfitzneriella rawlinsi sp. n. (unscaled). 13, Legs. 13a, Kozloviella viazmenskyi sp. n.; 13b, P. olafi sp. n.; 13c, P. rawlinsi sp. n.; 13d, P. titarenkoi sp. n.; 13e, P. yuliyakovalevae sp. n. 14, Metatibia. 14a, P. antonkozlovi sp. n.; 14b, P. rawlinsi sp. n.; 14c, P. titarenkoi; 14d, P. yuliyakovalevae sp. n.; 14e, P. antonkozlovi sp. n.; 14f, Dugdaleiella monticola. Photos: John Grehan (Figs 12, 14e, 14f), Carlos Mielke (Figs 13,14a–d). Scale 1 mm." httpUri="https://zenodo.org/record/1451066/files/figure.png" pageId="8" pageNumber="9">Fig. 14a</figureCitation>
) supporting elongate androconial scales (
<figureCitation id="133A7ADCFF90FFFCCB0EFE4E46CFFE4F" box="[603,699,476,501]" captionStart="FIGURES 12–14. 12" captionStartId="9.[151,264,1172,1195]" captionTargetBox="[150,1436,193,1151]" captionTargetId="figure@9.[150,1436,193,1151]" captionTargetPageId="9" captionText="FIGURES 12–14. 12, Antenna. Pfitzneriella rawlinsi sp. n. (unscaled). 13, Legs. 13a, Kozloviella viazmenskyi sp. n.; 13b, P. olafi sp. n.; 13c, P. rawlinsi sp. n.; 13d, P. titarenkoi sp. n.; 13e, P. yuliyakovalevae sp. n. 14, Metatibia. 14a, P. antonkozlovi sp. n.; 14b, P. rawlinsi sp. n.; 14c, P. titarenkoi; 14d, P. yuliyakovalevae sp. n.; 14e, P. antonkozlovi sp. n.; 14f, Dugdaleiella monticola. Photos: John Grehan (Figs 12, 14e, 14f), Carlos Mielke (Figs 13,14a–d). Scale 1 mm." httpUri="https://zenodo.org/record/1451066/files/figure.png" pageId="8" pageNumber="9">Fig. 14e</figureCitation>
). FW costal margin slightly convex; outer margin weakly convex with extended shallow curve to anal margin. Venation hepialine (
<bibRefCitation id="EF901BA8FF90FFFCCA27FE6D4042FDA3" box="[882,1078,511,536]" pageId="8" pageNumber="9" refId="ref14701" refString="Dumbleton, L. J. (1966) Genitalia, classification and zoogeography of the New Zealand Hepialidae (Lepidoptera). New Zealand Journal of Science, 9, 920 - 981." type="journal article">Dumbleton 1966</bibRefCitation>
), base of FW anal vein strongly curved anteriorly, distally curved posteriorly (
<figureCitation id="133A7ADCFF90FFFCCBFDFDB64779FD86" box="[680,781,548,573]" captionStart="FIGURE 10" captionStartId="6.[151,250,1649,1671]" captionTargetBox="[152,1437,195,1610]" captionTargetId="figure@6.[152,1437,195,1610]" captionTargetPageId="6" captionText="FIGURE 10. Wing venation (male unless otherwise specified). 10a, Kozloviella viazmenskyi sp. n.; 10b, P. antonkozlovi sp. n.; 10c, P. olafi sp. n.; 10d, P. rawlinsi sp. n.; 10e, P. remota (from photo only, venation incomplete due to damaged condition of HT); 10f, P. yuliyakovalevae sp. n.; 10g, P. yuliyakovalevae sp. n. (female); 10h, Dugdaleiella monticola." httpUri="https://zenodo.org/record/1451062/files/figure.png" pageId="8" pageNumber="9">Fig. 10b</figureCitation>
). FW dorsal ground colour a mix of diffuse pale orange brown and black markings; cubital region with longitudinal curved band of pale orange with anterior edge subtending black patch ending at CuA2, and posterior edge with inner black line and outer whitish line; outer terminus of curved band continued between veins as a curved, broken band to base of cell between M1 and M2; areas of black shading predominant between anal margin and CuA2, and distally between curved band and outer margin below M2 (
