556 lines
89 KiB
XML
556 lines
89 KiB
XML
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<mods:title id="4B5C78214C4DF86B5C697CB131E71D42">Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura)</mods:title>
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<mods:namePart id="1ED9D4D96CF48416B7D960C46B2CDC34">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="57454617AF8F69BCC69CFA4D620DB760">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore. peterng @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 5946 - 0608</mods:affiliation>
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<mods:affiliation id="B7CD1A6E149087A79FB64CC7DF1CDE42">Research Center for Marine and Land Bioindustry, National Research and Innovation Agency, Dusun Teluk Kodek, Pemenang, Lombok Utara 83756, NTB, Indonesia. dwilistyo @ yahoo. com; https: // orcid. org / 0000 - 0003 - 4342 - 9045</mods:affiliation>
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<mods:title id="E73B84EC8CFA035839494B291A31118C">Zootaxa</mods:title>
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<taxonomicName id="4C394D587E04732947C6FC82F6D9FCE3" ID-CoL="7P7TN" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[151,433,799,826]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
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<emphasis id="B94DEAC97E04732947C6FC82F671FCE0" bold="true" box="[151,281,799,826]" italics="true" pageId="1" pageNumber="429">Lophoplax</emphasis>
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<bibRefCitation id="EFA84B2A7E0473294671FC82F6D9FCE3" author="Tesch, J. J." box="[288,433,799,825]" pageId="1" pageNumber="429" pagination="149 - 295" refId="ref18770" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
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<emphasis id="B94DEAC97E04732947C6FCF9F66BFCA6" box="[151,259,868,892]" italics="true" pageId="1" pageNumber="429">Lophoplax</emphasis>
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;
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<treatmentCitation id="0A9810CA7E0473294695FCF9F514FCA6" author="Takeda, M." box="[452,636,868,892]" page="120" pageId="1" pageNumber="429" year="1977">
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<bibRefCitation id="EFA84B2A7E0473294695FCF9F514FCA6" author="Takeda, M." box="[452,636,868,892]" pageId="1" pageNumber="429" pagination="120 - 123" refId="ref18577" refString="Takeda, M. (1977) Rediscovery of Lophoplax sculpta (Stimpson, 1858) (Crustacea, Decapoda, Brachyura). Zoological Magazine, 86, 120 - 123." type="journal article" year="1977">Takeda 1977: 120</bibRefCitation>
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;
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<treatmentCitation id="0A9810CA7E04732945D7FCF9F47DFCA6" author="Ng, P. K. L." box="[646,789,868,892]" page="100" pageId="1" pageNumber="429" year="1987">
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<bibRefCitation id="EFA84B2A7E04732945D7FCF9F47DFCA6" author="Ng, P. K. L." box="[646,789,868,892]" pageId="1" pageNumber="429" pagination="69 - 111" refId="ref16944" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). Indo-Malayan Zoology, 1987, 4 (1), 69 - 111, pl. 1." type="journal article" year="1987">Ng 1987: 100</bibRefCitation>
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;
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<treatmentCitation id="0A9810CA7E047329444EFCF9F351FCA1" author="Poore, G. C. B. & Ahyong, S. T." box="[799,1081,867,892]" page="667" pageId="1" pageNumber="429" year="2023">
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<bibRefCitation id="EFA84B2A7E047329444EFCF9F351FCA1" author="Poore, G. C. B. & Ahyong, S. T." box="[799,1081,867,892]" pageId="1" pageNumber="429" refId="ref17985" refString="Poore, G. C. B. & Ahyong, S. T. (2023) Marine decapod Crustacea: a guide to the families and genera of the world. CSIRO Publishing, Collingwood, 916 pp. https: // doi. org / 10.1071 / 9781486311798" type="book" year="2023">Poore & Ahyong 2023: 667</bibRefCitation>
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, 670, 680, fig. 14.112i.
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<taxonomicName id="4C394D587E04732947C6FC1FF6F9FC40" ID-CoL="dc0a7f74-e1d5-48d7-a61a-06886f65fe7a" authority="Deb, 1989" authorityName="Deb" authorityYear="1989" box="[151,401,898,922]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
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<emphasis id="B94DEAC97E04732947C6FC1FF649FC40" box="[151,289,898,922]" italics="true" pageId="1" pageNumber="429">Myopilumnus</emphasis>
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<bibRefCitation id="EFA84B2A7E0473294676FC1FF6F9FC40" author="Deb, M." box="[295,401,898,922]" pageId="1" pageNumber="429" pagination="113 - 116" refId="ref15100" refString="Deb, M. (1989) Myopilumnus adamanicus n. gen., n. sp., a xanthid crab from Andamans. Journal of the Andaman Science Association, 5 (2), 113 - 116." type="journal article" year="1989">Deb, 1989</bibRefCitation>
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</taxonomicName>
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(
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species
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<taxonomicName id="4C394D587E0473294570FC1FF4C3FC40" ID-CoL="84G2B" authority="Deb, 1989" authorityName="Deb" authorityYear="1989" box="[545,939,898,922]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="andamanicus">
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<emphasis id="B94DEAC97E0473294570FC1FF450FC40" box="[545,824,898,922]" italics="true" pageId="1" pageNumber="429">Myopilumnus andamanicus</emphasis>
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<bibRefCitation id="EFA84B2A7E047329446EFC1FF4C3FC40" author="Deb, M." box="[831,939,898,922]" pageId="1" pageNumber="429" pagination="113 - 116" refId="ref15100" refString="Deb, M. (1989) Myopilumnus adamanicus n. gen., n. sp., a xanthid crab from Andamans. Journal of the Andaman Science Association, 5 (2), 113 - 116." type="journal article" year="1989">Deb, 1989</bibRefCitation>
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, by monotypy; gender masculine)
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<emphasis id="B94DEAC97E04732947C6FC54F643FC38" bold="true" box="[151,299,968,995]" pageId="1" pageNumber="429">
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<typeStatus id="548288797E04732947C6FC54F7BAFC39" box="[151,210,969,995]" pageId="1" pageNumber="429">Type</typeStatus>
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species
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</emphasis>
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.
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<taxonomicName id="4C394D587E0473294668FC55F5DCFC39" ID-CoL="3W3WR" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[313,692,968,995]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="bicristata">
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<emphasis id="B94DEAC97E0473294668FC55F54BFC38" box="[313,547,968,994]" italics="true" pageId="1" pageNumber="429">Lophoplax bicristata</emphasis>
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<bibRefCitation id="EFA84B2A7E047329457BFC55F5DCFC39" author="Tesch, J. J." box="[554,692,968,995]" pageId="1" pageNumber="429" pagination="149 - 295" refId="ref18770" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
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</taxonomicName>
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, by original designation and monotypy; gender feminine.
