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<document id="E372D4FA7413CB7E520927C8E8736BD6" ID-DOI="10.15298/rusentj.31.3.12" ID-ISSN="0132-8069" ID-Zenodo-Dep="13177978" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722593438979" checkinUser="felipe" docAuthor="Stuke, J. H." docDate="2022" docId="038E3646FFA6FF92FECACE47B2E8AF38" docLanguage="en" docName="RussEntomolJ.31.3.297-311.pdf" docOrigin="Russian Entomological Journal 31 (3)" docSource="http://dx.doi.org/10.15298/rusentj.31.3.12" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Allotrichoma tskarishdidae Stuke 2022, sp.n." docType="treatment" docVersion="2" lastPageNumber="306" masterDocId="FFB74E3EFFA1FF9BFFC1CA60B356AC5F" masterDocTitle="Faunistic and taxonomic remarks on European Allotrichoma Becker, 1896 (Diptera: Ephydridae) with the description of a new species from Georgia" masterLastPageNumber="311" masterPageNumber="297" pageNumber="304" updateTime="1722632869847" updateUser="ExternalLinkService">
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<mods:title id="6BFF13641F35F1B642C97635E655FA54">Faunistic and taxonomic remarks on European Allotrichoma Becker, 1896 (Diptera: Ephydridae) with the description of a new species from Georgia</mods:title>
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<emphasis id="B9535B42FFA6FF9CFECACE47B179A860" box="[267,559,1063,1087]" italics="true" pageId="7" pageNumber="304">Allotrichoma tskarishdidae</emphasis>
Stuke
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,
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<figureCitation id="131C9BD5FFA6FF9CFE63CE24B2A2A803" box="[418,500,1092,1116]" captionStart="Figs 912" captionStartId="6.[226,263,1969,1988]" captionTargetBox="[196,1394,646,1944]" captionTargetId="figure-321@6.[196,1399,646,1945]" captionTargetPageId="6" captionText="Figs 912. Epandrium, cercus, surstylus of European Allotrichoma, dorsal view (above) and lateral view (below): 9 — sicanum Canzoneri, 1980; 10 — strandi Duda, 1942; 11 — trispinum Becker, 1896; 12 — tskarishdidae sp.n. Рис. 1—12. Эпандрий, церки и сурстили европейских видов Allotrichoma, сверху (верхний рис.) и сбоку(ниЖний рис.): 9 — sicanum Canzoneri, 1980; 10 — strandi Duda, 1942; 11 — trispinum Becker, 1896; 12 — tskarishdidae sp.n." figureDoi="http://doi.org/10.5281/zenodo.13177988" httpUri="https://zenodo.org/record/13177988/files/figure.png" pageId="7" pageNumber="304">Figs 12</figureCitation>
,
<figureCitation id="131C9BD5FFA6FF9CFDC1CE24B148A803" box="[512,542,1092,1116]" captionStart="Figs 1324" captionStartId="8.[226,263,1801,1820]" captionText="Figs 1324. Male process of sternite 5 of European Allotrichoma, lateral view: 13 — bezzii Becker, 1896; 14 — bifidum Papp, 1974; 15 — filiforme Becker, 1896; 16 — laterale (Loew, 1860); 17 — leotoni Vitte, 1992; 18 — picenum Canzoneri, Rampini, 1990; 19 — quadripectinatum (Becker, 1903); 20 — schumanni Papp, 1974; 21 — sicanum Canzoneri, 1980; 22 — strandi Duda, 1942; 23 — trispinum Becker, 1896; 24 — tskarishdidae sp.n.; ca S5 — central arm of process of sternite 5; la S5 — lateral arm of process of sternite 5; lp S5 — lateral plate of process of sternite 5; p S5 — process of sternite 5; te S5 — tringular extension of sternite 5. Рис. 1324. Отростки 5 стернита самцов европейских видов Allotrichoma, сбоку: 13 — bezzii Becker, 1896; 14 — bifidum Papp, 1974; 15 — filiforme Becker, 1896; 16 — laterale (Loew, 1860); 17 — leotoni Vitte, 1992; 18 — picenum Canzoneri, Rampini, 1990; 19 — quadripectinatum (Becker, 1903); 20 — schumanni Papp, 1974; 21 — sicanum Canzoneri, 1980; 22 — strandi Duda, 1942; 23 — trispinum Becker, 1896; 24 — tskarishdidae sp.n.; ca S5 — центральный выступ отростка 5-го стернита; la S5 — латеральный выступ отростка 5-го стернита; lp S5 — латеральнаЯ пластинка отростка 5-го стернита; p S5 — отросток 5-го стернита; te S5 — треугольное расширение 5-го стернита." pageId="7" pageNumber="304">24</figureCitation>
.
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leg. / 2473 [cross written]”; (2) “
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<emphasis id="B9535B42FFA6FF9CFE73CEF9B1C9A8F3" box="[434,671,1177,1196]" italics="true" pageId="7" pageNumber="304">Allotrichoma tskarishdidae</emphasis>
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/ sp.n.
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det. 2021”.
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specimen is pinned using a minuten and is in excellent condition.
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abdomen is dissected, macerated and stored in a glycerine microvial pinned underneath the specimen.
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will be preserved in the collection of the
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für
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<collectorName id="26D2E286FFA6FF9CFEFBCF71B29CA97B" box="[314,458,1297,1316]" pageId="7" pageNumber="304">Leibniz Institute</collectorName>
for
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and
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,
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,
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(
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).
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N Kirach-Mughanlo
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]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33DD4DBFFA6FF9CFF23CFC1B670AA88" pageId="7" pageNumber="304" type="description">
<paragraph id="8B988750FFA6FF9CFF23CFC1B1EDA98E" blockId="7.[189,776,1063,2076]" pageId="7" pageNumber="304">
DESCRIPTION
<typeStatus id="549C39F2FFA6FF9CFE4CCFC1B15BA9E9" box="[397,525,1441,1462]" pageId="7" pageNumber="304" type="holotype">HOLOTYPE</typeStatus>
(
<specimenCount id="9D214CD9FFA6FF9CFDE3CFFEB16EA9E9" box="[546,568,1438,1462]" count="1" pageId="7" pageNumber="304" type="male"></specimenCount>
). Length about
<quantity id="4CDF2AB5FFA6FF9CFD29CFC1B3B3A98E" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="304" unit="mm" value="1.4">1.4 mm</quantity>
. Wing length =
<quantity id="4CDF2AB5FFA6FF9CFE43CFDCB299A98E" box="[386,463,1468,1489]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="304" unit="mm" value="1.2">1.2 mm</quantity>
. Head height =
<quantity id="4CDF2AB5FFA6FF9CFDABCFDCB1E1A98E" box="[618,695,1468,1489]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="304" unit="mm" value="0.4">0.4 mm</quantity>
.