<figureCitation id="133A7ADCFF90FFFCC83CFD7245BFFD42" box="[361,459,736,761]" captionStart="FIGURE 11" captionStartId="7.[151,250,1429,1451]" captionTargetBox="[150,1436,810,1407]" captionTargetId="figure@7.[150,1436,810,1407]" captionTargetPageId="7" captionText="FIGURE 11. Wing markings. 11a; Kozloviella viazmenskyi sp. n., 11b; Pfitzneriella antonkozlovi sp. n.; 11c, P. olafi sp. n.; 11d, P. rawlinsi sp. n.; 11e, P. remota; 11f, P. yuliyakovalevae sp. n. Blue line—edge of white banding." httpUri="https://zenodo.org/record/1451064/files/figure.png" pageId="8" pageNumber="9">Fig. 11b</figureCitation>
); outer margin with submarginal row of ovoid pale whitish brown sequentially edged with black and white lines, marginal band of black subtending orange-brown fringe tipped with black scales in line with terminus of veins, most noticeably between CuA2 and apex. FW ventral costal pocket present; ground colour greyish brown. HW dorsal greyish brown with pale orange-brown shading near apex and along outer fringe. HW ventral greyish brown except for orange-brown scales between Sc and costal margin, and marginal fringe as in FW.
</paragraph>
<paragraph id="8BBE6659FF90FFFCC992FC0840ADFB15" blockId="8.[151,1437,151,2027]" pageId="8" pageNumber="9">
<emphasis id="B975BA4BFF90FFFCC992FC084545FC08" box="[199,305,922,947]" italics="true" pageId="8" pageNumber="9">Abdomen</emphasis>
. Greyish brown. Tergites and sternites weakly sclerotized. Tergosternal connection with elongate tergosternal bar, elongate lateral tergal brace and short dorsal brace (damaged), tergal knob absent. Tergum II with thin lateral ridge, extending antero-medially to anterior margin by curing around outer and then anterior edge of the anterior lateral tuberculate plate; sternum II sub-rectangular, anterior margin broad V-shape to lateral margin, anterior lateral arms short, point of tergosternal attachment with short longitudinal ridge extending posteriorly. Sternum VII longer than wide, lateral margins convex. Tergum VIII subsquare, narrowing to rounded posterior margin, sternum VIII rectangular, wider than long, lateral margins slightly concave, posterior margin with slightly projecting lobe at lateral corners; margin forming a shallow V-shaped notch at median (
<figureCitation id="133A7ADCFF90FFFCCD3FFB0740B8FB15" box="[1130,1228,1173,1198]" captionStart="FIGURES 15–16" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[197,1386,199,1851]" captionTargetId="figure@10.[197,1386,199,1851]" captionTargetPageId="10" captionText="FIGURES 15–16. Tergosternal connection. 15a, Kozloviella viazmenskyi sp. n.; 15b, Pfitzneriella yuliyakovalevae sp. n.; 15c, P. rawlinsi sp. n.; 15d, Dugdaleiella monticola. 16, Abdomen (males unless otherwise specified). 16a, K. viazmenskyi sp. n.; 16b, P. antonkozlovi sp. n.; 16c, P. olafi sp. n.; 16d, P. rawlinsi sp. n.; 16e, P. titarenkoi sp. n.; 16f, P. yuliyakovalevae sp. n.; 16g, P. yuliyakovalevae sp. n. (female); 16h–16j, Dugdaleiella monticola. 16h, tergum II; 16i, sternum II; 16j, tergum and sternum VIII. Photos: John Grehan (Figs 15d, 16h–16j), Carlos Mielke (Figs 15a–15c, 16a–16g)." httpUri="https://zenodo.org/record/1451068/files/figure.png" pageId="8" pageNumber="9">Fig. 16b</figureCitation>
).