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<paragraph id="8B8636DB7E0473294796FC70F53CFA24" blockId="1.[151,1437,968,2039]" pageId="1" pageNumber="429">
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<emphasis id="B94DEAC97E0473294796FC70F65CFBDD" bold="true" box="[199,308,1005,1031]" pageId="1" pageNumber="429">Remarks</emphasis>
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. The presence of prominent areolets on the carapace is a character found in all species of
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<taxonomicName id="4C394D587E0473294275FC71F2F4FBDC" authorityName="Tesch" authorityYear="1918" box="[1316,1436,1004,1030]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
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<emphasis id="B94DEAC97E0473294275FC71F2F4FBDC" box="[1316,1436,1004,1030]" italics="true" pageId="1" pageNumber="429">Lophoplax</emphasis>
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</taxonomicName>
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||
but the number, size and strength vary considerably. In the type species,
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||
<taxonomicName id="4C394D587E0473294486FB8CF309FBF0" authorityName="Tesch" authorityYear="1918" box="[983,1121,1040,1066]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="bicristata">
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<emphasis id="B94DEAC97E0473294486FB8CF309FBF0" box="[983,1121,1040,1066]" italics="true" pageId="1" pageNumber="429">L. bicristata</emphasis>
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</taxonomicName>
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, the transverse protogastric areolet is especially distinct, with the epigastric areolet relatively low and not prominent, with all the other areolets not obvious, notably, the hepatic one is only represented by a tubercle. In addition, the dorsal surface of the carpus of the cheliped has one raised plate, without areolets (cf.
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<bibRefCitation id="EFA84B2A7E0473294461FBE1F4DEFB4D" author="Tesch, J. J." box="[816,950,1148,1175]" pageId="1" pageNumber="429" pagination="149 - 295" refId="ref18770" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch 1918</bibRefCitation>
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||
: pl. 12 fig. 2; unpublished photographs of
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<typeStatus id="548288797E04732947C6FB3CF786FB61" box="[151,238,1185,1211]" pageId="1" pageNumber="429" type="syntype">syntype</typeStatus>
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||
male). On the other hand, the other species included in
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<taxonomicName id="4C394D587E0473294425FB3DF484FB60" authorityName="Tesch" authorityYear="1918" box="[884,1004,1184,1210]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
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<emphasis id="B94DEAC97E0473294425FB3DF484FB60" box="[884,1004,1184,1210]" italics="true" pageId="1" pageNumber="429">Lophoplax</emphasis>
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||
</taxonomicName>
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by
|
||
<bibRefCitation id="EFA84B2A7E0473294348FB3DF3D8FB61" author="Tesch, J. J." box="[1049,1200,1184,1211]" pageId="1" pageNumber="429" pagination="149 - 295" refId="ref18770" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch (1918)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C394D587E04732943EFFB3CF246FB60" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1214,1326,1184,1210]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis id="B94DEAC97E04732943EFFB3CF246FB60" box="[1214,1326,1184,1210]" italics="true" pageId="1" pageNumber="429">L. sculpta</emphasis>
|
||
</taxonomicName>
|
||
, has well developed and very prominent smooth epigastric, protogastric and hepatic areolets (
|
||
<figureCitation id="13022A5E7E047329431EFB59F3A4FB05" box="[1103,1228,1220,1247]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="1" pageNumber="429">Fig. 3A–C</figureCitation>
|
||
). The same is true for
|
||
<taxonomicName id="4C394D587E04732947EEFB74F65AFAD8" authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[191,306,1256,1282]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="sordida" status="sp. nov.">
|
||
<emphasis id="B94DEAC97E04732947EEFB74F65AFAD8" box="[191,306,1256,1282]" italics="true" pageId="1" pageNumber="429">L. sordida</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAC97E0473294668FB74F610FAD9" bold="true" box="[313,376,1257,1283]" pageId="1" pageNumber="429">
|
||
<taxonomicNameLabel id="A27E57B27E0473294668FB74F610FAD9" box="[313,376,1257,1283]" pageId="1" pageNumber="429" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C394D587E04732946E1FB74F544FAD8" authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[432,556,1257,1282]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="pannosa" status="sp. nov.">
|
||
<emphasis id="B94DEAC97E04732946E1FB74F544FAD8" box="[432,556,1257,1282]" italics="true" pageId="1" pageNumber="429">L. pannosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAC97E0473294565FB74F51BFAD9" bold="true" box="[564,627,1257,1283]" pageId="1" pageNumber="429">
|
||
<taxonomicNameLabel id="A27E57B27E0473294565FB74F51BFAD9" box="[564,627,1257,1283]" pageId="1" pageNumber="429" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13022A5E7E04732945D2FB75F46AFAD9" box="[643,770,1256,1283]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="1" pageNumber="429">Figs. 3D–F</figureCitation>
|
||
,
|
||
<figureCitation id="13022A5E7E047329445FFB74F45DFAD9" box="[782,821,1257,1283]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="1" pageNumber="429">4D</figureCitation>
|
||
). In
|
||
<taxonomicName id="4C394D587E0473294436FB74F498FAD8" authorityName="Sakai" authorityYear="1935" box="[871,1008,1256,1282]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="takakurai">
|
||
<emphasis id="B94DEAC97E0473294436FB74F498FAD8" box="[871,1008,1256,1282]" italics="true" pageId="1" pageNumber="429">L. takakurai</emphasis>
|
||
</taxonomicName>
|
||
, there are no prominent areolets, with the gastric regions only possessing clusters of granules (cf.
|
||
<bibRefCitation id="EFA84B2A7E0473294468FA91F4D3FAFD" author="Sakai, T." box="[825,955,1292,1319]" pageId="1" pageNumber="429" pagination="63 - 88" refId="ref18159" refString="Sakai, T. (1935) New or Rare Species of Brachyura, Collected by the " Misago " during the Zoological Survey Around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32), 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai 1935</bibRefCitation>
|
||
: pl. 7 fig. 2;
|
||
<bibRefCitation id="EFA84B2A7E047329431CFA91F3A7FAFC" author="Sakai, T." box="[1101,1231,1292,1318]" pageId="1" pageNumber="429" refId="ref18220" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. In three volumes. Kodansha Ltd., Tokyo, xxix + 773 pp. (English text), 461 pp. (Japanese text), 16 pp, 251 pls. (plates volume)." type="book" year="1976">Sakai 1976</bibRefCitation>
|
||
: pl. 191 fig. 4). In
|
||
<taxonomicName id="4C394D587E04732947C6FAACF630FA90" authorityName="Takeda & Kurata" authorityYear="1984" box="[151,344,1328,1354]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="sextuberculata">
|
||
<emphasis id="B94DEAC97E04732947C6FAACF630FA90" box="[151,344,1328,1354]" italics="true" pageId="1" pageNumber="429">L. sextuberculata</emphasis>
|
||
</taxonomicName>
|
||
, the epigastric, protogastric and hepatic areolets are present but smaller and arranged more or less in a subtransverse row (
|
||
<bibRefCitation id="EFA84B2A7E04732946F2FAC9F5C2FAB5" author="Takeda, M. & Kurata, Y." box="[419,682,1364,1391]" pageId="1" pageNumber="429" pagination="195 - 202" refId="ref18657" refString="Takeda, M. & Kurata, Y. (1984) Crabs of the Ogasawara Islands. VII. Third Report on the Species Obtained from Stomachs of Fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4), 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda & Kurata 1984</bibRefCitation>
|
||
: fig. 14), but this condition varies, with areolets varying in number and positioning (
|
||
<bibRefCitation id="EFA84B2A7E0473294602FAE5F5EAFA49" author="Takeda, M. & Marumura, M." box="[339,642,1400,1427]" pageId="1" pageNumber="429" pagination="87 - 91" refId="ref18718" refString="Takeda, M. & Marumura, M. (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2), 87 - 91." type="journal article" year="1995">Takeda & Marumura 1995</bibRefCitation>
|
||
: fig. 1).
