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<document id="4B9DE2274A389F80065D5E3C81733469" ID-DOI="10.4314/met.v34i1.6" ID-ISSN="2307-5031" ID-Zenodo-Dep="14119408" ID-ZooBank="0F510750-563D-4E6D-BE9F-F901EE02A422" checkinTime="1731462156187" checkinUser="felipe" docAuthor="Bouyer, Thierry" docDate="2023" docId="039087D1C252FF94168AFED4FC3D64CF" docLanguage="en" docName="Metamorphosis.34.59-78.pdf" docOrigin="Metamorphosis 34 (1)" docSource="http://dx.doi.org/10.4314/met.v34i1.6" docStyle="DocumentStyle:73CBAE06DE6A380ACC54A94377F5FCBA.1:Metamorphosis.2021-.journal_article" docStyleId="73CBAE06DE6A380ACC54A94377F5FCBA" docStyleName="Metamorphosis.2021-.journal_article" docStyleVersion="1" docTitle="Setechin Bouyer 2023, gen. nov." docType="treatment" docUuid="88B3DA93-55C1-4400-9700-8E1EFA342F01" docUuidSource="ZooBank" docVersion="2" lastPageNumber="67" masterDocId="FFA9FFA9C25AFF9C160EFF85FF846037" masterDocTitle="The genera of the tribe Charaxini (Papilionoidea, Nymphalidae, Charaxinae)" masterLastPageNumber="78" masterPageNumber="59" pageNumber="67" updateTime="1731462803107" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="239A0215E1D3AC9208292427C86D5E93">The genera of the tribe Charaxini (Papilionoidea, Nymphalidae, Charaxinae)</mods:title>
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<mods:title id="AA699C619513FC057167B93430792958">Metamorphosis</mods:title>
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<mods:date id="C190B4EEF51C997FCF6D182873C58F0B">2023</mods:date>
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<treatment id="039087D1C252FF94168AFED4FC3D64CF" LSID="urn:lsid:zoobank.org:act:88B3DA93-55C1-4400-9700-8E1EFA342F01" httpUri="http://treatment.plazi.org/id/039087D1C252FF94168AFED4FC3D64CF" lastPageNumber="67" pageId="8" pageNumber="67">
<subSubSection id="C323654CC252FF94168AFED4FD6C61B5" pageId="8" pageNumber="67" type="nomenclature">
<paragraph id="8B8636C7C252FF94168AFED4FE1A615E" blockId="8.[132,744,337,386]" box="[132,414,337,361]" pageId="8" pageNumber="67">
<heading id="D0CE81ABC252FF94168AFED4FE1A615E" box="[132,414,337,361]" fontSize="10" level="2" pageId="8" pageNumber="67" reason="2">
<emphasis id="B94DEAD5C252FF94168AFED4FE1A615E" bold="true" box="[132,414,337,361]" pageId="8" pageNumber="67">
Genus
<taxonomicName id="4C394D44C252FF9416DDFED4FEB6615E" authority="Bouyer, 2023" authorityName="Bouyer" authorityYear="2023" box="[211,306,337,361]" class="Insecta" family="Nymphalidae" genus="Setechin" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94DEAD5C252FF9416DDFED4FEB6615E" bold="true" box="[211,306,337,361]" italics="true" pageId="8" pageNumber="67">Setechin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E57AEC252FF941737FED4FE1A615E" box="[313,414,337,361]" pageId="8" pageNumber="67" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8636C7C252FF94168AFEEBFD6C61B5" blockId="8.[132,744,337,386]" box="[132,744,366,386]" pageId="8" pageNumber="67">
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urn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection id="C323654CC252FF9416CEFE24FC3D64CF" pageId="8" pageNumber="67" type="materials_examined">
<paragraph id="8B8636C7C252FF9416CEFE24FD70618F" blockId="8.[132,756,416,471]" box="[192,756,416,441]" pageId="8" pageNumber="67">
<typeStatus id="54828865C252FF9416CEFE24FF73618E" box="[192,247,417,441]" pageId="8" pageNumber="67">Type</typeStatus>