</paragraph>
<paragraph id="8BBE6659FF90FFFCC992FB28448BFA75" blockId="8.[151,1437,151,2027]" pageId="8" pageNumber="9">
<emphasis id="B975BA4BFF90FFFCC992FB284504FB68" box="[199,368,1210,1235]" italics="true" pageId="8" pageNumber="9">Male genitalia</emphasis>
(
<figureCitation id="133A7ADCFF90FFFCC8D6FB2845ADFB69" box="[387,473,1210,1235]" captionStart="FIGURES 17–25" captionStartId="11.[151,264,1644,1667]" captionTargetBox="[151,1435,193,1629]" captionTargetId="figure@11.[151,1435,193,1629]" captionTargetPageId="11" captionText="FIGURES 17–25. Genitalia (male unless otherwise specified). 17, Kozloviella viazmenskyi sp. n. HT. 17a, ventral; 17b, lateral. 18, Pfitzneriella antonkozlovi sp. n. HT, ventral. 19, P. olafi sp. n. HT, ventral. 20, P. rawlinsi sp. n. HT, ventral. 21, P. remota HT (redrawn from Viette (1951a)). 22, P. titarenkoi sp. n. HT, ventral. 23, P. yuliyakovalevae sp. n. 23a, ventral; 23b, lateral. 24, P. yuliyakovalevae sp. n. PT female. 24a, posterior; 24b, lateral. 25, Dugdaleiella monticola. 25a, LT (ZMHB); 25b, Cotopaxi, Parque Nacional Cotopaxi. Dissection JRG 136 (CMNH). sa = saccus, fs = fultura superior, fi = fultura inferior, as = apodemal suture. Scale 1 mm (when present). Photos: Carlos Mielke (Figs 17–20, 22–25b)." httpUri="https://zenodo.org/record/1451070/files/figure.png" pageId="8" pageNumber="9">Fig. 18</figureCitation>
). Tegumen not distinct from pseudotegumen. Saccus forming a shallow ring with raised posterior margins lateral to valva, apodemal suture with pair of short triangular projections separated at median by deep central notch. Tergal lobes absent. Pseudotegumen as a pair of triangular plates, wide dorsally, narrowing to pointed apex ventrally, convex adjacent to membranous phallus, not fused medially. Valva distally lobate, pointed end facing ventro-posteriorly; inner and distal surfaces setose. Fultura superior membranous half distance from pseudotegumen, then sclerotized to broad fultura inferior, width twice length, anterior and lateral margins rounded, posterior broad shallow V-shape from posterior corners to median. Phallus membranous without cornutus.
</paragraph>
<paragraph id="8BBE6659FF90FFFCC992FA4845E4FA48" blockId="8.[151,1437,151,2027]" box="[199,400,1498,1523]" pageId="8" pageNumber="9">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF90FFFCC992FA6F466EF955" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BBE6659FF90FFFCC992FA6F4779F981" blockId="8.[151,1437,151,2027]" pageId="8" pageNumber="9">
<emphasis id="B975BA4BFF90FFFCC992FA6F452DF9AD" bold="true" box="[199,345,1533,1558]" pageId="8" pageNumber="9">Distribution</emphasis>
. Collected in a mountain valley adjacent to forest at 
<quantity id="4CF9CBBCFF90FFFCCAFCFA6C4072F9AD" box="[937,1030,1534,1558]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="8" pageNumber="9" unit="m" value="3720.0">3,720 m</quantity>
(
<figureCitation id="133A7ADCFF90FFFCCD40FA6F4001F9AD" box="[1045,1141,1533,1558]" captionStart="FIGURE 26" captionStartId="14.[151,250,1674,1696]" captionTargetBox="[152,1435,266,1652]" captionTargetId="figure@14.[152,1435,195,1652]" captionTargetPageId="14" captionText="FIGURE 26. Collection localities. 26a–26b, Kozloviella viazmenskyi sp. n. Peru, Cerro Machete, 2,050 m; 26c, Pfitzneriella antonkozlovi sp. n., Peru, Marañon Province, 3,270 m; 26d, P. titarenkoi, sp. n. and P. yuliyakovalevae sp. n., Peru, Pampa Hermosa district, 3,380 m; 26e–26f, P. yuliyakovalevae sp. n., Peru, Pampa Hermosa district, 2,960 m. With permission by Anton Kozlov and Yuliya Kovaleva, foreground 26b, 26e, Anton Kozlov and 26f, Yuliya Kovaleva. Photos: Anton Kozlov." httpUri="https://zenodo.org/record/1451072/files/figure.png" pageId="8" pageNumber="9">Fig. 26c</figureCitation>
). 