|
||
<taxonomicName id="4C394D587E047329458FFAE5F48BFA48" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[734,995,1400,1426]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis id="B94DEAC97E047329458FFAE5F48BFA48" box="[734,995,1400,1426]" italics="true" pageId="1" pageNumber="429">Lophoplax vermiculata</emphasis>
|
||
</taxonomicName>
|
||
has all the areolets of
|
||
<taxonomicName id="4C394D587E047329438FFAE4F224FA48" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1246,1356,1400,1426]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis id="B94DEAC97E047329438FFAE4F224FA48" box="[1246,1356,1400,1426]" italics="true" pageId="1" pageNumber="429">L. sculpta</emphasis>
|
||
</taxonomicName>
|
||
except that these are prominently rugose and eroded, very different from the almost smooth surfaces of
|
||
<taxonomicName id="4C394D587E04732943BCFA00F236FA6C" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1261,1374,1436,1462]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis id="B94DEAC97E04732943BCFA00F236FA6C" box="[1261,1374,1436,1462]" italics="true" pageId="1" pageNumber="429">L. sculpta</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13022A5E7E047329423EFA01F7B7FA01" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="1" pageNumber="429">Fig. 4A–C</figureCitation>
|
||
). The same is true of the areolets on the carpus of the cheliped in
|
||
<taxonomicName id="4C394D587E0473294491FA5CF30CFA00" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[960,1124,1472,1498]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis id="B94DEAC97E0473294491FA5CF30CFA00" box="[960,1124,1472,1498]" italics="true" pageId="1" pageNumber="429">L. vermiculata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C394D587E04732943CBFA5CF27FFA00" authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[1178,1303,1473,1498]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="pannosa" status="sp. nov.">
|
||
<emphasis id="B94DEAC97E04732943CBFA5CF27FFA00" box="[1178,1303,1473,1498]" italics="true" pageId="1" pageNumber="429">L. pannosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAC97E047329424FFA5CF235FA01" bold="true" box="[1310,1373,1473,1499]" pageId="1" pageNumber="429">
|
||
<taxonomicNameLabel id="A27E57B27E047329424FFA5CF235FA01" box="[1310,1373,1473,1499]" pageId="1" pageNumber="429" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, they are eroded and not smooth (
|
||
<figureCitation id="13022A5E7E0473294681FA79F520FA24" box="[464,584,1508,1534]" captionStart="FIGURE 6" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,813,1703]" captionTargetId="figure-327@11.[151,1436,813,1703]" captionTargetPageId="11" captionText="FIGURE 6. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu.A, B, D, E, dorsal view of right cheliped; C, dorsal view of left cheliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663784" httpUri="https://zenodo.org/record/7663784/files/figure.png" pageId="1" pageNumber="429">Fig. 6D–F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8636DB7E0473294796F994F64FF99D" blockId="1.[151,1437,968,2039]" pageId="1" pageNumber="429">
|
||
In
|
||
<taxonomicName id="4C394D587E04732947B4F994F6C8F9F8" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[229,416,1544,1570]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="symmetrinuda">
|
||
<emphasis id="B94DEAC97E04732947B4F994F6C8F9F8" box="[229,416,1544,1570]" italics="true" pageId="1" pageNumber="429">L. symmetrinuda</emphasis>
|
||
</taxonomicName>
|
||
, the areolets are smooth and prominent, being relatively even larger than those of
|
||
<taxonomicName id="4C394D587E047329427EF994F2F4F9F8" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1327,1436,1544,1570]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis id="B94DEAC97E047329427EF994F2F4F9F8" box="[1327,1436,1544,1570]" italics="true" pageId="1" pageNumber="429">L. sculpta</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13022A5E7E04732947CEF9B1F673F99D" box="[159,283,1580,1607]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="1" pageNumber="429">Fig. 3A–C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8636DB7E0473294796F9CDF655F850" blockId="1.[151,1437,968,2039]" pageId="1" pageNumber="429">
|
||
All but one of the
|
||
<taxonomicName id="4C394D587E04732946C9F9CDF500F9B0" box="[408,616,1616,1642]" pageId="1" pageNumber="429">
|
||
<emphasis id="B94DEAC97E04732946C9F9CDF578F9B0" box="[408,528,1616,1642]" italics="true" pageId="1" pageNumber="429">Lophoplax</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
have a prominent raised and rounded longitudinal ridge on the posterior part of sub-branchial region which is an extension of the posterior carapace margin (
|
||
<figureCitation id="13022A5E7E047329431FF9E9F3F5F955" box="[1102,1181,1652,1679]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="1" pageNumber="429">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="13022A5E7E04732943FBF9E8F3D3F955" box="[1194,1211,1653,1679]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="1" pageNumber="429">4</figureCitation>
|
||
). It forms a channel with the carapace margin (
|
||
<figureCitation id="13022A5E7E047329469BF905F54EF969" box="[458,550,1688,1715]" captionStart="FIGURE 9" captionStartId="14.[151,250,1509,1533]" captionTargetBox="[176,1410,181,1485]" captionTargetId="figure-17@14.[176,1410,181,1485]" captionTargetPageId="14" captionText="FIGURE 9. A–D, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia. A, right lateral view of posterior part of sub-branchial region; B, E–G, anterior male thoracic sternum and pleon; C, male pleonal somites 1–4; D, male pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.7663797" httpUri="https://zenodo.org/record/7663797/files/figure.png" pageId="1" pageNumber="429">Fig. 9A</figureCitation>
|
||
). This structure on the posterior part of the sub-branchial region is similar to the condition reported in
|
||
<taxonomicName id="4C394D587E04732946ECF920F447F90D" authority="Ng & Chia, 1994" authorityName="Ng & Chia" authorityYear="1994" box="[445,815,1724,1751]" class="Malacostraca" family="Xanthidae" genus="Antrocarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94DEAC97E04732946ECF920F533F90D" box="[445,603,1725,1751]" italics="true" pageId="1" pageNumber="429">Antrocarcinus</emphasis>
|
||
<bibRefCitation id="EFA84B2A7E0473294534F920F447F90D" author="Ng, P. K. L. & Chia, D. G. B." box="[613,815,1724,1751]" pageId="1" pageNumber="429" pagination="701 - 730" refId="ref17121" refString="Ng, P. K. L. & Chia, D. G. B. (1994) The genus Glyptocarcinus Takeda, 1973, with descriptions of a new subfamily, two new genera and two new species from New Caledonia (Crustacea: Decapoda: Brachyura: Xanthidae). Raffles Bulletin of Zoology, 42 (3), 701 - 730." type="journal article" year="1994">Ng & Chia, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
(Antrocarcinidae
|
||
