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<mods:title id="CE5636833AB432C578E21F25ECACBCBD">Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura)</mods:title>
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<heading id="D0CE81B77E15733847C6F89FF5A0F8C6" bold="true" box="[151,712,1794,1821]" fontSize="11" level="1" pageId="16" pageNumber="444" reason="1">
<taxonomicName id="4C394D587E15733847C6F89FF5A0F8C6" ID-CoL="dd79bc86-eb9c-400a-b43e-fc2a93bff2d9" authority="(Edmondson, 1951)" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[151,712,1794,1821]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="444" phylum="Arthropoda" rank="species" species="symmetrinuda">
<emphasis id="B94DEAC97E15733847C6F89FF5A0F8C6" bold="true" box="[151,712,1794,1821]" pageId="16" pageNumber="444">
<emphasis id="B94DEAC97E15733847C6F89FF6A6F8C7" bold="true" box="[151,462,1794,1821]" italics="true" pageId="16" pageNumber="444">Lophoplax symmetrinuda</emphasis>
(
<bibRefCitation id="EFA84B2A7E157338468CF89FF5D7F8C6" author="Edmondson, C. H." box="[477,703,1794,1820]" pageId="16" pageNumber="444" pagination="183 - 243" refId="ref15227" refString="Edmondson, C. H. (1951) Some Central Pacific Crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
)
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<paragraph id="8B8636DB7E15733847C6F8BAF792F89B" blockId="16.[151,712,1794,1857]" box="[151,250,1831,1857]" pageId="16" pageNumber="444">
(
<figureCitation id="13022A5E7E15733847CEF8BAF799F89B" box="[159,241,1831,1857]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="16" pageNumber="444">Fig. 12</figureCitation>
)
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<subSubSection id="C32365507E15733847C6F8F6F374F81B" pageId="16" pageNumber="444" type="reference_group">
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<treatmentCitationGroup id="AB2911F57E15733847C6F8F6F3F9F87B" pageId="16" pageNumber="444">
<treatmentCitation id="0A9810CA7E15733847C6F8F6F466F859" author="Edmondson, C. H." box="[151,782,1899,1923]" page="233" pageId="16" pageNumber="444" year="1951">
<taxonomicName id="4C394D587E15733847C6F8F6F466F859" ID-CoL="4NV9Q" authority="Edmondson, 1951: 233" authorityName="Edmondson" authorityPageNumber="233" authorityYear="1951" box="[151,782,1899,1923]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="444" phylum="Arthropoda" rank="species" species="symmetrinudus">
<emphasis id="B94DEAC97E15733847C6F8F6F570F859" box="[151,536,1899,1923]" italics="true" pageId="16" pageNumber="444">Pseudocryptocoeloma symmetrinudus</emphasis>
<bibRefCitation id="EFA84B2A7E157338454EF8F6F466F859" author="Edmondson, C. H." box="[543,782,1899,1923]" pageId="16" pageNumber="444" pagination="183 - 243" refId="ref15227" refString="Edmondson, C. H. (1951) Some Central Pacific Crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951: 233</bibRefCitation>
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, fig. 34;
<treatmentCitation id="0A9810CA7E1573384438F8F6F379F859" author="Serene, R." box="[873,1041,1899,1923]" page="86" pageId="16" pageNumber="444" year="1968">
<bibRefCitation id="EFA84B2A7E1573384438F8F6F379F859" author="Serene, R." box="[873,1041,1899,1923]" pageId="16" pageNumber="444" pagination="33 - 120" refId="ref18274" refString="Serene, R. (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a Check List of the Non-planctonic Marine Fauna of South East Asia. Special Publication of the Singapore National Academy of Science, No. 1, 33 - 120." type="book chapter" year="1968">Serène 1968: 86</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9810CA7E157338434DF8F1F3F6F859" author="Ng, P. K. L." box="[1052,1182,1899,1924]" page="79" pageId="16" pageNumber="444" year="1987">
<bibRefCitation id="EFA84B2A7E157338434DF8F1F3F6F859" author="Ng, P. K. L." box="[1052,1182,1899,1924]" pageId="16" pageNumber="444" pagination="69 - 111" refId="ref16944" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). Indo-Malayan Zoology, 1987, 4 (1), 69 - 111, pl. 1." type="journal article" year="1987">Ng 1987: 79</bibRefCitation>