<collectingRegion id="49C5A8BBFF90FFFCCDDCFA6F40AFF9AD" box="[1161,1243,1533,1558]" country="Sri Lanka" name="Central" pageId="8" pageNumber="9">Central</collectingRegion>
eastern Peruvian Andes, known from the 
<typeStatus id="54BAD8FBFF90FFFCC8F3F9B145A2F980" box="[422,470,1571,1595]" pageId="8" pageNumber="9">type</typeStatus>
locality only (
<figureCitation id="133A7ADCFF90FFFCCB2CF9B04771F981" box="[633,773,1570,1595]" captionStart="FIGURES 27–29" captionStartId="20.[151,264,1710,1733]" captionTargetBox="[156,1435,363,1685]" captionTargetId="figure@20.[151,1436,363,1690]" captionTargetPageId="20" captionText="FIGURES 27–29. Distribution. 27, Dugdaleiella gen. n., Kozloviella gen. n., and Pfitzneriella locality records. 27a, Ecuador and Peru. Tectonic boundaries approximate; 27b, Dugdaleiella gen. n. and P. lucicola in Ecuador. 28, Locality records of Peruvian Pfitzneriella in relation to eastern Andean montane forests (modified from Young &amp; León (2000) with permission by Kenneth Young). 29, Principal groups of Hepialidae in Central and South America. Phassus/Schausiana/Phthius (solid line), cibyrine genera of Hepialidae (dotted line), Dugdaleiella gen. n./Kozloviella gen. n./Pfitzneriella (pale green), Uiridigigas (red), and southern Andean genera (blue)." httpUri="https://zenodo.org/record/1451074/files/figure.png" pageId="8" pageNumber="9">Figs 27a, 28</figureCitation>
)
</paragraph>
<paragraph id="8BBE6659FF90FFFCC992F9D7466EF955" blockId="8.[151,1437,151,2027]" pageId="8" pageNumber="9">
<emphasis id="B975BA4BFF90FFFCC992F9D74557F9E5" bold="true" box="[199,291,1605,1630]" pageId="8" pageNumber="9">Habitat</emphasis>
. Flight record at the beginning of February coincides with wettest month of the year with an average monthly rainfall of 
<quantity id="4CF9CBBCFF90FFFCC827F9F845B5F939" box="[370,449,1642,1667]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="9" unit="mm" value="60.0">60 mm</quantity>
(World 
<bibRefCitation id="EF901BA8FF90FFFCCB4BF9F84748F939" box="[542,828,1642,1667]" pageId="8" pageNumber="9" refId="ref16676" refString="World Weather and Climate (2018) World Weather &amp; Climate Information. Available from: https: // weather-and-climate. com / average-monthly-Rainfall-Temperature-Sunshine, Huanuco, Peru (accessed 20 July 2018)" type="url">Weather &amp; Climate 2018</bibRefCitation>
). At an elevation of 
<quantity id="4CF9CBBCFF90FFFCCD75F9F84009F939" box="[1056,1149,1642,1666]" metricMagnitude="3" metricUnit="m" metricValue="3.27" pageId="8" pageNumber="9" unit="m" value="3270.0">3,270 m</quantity>
this record is close to the tree line of about 
<quantity id="4CF9CBBCFF90FFFCC820F91F4656F91D" box="[373,546,1677,1702]" metricMagnitude="3" metricUnit="m" metricValue="3.8" metricValueMax="4.0" metricValueMin="3.6" pageId="8" pageNumber="9" unit="m" value="3800.0" valueMax="4000.0" valueMin="3600.0">3,600-4,000 m</quantity>