<bibRefCitation id="EFA84B2A7E0473294351F920F3A2F90D" author="Ng, P. K. L. & Chia, D. G. B." box="[1024,1226,1724,1751]" pageId="1" pageNumber="429" pagination="701 - 730" refId="ref17121" refString="Ng, P. K. L. & Chia, D. G. B. (1994) The genus Glyptocarcinus Takeda, 1973, with descriptions of a new subfamily, two new genera and two new species from New Caledonia (Crustacea: Decapoda: Brachyura: Xanthidae). Raffles Bulletin of Zoology, 42 (3), 701 - 730." type="journal article" year="1994">Ng & Chia, 1994</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="4C394D587E047329425EF921F672F921" authority="Ward, 1933" authorityName="Ward" authorityYear="1933" class="Malacostraca" family="Planopilumnidae" genus="Rathbunaria" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94DEAC97E047329425EF921F2F5F90C" box="[1295,1437,1724,1750]" italics="true" pageId="1" pageNumber="429">Rathbunaria</emphasis>
|
||
<bibRefCitation id="EFA84B2A7E04732947C6F97DF672F921" author="Ward, M." box="[151,282,1760,1787]" pageId="1" pageNumber="429" pagination="377 - 394" refId="ref18964" refString="Ward, M. (1933) New genera and species of marine Decapod Brachyura. From the coasts of New South Wales and Queensland. Australian Zoologist, 7, 377 - 394, pls. 21 - 23." type="journal article" year="1933">Ward, 1933</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C394D587E0473294678F97DF5ECF921" authority="Serene, 1984" authorityName="Serene" authorityYear="1984" box="[297,644,1760,1787]" class="Malacostraca" family="Planopilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="family">
|
||
Planopilumnidae
|
||
<bibRefCitation id="EFA84B2A7E04732946A1F97CF5ECF921" author="Serene, R." box="[496,644,1761,1787]" pageId="1" pageNumber="429" pagination="1 - 400" refId="ref18324" refString="Serene, R. (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge. Xanthoidea: Xanthidae et Trapeziidae. Addendum Carpiliidae et Menippidae - A. Crosnier. Faune Tropicale, 24, 1 - 400, pls. 1 - 48." type="journal article" year="1984">Serène, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
) (see
|
||
<bibRefCitation id="EFA84B2A7E0473294598F97CF4E0F921" author="Ng, P. K. L. & Chia, D. G. B." box="[713,904,1760,1787]" pageId="1" pageNumber="429" pagination="701 - 730" refId="ref17121" refString="Ng, P. K. L. & Chia, D. G. B. (1994) The genus Glyptocarcinus Takeda, 1973, with descriptions of a new subfamily, two new genera and two new species from New Caledonia (Crustacea: Decapoda: Brachyura: Xanthidae). Raffles Bulletin of Zoology, 42 (3), 701 - 730." type="journal article" year="1994">Ng & Chia 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA84B2A7E04732944C5F97CF493F921" author="Ng, P. K. L." box="[916,1019,1761,1787]" pageId="1" pageNumber="429" pagination="33 - 61" refId="ref17056" refString="Ng, P. K. L. (2010) On the Planopilumnidae Serene, 1984 (Crustacea: Brachyura: Pseudozioidea), with diagnoses of two new pilumnoid genera for species previously assigned to Planopilumnus Balss, 1933. Zootaxa, 2392 (1), 33 - 61. https: // doi. org / 10.11646 / zootaxa. 2392.1.2" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFA84B2A7E047329435FF97CF3B5F921" author="Ng, P. K. L. & Chia, D. G. B." box="[1038,1245,1760,1787]" pageId="1" pageNumber="429" pagination="701 - 730" refId="ref17121" refString="Ng, P. K. L. & Chia, D. G. B. (1994) The genus Glyptocarcinus Takeda, 1973, with descriptions of a new subfamily, two new genera and two new species from New Caledonia (Crustacea: Decapoda: Brachyura: Xanthidae). Raffles Bulletin of Zoology, 42 (3), 701 - 730." type="journal article" year="1994">Ng & Chia (1994)</bibRefCitation>
|
||
suggested that it may assist in respiration when the animals are partially covered by silt and sand. The condition in these species of
|
||
<taxonomicName id="4C394D587E04732947C6F8B5F667F898" authorityName="Tesch" authorityYear="1918" box="[151,271,1832,1858]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94DEAC97E04732947C6F8B5F667F898" box="[151,271,1832,1858]" italics="true" pageId="1" pageNumber="429">Lophoplax</emphasis>
|
||
</taxonomicName>
|
||
, however, is not as pronounced as those of the above genera. Only in
|
||
<taxonomicName id="4C394D587E0473294346F8B4F3F7F898" authorityName="Tesch" authorityYear="1918" box="[1047,1183,1832,1858]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="bicristata">
|
||
<emphasis id="B94DEAC97E0473294346F8B4F3F7F898" box="[1047,1183,1832,1858]" italics="true" pageId="1" pageNumber="429">L. bicristata</emphasis>
|
||
</taxonomicName>
|
||
is the intestinal region not distinctly raised, with the area appearing almost flat (cf.
|
||
<bibRefCitation id="EFA84B2A7E0473294411F8D1F4ACF8BD" author="Tesch, J. J." box="[832,964,1868,1895]" pageId="1" pageNumber="429" pagination="149 - 295" refId="ref18770" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch 1918</bibRefCitation>
|
||
: pl. 12 fig. 2; unpublished photographs of
|
||
<typeStatus id="548288797E04732947C6F8ECF786F851" box="[151,238,1905,1931]" pageId="1" pageNumber="429" type="syntype">syntype</typeStatus>
|
||
male).
|
||
</paragraph>
|
||
<paragraph id="8B8636DB7E04732A4796F809F3F8FE77" blockId="1.[151,1437,968,2039]" lastBlockId="2.[151,1437,150,1257]" lastPageId="2" lastPageNumber="430" pageId="1" pageNumber="429">
|
||
<taxonomicName id="4C394D587E0473294796F809F657F874" authorityName="Tesch" authorityYear="1918" box="[199,319,1940,1966]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94DEAC97E0473294796F809F657F874" box="[199,319,1940,1966]" italics="true" pageId="1" pageNumber="429">Lophoplax</emphasis>
|
||
</taxonomicName>
|
||
can also be approximately divided into two groups of species on the basis of their adult ambulatory legs. The species in the first group (with
|
||
<taxonomicName id="4C394D587E0473294527F824F46AF808" authorityName="Tesch" authorityYear="1918" box="[630,770,1976,2002]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="bicristata">
|
||
<emphasis id="B94DEAC97E0473294527F824F46AF808" box="[630,770,1976,2002]" italics="true" pageId="1" pageNumber="429">L. bicristata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D587E0473294442F824F4BFF808" authorityName="Takeda & Kurata" authorityYear="1984" box="[787,983,1976,2002]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="sextuberculata">
|
||
<emphasis id="B94DEAC97E0473294442F824F4BFF808" box="[787,983,1976,2002]" italics="true" pageId="1" pageNumber="429">L. sextuberculata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C394D587E0473294343F824F3F6F808" authorityName="Sakai" authorityYear="1935" box="[1042,1182,1976,2002]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="takakurai">
|
||
<emphasis id="B94DEAC97E0473294343F824F3F6F808" box="[1042,1182,1976,2002]" italics="true" pageId="1" pageNumber="429">L. takakurai</emphasis>
|
||
</taxonomicName>
|
||
) have proportionately longer legs with the merus slender and more elongate (cf.