species of the genus:
<taxonomicName id="4C394D44C252FF9417EFFE25FD20618F" authorityName="Grose-Smith" authorityYear="1883" box="[481,676,416,440]" class="Insecta" family="Nymphalidae" genus="Charaxes" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="nichetes">
<emphasis id="B94DEAD5C252FF9417EFFE25FD20618F" box="[481,676,416,440]" italics="true" pageId="8" pageNumber="67">Charaxes nichetes</emphasis>
</taxonomicName>
Grose-
</paragraph>
<paragraph id="8B8636C7C252FF94168AFE3AFE8A61E0" blockId="8.[132,756,416,471]" box="[132,270,447,471]" pageId="8" pageNumber="67">Smith, 1883.</paragraph>
<paragraph id="8B8636C7C252FF94168AFE79FD666204" blockId="8.[132,755,508,563]" pageId="8" pageNumber="67">
Corresponds to the
<taxonomicName id="4C394D44C252FF94175AFE79FE2F6223" authorityName="Grose-Smith" authorityYear="1883" box="[340,427,508,532]" class="Insecta" family="Nymphalidae" genus="Charaxes" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="nichetes">
<emphasis id="B94DEAD5C252FF94175AFE79FE2F6223" box="[340,427,508,532]" italics="true" pageId="8" pageNumber="67">nichetes</emphasis>
</taxonomicName>
-group of
<bibRefCitation id="EFA84B36C252FF94141BFE79FD776223" author="HENNING, S. F." box="[533,755,508,532]" pageId="8" pageNumber="67" refId="ref8385" refString="HENNING, S. F. 1989. The Charaxinae Butterflies of Africa. Aloe Books, Johannesburg. 457 p." type="book" year="1989">Henning (1989: 380)</bibRefCitation>
and species-group Nichetes of
<bibRefCitation id="EFA84B36C252FF9417C3FD9EFD5A6204" author="ADUSE-POKU, K. &amp; VINGERHOEDT, E. &amp; WAHLBERG, N." box="[461,734,539,563]" pageId="8" pageNumber="67" pagination="463 - 478" refId="ref7924" refString="ADUSE-POKU, K., VINGERHOEDT, E., &amp; WAHLBERG, N. 2009. Out-of-Africa again: A phylogenetic hypothesis of the genus Charaxes (Lepidoptera: Nymphalidae) based on five gene regions. Molecular Phylogenetics and Evolution 53 (2): 463 - 478." type="journal article" year="2009">
Aduse-Poku
<emphasis id="B94DEAD5C252FF941456FD99FD0A6204" box="[600,654,539,563]" italics="true" pageId="8" pageNumber="67">et al.</emphasis>
(2009)
</bibRefCitation>
.
</paragraph>
<paragraph id="8B8636C7C252FF94168AFDD6FE4D62BE" blockId="8.[132,756,594,649]" pageId="8" pageNumber="67">
<emphasis id="B94DEAD5C252FF94168AFDD6FEC0625D" box="[132,324,595,618]" italics="true" pageId="8" pageNumber="67">Derivatio nominis</emphasis>
.
<taxonomicName id="4C394D44C252FF94175CFDD7FE28625D" authorityName="Bouyer" authorityYear="2023" box="[338,428,594,618]" class="Insecta" family="Nymphalidae" genus="Setechin" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEAD5C252FF94175CFDD7FE28625D" box="[338,428,594,618]" italics="true" pageId="8" pageNumber="67">Setechin</emphasis>
</taxonomicName>
is the anagram of
<taxonomicName id="4C394D44C252FF941461FDD7FD42625D" authorityName="Grose-Smith" authorityYear="1883" box="[623,710,594,618]" class="Insecta" family="Nymphalidae" genus="Charaxes" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="nichetes">
<emphasis id="B94DEAD5C252FF941461FDD7FD42625D" box="[623,710,594,618]" italics="true" pageId="8" pageNumber="67">nichetes</emphasis>
</taxonomicName>
, the
<typeStatus id="54828865C252FF94168AFDF4FF3662BE" box="[132,178,625,649]" pageId="8" pageNumber="67">type</typeStatus>
species of the new genus.