, although the precise limits in the local region are obscured by extensive conversion to farmland at higher elevations. The 
<typeStatus id="54BAD8FBFF90FFFCCB9BF921468AF970" box="[718,766,1715,1739]" pageId="8" pageNumber="9">type</typeStatus>
locality is close to the upper elevation limits of relatively continuous forest cover (
<figureCitation id="133A7ADCFF90FFFCC8F8F947467AF955" box="[429,526,1749,1774]" captionStart="FIGURE 26" captionStartId="14.[151,250,1674,1696]" captionTargetBox="[152,1435,266,1652]" captionTargetId="figure@14.[152,1435,195,1652]" captionTargetPageId="14" captionText="FIGURE 26. Collection localities. 26a–26b, Kozloviella viazmenskyi sp. n. Peru, Cerro Machete, 2,050 m; 26c, Pfitzneriella antonkozlovi sp. n., Peru, Marañon Province, 3,270 m; 26d, P. titarenkoi, sp. n. and P. yuliyakovalevae sp. n., Peru, Pampa Hermosa district, 3,380 m; 26e–26f, P. yuliyakovalevae sp. n., Peru, Pampa Hermosa district, 2,960 m. With permission by Anton Kozlov and Yuliya Kovaleva, foreground 26b, 26e, Anton Kozlov and 26f, Yuliya Kovaleva. Photos: Anton Kozlov." httpUri="https://zenodo.org/record/1451072/files/figure.png" pageId="8" pageNumber="9">Fig. 26c</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF90FFFCC992F9684740F8A8" box="[199,820,1786,1811]" pageId="8" pageNumber="9" type="biology_ecology">
<paragraph id="8BBE6659FF90FFFCC992F9684740F8A8" blockId="8.[151,1437,151,2027]" box="[199,820,1786,1811]" pageId="8" pageNumber="9">
<emphasis id="B975BA4BFF90FFFCC992F9684530F8A8" bold="true" box="[199,324,1786,1811]" pageId="8" pageNumber="9">Host plant</emphasis>
. Larval habits and host plants are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF90FFFCC992F88F469FF8E0" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8BBE6659FF90FFFCC992F88F469FF8E0" blockId="8.[151,1437,151,2027]" pageId="8" pageNumber="9">
<emphasis id="B975BA4BFF90FFFCC992F88F4532F88D" bold="true" box="[199,326,1821,1846]" pageId="8" pageNumber="9">Etymology</emphasis>
. The new species is named after Anton Olegovich Kozlov (
<collectingRegion id="49C5A8BBFF90FFFCCD76F88C40F2F88D" box="[1059,1158,1822,1846]" country="Russia" name="Moscow" pageId="8" pageNumber="9">Moscow</collectingRegion>
, 
<collectingCountry id="F31626C9FF90FFFCCDC2F88F4092F88D" box="[1175,1254,1821,1846]" name="Russia" pageId="8" pageNumber="9">Russia</collectingCountry>
), for collecting specimens of 
<taxonomicName id="4C011DDAFF90FFFCC864F8D045DEF8E0" authorityName="Stephens" authorityYear="1829" box="[305,426,1858,1883]" class="Insecta" family="Hepialidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
resulting in this description.