|
||
<bibRefCitation id="EFA84B2A7E0473294445F841F4FDF82D" author="Tesch, J. J." box="[788,917,2012,2039]" pageId="1" pageNumber="429" pagination="149 - 295" refId="ref18770" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch 1918</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA84B2A7E04732944CEF841F376F82D" author="Sakai, T." box="[927,1054,2012,2039]" pageId="1" pageNumber="429" pagination="63 - 88" refId="ref18159" refString="Sakai, T. (1935) New or Rare Species of Brachyura, Collected by the " Misago " during the Zoological Survey Around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32), 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai 1935</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA84B2A7E0473294379F841F24EF82D" author="Takeda, M. & Kurata, Y." box="[1064,1318,2012,2039]" pageId="1" pageNumber="429" pagination="195 - 202" refId="ref18657" refString="Takeda, M. & Kurata, Y. (1984) Crabs of the Ogasawara Islands. VII. Third Report on the Species Obtained from Stomachs of Fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4), 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda & Kurata 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA84B2A7E04732A4261F841F626FF6B" author="Takeda, M. & Marumura, M." lastPageId="2" lastPageNumber="430" pageId="1" pageNumber="429" pagination="87 - 91" refId="ref18718" refString="Takeda, M. & Marumura, M. (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2), 87 - 91." type="journal article" year="1995">Takeda & Marumura 1995</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA84B2A7E07732A4609FF0AF5EEFF6B" author="Marumura, M. & Kosaka, A." box="[344,646,151,177]" pageId="2" pageNumber="430" pagination="1 - 8" refId="ref16330" refString="Marumura, M. & Kosaka, A. (2003) Catalogue of Brachyuran and Anomuran crabs Collection Donated by the late Mr. Seiji Nagai to the Wakayama Prefectural Museum of Natural History. Wakayama Prefectural Museum of Natural History, Kainan, Wakayama Prefecture, 73 pp., pls. 1 - 8." type="book chapter" year="2003">Marumura & Kosaka 2003</bibRefCitation>
|
||
). The figure of
|
||
<taxonomicName id="4C394D587E07732A447CFF0AF4DCFF6B" authorityName="Tesch" authorityYear="1918" box="[813,948,151,177]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="bicristata">
|
||
<emphasis id="B94DEAC97E07732A447CFF0AF4DCFF6B" box="[813,948,151,177]" italics="true" pageId="2" pageNumber="430">L. bicristata</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="EFA84B2A7E07732A4484FF0AF309FF6B" author="Tesch, J. J." box="[981,1121,151,177]" pageId="2" pageNumber="430" pagination="149 - 295" refId="ref18770" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch (1918</bibRefCitation>
|
||
: pl. 12 fig. 2) is accurate; the authors have examined photographs of a
|
||
<typeStatus id="548288797E07732A4531FF26F5DFFF0F" box="[608,695,187,213]" pageId="2" pageNumber="430" type="syntype">syntype</typeStatus>
|
||
male in Leiden and the leg characters agree. Species in the second group (with
|
||
<taxonomicName id="4C394D587E07732A4675FF42F6FBFF23" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[292,403,223,249]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis id="B94DEAC97E07732A4675FF42F6FBFF23" box="[292,403,223,249]" italics="true" pageId="2" pageNumber="430">L. sculpta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D587E07732A46F3FF42F523FF23" baseAuthorityName="Deb" baseAuthorityYear="1989" box="[418,587,223,249]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis id="B94DEAC97E07732A46F3FF42F523FF23" box="[418,587,223,249]" italics="true" pageId="2" pageNumber="430">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D587E07732A450BFF42F5A6FF23" authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[602,718,223,249]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sordida" status="sp. nov.">
|
||
<emphasis id="B94DEAC97E07732A450BFF42F5A6FF23" box="[602,718,223,249]" italics="true" pageId="2" pageNumber="430">L. sordida</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAC97E07732A4584FF42F47DFF23" bold="true" box="[725,789,223,249]" pageId="2" pageNumber="430">
|
||
<taxonomicNameLabel id="A27E57B27E07732A4584FF42F47DFF23" box="[725,789,223,249]" pageId="2" pageNumber="430" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C394D587E07732A4475FF42F4CAFF22" authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[804,930,223,248]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="pannosa" status="sp. nov.">
|
||
<emphasis id="B94DEAC97E07732A4475FF42F4CAFF22" box="[804,930,223,248]" italics="true" pageId="2" pageNumber="430">L. pannosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAC97E07732A44FBFF42F482FF23" bold="true" box="[938,1002,223,249]" pageId="2" pageNumber="430">
|
||
<taxonomicNameLabel id="A27E57B27E07732A44FBFF42F482FF23" box="[938,1002,223,249]" pageId="2" pageNumber="430" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C394D587E07732A44A8FF42F3F5FF23" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1017,1181,223,249]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis id="B94DEAC97E07732A44A8FF42F3F5FF23" box="[1017,1181,223,249]" italics="true" pageId="2" pageNumber="430">L. vermiculata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C394D587E07732A4387FF42F2FBFF23" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[1238,1427,223,249]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="symmetrinuda">
|
||
<emphasis id="B94DEAC97E07732A4387FF42F2FBFF23" box="[1238,1427,223,249]" italics="true" pageId="2" pageNumber="430">L. symmetrinuda</emphasis>
|
||
</taxonomicName>
|
||
) have proportionately shorter legs, with the merus distinctly wider and stouter (
|
||
<figureCitation id="13022A5E7E07732A4340FE9EF3FEFEC7" box="[1041,1174,259,285]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; C–F, left chela; G–L, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="2" pageNumber="430">Figs. 7G–L</figureCitation>
|
||
,
|
||
<figureCitation id="13022A5E7E07732A43F3FE9EF3BDFEC7" box="[1186,1237,259,285]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="2" pageNumber="430">12C</figureCitation>
|
||
). The ambulatory legs of
|
||
<taxonomicName id="4C394D587E07732A47BCFEBAF6C3FE9B" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[237,427,295,321]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="symmetrinuda">
|
||
<emphasis id="B94DEAC97E07732A47BCFEBAF6C3FE9B" box="[237,427,295,321]" italics="true" pageId="2" pageNumber="430">L. symmetrinuda</emphasis>
|
||
</taxonomicName>
|
||
are very characteristic, being much stouter and shorter than in all the other species (