</paragraph>
<paragraph id="8B988750FFA6FF9CFF23CFB6B19BA443" blockId="7.[189,776,1063,2076]" pageId="7" pageNumber="304">Head black. Gena-eye-ratio (in lateral view genal height measured at the maximum eye height: eye height) = 0.2. Antenna black, slightly silver dusted. Arista black, with 4 long branches. Eye red brown, all ommatidia about same size, no ommatrichia recognized. Eye height-length-ratio (in lateral view maximum eye height: maximum eye length) = 1.3. Ocelli forming an almost isosceles triangle. Frons silver grey to brown dusted with fronto-orbital plate and lunule densely silver dusted. Frontal triangle indistinct, large, reaching to ptilinal suture. Face not protruding, silver dusted. Facial ratio (distance between the oral margin and the dorsum of the ptilinal fissure: narrowest distance between the compound eyes across the face) = 1.0. Clypeus black, short, slightly silver grey dusted. Gena silver dusted Occiput silver grey dusted. Palpus black. Proboscis inconspicuous. Chaetotaxy: large ocellar seta inserted lateral to anterior ocellus; no setulae between ocelli; 1 large proclinate postocellar seta; 1 outer and 1 inner vertical seta of about the same size; no paravertical seta; regularly arranged line of postocular setae; 1 proclinate and 1 reclinate fronto-orbital setae; no frontal setulae; 2 large inclinate facial setae and 2 minute facial setulae; 1 prominent genal seta and 34 genal setulae.</paragraph>
<paragraph id="8B988750FFA6FF9CFC93CB5BB664A884" blockId="7.[812,1400,315,2071]" pageId="7" pageNumber="304">
Thorax covered with brown to silver grey dusting. Scutum with lines of short black setulae. Scutum and scutellum grey dusted; scutum with a central stripe and 2 sublateral stripes of distinct brown dusting. Anepisternum and katepisternum with scattered setulae. Pleura silver grey dusted with a brown spot dorsally at the anepisternum and dorsally at the notopleuron. Chaetotaxy: 24 lines of acrostichal setae; 0+1 dorsocentral setae; 1 prescutellar seta; 1 presutural supra alar seta; 1 postpronotal seta; 2 notopleural seta, posterior inserted above level of anterior; 1 postalar seta; 1 large apical, 1 large lateral, 1 small lateral and 2 small central scutellar setae; 1 posterodorsal katepisternal seta; 2 setae at posterior margin of anepisternum. Wing hyaline, completely covered with microtrichia. Veins light brown. Costa without seta between subcostal break and R
<subScript id="17A38515FFA6FF9CFCFDC8A0B007AE93" attach="left" box="[828,849,704,716]" fontSize="5" pageId="7" pageNumber="304">4+5</subScript>
. Alula small, with long brown setulae at its hind margin. Costal index I (straight line distance between the apices of R 1 and R
<subScript id="17A38515FFA6FF9CFCA2C895B02EAF5E" attach="left" box="[867,888,757,769]" fontSize="5" pageId="7" pageNumber="304">2+3</subScript>
[section 2 of Costa]: straight line distance between the apices of R
<subScript id="17A38515FFA6FF9CFC54C970B0FCAF43" attach="left" box="[917,938,784,796]" fontSize="5" pageId="7" pageNumber="304">2+3</subScript>
and R
<subScript id="17A38515FFA6FF9CFC26C970B0AAAF43" attach="left" box="[999,1020,784,796]" fontSize="5" pageId="7" pageNumber="304">4+5</subScript>
[section 3 of Costa]) = 2.6. Costal index II (straight line distance between the apices of R
<subScript id="17A38515FFA6FF9CFAC9C94AB64BAF69" attach="left" box="[1288,1309,810,822]" fontSize="5" pageId="7" pageNumber="304">4+5</subScript>
and R
<subScript id="17A38515FFA6FF9CFAA0C94BB620AF68" attach="left" box="[1377,1398,811,823]" fontSize="5" pageId="7" pageNumber="304">2+3</subScript>
[section 3 of Costa]: straight line distance between the apices of Media and R
<subScript id="17A38515FFA6FF9CFC05C900B08FAF33" attach="left" box="[964,985,864,876]" fontSize="5" pageId="7" pageNumber="304">4+5</subScript>
[section 4 of Costa]) = 2.5. R
<subScript id="17A38515FFA6FF9CFB3FC900B645AF33" attach="left" box="[1278,1299,864,876]" fontSize="5" pageId="7" pageNumber="304">4+5</subScript>
vein-ratio (straight line distance along vein R
<subScript id="17A38515FFA6FF9CFBBEC91AB7C2AFD9" attach="left" box="[1151,1172,890,902]" fontSize="5" pageId="7" pageNumber="304">4+5</subScript>
between crossvein r-m and branch of R
<subScript id="17A38515FFA6FF9CFC02C9F5B08EAFFE" attach="left" box="[963,984,917,929]" fontSize="5" pageId="7" pageNumber="304">4+5</subScript>
and R
<subScript id="17A38515FFA6FF9CFBD7C9F5B77DAFFE" attach="left" box="[1046,1067,917,929]" fontSize="5" pageId="7" pageNumber="304">2+3</subScript>
[section 1 of R
<subScript id="17A38515FFA6FF9CFB7DC9F5B787AFFE" attach="left" box="[1212,1233,917,929]" fontSize="5" pageId="7" pageNumber="304">4+5</subScript>
]: distance apicad of r-m [section 2 of R
<subScript id="17A38515FFA6FF9CFBC9C9D0B74BAFE3" attach="left" box="[1032,1053,944,956]" fontSize="5" pageId="7" pageNumber="304">4+5</subScript>
]) = 0.1. M vein-ratio (straight line distance along vein M between crossveins dm-cu and r-m [section 1 of M]: distance apicad of dm-cu [section 2 of M]) = 0.4. Costa reaching apex of media. Radial vein R
<subScript id="17A38515FFA6FF9CFAD8CE60B678A853" attach="left" box="[1305,1326,1024,1036]" fontSize="5" pageId="7" pageNumber="304">4+5</subScript>
almost straight. Haltere whitish with light brown base. Legs black to brown with bases and apices of tibiae yellow brown and three basal tarsomeres yellow. Legs silver grey dusted with the exception of hind side of shining hind femur and hind tibia. Legs covered with short black setulae. Hind metatarsus ventrally with dense golden setulae.Fore femur posterodorsally with 56 black setae; middle femur anterodorsally with 4 setae. Metatarsus IItibia II-ratio (length metatarsus 2: length tibia 2) = 0.4.