</paragraph>
<paragraph id="8B8636C7C252FF941531FF12FA2B6098" blockId="8.[831,1455,155,205]" box="[831,1455,151,175]" pageId="8" pageNumber="67">pointed when being short, and spatulate when being</paragraph>
<paragraph id="8B8636C7C252FF941531FF30FC2560FA" blockId="8.[831,1455,155,205]" box="[831,929,181,205]" pageId="8" pageNumber="67">elongate.</paragraph>
<paragraph id="8B8636C7C252FF941531FF69FB566175" blockId="8.[831,1454,236,323]" pageId="8" pageNumber="67">
Shape of the forewing elongated with the outer edge very sinuate and the apex clearly rounded (reminiscent of certain species of the genus
<taxonomicName id="4C394D44C252FF941262FEAFFB466175" authorityName="Bouyer" authorityYear="2023" box="[1132,1218,298,322]" class="Hydrozoa" family="Laodiceidae" genus="Laodice" kingdom="Animalia" order="Leptothecata" pageId="8" pageNumber="67" phylum="Cnidaria" rank="genus">
<emphasis id="B94DEAD5C252FF941262FEAFFB466175" box="[1132,1218,298,322]" italics="true" pageId="8" pageNumber="67">Laodice</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B8636C7C252FF941531FEE4FA0F62BA" blockId="8.[831,1456,353,653]" pageId="8" pageNumber="67">The brown or black markings on the upperside are composed of a cellular spot (sometimes discreet), a marked discocellular stroke, a discal series of more or less Sshaped interveinal spots, a postdiscal band (or submarginal) chevrons and a separate or contiguous marginal band at the edge of the wing of more or less conspicuous spots. The hindwing patterns are essentially composed of a discal band and a series of submarginal interveinal circles. The markings are more or less scattered, sometimes merged (depending on climatic conditions).</paragraph>
<paragraph id="8B8636C7C252FF941531FD29FA836335" blockId="8.[831,1456,684,770]" pageId="8" pageNumber="67">The underside is shiny with markings in fine reddishbrown lines. The two wings are crossed by a postdiscal dark band slightly straight and continuous.</paragraph>
<caption id="DF46664FC252FF94168AFC7AFE78647B" ID-DOI="http://doi.org/10.5281/zenodo.14119450" ID-Zenodo-Dep="14119450" httpUri="https://zenodo.org/record/14119450/files/figure.png" pageId="8" pageNumber="67" startId="8.[132,200,1023,1044]" targetBox="[132,764,684,969]" targetPageId="8" targetType="figure">
<paragraph id="8B8636C7C252FF94168AFC7AFE78647B" blockId="8.[132,737,1023,1100]" pageId="8" pageNumber="67">
<emphasis id="B94DEAD5C252FF94168AFC7AFDE56423" bold="true" box="[132,609,1023,1044]" pageId="8" pageNumber="67">
Figure 17 Type species of the genus
<taxonomicName id="4C394D44C252FF941408FC7AFDDF6423" authorityName="Bouyer" authorityYear="2023" box="[518,603,1023,1044]" class="Insecta" family="Nymphalidae" genus="Setechin" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEAD5C252FF941408FC7AFDDF6423" bold="true" box="[518,603,1023,1044]" italics="true" pageId="8" pageNumber="67">Setechin</emphasis>
</taxonomicName>
.
</emphasis>
<taxonomicName id="4C394D44C252FF941469FC7AFF576407" authorityName="Bouyer" authorityYear="2023" baseAuthorityName="Grose-Smith" baseAuthorityYear="1883" class="Insecta" family="Nymphalidae" genus="Setechin" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="nichetes">
<emphasis id="B94DEAD5C252FF941469FC7AFF576407" italics="true" pageId="8" pageNumber="67">Setechin nichetes</emphasis>
</taxonomicName>
male upperside on left and underside on right (Gabon,
<emphasis id="B94DEAD5C252FF94168AFBB2FF13647B" box="[132,151,1079,1100]" italics="true" pageId="8" pageNumber="67">in</emphasis>
coll. G. Faravel). FW length 34 mm.
</paragraph>
</caption>
<caption id="DF46664FC252FF94168AFA3AFF34663C" ID-DOI="http://doi.org/10.5281/zenodo.14119454" ID-Zenodo-Dep="14119454" httpUri="https://zenodo.org/record/14119454/files/figure.png" pageId="8" pageNumber="67" startId="8.[132,200,1471,1492]" targetBox="[270,618,1128,1436]" targetPageId="8" targetType="figure">
<paragraph id="8B8636C7C252FF94168AFA3AFF34663C" blockId="8.[132,745,1471,1547]" pageId="8" pageNumber="67">
<emphasis id="B94DEAD5C252FF94168AFA3AFDC965E3" bold="true" box="[132,589,1471,1492]" pageId="8" pageNumber="67">
Figure 18
<taxonomicName id="4C394D44C252FF9416F0FA3AFE2F65E3" authorityName="Bouyer" authorityYear="2023" baseAuthorityName="Grose-Smith" baseAuthorityYear="1883" box="[254,427,1471,1492]" class="Insecta" family="Nymphalidae" genus="Setechin" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="nichetes">
<emphasis id="B94DEAD5C252FF9416F0FA3AFE2F65E3" bold="true" box="[254,427,1471,1492]" italics="true" pageId="8" pageNumber="67">Setechin nichetes</emphasis>
</taxonomicName>
male upperside
</emphasis>
(D.R. Congo, Haut-Katanga, Kibomboma,
<emphasis id="B94DEAD5C252FF941795FA5EFE2965C7" box="[411,429,1499,1520]" italics="true" pageId="8" pageNumber="67">in</emphasis>
coll. Th. Bouyer). FW length 33 mm.