|
||
<figureCitation id="13022A5E7E07732A423EFEBAF7A2FEBF" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="2" pageNumber="430">Fig. 12C</figureCitation>
|
||
). The adult ambulatory legs of
|
||
<taxonomicName id="4C394D587E07732A4577FED6F5A1FEBF" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[550,713,331,357]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis id="B94DEAC97E07732A4577FED6F5A1FEBF" box="[550,713,331,357]" italics="true" pageId="2" pageNumber="430">L. vermiculata</emphasis>
|
||
</taxonomicName>
|
||
are intermediate in length to
|
||
<taxonomicName id="4C394D587E07732A4340FED6F3A5FEBF" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[1041,1229,331,357]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="symmetrinuda">
|
||
<emphasis id="B94DEAC97E07732A4340FED6F3A5FEBF" box="[1041,1229,331,357]" italics="true" pageId="2" pageNumber="430">L. symmetrinuda</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C394D587E07732A425BFED6F210FEBF" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1290,1400,331,357]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis id="B94DEAC97E07732A425BFED6F210FEBF" box="[1290,1400,331,357]" italics="true" pageId="2" pageNumber="430">L. sculpta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D587E07732A42D7FED6F649FE53" baseAuthorityName="Deb" baseAuthorityYear="1989" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis id="B94DEAC97E07732A42D7FED6F649FE53" italics="true" pageId="2" pageNumber="430">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D587E07732A4661FEF2F6CBFE53" authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[304,419,367,393]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sordida" status="sp. nov.">
|
||
<emphasis id="B94DEAC97E07732A4661FEF2F6CBFE53" box="[304,419,367,393]" italics="true" pageId="2" pageNumber="430">L. sordida</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAC97E07732A46FAFEF2F682FE53" bold="true" box="[427,490,367,393]" pageId="2" pageNumber="430">
|
||
<taxonomicNameLabel id="A27E57B27E07732A46FAFEF2F682FE53" box="[427,490,367,393]" pageId="2" pageNumber="430" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C394D587E07732A4573FEF2F5F7FE52" authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[546,671,367,392]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="pannosa" status="sp. nov.">
|
||
<emphasis id="B94DEAC97E07732A4573FEF2F5F7FE52" box="[546,671,367,392]" italics="true" pageId="2" pageNumber="430">L. pannosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAC97E07732A45F6FEF2F58EFE53" bold="true" box="[679,742,367,393]" pageId="2" pageNumber="430">
|
||
<taxonomicNameLabel id="A27E57B27E07732A45F6FEF2F58EFE53" box="[679,742,367,393]" pageId="2" pageNumber="430" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13022A5E7E07732A45A7FEF2F42FFE53" box="[758,839,367,393]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; C–F, left chela; G–L, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="2" pageNumber="430">Fig. 7J</figureCitation>
|
||
). It must be noted, however, that subadult specimens of
|
||
<taxonomicName id="4C394D587E07732A47E7FE0EF631FE77" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[182,345,403,429]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis id="B94DEAC97E07732A47E7FE0EF631FE77" box="[182,345,403,429]" italics="true" pageId="2" pageNumber="430">L. vermiculata</emphasis>
|
||
</taxonomicName>
|
||
have proportionately more slender and longer ambulatory legs (
|
||
<figureCitation id="13022A5E7E07732A4378FE0EF3EBFE77" box="[1065,1155,403,429]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; C–F, left chela; G–L, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="2" pageNumber="430">Fig. 7K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8636DB7E07732A4796FE2AF36BFD2B" blockId="2.[151,1437,150,1257]" pageId="2" pageNumber="430">
|
||
The carapace shape of most of the species varies from almost quadrate (e.g.,
|
||
<taxonomicName id="4C394D587E07732A4370FE2AF3C1FE0B" authorityName="Tesch" authorityYear="1918" box="[1057,1193,439,465]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="bicristata">
|
||
<emphasis id="B94DEAC97E07732A4370FE2AF3C1FE0B" box="[1057,1193,439,465]" italics="true" pageId="2" pageNumber="430">L. bicristata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C394D587E07732A438EFE2AF225FE0B" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[1247,1357,439,465]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis id="B94DEAC97E07732A438EFE2AF225FE0B" box="[1247,1357,439,465]" italics="true" pageId="2" pageNumber="430">L. sculpta</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFA84B2A7E07732A420AFE2AF7BBFE2F" author="Tesch, J. J." pageId="2" pageNumber="430" pagination="149 - 295" refId="ref18770" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch 1918</bibRefCitation>
|
||
: pl. 12 fig. 2;
|
||
<figureCitation id="13022A5E7E07732A4625FE46F687FE2F" box="[372,495,475,501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="2" pageNumber="430">Fig. 3A–C</figureCitation>
|
||
), to transversely hexagonal or subquadrate (e.g.,
|
||
<taxonomicName id="4C394D587E07732A434AFE46F3F0FE2E" authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[1051,1176,475,500]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="pannosa" status="sp. nov.">
|
||
<emphasis id="B94DEAC97E07732A434AFE46F3F0FE2E" box="[1051,1176,475,500]" italics="true" pageId="2" pageNumber="430">L. pannosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAC97E07732A43F1FE46F388FE2F" bold="true" box="[1184,1248,475,501]" pageId="2" pageNumber="430">
|
||
<taxonomicNameLabel id="A27E57B27E07732A43F1FE46F388FE2F" box="[1184,1248,475,501]" pageId="2" pageNumber="430" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="13022A5E7E07732A43BEFE46F221FE2F" box="[1263,1353,475,501]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="2" pageNumber="430">Fig. 4D</figureCitation>
|
||
), being proportionately widest in
|
||
<taxonomicName id="4C394D587E07732A46E8FE62F535FDC3" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[441,605,511,537]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis id="B94DEAC97E07732A46E8FE62F535FDC3" box="[441,605,511,537]" italics="true" pageId="2" pageNumber="430">L. vermiculata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13022A5E7E07732A453CFE62F580FDC3" box="[621,744,511,537]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="2" pageNumber="430">Fig. 4A, C</figureCitation>
|
||
), with all of them possessing well defined anterolateral teeth.