</paragraph>
<paragraph id="8B988750FFA6FF9CFC93CE81B7A5AAE3" blockId="7.[812,1400,315,2071]" pageId="7" pageNumber="304">
Abdomen black. Tergites grey dusted and covered with black setulae. Tergite III-IV-ratio (length tergite 3 medially: length tergite 4 medially) = 0.8. Tergite IV-V-ratio (length tergite 4 medially: length tergite 5 medially) = 0.5. Sternites 25 each with 25 black setulae. No process of sternite 5 (“medial process at 5th sternal flap” sensu
<bibRefCitation id="EFB6FAA1FFA6FF9CFB19CF06B0EDA9C9" author="Mathis W. N. &amp; Zatwarnicki T." pageId="7" pageNumber="304" pagination="1 - 101" refId="ref13183" refString="Mathis W. N., Zatwarnicki T. 2012. Revision of New World Species of the Shorefly Subgenus Allotrichoma Becker of the Genus Allotrichoma with Description of the Subgenus Neotrichoma (Diptera, Ephydridae, Hecamedini) // Zookeys. Vol. 161. P. 1 - 101." type="book chapter" year="2012">Mathis and Zatwarnicki 2012</bibRefCitation>
) but with a pair of triangular extensions and base of sternite 5 with long posterior directed setae (
<figureCitation id="131C9BD5FFA6FF9CFAE2CFFBB63AA9EF" box="[1315,1388,1435,1457]" captionStart="Figs 1324" captionStartId="8.[226,263,1801,1820]" captionText="Figs 1324. Male process of sternite 5 of European Allotrichoma, lateral view: 13 — bezzii Becker, 1896; 14 — bifidum Papp, 1974; 15 — filiforme Becker, 1896; 16 — laterale (Loew, 1860); 17 — leotoni Vitte, 1992; 18 — picenum Canzoneri, Rampini, 1990; 19 — quadripectinatum (Becker, 1903); 20 — schumanni Papp, 1974; 21 — sicanum Canzoneri, 1980; 22 — strandi Duda, 1942; 23 — trispinum Becker, 1896; 24 — tskarishdidae sp.n.; ca S5 — central arm of process of sternite 5; la S5 — lateral arm of process of sternite 5; lp S5 — lateral plate of process of sternite 5; p S5 — process of sternite 5; te S5 — tringular extension of sternite 5. Рис. 1324. Отростки 5 стернита самцов европейских видов Allotrichoma, сбоку: 13 — bezzii Becker, 1896; 14 — bifidum Papp, 1974; 15 — filiforme Becker, 1896; 16 — laterale (Loew, 1860); 17 — leotoni Vitte, 1992; 18 — picenum Canzoneri, Rampini, 1990; 19 — quadripectinatum (Becker, 1903); 20 — schumanni Papp, 1974; 21 — sicanum Canzoneri, 1980; 22 — strandi Duda, 1942; 23 — trispinum Becker, 1896; 24 — tskarishdidae sp.n.; ca S5 — центральный выступ отростка 5-го стернита; la S5 — латеральный выступ отростка 5-го стернита; lp S5 — латеральнаЯ пластинка отростка 5-го стернита; p S5 — отросток 5-го стернита; te S5 — треугольное расширение 5-го стернита." pageId="7" pageNumber="304">Fig. 24</figureCitation>
). Cercus-epandrium-surstylus-complex as shown in
<figureCitation id="131C9BD5FFA6FF9CFAE6CFD6B625A994" box="[1319,1395,1462,1484]" captionStart="Figs 912" captionStartId="6.[226,263,1969,1988]" captionTargetBox="[196,1394,646,1944]" captionTargetId="figure-321@6.[196,1399,646,1945]" captionTargetPageId="6" captionText="Figs 912. Epandrium, cercus, surstylus of European Allotrichoma, dorsal view (above) and lateral view (below): 9 — sicanum Canzoneri, 1980; 10 — strandi Duda, 1942; 11 — trispinum Becker, 1896; 12 — tskarishdidae sp.n. Рис. 1—12. Эпандрий, церки и сурстили европейских видов Allotrichoma, сверху (верхний рис.) и сбоку(ниЖний рис.): 9 — sicanum Canzoneri, 1980; 10 — strandi Duda, 1942; 11 — trispinum Becker, 1896; 12 — tskarishdidae sp.n." figureDoi="http://doi.org/10.5281/zenodo.13177988" httpUri="https://zenodo.org/record/13177988/files/figure.png" pageId="7" pageNumber="304">Fig. 12</figureCitation>
. Epandrium with 2 strong setae. Cercus with 2 large apical inwards curved setae and 3 smaller, subapical, ventral, upwards curved setae. Surstylus completely fused with epandrium and not distinct. Its position might be marked by a row of ventrally directed setulae (
<figureCitation id="131C9BD5FFA6FF9CFBEDCC5BB722AA0F" box="[1068,1140,1595,1617]" captionStart="Figs 912" captionStartId="6.[226,263,1969,1988]" captionTargetBox="[196,1394,646,1944]" captionTargetId="figure-321@6.[196,1399,646,1945]" captionTargetPageId="6" captionText="Figs 912. Epandrium, cercus, surstylus of European Allotrichoma, dorsal view (above) and lateral view (below): 9 — sicanum Canzoneri, 1980; 10 — strandi Duda, 1942; 11 — trispinum Becker, 1896; 12 — tskarishdidae sp.n. Рис. 1—12. Эпандрий, церки и сурстили европейских видов Allotrichoma, сверху (верхний рис.) и сбоку(ниЖний рис.): 9 — sicanum Canzoneri, 1980; 10 — strandi Duda, 1942; 11 — trispinum Becker, 1896; 12 — tskarishdidae sp.n." figureDoi="http://doi.org/10.5281/zenodo.13177988" httpUri="https://zenodo.org/record/13177988/files/figure.png" pageId="7" pageNumber="304">Fig. 12</figureCitation>
). Postgonite long, straight, with broad base, with two lateral directed small setulae and no obvious seta at its base. Phallus apodeme triangular with the muscle attachment slightly backwards directed. Phallus elongated and without any obvious characters.
</paragraph>
<paragraph id="8B988750FFA6FF9CFC93CCA1B670AA88" blockId="7.[812,1400,315,2071]" box="[850,1318,1729,1751]" pageId="7" pageNumber="304">Female: Females cannot currently be identified.</paragraph>
</subSubSection>
<subSubSection id="C33DD4DBFFA6FF92FC93CCBBB054AE29" lastPageId="9" lastPageNumber="306" pageId="7" pageNumber="304" type="diagnosis">
<paragraph id="8B988750FFA6FF9CFC93CCBBB620A448" blockId="7.[812,1400,315,2071]" pageId="7" pageNumber="304">
DIAGNOSIS:
<taxonomicName id="4C27FCD3FFA6FF9CFC22CCBBB7BEAAAF" authorityName="Stuke" authorityYear="2022" box="[995,1256,1755,1776]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="304" phylum="Arthropoda" rank="species" species="tskarishdidae" status="sp. nov.">
<emphasis id="B9535B42FFA6FF9CFC22CCBBB7BEAAAF" box="[995,1256,1755,1776]" italics="true" pageId="7" pageNumber="304">Allotrichoma tskarishdidae</emphasis>
</taxonomicName>
<emphasis id="B9535B42FFA6FF9CFB32CCBBB675AAAF" bold="true" box="[1267,1315,1755,1776]" pageId="7" pageNumber="304">
<taxonomicNameLabel id="A260E639FFA6FF9CFB32CCBBB675AAAF" box="[1267,1315,1755,1776]" pageId="7" pageNumber="304" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