</treatmentCitation>
, 97;
<treatmentCitation id="0A9810CA7E157338439EF8F1F2F0F859" author="Ng, P. K. L. &amp; D. Guinot &amp; P. J. F. Davie" box="[1231,1432,1899,1924]" page="144" pageId="16" pageNumber="444" year="2008">
<bibRefCitation id="EFA84B2A7E157338439EF8F1F2F0F859" author="Ng, P. K. L. &amp; D. Guinot &amp; P. J. F. Davie" box="[1231,1432,1899,1924]" pageId="16" pageNumber="444" pagination="1 - 286" refId="ref17397" refString="Ng, P. K. L., D. Guinot &amp; P. J. F. Davie, 2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="B94DEAC97E15733843A4F8F1F241F859" box="[1269,1321,1899,1923]" italics="true" pageId="16" pageNumber="444">et al.</emphasis>
2008: 144
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9810CA7E1573384796F817F561F878" author="Marumura, M. &amp; Takeda, M." box="[199,521,1930,1954]" page="192" pageId="16" pageNumber="444" year="2012">
<bibRefCitation id="EFA84B2A7E1573384796F817F561F878" author="Marumura, M. &amp; Takeda, M." box="[199,521,1930,1954]" pageId="16" pageNumber="444" pagination="189 - 197" refId="ref16388" refString="Marumura, M. &amp; Takeda, M. (2012) Crab specimens in the Wakayama Prefectural Museum of Nature (Nagai Collection) taxonomic study III. New record species in Japan (3). Teikyo Heisei University Bulletin, 23 (1), 189 - 197." type="journal article" year="2012">Marumura &amp; Takeda 2012: 192</bibRefCitation>
</treatmentCitation>
, figs. 2A, 3, 4;
<treatmentCitation id="0A9810CA7E15733845F4F817F416F878" author="Maenosono, T." box="[677,894,1930,1954]" page="32" pageId="16" pageNumber="444" year="2019">
<bibRefCitation id="EFA84B2A7E15733845F4F817F416F878" author="Maenosono, T." box="[677,894,1930,1954]" pageId="16" pageNumber="444" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019: 32</bibRefCitation>
</treatmentCitation>
, figs. 8B, 10, 13B, 14K, L.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8636DB7E15733847C6F834F374F81B" blockId="16.[151,1436,1899,1985]" box="[151,1052,1960,1985]" pageId="16" pageNumber="444">
<treatmentCitationGroup id="AB2911F57E15733847C6F834F374F81B" box="[151,1052,1960,1985]" pageId="16" pageNumber="444">
<emphasis id="B94DEAC97E15733847C6F834F6D2F81B" box="[151,442,1961,1985]" italics="true" pageId="16" pageNumber="444">
<taxonomicName id="4C394D587E15733847C6F834F6C8F81B" authority="(Edmondson, 1951)" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[151,416,1961,1985]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="444" phylum="Arthropoda" rank="species" species="symmetrinudus">Lophoplax symmetrinudus</taxonomicName>
</emphasis>
<treatmentCitation id="0A9810CA7E15733846EBF834F5BFF81B" author="Takeda, M. &amp; Kurata, Y." box="[442,727,1961,1985]" page="201" pageId="16" pageNumber="444" year="1984">
<bibRefCitation id="EFA84B2A7E15733846EBF834F5BFF81B" author="Takeda, M. &amp; Kurata, Y." box="[442,727,1961,1985]" pageId="16" pageNumber="444" pagination="195 - 202" refId="ref18657" refString="Takeda, M. &amp; Kurata, Y. (1984) Crabs of the Ogasawara Islands. VII. Third Report on the Species Obtained from Stomachs of Fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4), 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata 1984: 201</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9810CA7E15733845B0F834F370F81B" author="Takeda, M. &amp; Marumura, M." box="[737,1048,1960,1985]" page="90" pageId="16" pageNumber="444" year="1995">
<bibRefCitation id="EFA84B2A7E15733845B0F834F370F81B" author="Takeda, M. &amp; Marumura, M." box="[737,1048,1960,1985]" pageId="16" pageNumber="444" pagination="87 - 91" refId="ref18718" refString="Takeda, M. &amp; Marumura, M. (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2), 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura 1995: 90</bibRefCitation>
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.