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF90FFFCC992F8F74588F850" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BBE6659FF90FFFCC992F8F74588F850" blockId="8.[151,1437,151,2027]" pageId="8" pageNumber="9">
<materialsCitation id="3B696C04FF90FFFCC992F8F74588F818" ID-GBIF-Occurrence="1914299825" collectorName="S. America" country="Peru" location="Huanoco" pageId="8" pageNumber="9" specimenCount="2" stateProvince="Huanuco" typeStatus="holotype">
<emphasis id="B975BA4BFF90FFFCC992F8F7451BF8C5" bold="true" box="[199,367,1893,1918]" pageId="8" pageNumber="9">
<typeStatus id="54BAD8FBFF90FFFCC992F8F74576F8C5" box="[199,258,1893,1918]" pageId="8" pageNumber="9">Type</typeStatus>
material
</emphasis>
. 
<typeStatus id="54BAD8FBFF90FFFCC82BF8F74591F8C5" box="[382,485,1893,1918]" pageId="8" pageNumber="9" type="holotype">
<emphasis id="B975BA4BFF90FFFCC82BF8F74591F8C5" box="[382,485,1893,1918]" italics="true" pageId="8" pageNumber="9">Holotype</emphasis>
</typeStatus>
male (with the following labels separated by forward slashes): / 
<collectorName id="26F4038FFF90FFFCCD97F8F44135F8C5" box="[1218,1345,1893,1918]" pageId="8" pageNumber="9">S. America</collectorName>
, 
<collectingCountry id="F31626C9FF90FFFCCC18F8F541ECF8C5" box="[1357,1432,1895,1918]" name="Peru" pageId="8" pageNumber="9">PERU</collectingCountry>
, 
<location id="8EDE3082FF90FFFCC9C2F8194488F819" LSID="urn:lsid:plazi:treatment:03A8D74FFF9FFFFCC9C2F9874588F850:8EDE3082FF90FFFCC9C2F8194488F819" box="[151,252,1931,1954]" country="Peru" name="Huanoco" pageId="8" pageNumber="9" stateProvince="Huanuco">Huanoco</location>
[
<emphasis id="B975BA4BFF90FFFCC844F819453CF819" box="[273,328,1931,1954]" italics="true" pageId="8" pageNumber="9">recte</emphasis>
<collectingRegion id="49C5A8BBFF90FFFCC801F81845C8F819" box="[340,444,1930,1954]" country="Peru" name="Huanuco" pageId="8" pageNumber="9">Huánuco</collectingRegion>
] reg.
</materialsCitation>
, 
<materialsCitation id="3B696C04FF90FFFCCB58F8194588F850" ID-GBIF-Occurrence="1914299828" collectingDate="2017-02-02" collectionCode="CMNH" collectorName="Kozlov &amp; Yu. Kovaleva &amp; R. Gortovannyi &amp; Dissection" country="Peru" elevation="3270" latitude="-8.68902" location="Maranon" longLatPrecision="1" longitude="-76.99518" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" stateProvince="Maranon" typeStatus="holotype">
<collectingRegion id="49C5A8BBFF90FFFCCB58F8194606F819" box="[525,626,1931,1954]" country="Brazil" name="Maranhao" pageId="8" pageNumber="9">Maranon</collectingRegion>
prov. 
<geoCoordinate id="EE35009EFF90FFFCCB96F818474AF818" box="[707,830,1930,1955]" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="1" value="-8.68902">08.68902S</geoCoordinate>
, 
<geoCoordinate id="EE35009EFF90FFFCCA1BF8184795F818" box="[846,993,1930,1955]" direction="west" orientation="longitude" pageId="8" pageNumber="9" precision="1" value="-76.99518">076.99518W</geoCoordinate>
, h= 
<elevation id="002C816AFF90FFFCCD5BF818402BF818" box="[1038,1119,1930,1955]" metricMagnitude="3" metricUnit="m" metricValue="3.27" pageId="8" pageNumber="9" unit="m" value="3270.0">
<quantity id="4CF9CBBCFF90FFFCCD5BF818402BF818" box="[1038,1119,1930,1955]" metricMagnitude="3" metricUnit="m" metricValue="3.27" pageId="8" pageNumber="9" unit="m" value="3270.0">3270m</quantity>
</elevation>
, 
<collectingDate id="EFFBB971FF90FFFCCD25F81840AEF819" box="[1136,1242,1930,1954]" pageId="8" pageNumber="9" value="2017-02-02">
<date id="FFBF4099FF90FFFCCD25F81840AEF819" box="[1136,1242,1930,1954]" pageId="8" pageNumber="9" value="2017-02-02">2/II/2017</date>
</collectingDate>
, A. 