belongs to a group of
<taxonomicName id="4C27FCD3FFA6FF9CFC6ACC96B727AB53" box="[939,1137,1782,1804]" pageId="7" pageNumber="304">
<emphasis id="B9535B42FFA6FF9CFC6ACC96B77EAB54" box="[939,1064,1782,1803]" italics="true" pageId="7" pageNumber="304">Allotrichoma</emphasis>
species
</taxonomicName>
that is easily recognised by this set of characters: (i) face only shallowly convex and not protruding nor with a tubercle; (ii) sternite 5 without distinct process (
<figureCitation id="131C9BD5FFA6FF9CFC40CD26B09FAB04" box="[897,969,1862,1884]" captionStart="Figs 1324" captionStartId="8.[226,263,1801,1820]" captionText="Figs 1324. Male process of sternite 5 of European Allotrichoma, lateral view: 13 — bezzii Becker, 1896; 14 — bifidum Papp, 1974; 15 — filiforme Becker, 1896; 16 — laterale (Loew, 1860); 17 — leotoni Vitte, 1992; 18 — picenum Canzoneri, Rampini, 1990; 19 — quadripectinatum (Becker, 1903); 20 — schumanni Papp, 1974; 21 — sicanum Canzoneri, 1980; 22 — strandi Duda, 1942; 23 — trispinum Becker, 1896; 24 — tskarishdidae sp.n.; ca S5 — central arm of process of sternite 5; la S5 — lateral arm of process of sternite 5; lp S5 — lateral plate of process of sternite 5; p S5 — process of sternite 5; te S5 — tringular extension of sternite 5. Рис. 1324. Отростки 5 стернита самцов европейских видов Allotrichoma, сбоку: 13 — bezzii Becker, 1896; 14 — bifidum Papp, 1974; 15 — filiforme Becker, 1896; 16 — laterale (Loew, 1860); 17 — leotoni Vitte, 1992; 18 — picenum Canzoneri, Rampini, 1990; 19 — quadripectinatum (Becker, 1903); 20 — schumanni Papp, 1974; 21 — sicanum Canzoneri, 1980; 22 — strandi Duda, 1942; 23 — trispinum Becker, 1896; 24 — tskarishdidae sp.n.; ca S5 — центральный выступ отростка 5-го стернита; la S5 — латеральный выступ отростка 5-го стернита; lp S5 — латеральнаЯ пластинка отростка 5-го стернита; p S5 — отросток 5-го стернита; te S5 — треугольное расширение 5-го стернита." pageId="7" pageNumber="304">Fig. 24</figureCitation>
); (iii) surstylus, cercus and epandrium fused. Four
<taxonomicName id="4C27FCD3FFA6FF9CFC9FCD01B772AB28" box="[862,1060,1889,1911]" pageId="7" pageNumber="304">
<emphasis id="B9535B42FFA6FF9CFC9FCD01B08DAB29" box="[862,987,1889,1910]" italics="true" pageId="7" pageNumber="304">Allotrichoma</emphasis>
species
</taxonomicName>
are known from the Palaearctic and from the Arabian Peninsula to belong to this
<taxonomicName id="4C27FCD3FFA6FF9CFB2BCD1BB019ABF3" baseAuthorityName="Becker" baseAuthorityYear="1903" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="304" phylum="Arthropoda" rank="species" species="quadripectinatum">
<emphasis id="B9535B42FFA6FF9CFB2BCD1BB019ABF3" italics="true" pageId="7" pageNumber="304">quadripectinatum</emphasis>
</taxonomicName>
-species group:
<taxonomicName id="4C27FCD3FFA6FF9CFC28CDF6B621ABF4" authority="Mathis et Zatwarnicki, 2017" authorityName="Mathis et Zatwarnicki" authorityYear="2017" box="[1001,1399,1942,1964]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="304" phylum="Arthropoda" rank="species" species="choanum">
<emphasis id="B9535B42FFA6FF9CFC28CDF6B70AABF4" box="[1001,1116,1942,1963]" italics="true" pageId="7" pageNumber="304">A. choanum</emphasis>
Mathis et Zatwarnicki, 2017
</taxonomicName>
(United Arabian Emirates), A.
<emphasis id="B9535B42FFA6FF9CFB9ECDD1B7D9AB99" box="[1119,1167,1969,1990]" italics="true" pageId="7" pageNumber="304">hatta</emphasis>
Mathis et Zatwarnicki, 2017 (United Arabian Emirates),
<emphasis id="B9535B42FFA6FF9CFBA9CDACB7F0ABBE" box="[1128,1190,1996,2017]" italics="true" pageId="7" pageNumber="304">tuareg</emphasis>
<bibRefCitation id="EFB6FAA1FFA6FF9CFB6DCDABB621ABBE" author="Giordani Soika A. G." box="[1196,1399,1995,2017]" pageId="7" pageNumber="304" pagination="95 - 117" refId="ref12952" refString="Giordani Soika A. G. 1956. Viaggi di A. Giordani Soika nel Sahara IX. Contributo allo studio del popolamento del Sahara Diptera Ephydridae // Bollettino del Museo Civico di Storia Naturale di Venezia. Vol. 9. P. 95 - 117." type="journal article" year="1956">Giordani Soika, 1956</bibRefCitation>
(
<collectingCountry id="F330C7C0FFA6FF9CFCF2CD86B028ABA3" box="[819,894,2022,2044]" name="Algeria" pageId="7" pageNumber="304">Algeria</collectingCountry>
,
<collectingCountry id="F330C7C0FFA6FF9CFC48CD86B0B6ABA4" box="[905,992,2022,2043]" name="Morocco" pageId="7" pageNumber="304">Morocco</collectingCountry>
) and
<taxonomicName id="4C27FCD3FFA6FF9CFBD8CD86B620ABA4" authority="(Becker, 1903)" baseAuthorityName="Becker" baseAuthorityYear="1903" box="[1049,1398,2022,2044]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="304" phylum="Arthropoda" rank="species" species="quadripectinatum">
<emphasis id="B9535B42FFA6FF9CFBD8CD86B788ABA4" box="[1049,1246,2022,2043]" italics="true" pageId="7" pageNumber="304">A. quadripectinatum</emphasis>
(
<bibRefCitation id="EFB6FAA1FFA6FF9CFB2DCD86B626ABA4" author="Becker T." box="[1260,1392,2022,2044]" pageId="7" pageNumber="304" pagination="67 - 195" refId="ref12402" refString="Becker T. 1903. Agyptische Dipteren (Fortsetzung und Schluss) // Mitteilungen aus dem Zoologischen Museum in Berlin Bd. 2. S. 67 - 195." type="book chapter" year="1903">Becker, 1903</bibRefCitation>
)
</taxonomicName>
[
<figureCitation id="131C9BD5FFA6FF9CFCF3C261B02AA449" box="[818,892,2049,2071]" captionStart="Fig" captionStartId="12.[226,257,1033,1052]" captionTargetBox="[189,1399,314,998]" captionTargetId="figure-132@12.[189,1399,314,998]" captionTargetPageId="12" captionText="Fig. 29. Records of Allotrichoma quadripectinatum [Becker, 1903] (based on the data presented in this paper and data from Becker [1903], Beschovski, Zatwarnicki [2002], El-Hawagry et al. [2018], Giordani Soika [1956, as A. bellicosum], Stuke [2012], Vite [1991, as A. bellicosum]). Рис. 29. Распространение Allotrichoma quadripectinatum [Becker, 1903] (по данным представленной статьи и работ Becker [1903], Beschovski, Zatwarnicki [2002], El-Hawagry et al. [2018], Giordani Soika [1956, как A. bellicosum], Stuke [2012], Vite [1991, как A. bellicosum])." figureDoi="http://doi.org/10.5281/zenodo.13178002" httpUri="https://zenodo.org/record/13178002/files/figure.png" pageId="7" pageNumber="304">Fig. 29</figureCitation>