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<caption id="DF4666537E14733947C6FA12F217FA16" ID-DOI="http://doi.org/10.5281/zenodo.7663805" ID-Zenodo-Dep="7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="17" pageNumber="445" startId="17.[151,250,1423,1447]" targetBox="[159,1422,188,1386]" targetPageId="17">
<paragraph id="8B8636DB7E14733947C6FA12F217FA16" blockId="17.[151,1436,1423,1484]" pageId="17" pageNumber="445">
<emphasis id="B94DEAC97E14733947C6FA12F649FA72" bold="true" box="[151,289,1423,1448]" pageId="17" pageNumber="445">FIGURE 12.</emphasis>
<taxonomicName id="4C394D587E1473394678FA0DF46AFA72" authority="(Edmondson, 1951)" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[297,770,1424,1448]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="445" phylum="Arthropoda" rank="species" species="symmetrinuda">
<emphasis id="B94DEAC97E1473394678FA0DF545FA72" box="[297,557,1424,1448]" italics="true" pageId="17" pageNumber="445">Lophoplax symmetrinuda</emphasis>
(
<bibRefCitation id="EFA84B2A7E147339456AFA0DF592FA72" author="Edmondson, C. H." box="[571,762,1424,1448]" pageId="17" pageNumber="445" pagination="183 - 243" refId="ref15227" refString="Edmondson, C. H. (1951) Some Central Pacific Crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
)
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, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon.
</paragraph>
</caption>
<subSubSection id="C32365507E14733947C6FA67F643F9E2" pageId="17" pageNumber="445" type="materials_examined">
<paragraph id="8B8636DB7E14733947C6FA67F643F9E2" blockId="17.[151,1437,1530,2024]" pageId="17" pageNumber="445">
<emphasis id="B94DEAC97E14733947C6FA67F657F9CE" bold="true" box="[151,319,1530,1556]" pageId="17" pageNumber="445">Type material</emphasis>
.
<materialsCitation id="3B513C867E147339461FFA67F643F9E2" ID-GBIF-Occurrence="4035852327" collectionCode="BPBM" collectorName="W. Harris" country="Samoa" location="Samoa" pageId="17" pageNumber="445" specimenCode="BPBM 5109" specimenCount="1" typeStatus="holotype">
<emphasis id="B94DEAC97E147339461FFA67F6DDF9CE" box="[334,437,1530,1556]" italics="true" pageId="17" pageNumber="445">
<typeStatus id="548288797E147339461FFA67F6DDF9CE" box="[334,437,1530,1556]" pageId="17" pageNumber="445" type="holotype">Holotype</typeStatus>
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: male (6.5 × 5.0 mm) (
<specimenCode id="DB9F9EA07E1473394591FA67F43BF9CF" box="[704,851,1530,1557]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="17" pageNumber="445" type="Museum">BPBM 5109</specimenCode>
), low tide, table reef, Siufaga, Tau,
<collectingCountry id="F32E764B7E14733943B8FA67F250F9CE" box="[1257,1336,1530,1556]" name="Samoa" pageId="17" pageNumber="445">Samoa</collectingCountry>
, coll.
<collectorName id="26CC530D7E147339422CFA67F788F9E2" pageId="17" pageNumber="445">W. Harris</collectorName>
, 1937.
</materialsCitation>
</paragraph>
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<subSubSection id="C32365507E14733A4796F9DFF3ABFE53" lastPageId="18" lastPageNumber="446" pageId="17" pageNumber="445" type="diagnosis">
<paragraph id="8B8636DB7E14733A4796F9DFF22FFE9B" blockId="17.[151,1437,1530,2024]" lastBlockId="18.[151,1437,151,1149]" lastPageId="18" lastPageNumber="446" pageId="17" pageNumber="445">
<emphasis id="B94DEAC97E1473394796F9DFF652F986" bold="true" box="[199,314,1602,1628]" pageId="17" pageNumber="445">Diagnosis</emphasis>
. Carapace transversely subovate (
<figureCitation id="13022A5E7E14733945EFF9DFF44FF986" box="[702,807,1602,1628]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="17" pageNumber="445">Fig. 12A</figureCitation>
;
<bibRefCitation id="EFA84B2A7E1473394465F9DFF30DF986" author="Marumura, M. &amp; Takeda, M." box="[820,1125,1602,1628]" pageId="17" pageNumber="445" pagination="189 - 197" refId="ref16388" refString="Marumura, M. &amp; Takeda, M. (2012) Crab specimens in the Wakayama Prefectural Museum of Nature (Nagai Collection) taxonomic study III. New record species in Japan (3). Teikyo Heisei University Bulletin, 23 (1), 189 - 197." type="journal article" year="2012">Marumura &amp; Takeda 2012</bibRefCitation>