|
||
<taxonomicName id="4C394D587E07732A47C6FDBEF6E9FDE7" authorityName="Tesch" authorityYear="1918" box="[151,385,547,573]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="bicristata">
|
||
<emphasis id="B94DEAC97E07732A47C6FDBEF6E9FDE7" box="[151,385,547,573]" italics="true" pageId="2" pageNumber="430">Lophoplax bicristata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D587E07732A46DEFDBEF695FDE7" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[399,509,547,573]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis id="B94DEAC97E07732A46DEFDBEF695FDE7" box="[399,509,547,573]" italics="true" pageId="2" pageNumber="430">L. sculpta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D587E07732A455AFDBEF5FCFDE7" authorityName="Sakai" authorityYear="1935" box="[523,660,547,573]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="takakurai">
|
||
<emphasis id="B94DEAC97E07732A455AFDBEF5FCFDE7" box="[523,660,547,573]" italics="true" pageId="2" pageNumber="430">L. takakurai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D587E07732A45F3FDBEF421FDE7" baseAuthorityName="Deb" baseAuthorityYear="1989" box="[674,841,547,573]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis id="B94DEAC97E07732A45F3FDBEF421FDE7" box="[674,841,547,573]" italics="true" pageId="2" pageNumber="430">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D587E07732A4406FDBEF4A2FDE7" authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[855,970,547,573]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sordida" status="sp. nov.">
|
||
<emphasis id="B94DEAC97E07732A4406FDBEF4A2FDE7" box="[855,970,547,573]" italics="true" pageId="2" pageNumber="430">L. sordida</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAC97E07732A4481FDBEF367FDE7" bold="true" box="[976,1039,547,573]" pageId="2" pageNumber="430">
|
||
<taxonomicNameLabel id="A27E57B27E07732A4481FDBEF367FDE7" box="[976,1039,547,573]" pageId="2" pageNumber="430" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C394D587E07732A4314FDBEF3AAFDE6" authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[1093,1218,547,572]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="pannosa" status="sp. nov.">
|
||
<emphasis id="B94DEAC97E07732A4314FDBEF3AAFDE6" box="[1093,1218,547,572]" italics="true" pageId="2" pageNumber="430">L. pannosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAC97E07732A4399FDBEF26FFDE7" bold="true" box="[1224,1287,547,573]" pageId="2" pageNumber="430">
|
||
<taxonomicNameLabel id="A27E57B27E07732A4399FDBEF26FFDE7" box="[1224,1287,547,573]" pageId="2" pageNumber="430" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
have distinct tubercles on the posterolateral margin of the carapace, just posterior of the last anterolateral tooth (e.g.,
|
||
<figureCitation id="13022A5E7E07732A4244FDDAF2F8FDBB" box="[1301,1424,583,609]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="2" pageNumber="430">Fig. 3A–C</figureCitation>
|
||
); being absent in
|
||
<taxonomicName id="4C394D587E07732A4603FDF6F57FFD5F" authorityName="Takeda & Kurata" authorityYear="1984" box="[338,535,619,645]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sextuberculata">
|
||
<emphasis id="B94DEAC97E07732A4603FDF6F57FFD5F" box="[338,535,619,645]" italics="true" pageId="2" pageNumber="430">L. sextuberculata</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="EFA84B2A7E07732A4503FDF6F408FD5F" author="Takeda, M. & Kurata, Y." box="[594,864,619,645]" pageId="2" pageNumber="430" pagination="195 - 202" refId="ref18657" refString="Takeda, M. & Kurata, Y. (1984) Crabs of the Ogasawara Islands. VII. Third Report on the Species Obtained from Stomachs of Fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4), 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda & Kurata 1984</bibRefCitation>
|
||
: fig. 14), and in adult
|
||
<taxonomicName id="4C394D587E07732A433AFDF6F27AFD5F" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1131,1298,619,645]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis id="B94DEAC97E07732A433AFDF6F27AFD5F" box="[1131,1298,619,645]" italics="true" pageId="2" pageNumber="430">L. vermiculata</emphasis>
|
||
</taxonomicName>
|
||
, only sharp granules or tubercles are present instead (
|
||
<figureCitation id="13022A5E7E07732A453DFD12F58FFD73" box="[620,743,655,681]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="2" pageNumber="430">Fig. 4A, C</figureCitation>
|
||
), being unarmed in subadult specimens (
|
||
<figureCitation id="13022A5E7E07732A43E7FD12F278FD73" box="[1206,1296,655,681]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="2" pageNumber="430">Fig. 4B</figureCitation>
|
||
).
|
||
<taxonomicName id="4C394D587E07732A4275FD12F65EFD17" baseAuthorityName="Edmondson" baseAuthorityYear="1951" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="symmetrinuda">
|
||
<emphasis id="B94DEAC97E07732A4275FD12F65EFD17" italics="true" pageId="2" pageNumber="430">Lophoplax symmetrinuda</emphasis>
|
||
</taxonomicName>
|
||
is in a group on its own, with the carapace transversely ovate, the anterolateral margin almost entire with only low rounded lobes and the posterolateral margin smooth (
|
||
<figureCitation id="13022A5E7E07732A44DFFD4AF49FFD2B" box="[910,1015,727,753]" captionStart="FIGURE 12" captionStartId="17.[151,250,1423,1447]" captionTargetBox="[159,1422,188,1386]" captionTargetId="figure-25@17.[151,1436,181,1400]" captionTargetPageId="17" captionText="FIGURE 12. Lophoplax symmetrinuda (Edmondson, 1951), holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace; B, left third maxilliped; C, left P5; E, left G1 (ventral view); F, left g1 (dorsal view); G, left G2; D, pleon." figureDoi="http://doi.org/10.5281/zenodo.7663805" httpUri="https://zenodo.org/record/7663805/files/figure.png" pageId="2" pageNumber="430">Fig. 12A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8636DB7E07732A4796FD66F352FC87" blockId="2.[151,1437,150,1257]" pageId="2" pageNumber="430">
|
||
The male anterior thoracic sternum in most of the species of
|
||
<taxonomicName id="4C394D587E07732A4425FD66F484FCCF" authorityName="Tesch" authorityYear="1918" box="[884,1004,763,789]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94DEAC97E07732A4425FD66F484FCCF" box="[884,1004,763,789]" italics="true" pageId="2" pageNumber="430">Lophoplax</emphasis>
|
||
</taxonomicName>
|
||
is relatively narrow transversely (e.g.,
|
||
<figureCitation id="13022A5E7E07732A47C6FC82F663FCE3" box="[151,267,799,825]" captionStart="FIGURE 9" captionStartId="14.[151,250,1509,1533]" captionTargetBox="[176,1410,181,1485]" captionTargetId="figure-17@14.[176,1410,181,1485]" captionTargetPageId="14" captionText="FIGURE 9. A–D, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia. A, right lateral view of posterior part of sub-branchial region; B, E–G, anterior male thoracic sternum and pleon; C, male pleonal somites 1–4; D, male pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.7663797" httpUri="https://zenodo.org/record/7663797/files/figure.png" pageId="2" pageNumber="430">Fig. 9B, E</figureCitation>
|
||
) but in adult
|
||
<taxonomicName id="4C394D587E07732A46F1FC82F52BFCE3" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[416,579,799,825]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis id="B94DEAC97E07732A46F1FC82F52BFCE3" box="[416,579,799,825]" italics="true" pageId="2" pageNumber="430">L. vermiculata</emphasis>
|
||
</taxonomicName>
|
||
, sternites 1–4 are distinctly wider (
|
||
<figureCitation id="13022A5E7E07732A449DFC82F34DFCE3" box="[972,1061,799,825]" captionStart="FIGURE 9" captionStartId="14.[151,250,1509,1533]" captionTargetBox="[176,1410,181,1485]" captionTargetId="figure-17@14.[176,1410,181,1485]" captionTargetPageId="14" captionText="FIGURE 9. A–D, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia. A, right lateral view of posterior part of sub-branchial region; B, E–G, anterior male thoracic sternum and pleon; C, male pleonal somites 1–4; D, male pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.7663797" httpUri="https://zenodo.org/record/7663797/files/figure.png" pageId="2" pageNumber="430">Fig. 9G</figureCitation>
|
||
), although in subadult males, it is relatively less obvious and more like those of other species of
|
||
<taxonomicName id="4C394D587E07732A4400FCDEF4A1FC87" authorityName="Tesch" authorityYear="1918" box="[849,969,835,861]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94DEAC97E07732A4400FCDEF4A1FC87" box="[849,969,835,861]" italics="true" pageId="2" pageNumber="430">Lophoplax</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13022A5E7E07732A4489FCDEF345FC87" box="[984,1069,835,861]" captionStart="FIGURE 9" captionStartId="14.[151,250,1509,1533]" captionTargetBox="[176,1410,181,1485]" captionTargetId="figure-17@14.[176,1410,181,1485]" captionTargetPageId="14" captionText="FIGURE 9. A–D, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia. A, right lateral view of posterior part of sub-branchial region; B, E–G, anterior male thoracic sternum and pleon; C, male pleonal somites 1–4; D, male pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.7663797" httpUri="https://zenodo.org/record/7663797/files/figure.png" pageId="2" pageNumber="430">Fig. 9F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8636DB7E07732A4796FCFAF6CEFC13" blockId="2.[151,1437,150,1257]" pageId="2" pageNumber="430">
|
||
The females of all the species are relatively consistent in their characters; the pleons being longitudinally ovate (e.g.