]. This
<taxonomicName id="4C27FCD3FFA6FF9CFC7CC261B733A449" baseAuthorityName="Becker" baseAuthorityYear="1903" box="[957,1125,2049,2070]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="304" phylum="Arthropoda" rank="species" species="quadripectinatum">
<emphasis id="B9535B42FFA6FF9CFC7CC261B733A449" box="[957,1125,2049,2070]" italics="true" pageId="7" pageNumber="304">quadripectinatum</emphasis>
</taxonomicName>
species group shares impor-
</paragraph>
<paragraph id="8B988750FFA9FF92FDAACA90B054AE29" blockId="8.[619,1398,240,265]" lastBlockId="9.[189,775,315,871]" lastPageId="9" lastPageNumber="306" pageId="8" pageNumber="305">
Remarks on European
<taxonomicName id="4C27FCD3FFA9FF93FCA0CA90B0B8AD57" authorityName="Bezzi Becker" authorityYear="1896" box="[865,1006,240,264]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B9535B42FFA9FF93FCA0CA90B0B8AD57" box="[865,1006,240,264]" italics="true" pageId="8" pageNumber="305">Allotrichoma</emphasis>
</taxonomicName>
305 tant characters with the
<emphasis id="B9535B42FFA8FF92FE74CB5BB2B5AD0F" box="[437,483,315,336]" italics="true" pageId="9" pageNumber="306">dyna</emphasis>
species-group as defined by
<bibRefCitation id="EFB6FAA1FFA8FF92FF7CCB36B2A3AD34" author="Mathis W. N. &amp; Zatwarnicki T." box="[189,501,342,364]" pageId="9" pageNumber="306" pagination="1 - 101" refId="ref13183" refString="Mathis W. N., Zatwarnicki T. 2012. Revision of New World Species of the Shorefly Subgenus Allotrichoma Becker of the Genus Allotrichoma with Description of the Subgenus Neotrichoma (Diptera, Ephydridae, Hecamedini) // Zookeys. Vol. 161. P. 1 - 101." type="book chapter" year="2012">Mathis and Zatwarnicki [2012]</bibRefCitation>
. With the set of characters given in their cladogram [
<bibRefCitation id="EFB6FAA1FFA8FF92FE0BCB11B055ADD9" author="Mathis W. N. &amp; Zatwarnicki T." box="[458,771,369,391]" pageId="9" pageNumber="306" pagination="1 - 101" refId="ref13183" refString="Mathis W. N., Zatwarnicki T. 2012. Revision of New World Species of the Shorefly Subgenus Allotrichoma Becker of the Genus Allotrichoma with Description of the Subgenus Neotrichoma (Diptera, Ephydridae, Hecamedini) // Zookeys. Vol. 161. P. 1 - 101." type="book chapter" year="2012">Mathis, Zatwarnicki, 2012: 10</bibRefCitation>
,
<figureCitation id="131C9BD5FFA8FF92FF7CCBEBB259ADFF" box="[189,271,395,417]" captionStart="Figs 14" captionStartId="3.[226,263,1921,1940]" captionTargetBox="[201,1398,707,1897]" captionTargetId="figure-236@3.[201,1398,707,1905]" captionTargetPageId="3" captionText="Figs 14. Epandrium, cercus, surstylus of European Allotrichoma, dorsal view (above) and lateral view (below): 1 — bezzii Becker, 1896; 2 — bifidum Papp, 1974; 3 — filiforme Becker, 1896; 4 — laterale (Loew, 1860); ce — cercus; ep — epandrium; pce — protruding part of cercus; su — surstylus. Рис. 1—12. Эпандрий, церки и сурстили европейских видов Allotrichoma, сверху (верхний рис.) и сбоку (ниЖний рис.): 1 — bezzii Becker, 1896; 2 — bifidum Papp, 1974; 3 — filiforme Becker, 1896; 4 — laterale (Loew, 1860); ce — церк; ep — Эпандрий; pce — выступаюЩаЯ часть церка; su — сурстиль." figureDoi="http://doi.org/10.5281/zenodo.13177982" httpUri="https://zenodo.org/record/13177982/files/figure.png" pageId="9" pageNumber="306">Figure 2</figureCitation>
] an assignment to one of their species-groups is not possible. To distinguish the species of the
<taxonomicName id="4C27FCD3FFA8FF92FD9ECBC6B051ADE4" baseAuthorityName="Becker" baseAuthorityYear="1903" box="[607,775,422,443]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="306" phylum="Arthropoda" rank="species" species="quadripectinatum">
<emphasis id="B9535B42FFA8FF92FD9ECBC6B051ADE4" box="[607,775,422,443]" italics="true" pageId="9" pageNumber="306">quadripectinatum</emphasis>
</taxonomicName>
species group the shape and setulation of the terminalia has to be taken into account.
<taxonomicName id="4C27FCD3FFA8FF92FE68CBBBB1E6ADAF" authorityName="Stuke" authorityYear="2022" box="[425,688,475,496]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="306" phylum="Arthropoda" rank="species" species="tskarishdidae" status="sp. nov.">
<emphasis id="B9535B42FFA8FF92FE68CBBBB1E6ADAF" box="[425,688,475,496]" italics="true" pageId="9" pageNumber="306">Allotrichoma tskarishdidae</emphasis>
</taxonomicName>
<emphasis id="B9535B42FFA8FF92FD7DCBBBB1BAADAF" bold="true" box="[700,748,475,496]" pageId="9" pageNumber="306">
<taxonomicNameLabel id="A260E639FFA8FF92FD7DCBBBB1BAADAF" box="[700,748,475,496]" pageId="9" pageNumber="306" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is distinguished from all other species by (iv) the presence of 2 dominant, moderately long and (in dorsal view) medially curved setae at the tip of the cercus and additional upcurved setae ventrally to these (
<figureCitation id="131C9BD5FFA8FF92FE63C826B2BFAE04" box="[418,489,582,604]" captionStart="Figs 912" captionStartId="6.[226,263,1969,1988]" captionTargetBox="[196,1394,646,1944]" captionTargetId="figure-321@6.[196,1399,646,1945]" captionTargetPageId="6" captionText="Figs 912. Epandrium, cercus, surstylus of European Allotrichoma, dorsal view (above) and lateral view (below): 9 — sicanum Canzoneri, 1980; 10 — strandi Duda, 1942; 11 — trispinum Becker, 1896; 12 — tskarishdidae sp.n. Рис. 1—12. Эпандрий, церки и сурстили европейских видов Allotrichoma, сверху (верхний рис.) и сбоку(ниЖний рис.): 9 — sicanum Canzoneri, 1980; 10 — strandi Duda, 1942; 11 — trispinum Becker, 1896; 12 — tskarishdidae sp.n." figureDoi="http://doi.org/10.5281/zenodo.13177988" httpUri="https://zenodo.org/record/13177988/files/figure.png" pageId="9" pageNumber="306">Fig. 12</figureCitation>
), (v) the lack of obvious setae at the postgonite and (vi) a finger like tip of cercus (