: fig. 3A;
<bibRefCitation id="EFA84B2A7E1473394383F9DFF2F1F986" author="Maenosono, T." box="[1234,1433,1602,1628]" pageId="17" pageNumber="445" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 10A); epigastric, protogastric, hepatic, cardiac and intestinal areolets all glabrous, prominent, smooth; epigastric areolets relatively wide, ovate, short (
<figureCitation id="13022A5E7E1473394515F917F5C5F97E" box="[580,685,1674,1700]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="17" pageNumber="445">Fig. 12A</figureCitation>
;
<bibRefCitation id="EFA84B2A7E14733945EBF917F483F97E" author="Marumura, M. &amp; Takeda, M." box="[698,1003,1674,1700]" pageId="17" pageNumber="445" pagination="189 - 197" refId="ref16388" refString="Marumura, M. &amp; Takeda, M. (2012) Crab specimens in the Wakayama Prefectural Museum of Nature (Nagai Collection) taxonomic study III. New record species in Japan (3). Teikyo Heisei University Bulletin, 23 (1), 189 - 197." type="journal article" year="2012">Marumura &amp; Takeda 2012</bibRefCitation>
: fig. 3A;
<bibRefCitation id="EFA84B2A7E1473394308F917F248F97E" author="Maenosono, T." box="[1113,1312,1674,1700]" pageId="17" pageNumber="445" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 10A); protogastric areolet prominent, transversely ovate positioned obliquely (
<figureCitation id="13022A5E7E1473394494F933F346F912" box="[965,1070,1710,1736]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="17" pageNumber="445">Fig. 12A</figureCitation>
;
<bibRefCitation id="EFA84B2A7E147339436AF933F202F912" author="Marumura, M. &amp; Takeda, M." box="[1083,1386,1710,1736]" pageId="17" pageNumber="445" pagination="189 - 197" refId="ref16388" refString="Marumura, M. &amp; Takeda, M. (2012) Crab specimens in the Wakayama Prefectural Museum of Nature (Nagai Collection) taxonomic study III. New record species in Japan (3). Teikyo Heisei University Bulletin, 23 (1), 189 - 197." type="journal article" year="2012">Marumura &amp; Takeda 2012</bibRefCitation>
: fig. 3A;
<bibRefCitation id="EFA84B2A7E1473394798F94FF6E7F936" author="Maenosono, T." box="[201,399,1746,1772]" pageId="17" pageNumber="445" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 10A); cardiac areolet medially separated, each element positioned obliquely (
<figureCitation id="13022A5E7E147339427EF94FF2F0F936" box="[1327,1432,1746,1772]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="17" pageNumber="445">Fig. 12A</figureCitation>
;
<bibRefCitation id="EFA84B2A7E14733947C6F96BF6A8F8CA" author="Marumura, M. &amp; Takeda, M." box="[151,448,1782,1808]" pageId="17" pageNumber="445" pagination="189 - 197" refId="ref16388" refString="Marumura, M. &amp; Takeda, M. (2012) Crab specimens in the Wakayama Prefectural Museum of Nature (Nagai Collection) taxonomic study III. New record species in Japan (3). Teikyo Heisei University Bulletin, 23 (1), 189 - 197." type="journal article" year="2012">Marumura &amp; Takeda 2012</bibRefCitation>
: fig. 3A;
<bibRefCitation id="EFA84B2A7E1473394574F96BF582F8CA" author="Maenosono, T." box="[549,746,1782,1808]" pageId="17" pageNumber="445" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 10A); hepatic areolet swollen, long, reaching only to base of second anterolateral tooth (
<figureCitation id="13022A5E7E14733946BFF887F530F8EE" box="[494,600,1818,1844]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="17" pageNumber="445">Fig. 12A</figureCitation>
;
<bibRefCitation id="EFA84B2A7E1473394537F887F446F8EE" author="Maenosono, T." box="[614,814,1818,1844]" pageId="17" pageNumber="445" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 10A); external orbital tooth wide, low (
<figureCitation id="13022A5E7E147339427FF887F2F0F8EE" box="[1326,1432,1818,1844]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="17" pageNumber="445">Fig. 12A</figureCitation>
;