|
||
<figureCitation id="13022A5E7E07732A479EFC16F64EFC7F" box="[207,294,907,933]" captionStart="FIGURE 8" captionStartId="13.[151,250,1105,1129]" captionTargetBox="[151,1435,181,1081]" captionTargetId="figure-25@13.[151,1435,181,1081]" captionTargetPageId="13" captionText="FIGURE 8. A, B, Lophoplax sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, female thoracic sternum and pleon; B–D, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7663793" httpUri="https://zenodo.org/record/7663793/files/figure.png" pageId="2" pageNumber="430">Fig. 8A</figureCitation>
|
||
), and the vulvae are large, ovate, obliquely positioned and occupy most of sternite 6 without any visible operculum (
|
||
<figureCitation id="13022A5E7E07732A464FFC32F6F1FC13" box="[286,409,943,969]" captionStart="FIGURE 8" captionStartId="13.[151,250,1105,1129]" captionTargetBox="[151,1435,181,1081]" captionTargetId="figure-25@13.[151,1435,181,1081]" captionTargetPageId="13" captionText="FIGURE 8. A, B, Lophoplax sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, female thoracic sternum and pleon; B–D, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7663793" httpUri="https://zenodo.org/record/7663793/files/figure.png" pageId="2" pageNumber="430">Fig. 8B–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8636DB7E07732A4796FC4EF312FB33" blockId="2.[151,1437,150,1257]" pageId="2" pageNumber="430">
|
||
In summary, the composition of
|
||
<taxonomicName id="4C394D587E07732A451AFC4EF5ABFC37" authorityName="Tesch" authorityYear="1918" box="[587,707,979,1005]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94DEAC97E07732A451AFC4EF5ABFC37" box="[587,707,979,1005]" italics="true" pageId="2" pageNumber="430">Lophoplax</emphasis>
|
||
</taxonomicName>
|
||
remains imperfect. Looking at the series of characters of the recognised species of
|
||
<taxonomicName id="4C394D587E07732A46DCFC6AF56DFBCB" authorityName="Tesch" authorityYear="1918" box="[397,517,1015,1041]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94DEAC97E07732A46DCFC6AF56DFBCB" box="[397,517,1015,1041]" italics="true" pageId="2" pageNumber="430">Lophoplax</emphasis>
|
||
</taxonomicName>
|
||
, it can be argued that the genus is heterogeneous, and should perhaps be separated into three or even four genera. Certainly, there are sufficient differences between
|
||
<taxonomicName id="4C394D587E07732A4349FB86F3F7FBEF" authorityName="Tesch" authorityYear="1918" box="[1048,1183,1051,1077]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="bicristata">
|
||
<emphasis id="B94DEAC97E07732A4349FB86F3F7FBEF" box="[1048,1183,1051,1077]" italics="true" pageId="2" pageNumber="430">L. bicristata</emphasis>
|
||
</taxonomicName>
|
||
(the
|
||
<typeStatus id="548288797E07732A4386FB86F26FFBEF" box="[1239,1287,1051,1077]" pageId="2" pageNumber="430">type</typeStatus>
|
||
species), and
|
||
<taxonomicName id="4C394D587E07732A47C6FBA2F66FFB83" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[151,263,1087,1113]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis id="B94DEAC97E07732A47C6FBA2F66FFB83" box="[151,263,1087,1113]" italics="true" pageId="2" pageNumber="430">L. sculpta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D587E07732A4647FBA2F6D6FB83" baseAuthorityName="Deb" baseAuthorityYear="1989" box="[278,446,1087,1113]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis id="B94DEAC97E07732A4647FBA2F6D6FB83" box="[278,446,1087,1113]" italics="true" pageId="2" pageNumber="430">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D587E07732A469FFBA2F529FB83" authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[462,577,1087,1113]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sordida" status="sp. nov.">
|
||
<emphasis id="B94DEAC97E07732A469FFBA2F529FB83" box="[462,577,1087,1113]" italics="true" pageId="2" pageNumber="430">L. sordida</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAC97E07732A4518FBA2F5E1FB83" bold="true" box="[585,649,1087,1113]" pageId="2" pageNumber="430">
|
||
<taxonomicNameLabel id="A27E57B27E07732A4518FBA2F5E1FB83" box="[585,649,1087,1113]" pageId="2" pageNumber="430" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C394D587E07732A4593FBA2F457FB82" authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[706,831,1087,1112]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="pannosa" status="sp. nov.">
|
||
<emphasis id="B94DEAC97E07732A4593FBA2F457FB82" box="[706,831,1087,1112]" italics="true" pageId="2" pageNumber="430">L. pannosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAC97E07732A4416FBA2F4EFFB83" bold="true" box="[839,903,1087,1113]" pageId="2" pageNumber="430">
|
||
<taxonomicNameLabel id="A27E57B27E07732A4416FBA2F4EFFB83" box="[839,903,1087,1113]" pageId="2" pageNumber="430" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, to treat the latter group as a separate genus. A case can also be made for
|
||
<taxonomicName id="4C394D587E07732A46E6FBFEF531FBA7" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[439,601,1123,1149]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis id="B94DEAC97E07732A46E6FBFEF531FBA7" box="[439,601,1123,1149]" italics="true" pageId="2" pageNumber="430">L. vermiculata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C394D587E07732A45DCFBFEF421FBA7" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[653,841,1123,1149]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="symmetrinuda">
|
||
<emphasis id="B94DEAC97E07732A45DCFBFEF421FBA7" box="[653,841,1123,1149]" italics="true" pageId="2" pageNumber="430">L. symmetrinuda</emphasis>
|
||
</taxonomicName>
|
||
to be recognised as two additional monotypic genera, especially the latter species which has many peculiar features. Recognising new genera now, however, may not be the best approach, especially since we have not had the opportunity to examine specimens of
|
||
<taxonomicName id="4C394D587E07732A4393FB36F224FB1F" authorityName="Sakai" authorityYear="1935" box="[1218,1356,1195,1221]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="takakurai">
|
||
<emphasis id="B94DEAC97E07732A4393FB36F224FB1F" box="[1218,1356,1195,1221]" italics="true" pageId="2" pageNumber="430">L. takakurai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C394D587E07732A42D7FB36F653FB33" authorityName="Takeda & Kurata" authorityYear="1984" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sextuberculata">
|
||
<emphasis id="B94DEAC97E07732A42D7FB36F653FB33" italics="true" pageId="2" pageNumber="430">L. sextuberculata</emphasis>
|
||
</taxonomicName>
|
||
. The resolution of the this matter will need to be deferred to another time.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |