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<document id="5AA0CF4A5A1B958953B3DC701C2C2E08" ID-DOI="10.5252/geodiversitas2024v46a15" ID-ISSN="1638-9395" ID-Zenodo-Dep="13985760" ID-ZooBank="urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1729758589633" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel &amp; Goret, Bernard" docDate="2024" docId="038E87DAFFCFE42FFF11FCC3FE1BF858" docLanguage="en" docName="geodiversitas.2024.46.15.pdf" docOrigin="Geodiversitas 46 (15)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Jsowerbya Merle, Pacaud, Ledon &amp; Goret, 2024, n. gen." docType="treatment" docVersion="5" lastPageNumber="534" masterDocId="FFB7FFA2FFE6E406FFB8FFD6FF81FFB3" masterDocTitle="New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe" masterLastPageNumber="551" masterPageNumber="495" pageNumber="534" updateTime="1729856536171" updateUser="valdenar" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="7EDDF20C21CF3CB9B62A776EB487D177">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:namePart id="FF504ADB3115DA3D49E776FB0DC97B2A">Merle, Didier</mods:namePart>
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<mods:namePart id="63A7C9B902527AB87A86C5B74920D94C">Ledon, Daniel</mods:namePart>
<mods:affiliation id="35540C0876E2F722C79CAC75AB3E0B92">Muséum national dhistoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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<mods:date id="985AA32A4471B253B756474BEE64CE36">2024</mods:date>
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<subSubSection id="C33D6547FFCFE42FFF11FCC3FD01FCE3" pageId="41" pageNumber="534" type="nomenclature">
<paragraph id="8B9836CCFFCFE42FFF11FCC3FD01FCE3" blockId="41.[169,737,789,848]" pageId="41" pageNumber="534">
<heading id="D0D081A0FFCFE42FFF11FCC3FD01FCE3" centered="true" fontSize="11" level="2" pageId="41" pageNumber="534" reason="2">
Informal group
<taxonomicName id="4C274D4FFFCFE42FFEE7FCC3FE49FC9C" ID-CoL="8VKK8" authorityName="Merle" authorityYear="2005" box="[351,456,789,815]" class="Gastropoda" family="Muricidae" genus="Jsowerbya" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFCFE42FFEE7FCC3FE49FC9C" box="[351,456,789,815]" italics="true" pageId="41" pageNumber="534">Jsowerbya</emphasis>
</taxonomicName>
,
<taxonomicName id="4C274D4FFFCFE42FFE6FFCC0FD08FC9C" authority="Merle" authorityName="Merle" authorityYear="2024" box="[471,649,789,815]" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFCFE42FFE6FFCC0FDBAFC9C" box="[471,571,790,815]" italics="true" pageId="41" pageNumber="534">Beyregrex</emphasis>
Merle
</taxonomicName>
, n. gen,
<taxonomicName id="4C274D4FFFCFE42FFEB4FCE3FDA7FCFC" authority="Merle" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[268,550,821,847]" class="Gastropoda" family="Muricidae" genus="Pseudotrophonopsis" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFCFE42FFEB4FCE3FE5BFCFC" box="[268,474,821,847]" italics="true" pageId="41" pageNumber="534">Pseudotrophonopsis</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFCFE42FFD8AFCE0FD01FCE3" box="[562,640,822,848]" pageId="41" pageNumber="534" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C33D6547FFCFE42FFF3CFCB9FDD1FBDC" pageId="41" pageNumber="534" type="diagnosis">
<paragraph id="8B9836CCFFCFE42FFF3CFCB9FEEBFB96" blockId="41.[132,775,878,1061]" pageId="41" pageNumber="534">
<smallCapsWord id="8D7EA010FFCFE42FFF3CFCB9FF77FC36" baselines="896,897" box="[132,246,879,902]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="41" pageNumber="534">DIAGNOSIS</smallCapsWord>
. — Shell medium sized for family, biconic. P6 not atrophied. IP, ADP, MP and ABP present. Poorly developed axial sculpture formed by slightly lamellose varices. Posterior sinus of sutural ramp shallow. Shoulder sinus of P1 reduced or missing. ID missing, D1 to D6 present. Columellar lip occasionally ornamented by 1-3 tubercles. Microstructure of three aragonite layers.
</paragraph>
<paragraph id="8B9836CCFFCFE42FFF3CFBEBFDD1FBDC" blockId="41.[131,775,1084,1135]" pageId="41" pageNumber="534">
<smallCapsWord id="8D7EA010FFCFE42FFF3CFBEBFF6CFBE0" baselines="1102,1103" box="[132,237,1085,1108]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Included" pageId="41" pageNumber="534">INCLUDED</smallCapsWord>
<smallCapsWord id="8D7EA010FFCFE42FFF4BFB94FEC0FBE0" baselines="1103" box="[243,321,1090,1107]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="genera" pageId="41" pageNumber="534">GENERA</smallCapsWord>
. —
<taxonomicName id="4C274D4FFFCFE42FFED3FBEAFDDAFBE7" authority="Merle, 2005 b" authorityName="Merle" authorityYear="2005" box="[363,603,1084,1108]" class="Gastropoda" family="Muricidae" genus="Jsowerbya" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFCFE42FFED3FBEAFE4BFBE7" box="[363,458,1084,1108]" italics="true" pageId="41" pageNumber="534">Jsowerbya</emphasis>
<bibRefCitation id="EFB64B3DFFCFE42FFE68FBEAFDDAFBE7" author="MERLE D." box="[464,603,1084,1108]" pageId="41" pageNumber="534" pagination="505 - 517" refId="ref46835" refString="MERLE D. 2005 b. - Jsowerbya, new genus of Muricidae (Mollusca Gastropoda) from the Eocene of the Paris (France) and Hampshire basins (England), with a phylogenetic assessment of its ocenebrine versus ergalataxine affinities. Geobios 38 (4): 505 - 517. https: // doi. org / 10.1016 / j. geobios. 2004.02.002" type="journal article" year="2005">Merle, 2005b</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C274D4FFFCFE42FFDDCFBEBFC82FBE7" authority="Merle" authorityName="Merle" authorityYear="2024" box="[612,771,1084,1108]" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFCFE42FFDDCFBEBFD3FFBE7" box="[612,702,1085,1108]" italics="true" pageId="41" pageNumber="534">Beyregrex</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFCFE42FFF3BFB81FF4BFBDC" box="[131,202,1111,1135]" pageId="41" pageNumber="534" rank="genus">n. gen.</taxonomicNameLabel>
and
<taxonomicName id="4C274D4FFFCFE42FFF47FB81FE7FFBDC" authority="Merle" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[255,510,1111,1135]" class="Gastropoda" family="Muricidae" genus="Pseudotrophonopsis" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFCFE42FFF47FB81FE38FBDC" box="[255,441,1111,1135]" italics="true" pageId="41" pageNumber="534">Pseudotrophonopsis</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFCFE42FFDB2FB81FDD1FBDC" box="[522,592,1111,1135]" pageId="41" pageNumber="534" rank="genus">n. gen.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C33D6547FFCFE42FFF3CFB42FE1BF858" pageId="41" pageNumber="534" type="discussion">
<paragraph id="8B9836CCFFCFE42FFF3CFB42FE8DFB1F" blockId="41.[131,777,1172,2028]" box="[132,268,1172,1198]" pageId="41" pageNumber="534">
<smallCapsWord id="8D7EA010FFCFE42FFF3CFB42FE8DFB1F" baselines="1191,1191" box="[132,268,1172,1198]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Discussion" pageId="41" pageNumber="534">DISCUSSION</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFCFE42FFF3CFB62FE1BF858" blockId="41.[131,777,1172,2028]" pageId="41" pageNumber="534">
This informal group includes basal taxa often erroneously attributed to the
<taxonomicName id="4C274D4FFFCFE42FFEF4FB02FE67FB5D" authorityName="Radwin &amp; d'Attilio" authorityYear="1971" box="[332,486,1236,1262]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="subFamily" subFamily="Muricopsinae">Muricopsinae</taxonomicName>
(synonym of
<taxonomicName id="4C274D4FFFCFE42FFD31FB02FD7FFB5D" authorityName="Keen" authorityYear="1971" box="[649,766,1236,1262]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="subFamily" subFamily="Aspellinae">Aspellinae</taxonomicName>