<bibRefCitation id="EFA84B2A7E14733947C6F8A3F635F882" author="Maenosono, T." box="[151,349,1854,1880]" pageId="17" pageNumber="445" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 10A, B); tips of external orbital and anterolateral teeth low, rounded (
<figureCitation id="13022A5E7E14733943F4F8A3F266F882" box="[1189,1294,1854,1880]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="17" pageNumber="445">Fig. 12A</figureCitation>
;
<bibRefCitation id="EFA84B2A7E147339424BF8A3F7BBF8A6" author="Maenosono, T." pageId="17" pageNumber="445" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 10A, B); junction between antero- and posterolateral margins demarcated by second anterolateral tooth without additional lobes or tubercles (
<figureCitation id="13022A5E7E14733945D8F81BF59AF87A" box="[649,754,1926,1952]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="17" pageNumber="445">Fig. 12A</figureCitation>
;
<bibRefCitation id="EFA84B2A7E14733945AEF81BF4AEF87A" author="Maenosono, T." box="[767,966,1926,1952]" pageId="17" pageNumber="445" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 10A, B); epistome relatively narrow longitudinally (
<bibRefCitation id="EFA84B2A7E1473394614F837F567F81E" author="Maenosono, T." box="[325,527,1962,1988]" pageId="17" pageNumber="445" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 10C); median lobe of posterior margin of the epistome broadly triangular, with lateral margins gently concave (
<bibRefCitation id="EFA84B2A7E147339456DF853F46DF832" author="Maenosono, T." box="[572,773,1998,2024]" pageId="17" pageNumber="445" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 10C). Third maxilliped with merus quadrate, ischium short (
<figureCitation id="13022A5E7E17733A478CFF0AF62BFF6B" box="[221,323,151,177]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="18" pageNumber="446">Fig. 12B</figureCitation>
). Dorsal and lateral surfaces of carpus of cheliped rugose; cheliped fingers not distinctly bent, pollex subparallel with the ventral margin of palm (
<bibRefCitation id="EFA84B2A7E17733A45D2FF26F422FF0F" author="Maenosono, T." box="[643,842,187,213]" pageId="18" pageNumber="446" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 10D). P2P5 very short, not elongate (
<figureCitation id="13022A5E7E17733A4278FF26F2E7FF0F" box="[1321,1423,187,213]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="18" pageNumber="446">Fig. 12C</figureCitation>
); outer surface of the P3P5 merus, carpus and propodus smooth (
<figureCitation id="13022A5E7E17733A443EFF42F4B0FF23" box="[879,984,223,249]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="18" pageNumber="446">Fig. 12A</figureCitation>
). Male pleonal somites 46 transversely wide, telson wider than long, semicircular (
<figureCitation id="13022A5E7E17733A45D3FE9EF583FEC7" box="[642,747,259,285]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="18" pageNumber="446">Fig. 12D</figureCitation>
;
<bibRefCitation id="EFA84B2A7E17733A45A9FE9EF4D6FEC7" author="Maenosono, T." box="[760,958,259,285]" pageId="18" pageNumber="446" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 13B). G1 slender, sinuous, distal part gently curved upwards (
<figureCitation id="13022A5E7E17733A46F7FEBAF540FE9B" box="[422,552,295,321]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="18" pageNumber="446">Fig. 12E, F</figureCitation>
;
<bibRefCitation id="EFA84B2A7E17733A4562FEBAF408FE9B" author="Marumura, M. &amp; Takeda, M." box="[563,864,295,321]" pageId="18" pageNumber="446" pagination="189 - 197" refId="ref16388" refString="Marumura, M. &amp; Takeda, M. (2012) Crab specimens in the Wakayama Prefectural Museum of Nature (Nagai Collection) taxonomic study III. New record species in Japan (3). Teikyo Heisei University Bulletin, 23 (1), 189 - 197." type="journal article" year="2012">Marumura &amp; Takeda 2012</bibRefCitation>
: fig. 4A, B;
<bibRefCitation id="EFA84B2A7E17733A44BBFEBAF3C7FE9B" author="Maenosono, T." box="[1002,1199,295,321]" pageId="18" pageNumber="446" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 14K, L).