<figureCitation id="131C9BD5FFA8FF92FD6EC801B1A0AE29" box="[687,758,609,631]" captionStart="Figs 912" captionStartId="6.[226,263,1969,1988]" captionTargetBox="[196,1394,646,1944]" captionTargetId="figure-321@6.[196,1399,646,1945]" captionTargetPageId="6" captionText="Figs 912. Epandrium, cercus, surstylus of European Allotrichoma, dorsal view (above) and lateral view (below): 9 — sicanum Canzoneri, 1980; 10 — strandi Duda, 1942; 11 — trispinum Becker, 1896; 12 — tskarishdidae sp.n. Рис. 1—12. Эпандрий, церки и сурстили европейских видов Allotrichoma, сверху (верхний рис.) и сбоку(ниЖний рис.): 9 — sicanum Canzoneri, 1980; 10 — strandi Duda, 1942; 11 — trispinum Becker, 1896; 12 — tskarishdidae sp.n." figureDoi="http://doi.org/10.5281/zenodo.13177988" httpUri="https://zenodo.org/record/13177988/files/figure.png" pageId="9" pageNumber="306">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<paragraph id="8B988750FFA9FF93FC86C87DB7EEAA8E" pageId="8" pageNumber="305">
<uuid id="FF81BD85FFA9FF93FC86C87DB7EEAA8E" pageId="8" pageNumber="305">14 15 13 16 18 17 21 19 22 20 23 24</uuid>
</paragraph>
<caption id="DF58D7D8FFA9FF93FF23CD69B2E1A44E" pageId="8" pageNumber="305" startId="8.[226,263,1801,1820]" targetType="figure">
<paragraph id="8B988750FFA9FF93FF23CD69B738AB23" blockId="8.[189,1399,1801,2065]" pageId="8" pageNumber="305">
Figs 1324. Male process of sternite 5 of European
<taxonomicName id="4C27FCD3FFA9FF93FD6DCD69B04BAB43" authorityName="Bezzi Becker" authorityYear="1896" box="[684,797,1801,1820]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B9535B42FFA9FF93FD6DCD69B04BAB43" box="[684,797,1801,1820]" italics="true" pageId="8" pageNumber="305">Allotrichoma</emphasis>
</taxonomicName>
, lateral view: 13 —
<emphasis id="B9535B42FFA9FF93FC14CD69B750AB43" box="[981,1030,1801,1820]" italics="true" pageId="8" pageNumber="305">bezzii</emphasis>
Becker, 1896; 14 —
<taxonomicName id="4C27FCD3FFA9FF93FB07CD69B625AB43" authority="Papp, 1974" authorityName="Papp" authorityYear="1974" box="[1222,1395,1801,1821]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="bifidum">
<emphasis id="B9535B42FFA9FF93FB07CD69B65EAB43" box="[1222,1288,1801,1820]" italics="true" pageId="8" pageNumber="305">bifidum</emphasis>
<bibRefCitation id="EFB6FAA1FFA9FF93FACFCD6AB625AB43" author="Papp L." box="[1294,1395,1801,1821]" pageId="8" pageNumber="305" pagination="403 - 409" refId="ref13235" refString="Papp L. 1974. New or little known species of Ephydridae and Sphaeroceridae from Hungary (Diptera: Acalyptratae) // Acta Zoologica Academiae Scientiarum Hungaricae. Vol. 20. P. 403 - 409." type="book chapter" year="1974">Papp, 1974</bibRefCitation>
</taxonomicName>
; 15 —
<taxonomicName id="4C27FCD3FFA9FF93FF36CD41B297AB6B" authority="Becker, 1896" authorityName="Becker" authorityYear="1896" box="[247,449,1825,1844]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="filiforme">
<emphasis id="B9535B42FFA9FF93FF36CD41B217AB6B" box="[247,321,1825,1844]" italics="true" pageId="8" pageNumber="305">filiforme</emphasis>
Becker, 1896
</taxonomicName>
; 16 —
<taxonomicName id="4C27FCD3FFA9FF93FDC4CD41B19CAB6B" authority="(Loew, 1860)" baseAuthorityName="Loew" baseAuthorityYear="1860" box="[517,714,1825,1844]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="laterale">
<emphasis id="B9535B42FFA9FF93FDC4CD41B11EAB6B" box="[517,584,1825,1844]" italics="true" pageId="8" pageNumber="305">laterale</emphasis>
(Loew, 1860)
</taxonomicName>
; 17 —
<taxonomicName id="4C27FCD3FFA9FF93FCD1CD41B0EFAB6B" authority="Vitte, 1992" authorityName="Vitte" authorityYear="1992" box="[784,953,1825,1844]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="leotoni">
<emphasis id="B9535B42FFA9FF93FCD1CD41B01AAB6B" box="[784,844,1825,1844]" italics="true" pageId="8" pageNumber="305">leotoni</emphasis>
<bibRefCitation id="EFB6FAA1FFA9FF93FC92CD41B0EFAB6B" author="Vitte B." box="[851,953,1825,1844]" pageId="8" pageNumber="305" pagination="255 - 258" refId="ref13844" refString="Vitte B. 1992. Une nouvelle espece du genre Allotrichoma Becker (Diptera, Brachycera) // Etude des Dipteres Ephydrides du Maroc. IV. Nouvelle revue d'entomologie. Vol. 9. P. 255 - 258." type="journal article" year="1992">Vitte, 1992</bibRefCitation>
</taxonomicName>
; 18 —
<taxonomicName id="4C27FCD3FFA9FF93FC3FCD42B66FAB6B" authority="Canzoneri, Rampini, 1990" authorityName="Canzoneri, Rampini" authorityYear="1990" box="[1022,1337,1825,1844]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="picenum">
<emphasis id="B9535B42FFA9FF93FC3FCD42B711AB6B" box="[1022,1095,1826,1844]" italics="true" pageId="8" pageNumber="305">picenum</emphasis>
<bibRefCitation id="EFB6FAA1FFA9FF93FB8ECD41B66FAB6B" author="Canzoneri S. &amp; Rampini L." box="[1103,1337,1825,1844]" pageId="8" pageNumber="305" pagination="43 - 46" refId="ref12633" refString="Canzoneri S., Rampini L. 1990. Osservazioni sul genere Allotrichoma Becker (Diptera, Ephydridae) // Lavori. Societa Veneziana di Scienze Naturali. Vol. 15. P. 43 - 46." type="journal article" year="1990">Canzoneri, Rampini, 1990</bibRefCitation>
</taxonomicName>
; 19 —
<taxonomicName id="4C27FCD3FFA9FF93FF7CCD59B289AB13" authority="(Becker, 1903)" baseAuthorityName="Becker" baseAuthorityYear="1903" box="[189,479,1849,1868]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="quadripectinatum">
<emphasis id="B9535B42FFA9FF93FF7CCD59B200AB13" box="[189,342,1849,1868]" italics="true" pageId="8" pageNumber="305">quadripectinatum</emphasis>
(
<bibRefCitation id="EFB6FAA1FFA9FF93FEA0CD59B28EAB13" author="Becker T." box="[353,472,1849,1868]" pageId="8" pageNumber="305" pagination="67 - 195" refId="ref12402" refString="Becker T. 1903. Agyptische Dipteren (Fortsetzung und Schluss) // Mitteilungen aus dem Zoologischen Museum in Berlin Bd. 2. S. 67 - 195." type="book chapter" year="1903">Becker, 1903</bibRefCitation>
)
</taxonomicName>
; 20 —
<taxonomicName id="4C27FCD3FFA9FF93FDDCCD59B1B5AB13" authority="Papp, 1974" authorityName="Papp" authorityYear="1974" box="[541,739,1849,1869]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="schumanni">
<emphasis id="B9535B42FFA9FF93FDDCCD59B12CAB13" box="[541,634,1849,1868]" italics="true" pageId="8" pageNumber="305">schumanni</emphasis>