) because of their biconic shape and the presence of 1-3 columellar tubercles or attributed to the
<taxonomicName id="4C274D4FFFCFE42FFDD3FAC5FC83FA9E" authorityName="Cossmann" authorityYear="1903" box="[619,770,1299,1325]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="subFamily" subFamily="Trophoninae">Trophoninae</taxonomicName>
, because of the more or less lamellose varices. In the phylogenetic analysis, they represent basal taxa (Clades G, H, I, J, L of Winclada [
<figureCitation id="131C2A49FFCFE42FFEC9FAA5FE32FA3E" box="[369,435,1395,1421]" captionStart="FIG" captionStartId="6.[134,143,1784,1801]" captionTargetBox="[134,618,235,1734]" captionTargetId="graphics-161@6.[134,618,235,1734]" captionTargetPageId="6" captionText="FIG. 1. — Strict Consensus tree of 7 equally parsimonious trees (length 107, ci 0.53, ri 0.78) generated by Winclada and computed using the option Heuristic Analysis (mult* max*), all characters non additive." figureDoi="http://doi.org/10.5281/zenodo.13985762" httpUri="https://zenodo.org/record/13985762/files/figure.png" pageId="41" pageNumber="534">Fig. 1</figureCitation>
; Appendix 3] and Clades 44, 43, 42, 41, 40 of PAUP [Appendix 5]). They essentially differ from members from the other subfamilies by having a combination of characters including a posterior sinus, a D6 denticle, occasional columellar tubercles, and by lacking the ID denticle. The primary cords appear late in ontogeny and IP is missing in the most basal species of this group (e.g.,
<taxonomicName id="4C274D4FFFCFE42FFEB1F985FD10F9DF" authority="(Carez, 1879)" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Carez" baseAuthorityYear="1879" box="[265,657,1618,1644]" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="sarroniensis" status="comb. nov.">
<emphasis id="B953EADEFFCFE42FFEB1F985FE6CF9DF" box="[265,493,1619,1644]" italics="true" pageId="41" pageNumber="534">Beyregrex sarroniensis</emphasis>
(
<bibRefCitation id="EFB64B3DFFCFE42FFE45F984FD09F9DF" author="CAREZ L." box="[509,648,1618,1644]" pageId="41" pageNumber="534" pagination="637 - 641" refId="ref43315" refString="CAREZ L. 1879. - Description d'especes nouvelles du bassin de Paris. Bulletin de la Societe geologique de France serie 3, 7: 637 - 641." type="journal article" year="1879">Carez, 1879</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFCFE42FFD24F985FD7EF9DF" box="[668,767,1618,1645]" pageId="41" pageNumber="534" rank="species">n. comb.</taxonomicNameLabel>
) from the Thanetian, whereas in the derived members (e.g., members of
<taxonomicName id="4C274D4FFFCFE42FFEABF944FD9EF91E" authority="Merle, 2005 b" authorityName="Merle" authorityYear="2005" box="[275,543,1682,1709]" class="Gastropoda" family="Muricidae" genus="Jsowerbya" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFCFE42FFEABF944FEFDF91F" box="[275,380,1682,1708]" italics="true" pageId="41" pageNumber="534">Jsowerbya</emphasis>
<bibRefCitation id="EFB64B3DFFCFE42FFE3CF944FD9EF91E" author="MERLE D." box="[388,543,1682,1709]" pageId="41" pageNumber="534" pagination="505 - 517" refId="ref46835" refString="MERLE D. 2005 b. - Jsowerbya, new genus of Muricidae (Mollusca Gastropoda) from the Eocene of the Paris (France) and Hampshire basins (England), with a phylogenetic assessment of its ocenebrine versus ergalataxine affinities. Geobios 38 (4): 505 - 517. https: // doi. org / 10.1016 / j. geobios. 2004.02.002" type="journal article" year="2005">Merle, 2005b</bibRefCitation>
</taxonomicName>
) they appear earlier, and the primary cord IP is often present. From an evolutionary point of view, members of this group are diversified from the late Paleocene to the middle Eocene with