<collectorName id="26F4038FFF90FFFCCC47F8184112F818" box="[1298,1382,1930,1955]" pageId="8" pageNumber="9">Kozlov</collectorName>
, 
<collectorName id="26F4038FFF90FFFCCC23F8194571F87D" pageId="8" pageNumber="9">Yu. Kovaleva</collectorName>
, 
<collectorName id="26F4038FFF90FFFCC845F83D45CAF87D" box="[272,446,1965,1990]" pageId="8" pageNumber="9">R. Gortovannyi</collectorName>
leg./ 
<typeStatus id="54BAD8FBFF90FFFCC8A9F83F4607F87D" box="[508,627,1965,1990]" pageId="8" pageNumber="9" type="holotype">Holotypus</typeStatus>
, 
<taxonomicName id="4C011DDAFF90FFFCCB28F83F40DFF87D" authority="♂, Grehan &amp; C. Mielke" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[637,1195,1965,1990]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="antonkozlovi">
<emphasis id="B975BA4BFF90FFFCCB28F83F47E9F87D" box="[637,925,1965,1990]" italics="true" pageId="8" pageNumber="9">Pfitzneriella antonkozlovi</emphasis>
<specimenCount id="9D07ADD0FF90FFFCCAF1F83D47CFF87D" box="[932,955,1967,1990]" pageId="8" pageNumber="9" type="male">♂</specimenCount>
, Grehan &amp; C. Mielke
</taxonomicName>
det. 2017/ 
<collectorName id="26F4038FFF90FFFCCC7DF83F41E8F87D" box="[1320,1436,1965,1990]" pageId="8" pageNumber="9">Dissection</collectorName>
JRG 259/ (
<collectionCode id="ED10FE9CFF90FFFCC85BF8404513F851" box="[270,359,2002,2026]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="8" pageNumber="9">CMNH</collectionCode>
). 
<figureCitation id="133A7ADCFF90FFFCC82CF840458CF850" box="[377,504,2002,2027]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1707,1730]" captionTargetBox="[151,1436,195,1660]" captionTargetId="figure@4.[151,1436,195,1660]" captionTargetPageId="4" captionText="FIGURES 1–7. Voucher specimens. 1, Kozloviella viazmenskyi gen. n., sp. n. male HT (CMNH). 1a, dorsal; 1b, ventral. 2, Pfitzneriella antonkozlovi sp. n. male HT (CMNH), 2a, dorsal; 2b, ventral. 3, P. olafi sp. n. male HT (MNHN), 3a, dorsal; 3b, ventral. 4, P. rawlinsi sp. n. male, 4a, HT dorsal (CMNH); 4b, PT dorsal; 4c, PT ventral (CGCM). 5, P. remota male HT dorsal (MNHN). 6, P. titarenkoi sp. n. male HT (CMNH), 6a, dorsal; 6b, ventral. 7, P. yuliyakovalevae sp. n. male HT (CMNH), 7a, dorsal; 7b, ventral. Female PT (CMNH); 7c, dorsal; 7d, ventral. Photos: Ernst Brockmann (Fig. 5), Jane Hyland (Fig. 4a), Anton Kozlov (Figs 1, 2, 6, 7), and Carlos Mielke (Figs 3, 4b, 4c)." httpUri="https://zenodo.org/record/1451058/files/figure.png" pageId="8" pageNumber="9">Figs 2a, 2b</figureCitation>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>