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<mods:title id="77D20E0443EB752C7C56CF99C92F4B8A">A re-examination of the enigmatic Russian tetrapod Phreatophasma aenigmaticum and its evolutionary implications</mods:title>
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<mods:namePart id="8CE4389E6F536CD5DC6601E69DE15FD7">Brocklehurst, Neil</mods:namePart>
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<mods:namePart id="64101E567A2AAB6FDF41AF9895F08E48">Fröbisch, Jörg</mods:namePart>
<mods:affiliation id="699D0421DD59A5CC0736ADB2D483A1AD">Jörg Fröbisch &amp; Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse 43, &amp; Institut für Biologie, Humboldt-Universität zu Berlin, Invalidenstrasse 110, 10115 Berlin, Germany &amp; Evolutionary Studies Institute &amp; School of Geosciences, University of the Witwatersrand, Private Bag 3,</mods:affiliation>
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<taxonomicName id="4C394D75FF54E853FFD369AD3E27FB12" authority="Efremov, 1954" authorityName="Efremov" authorityYear="1954" box="[124,612,1212,1236]" family="Caseidae" genus="Phreatophasma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="1" pageNumber="88" phylum="Chordata" rank="species" species="aenigmaticum">
<emphasis id="B94DEAE4FF54E853FFD369AD3D82FB15" box="[124,449,1212,1236]" italics="true" pageId="1" pageNumber="88">Phreatophasma aenigmaticum</emphasis>
<bibRefCitation id="EFA84B07FF54E853FE6869AD3E27FB12" author="Efremov, I. A." box="[455,612,1212,1235]" pageId="1" pageNumber="88" pagination="1 - 146" refId="ref4629" refString="Efremov, I. A.: The fauna of terrestrial vertebrates in the Permian copper sandstones of the western Cis-Urals. Transactions of the Paleontological Institute, Academy of Science USSR, 56, 1 - 146, 1954." type="journal article" year="1954">Efremov, 1954</bibRefCitation>
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Diagnosis: distinguished from other members of
<taxonomicName id="4C394D75FF54E853FD1469ED3CE2FAF2" authorityName="Williston" authorityYear="1911" family="Caseidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="1" pageNumber="88" phylum="Chordata" rank="family">Caseidae</taxonomicName>
by the lack of a prominent, rugose mound on the dorsal surface and by the almost identical distal extent of the condyles of the femur.
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<typeStatus id="54828854FF54E853FFD3688D3CA0FA72" box="[124,227,1436,1459]" pageId="1" pageNumber="88" type="holotype">Holotype</typeStatus>
: PIN 294/25, right femur (
<figureCitation id="13022A73FF54E853FD8B688D3E29FA72" box="[548,618,1436,1459]" captionStart="Figure 1" captionStartId="1.[819,886,887,909]" captionTargetBox="[829,1446,183,854]" captionTargetId="figure-372@1.[827,1454,177,856]" captionTargetPageId="1" captionText="Figure 1. PIN 294/25, cast of the holotype of Phreatophasma aenigmaticum. (a) Ventral view; (b) dorsal view. Abbreviations: IF intertrochanteric fossa, AC anterior condyle, PC posterior condyle." figureDoi="http://doi.org/10.5281/zenodo.13751120" httpUri="https://zenodo.org/record/13751120/files/figure.png" pageId="1" pageNumber="88">Fig. 1</figureCitation>
). This specimen is a cast; the original has been lost.
</paragraph>
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Locality and horizon: Santagulov Mine,
<collectingRegion id="49FDF814FF54E853FDC868EA3C9EF9F3" country="Russia" name="Bashkortostan" pageId="1" pageNumber="88">Bashkortostan Province</collectingRegion>
,
<collectingCountry id="F32E7666FF54E853FF566B0A3D00F9F3" box="[249,323,1563,1586]" name="Russia" pageId="1" pageNumber="88">Russia</collectingCountry>
. Golyusherma Subassemblage, early Kazanian age (
<bibRefCitation id="EFA84B07FF54E853FEB06B2A3D80F993" author="Golubev, V." box="[287,451,1595,1618]" pageId="1" pageNumber="88" pagination="211 - 224" refId="ref4766" refString="Golubev, V.: The faunal assemblages of Permian terrestrial vertebrates from Eastern Europe, Paleontol. J., 34, 211 - 224, 2000." type="journal article" year="2000">Golubev, 2000</bibRefCitation>
), thought to be equivalent to the Roadian (
<bibRefCitation id="EFA84B07FF54E853FEA56B4A3DDCF9B3" author="Benton, M. J." box="[266,415,1627,1650]" pageId="1" pageNumber="88" pagination="339 - 342" refId="ref4203" refString="Benton, M. J.: No gap in the Middle Permian record of terrestrial vertebrates, Geology, 40, 339 - 342, 2012." type="journal article" year="2012">Benton, 2012</bibRefCitation>
).
</paragraph>
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