<taxonomicName id="4C274D4FFFCFE42FFF3CF8C4FEB4F89F" authority="Merle" authorityName="Merle" authorityYear="2024" box="[132,309,1810,1836]" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFCFE42FFF3CF8C4FF69F898" box="[132,232,1810,1835]" italics="true" pageId="41" pageNumber="534">Beyregrex</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFCFE42FFEF9F8C4FE0FF89F" box="[321,398,1810,1836]" pageId="41" pageNumber="534" rank="genus">n. gen.</taxonomicNameLabel>
and
<taxonomicName id="4C274D4FFFCFE42FFE7DF8C4FDAFF89F" authorityName="Merle" authorityYear="2005" box="[453,558,1810,1836]" class="Gastropoda" family="Muricidae" genus="Jsowerbya" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFCFE42FFE7DF8C4FDAFF89F" box="[453,558,1810,1836]" italics="true" pageId="41" pageNumber="534">Jsowerbya</emphasis>
</taxonomicName>
. The decline of the group begins at the Priabonian in which only
<taxonomicName id="4C274D4FFFCFE42FFD19F8E4FF48F8D8" authority="Merle" authorityName="Merle" authorityYear="2024" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFCFE42FFD19F8E4FC86F8F8" box="[673,775,1842,1867]" italics="true" pageId="41" pageNumber="534">Beyregrex</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFCFE42FFF6FF884FEA6F8DF" box="[215,295,1874,1900]" pageId="41" pageNumber="534" rank="genus">n. gen.</taxonomicNameLabel>
and
<taxonomicName id="4C274D4FFFCFE42FFEDDF887FD01F8D8" authority="Merle" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[357,640,1873,1899]" class="Gastropoda" family="Muricidae" genus="Pseudotrophonopsis" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFCFE42FFEDDF887FDB2F8D8" box="[357,563,1873,1899]" italics="true" pageId="41" pageNumber="534">Pseudotrophonopsis</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFCFE42FFD36F884FD5CF8DF" box="[654,733,1874,1900]" pageId="41" pageNumber="534" rank="genus">n. gen.</taxonomicNameLabel>
are recorded.
<taxonomicName id="4C274D4FFFCFE42FFF48F8A4FE1BF838" authority="Merle" authorityName="Merle" authorityYear="2024" box="[240,410,1905,1931]" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFCFE42FFF48F8A4FED0F838" box="[240,337,1906,1931]" italics="true" pageId="41" pageNumber="534">Beyregrex</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFCFE42FFE1CF8A4FE6FF83F" box="[420,494,1906,1932]" pageId="41" pageNumber="534" rank="genus">n. gen.</taxonomicNameLabel>
survives after the Eocene/ Oligocene climatic crisis and is known in the Early Miocene (Burdigalian) with
<taxonomicName id="4C274D4FFFCFE42FFE2AF864FEA0F858" authority="(Cossmann &amp; Peyrot, 1924)" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Cossmann &amp; Peyrot" baseAuthorityYear="1924" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="cedillatus" status="comb. nov.">
<emphasis id="B953EADEFFCFE42FFE2AF864FDE3F878" box="[402,610,1969,1995]" italics="true" pageId="41" pageNumber="534">Beyregrex cedillatus</emphasis>
(
<bibRefCitation id="EFB64B3DFFCFE42FFDC9F867FE98F858" author="COSSMANN M. &amp; PEYROT A." pageId="41" pageNumber="534" pagination="71 - 144" refId="ref43770" refString="COSSMANN M. &amp; PEYROT A. 1924. - Conchologie neogenique de l'Aquitaine (Species n ° 707 - 762). Actes de la Societe linneenne de Bordeaux 75 (2): 71 - 144." type="journal article" year="1924">Cossmann &amp; Peyrot, 1924</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFCFE42FFE96F804FE0EF858" box="[302,399,2001,2028]" pageId="41" pageNumber="534" rank="species">n. comb</taxonomicNameLabel>
..
</paragraph>
</subSubSection>
</treatment>
</document>