</paragraph>
<paragraph id="8B8636DB7E17733A4796FED7F3ABFE53" blockId="18.[151,1437,151,1149]" pageId="18" pageNumber="446">
<emphasis id="B94DEAC97E17733A4796FED7F60EFEBE" bold="true" box="[199,358,330,356]" pageId="18" pageNumber="446">Colour in life</emphasis>
. Background colour white; setae brown; epigastric, protogastric and hepatic areolets bright red; cardiac areolets pale red to red (
<bibRefCitation id="EFA84B2A7E17733A46ADFEF2F443FE53" author="Marumura, M. &amp; Takeda, M." box="[508,811,367,393]" pageId="18" pageNumber="446" pagination="189 - 197" refId="ref16388" refString="Marumura, M. &amp; Takeda, M. (2012) Crab specimens in the Wakayama Prefectural Museum of Nature (Nagai Collection) taxonomic study III. New record species in Japan (3). Teikyo Heisei University Bulletin, 23 (1), 189 - 197." type="journal article" year="2012">Marumura &amp; Takeda 2012</bibRefCitation>
: fig. 2A;
<bibRefCitation id="EFA84B2A7E17733A44C5FEF2F332FE53" author="Maenosono, T." box="[916,1114,367,393]" pageId="18" pageNumber="446" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 8B).
</paragraph>
</subSubSection>
<subSubSection id="C32365507E17733A4796FE0EF464FB83" pageId="18" pageNumber="446" type="discussion">
<paragraph id="8B8636DB7E17733A4796FE0EF5A3FD2B" blockId="18.[151,1437,151,1149]" pageId="18" pageNumber="446">
<emphasis id="B94DEAC97E17733A4796FE0EF65CFE77" bold="true" box="[199,308,403,429]" pageId="18" pageNumber="446">Remarks</emphasis>
.
<taxonomicName id="4C394D587E17733A466EFE0EF535FE77" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[319,605,403,429]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="446" phylum="Arthropoda" rank="species" species="symmetrinuda">
<emphasis id="B94DEAC97E17733A466EFE0EF535FE77" box="[319,605,403,429]" italics="true" pageId="18" pageNumber="446">Lophoplax symmetrinuda</emphasis>
</taxonomicName>
is arguably the most distinctive member of the genus
<taxonomicName id="4C394D587E17733A43F1FE0EF270FE77" authorityName="Tesch" authorityYear="1918" box="[1184,1304,403,429]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEAC97E17733A43F1FE0EF270FE77" box="[1184,1304,403,429]" italics="true" pageId="18" pageNumber="446">Lophoplax</emphasis>
</taxonomicName>
as currently diagnosed: it differs markedly from all congeners in its transversely subovate carapace (
<figureCitation id="13022A5E7E17733A43DFFE2AF391FE0B" box="[1166,1273,439,465]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="18" pageNumber="446">Fig. 12A</figureCitation>
;
<bibRefCitation id="EFA84B2A7E17733A4256FE2AF644FE2F" author="Marumura, M. &amp; Takeda, M." pageId="18" pageNumber="446" pagination="189 - 197" refId="ref16388" refString="Marumura, M. &amp; Takeda, M. (2012) Crab specimens in the Wakayama Prefectural Museum of Nature (Nagai Collection) taxonomic study III. New record species in Japan (3). Teikyo Heisei University Bulletin, 23 (1), 189 - 197." type="journal article" year="2012">Marumura &amp; Takeda 2012</bibRefCitation>
: fig. 3A;
<bibRefCitation id="EFA84B2A7E17733A46C9FE46F537FE2F" author="Maenosono, T." box="[408,607,475,501]" pageId="18" pageNumber="446" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 10A); the wide and low external orbital and anterolateral teeth (
<figureCitation id="13022A5E7E17733A423EFE46F7A3FDC3" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="18" pageNumber="446">Fig. 12A</figureCitation>
;
<bibRefCitation id="EFA84B2A7E17733A478BFE62F6CCFDC3" author="Maenosono, T." box="[218,420,511,537]" pageId="18" pageNumber="446" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 8A, B); the absence of tubercles or lobes at the junction between the antero- and posterolateral margins (
<figureCitation id="13022A5E7E17733A46CDFDBEF56AFDE7" box="[412,514,547,573]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="18" pageNumber="446">Fig. 12A</figureCitation>
;
<bibRefCitation id="EFA84B2A7E17733A455DFDBEF5B8FDE7" author="Maenosono, T." box="[524,720,547,573]" pageId="18" pageNumber="446" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 8A, B); the proportionately shorter merus and ischium of the third maxilliped (