<bibRefCitation id="EFB6FAA1FFA9FF93FDBECD5AB1B5AB13" author="Papp L." box="[639,739,1849,1869]" pageId="8" pageNumber="305" pagination="403 - 409" refId="ref13235" refString="Papp L. 1974. New or little known species of Ephydridae and Sphaeroceridae from Hungary (Diptera: Acalyptratae) // Acta Zoologica Academiae Scientiarum Hungaricae. Vol. 20. P. 403 - 409." type="book chapter" year="1974">Papp, 1974</bibRefCitation>
</taxonomicName>
; 21 —
<taxonomicName id="4C27FCD3FFA9FF93FCE0CD5AB0A9AB13" authority="Canzoneri, 1980" authorityName="Canzoneri" authorityYear="1980" box="[801,1023,1849,1868]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="sicanum">
<emphasis id="B9535B42FFA9FF93FCE0CD5AB03FAB13" box="[801,873,1850,1868]" italics="true" pageId="8" pageNumber="305">sicanum</emphasis>
<bibRefCitation id="EFB6FAA1FFA9FF93FCAFCD59B0A9AB13" author="Canzoneri S." box="[878,1023,1849,1868]" pageId="8" pageNumber="305" pagination="9 - 11" refId="ref12599" refString="Canzoneri S. 1980. Tre nuove specie di Sicilia (Diptera, Ephydridae) // Lavori. Societa Veneziana di Scienze Naturali. Vol. 5. P. 9 - 11." type="journal article" year="1980">Canzoneri, 1980</bibRefCitation>
</taxonomicName>
; 22 —
<taxonomicName id="4C27FCD3FFA9FF93FBFCCD59B7B0AB13" authority="Duda, 1942" authorityName="Duda" authorityYear="1942" box="[1085,1254,1849,1868]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="strandi">
<emphasis id="B9535B42FFA9FF93FBFCCD59B72CAB13" box="[1085,1146,1849,1868]" italics="true" pageId="8" pageNumber="305">strandi</emphasis>
Duda, 1942
</taxonomicName>
; 23 —
<taxonomicName id="4C27FCD3FFA9FF93FAE5CD5AB265AB3B" authority="Becker, 1896" authorityName="Becker. An" authorityYear="1896" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="trispinum">
<emphasis id="B9535B42FFA9FF93FAE5CD5AB620AB13" box="[1316,1398,1850,1868]" italics="true" pageId="8" pageNumber="305">trispinum</emphasis>
Becker, 1896
</taxonomicName>
; 24 —
<taxonomicName id="4C27FCD3FFA9FF93FEB3CD31B2B0AB3B" authorityName="Stuke" authorityYear="2022" box="[370,486,1873,1892]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="tskarishdidae" status="sp. nov.">
<emphasis id="B9535B42FFA9FF93FEB3CD31B2B0AB3B" box="[370,486,1873,1892]" italics="true" pageId="8" pageNumber="305">tskarishdidae</emphasis>
</taxonomicName>
<emphasis id="B9535B42FFA9FF93FE2ACD31B140AB3B" bold="true" box="[491,534,1873,1892]" pageId="8" pageNumber="305">
<taxonomicNameLabel id="A260E639FFA9FF93FE2ACD31B140AB3B" box="[491,534,1873,1892]" pageId="8" pageNumber="305" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; ca S5 — central arm of process of sternite 5; la S5 — lateral arm of process of sternite 5; lp S5 — lateral plate of process of sternite 5; p S5 — process of sternite 5; te S5 — tringular extension of sternite 5.
</paragraph>
<paragraph id="8B988750FFA9FF93FF23CDE6B2E1A44E" blockId="8.[189,1399,1801,2065]" pageId="8" pageNumber="305">
Рис. 1324. Отростки 5 стернита самцов европейских видов
<taxonomicName id="4C27FCD3FFA9FF93FCD6CDE5B0DEABC7" authorityName="Bezzi Becker" authorityYear="1896" box="[791,904,1925,1944]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B9535B42FFA9FF93FCD6CDE5B0DEABC7" box="[791,904,1925,1944]" italics="true" pageId="8" pageNumber="305">Allotrichoma</emphasis>
</taxonomicName>
, сбоку: 13 —
<emphasis id="B9535B42FFA9FF93FBCCCDE5B768ABC7" box="[1037,1086,1925,1944]" italics="true" pageId="8" pageNumber="305">bezzii</emphasis>
Becker, 1896; 14 —
<taxonomicName id="4C27FCD3FFA9FF93FB3CCDE5B3BDABEF" authority="Papp, 1974" authorityName="Papp" authorityYear="1974" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="bifidum">
<emphasis id="B9535B42FFA9FF93FB3CCDE5B669ABC7" box="[1277,1343,1925,1944]" italics="true" pageId="8" pageNumber="305">bifidum</emphasis>
<bibRefCitation id="EFB6FAA1FFA9FF93FA87CDE6B3BDABEF" author="Papp L." pageId="8" pageNumber="305" pagination="403 - 409" refId="ref13235" refString="Papp L. 1974. New or little known species of Ephydridae and Sphaeroceridae from Hungary (Diptera: Acalyptratae) // Acta Zoologica Academiae Scientiarum Hungaricae. Vol. 20. P. 403 - 409." type="book chapter" year="1974">Papp, 1974</bibRefCitation>
</taxonomicName>
; 15 —
<taxonomicName id="4C27FCD3FFA9FF93FEEFCDFDB2AFABEF" authority="Becker, 1896" authorityName="Becker" authorityYear="1896" box="[302,505,1949,1968]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="filiforme">
<emphasis id="B9535B42FFA9FF93FEEFCDFDB22CABEF" box="[302,378,1949,1968]" italics="true" pageId="8" pageNumber="305">filiforme</emphasis>
Becker, 1896
</taxonomicName>
; 16 —
<taxonomicName id="4C27FCD3FFA9FF93FDFCCDFDB055ABEF" authority="(Loew, 1860)" baseAuthorityName="Loew" baseAuthorityYear="1860" box="[573,771,1949,1968]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="laterale">
<emphasis id="B9535B42FFA9FF93FDFCCDFDB1D7ABEF" box="[573,641,1949,1968]" italics="true" pageId="8" pageNumber="305">laterale</emphasis>
(Loew, 1860)
</taxonomicName>
; 17 —
<taxonomicName id="4C27FCD3FFA9FF93FC86CDFDB0A7ABEF" authority="Vitte, 1992" authorityName="Vitte" authorityYear="1992" box="[839,1009,1949,1968]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="leotoni">
<emphasis id="B9535B42FFA9FF93FC86CDFDB0D2ABEF" box="[839,900,1949,1968]" italics="true" pageId="8" pageNumber="305">leotoni</emphasis>
<bibRefCitation id="EFB6FAA1FFA9FF93FC4ACDFDB0A7ABEF" author="Vitte B." box="[907,1009,1949,1968]" pageId="8" pageNumber="305" pagination="255 - 258" refId="ref13844" refString="Vitte B. 1992. Une nouvelle espece du genre Allotrichoma Becker (Diptera, Brachycera) // Etude des Dipteres Ephydrides du Maroc. IV. Nouvelle revue d'entomologie. Vol. 9. P. 255 - 258." type="journal article" year="1992">Vitte, 1992</bibRefCitation>
</taxonomicName>
; 18 —
<taxonomicName id="4C27FCD3FFA9FF93FBF4CDFEB624ABEF" authority="Canzoneri, Rampini, 1990" authorityName="Canzoneri, Rampini" authorityYear="1990" box="[1077,1394,1949,1968]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="picenum">