</paragraph>
</caption>
<caption id="DF46664FC252FF94168AF806FE306783" ID-DOI="http://doi.org/10.5281/zenodo.14119456" ID-Zenodo-Dep="14119456" httpUri="https://zenodo.org/record/14119456/files/figure.png" pageId="8" pageNumber="67" startId="8.[132,200,1923,1944]" targetBox="[252,636,1576,1890]" targetPageId="8" targetType="figure">
<paragraph id="8B8636C7C252FF94168AF806FE306783" blockId="8.[132,757,1923,1972]" pageId="8" pageNumber="67">
<emphasis id="B94DEAD5C252FF94168AF806FF6367AF" bold="true" box="[132,231,1923,1944]" pageId="8" pageNumber="67">Figure 19</emphasis>
<taxonomicName id="4C394D44C252FF94170FF806FE2C67AE" authorityName="Bouyer" authorityYear="2023" baseAuthorityName="Grose-Smith" baseAuthorityYear="1883" box="[257,424,1923,1945]" class="Insecta" family="Nymphalidae" genus="Setechin" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="nichetes">
<emphasis id="B94DEAD5C252FF94170FF806FE2C67AE" box="[257,424,1923,1945]" italics="true" pageId="8" pageNumber="67">Setechin nichetes</emphasis>
</taxonomicName>
female upperside (Gabon,
<emphasis id="B94DEAD5C252FF9414BFF806FD4067AF" box="[689,708,1923,1944]" italics="true" pageId="8" pageNumber="67">in</emphasis>
coll. G. Faravel). FW length 42 mm.
</paragraph>
</caption>
<paragraph id="8B8636C7C252FF94168AF856FBFE63C6" blockId="8.[132,755,2003,2088]" lastBlockId="8.[831,1455,801,1009]" pageId="8" pageNumber="67">
A basically orange
<taxonomicName id="4C394D44C252FF94175CF856FE3F67DC" box="[338,443,2003,2027]" class="Insecta" family="Nymphalidae" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="tribe" tribe="Charaxini">Charaxini</taxonomicName>
with black or brown patterns on the upperside and underside. Hindwings not caudate except for some females with a tail at M3. This tail is The individual variation is strongly subject to climatic or seasonal conditions with the dark pattern on the upperside more extensive as the conditions get more humid. Females are caudate or not, sometimes even within a same population. These are dichromatic black and orange, sometimes trichromatic with the postdiscal orange part lighter, sometimes yellowish.
</paragraph>
<paragraph id="8B8636C7C252FF941531FB95FCFC6495" blockId="8.[831,1456,1040,1187]" pageId="8" pageNumber="67">
Early stages. The genus is linked to
<taxonomicName id="4C394D44C252FF9412E7FB94FAB9641F" box="[1257,1341,1041,1064]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="genus">
<emphasis id="B94DEAD5C252FF9412E7FB94FAB9641F" box="[1257,1341,1041,1064]" italics="true" pageId="8" pageNumber="67">Uapaca</emphasis>
</taxonomicName>
foodplant (Euphorbiacae):
<taxonomicName id="4C394D44C252FF9415F6FBB5FA986470" authority="Mull. Arg." authorityName="Mull. Arg." box="[1016,1308,1071,1095]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="species" species="nitida">
<emphasis id="B94DEAD5C252FF9415F6FBB5FB136470" box="[1016,1175,1071,1095]" italics="true" pageId="8" pageNumber="67">Uapaca nitida</emphasis>
Müll. Arg.
</taxonomicName>
and
<taxonomicName id="4C394D44C252FF941355FBB5FBAC6452" authority="Mull. Arg." authorityName="Mull. Arg." class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="species" species="kirkiana">
<emphasis id="B94DEAD5C252FF941355FBB5FC1C6452" italics="true" pageId="8" pageNumber="67">Uapaca kirkiana</emphasis>
Müll. Arg.
</taxonomicName>
Description of early stages and illustration of final instar larva
<emphasis id="B94DEAD5C252FF9412ABFBE8FB3E64B3" box="[1189,1210,1133,1156]" italics="true" pageId="8" pageNumber="67">in</emphasis>
<bibRefCitation id="EFA84B36C252FF9412C8FBE9FCF56494" author="HENNING, S. F." pageId="8" pageNumber="67" refId="ref8385" refString="HENNING, S. F. 1989. The Charaxinae Butterflies of Africa. Aloe Books, Johannesburg. 457 p." type="book" year="1989">Henning (1989: 382- 383)</bibRefCitation>
.
</paragraph>
<paragraph id="8B8636C7C252FF941531FB47FC3D64CF" blockId="8.[831,1455,1218,1272]" pageId="8" pageNumber="67">The genus contains a single species which is divided into 6 subspecies.</paragraph>
</subSubSection>
</treatment>
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