<figureCitation id="13022A5E7E17733A4608FDDAF6D7FDBB" box="[345,447,583,609]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="18" pageNumber="446">Fig. 12B</figureCitation>
); the comparatively shorter ambulatory legs, especially the merus (
<figureCitation id="13022A5E7E17733A43F8FDDAF266FDBB" box="[1193,1294,583,609]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="18" pageNumber="446">Fig. 12C</figureCitation>
;
<bibRefCitation id="EFA84B2A7E17733A424BFDDAF7BBFD5F" author="Maenosono, T." pageId="18" pageNumber="446" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 8B); and the distinctly wider male pleon (
<figureCitation id="13022A5E7E17733A4585FDF6F453FD5F" box="[724,827,619,645]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="18" pageNumber="446">Fig. 12D</figureCitation>
;
<bibRefCitation id="EFA84B2A7E17733A4417FDF6F318FD5F" author="Marumura, M. &amp; Takeda, M." box="[838,1136,619,645]" pageId="18" pageNumber="446" pagination="189 - 197" refId="ref16388" refString="Marumura, M. &amp; Takeda, M. (2012) Crab specimens in the Wakayama Prefectural Museum of Nature (Nagai Collection) taxonomic study III. New record species in Japan (3). Teikyo Heisei University Bulletin, 23 (1), 189 - 197." type="journal article" year="2012">Marumura &amp; Takeda 2012</bibRefCitation>
: fig. 4C;
<bibRefCitation id="EFA84B2A7E17733A4385FDF6F2F1FD5F" author="Maenosono, T." box="[1236,1433,619,645]" pageId="18" pageNumber="446" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
: fig. 8B). Despite these characters, the presence and arrangement of the gastric, cardiac and hepatic lobes agrees well with other
<taxonomicName id="4C394D587E17733A4641FD2EF68BFD17" box="[272,483,691,717]" pageId="18" pageNumber="446">
<emphasis id="B94DEAC97E17733A4641FD2EF6E0FD17" box="[272,392,691,717]" italics="true" pageId="18" pageNumber="446">Lophoplax</emphasis>
species
</taxonomicName>
, and as such we retain it in this genus for the time being, as first suggested by
<bibRefCitation id="EFA84B2A7E17733A421DFD2EF63AFD2B" author="Takeda, M. &amp; Kurata, Y." pageId="18" pageNumber="446" pagination="195 - 202" refId="ref18657" refString="Takeda, M. &amp; Kurata, Y. (1984) Crabs of the Ogasawara Islands. VII. Third Report on the Species Obtained from Stomachs of Fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4), 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata (1984)</bibRefCitation>
and
<bibRefCitation id="EFA84B2A7E17733A46D9FD4AF5A0FD2B" author="Takeda, M. &amp; Marumura, M." box="[392,712,726,753]" pageId="18" pageNumber="446" pagination="87 - 91" refId="ref18718" refString="Takeda, M. &amp; Marumura, M. (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2), 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura (1995)</bibRefCitation>
.
</paragraph>
<paragraph id="8B8636DB7E17733A4796FD66F3E7FCE3" blockId="18.[151,1437,151,1149]" pageId="18" pageNumber="446">
The original figures of the species by
<bibRefCitation id="EFA84B2A7E17733A4522FD66F42EFCCF" author="Edmondson, C. H." box="[627,838,762,789]" pageId="18" pageNumber="446" pagination="183 - 243" refId="ref15227" refString="Edmondson, C. H. (1951) Some Central Pacific Crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson (1951</bibRefCitation>
: fig. 34ad) are inaccurate, the carapace in particular appearing quadrate rather than ovate and the dentition being incorrectly depicted (
<figureCitation id="13022A5E7E17733A437FFC82F3E9FCE3" box="[1070,1153,799,825]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="18" pageNumber="446">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8B8636DB7E17733A4796FCDEF436FBCB" blockId="18.[151,1437,151,1149]" pageId="18" pageNumber="446">
<bibRefCitation id="EFA84B2A7E17733A4796FCDEF6BDFC87" author="Edmondson, C. H." box="[199,469,834,861]" pageId="18" pageNumber="446" pagination="183 - 243" refId="ref15227" refString="Edmondson, C. H. (1951) Some Central Pacific Crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson (1951: 235)</bibRefCitation>
comments on why it should be in
<taxonomicName id="4C394D587E17733A441FFCDEF32FFC87" authorityName="Ward" authorityYear="1936" box="[846,1095,835,861]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEAC97E17733A441FFCDEF32FFC87" box="[846,1095,835,861]" italics="true" pageId="18" pageNumber="446">Pseudocryptocoeloma</emphasis>
</taxonomicName>
are brief and non-informative: “The genus
<taxonomicName id="4C394D587E17733A464EFCFAF570FC5B" authorityName="Ward" authorityYear="1936" box="[287,536,871,897]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEAC97E17733A464EFCFAF570FC5B" box="[287,536,871,897]" italics="true" pageId="18" pageNumber="446">Pseudocryptocoeloma</emphasis>
</taxonomicName>
was established by Ward in 1936 to provide for the species
<taxonomicName id="4C394D587E17733A43EAFCFAF24AFC5A" authorityName="Ward" authorityYear="1936" box="[1211,1314,871,896]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="446" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B94DEAC97E17733A43EAFCFAF24AFC5A" box="[1211,1314,871,896]" italics="true" pageId="18" pageNumber="446">P. parvus</emphasis>
</taxonomicName>
, described from Lindeman Island,
<collectingRegion id="49FDF8397E17733A46CAFC16F54BFC7F" box="[411,547,907,933]" country="Australia" name="Queensland" pageId="18" pageNumber="446">Queensland</collectingRegion>
. The new species differs from the Australian form in the arrangement of the nude areas of the carapace. In
<taxonomicName id="4C394D587E17733A46E2FC32F573FC12" authorityName="Ward" authorityYear="1936" box="[435,539,943,968]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="446" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B94DEAC97E17733A46E2FC32F573FC12" box="[435,539,943,968]" italics="true" pageId="18" pageNumber="446">P. parvus</emphasis>
</taxonomicName>
the entire median and posterior portions of the carapace are bare.”
<bibRefCitation id="EFA84B2A7E17733A424BFC32F78AFC37" author="Maenosono, T." pageId="18" pageNumber="446" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono (2019)</bibRefCitation>
described and figured the species at length and in the present examination of the
<typeStatus id="548288797E17733A4327FC4EF3B0FC37" box="[1142,1240,979,1005]" pageId="18" pageNumber="446" type="holotype">holotype</typeStatus>
male agrees well with his account, notably in the form of the male pleon and G1.
</paragraph>
<paragraph id="8B8636DB7E17733A4796FB87F464FB83" blockId="18.[151,1437,151,1149]" pageId="18" pageNumber="446">
<emphasis id="B94DEAC97E17733A4796FB87F64BFBEE" bold="true" box="[199,291,1050,1076]" pageId="18" pageNumber="446">Habitat</emphasis>
.
<materialsCitation id="3B513C867E17733A4661FB86F460FB83" ID-GBIF-Occurrence="4035852310" country="Japan" location="Maenoson" pageId="18" pageNumber="446" specimenCount="1">
The
<typeStatus id="548288797E17733A4633FB86F6FAFBEF" box="[354,402,1051,1077]" pageId="18" pageNumber="446">type</typeStatus>
male was collected on a table reef in the intertidal zone of a reef.
<location id="8EE660007E17733A433AFB86F388FBEF" LSID="urn:lsid:plazi:treatment:039087CD7E15733A47C6F89FF40AFBA7:8EE660007E17733A433AFB86F388FBEF" box="[1131,1248,1051,1077]" country="Japan" name="Maenoson" pageId="18" pageNumber="446">Maenoson</location>
(2019) collected the species from intertidal reef habitats as well in
<collectingCountry id="F32E764B7E17733A4594FBA2F460FB83" box="[709,776,1087,1113]" name="Japan" pageId="18" pageNumber="446">Japan</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32365507E17733A4796FBFFF40AFBA7" box="[199,866,1122,1149]" pageId="18" pageNumber="446" type="distribution">
<paragraph id="8B8636DB7E17733A4796FBFFF40AFBA7" blockId="18.[151,1437,151,1149]" box="[199,866,1122,1149]" pageId="18" pageNumber="446">
<emphasis id="B94DEAC97E17733A4796FBFFF631FBA6" bold="true" box="[199,345,1122,1148]" pageId="18" pageNumber="446">Distribution</emphasis>
. Known from
<collectingCountry id="F32E764B7E17733A46AFFBFEF522FBA7" box="[510,586,1123,1149]" name="Samoa" pageId="18" pageNumber="446">Samoa</collectingCountry>
and the Ryukyus,
<collectingCountry id="F32E764B7E17733A444AFBFEF436FBA7" box="[795,862,1123,1149]" name="Japan" pageId="18" pageNumber="446">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>