<emphasis id="B9535B42FFA9FF93FBF4CDFEB729ABEF" box="[1077,1151,1950,1968]" italics="true" pageId="8" pageNumber="305">picenum</emphasis>
<bibRefCitation id="EFB6FAA1FFA9FF93FB47CDFDB624ABEF" author="Canzoneri S. &amp; Rampini L." box="[1158,1394,1949,1968]" pageId="8" pageNumber="305" pagination="43 - 46" refId="ref12633" refString="Canzoneri S., Rampini L. 1990. Osservazioni sul genere Allotrichoma Becker (Diptera, Ephydridae) // Lavori. Societa Veneziana di Scienze Naturali. Vol. 15. P. 43 - 46." type="journal article" year="1990">Canzoneri, Rampini, 1990</bibRefCitation>
</taxonomicName>
; 19 —
<taxonomicName id="4C27FCD3FFA9FF93FF37CDD5B14AAB97" authority="(Becker, 1903)" baseAuthorityName="Becker" baseAuthorityYear="1903" box="[246,540,1973,1992]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="quadripectinatum">
<emphasis id="B9535B42FFA9FF93FF37CDD5B2D9AB97" box="[246,399,1973,1992]" italics="true" pageId="8" pageNumber="305">quadripectinatum</emphasis>
(
<bibRefCitation id="EFB6FAA1FFA9FF93FE5DCDD5B143AB97" author="Becker T." box="[412,533,1973,1992]" pageId="8" pageNumber="305" pagination="67 - 195" refId="ref12402" refString="Becker T. 1903. Agyptische Dipteren (Fortsetzung und Schluss) // Mitteilungen aus dem Zoologischen Museum in Berlin Bd. 2. S. 67 - 195." type="book chapter" year="1903">Becker, 1903</bibRefCitation>
)
</taxonomicName>
; 20 —
<taxonomicName id="4C27FCD3FFA9FF93FD9FCDD5B07EAB97" authority="Papp, 1974" authorityName="Papp" authorityYear="1974" box="[606,808,1973,1993]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="schumanni">
<emphasis id="B9535B42FFA9FF93FD9FCDD5B1EDAB97" box="[606,699,1973,1992]" italics="true" pageId="8" pageNumber="305">schumanni</emphasis>
<bibRefCitation id="EFB6FAA1FFA9FF93FD03CDD6B07EAB97" author="Papp L." box="[706,808,1973,1993]" pageId="8" pageNumber="305" pagination="403 - 409" refId="ref13235" refString="Papp L. 1974. New or little known species of Ephydridae and Sphaeroceridae from Hungary (Diptera: Acalyptratae) // Acta Zoologica Academiae Scientiarum Hungaricae. Vol. 20. P. 403 - 409." type="book chapter" year="1974">Papp, 1974</bibRefCitation>
</taxonomicName>
; 21 —
<taxonomicName id="4C27FCD3FFA9FF93FCABCDD6B71AAB97" authority="Canzoneri, 1980" authorityName="Canzoneri" authorityYear="1980" box="[874,1100,1973,1992]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="sicanum">
<emphasis id="B9535B42FFA9FF93FCABCDD6B0E4AB97" box="[874,946,1974,1992]" italics="true" pageId="8" pageNumber="305">sicanum</emphasis>
<bibRefCitation id="EFB6FAA1FFA9FF93FC78CDD5B71AAB97" author="Canzoneri S." box="[953,1100,1973,1992]" pageId="8" pageNumber="305" pagination="9 - 11" refId="ref12599" refString="Canzoneri S. 1980. Tre nuove specie di Sicilia (Diptera, Ephydridae) // Lavori. Societa Veneziana di Scienze Naturali. Vol. 5. P. 9 - 11." type="journal article" year="1980">Canzoneri, 1980</bibRefCitation>
</taxonomicName>
; 22 —
<taxonomicName id="4C27FCD3FFA9FF93FB4FCDD5B66DAB97" authority="Duda, 1942" authorityName="Duda" authorityYear="1942" box="[1166,1339,1973,1992]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="strandi">
<emphasis id="B9535B42FFA9FF93FB4FCDD5B79DAB97" box="[1166,1227,1973,1992]" italics="true" pageId="8" pageNumber="305">strandi</emphasis>
Duda, 1942
</taxonomicName>
; 23 —
<taxonomicName id="4C27FCD3FFA9FF93FF7CCDAEB2DDABBF" authority="Becker, 1896" authorityName="Becker. An" authorityYear="1896" box="[189,395,1997,2016]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="trispinum">
<emphasis id="B9535B42FFA9FF93FF7CCDAEB259ABBF" box="[189,271,1998,2016]" italics="true" pageId="8" pageNumber="305">trispinum</emphasis>
Becker, 1896
</taxonomicName>
; 24 —
<taxonomicName id="4C27FCD3FFA9FF93FE08CDADB16BABBF" authorityName="Stuke" authorityYear="2022" box="[457,573,1997,2016]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="tskarishdidae" status="sp. nov.">
<emphasis id="B9535B42FFA9FF93FE08CDADB16BABBF" box="[457,573,1997,2016]" italics="true" pageId="8" pageNumber="305">tskarishdidae</emphasis>
</taxonomicName>
<emphasis id="B9535B42FFA9FF93FD82CDADB138ABBF" bold="true" box="[579,622,1997,2016]" pageId="8" pageNumber="305">
<taxonomicNameLabel id="A260E639FFA9FF93FD82CDADB138ABBF" box="[579,622,1997,2016]" pageId="8" pageNumber="305" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; ca S5 — центральный выступ отростка 5-го стернита; la S5 — латеральный выступ отростка 5-го стернита; lp S5 — латеральнаЯ пластинка отростка 5-го стернита; p S5 — отросток 5-го стернита; te S5 — треугольное расширение 5-го стернита.
</paragraph>
</caption>
<subSubSection id="C33DD4DBFFA8FF92FF23C81BB1E8AE98" pageId="9" pageNumber="306" type="etymology">
<paragraph id="8B988750FFA8FF92FF23C81BB1E8AE98" blockId="9.[189,775,315,871]" pageId="9" pageNumber="306">
ETYMOLOGY:
<taxonomicName id="4C27FCD3FFA8FF92FE47C81BB1DAAECF" authorityName="Stuke" authorityYear="2022" box="[390,652,635,656]" class="Insecta" family="Ephydridae" genus="Allotrichoma" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="306" phylum="Arthropoda" rank="species" species="tskarishdidae" status="sp. nov.">
<emphasis id="B9535B42FFA8FF92FE47C81BB1DAAECF" box="[390,652,635,656]" italics="true" pageId="9" pageNumber="306">Allotrichoma tskarishdidae</emphasis>
</taxonomicName>
<emphasis id="B9535B42FFA8FF92FD50C81BB194AECF" bold="true" box="[657,706,635,656]" pageId="9" pageNumber="306">
<taxonomicNameLabel id="A260E639FFA8FF92FD50C81BB194AECF" box="[657,706,635,656]" pageId="9" pageNumber="306" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is dedicated to Tskarishdida (-). In Georgian myths Tskarishdida is a half-fish and half-woman deity of rivers and lakes.
</paragraph>
</subSubSection>
<subSubSection id="C33DD4DBFFA8FF92FF23C8ABB2DDAF6F" pageId="9" pageNumber="306" type="distribution">
<paragraph id="8B988750FFA8FF92FF23C8ABB2DDAF6F" blockId="9.[189,775,315,871]" pageId="9" pageNumber="306">
DISTRIBUTION: The records of this species are confined to a small area of the Kura river and its tributary. All locations are situated in
<collectingCountry id="F330C7C0FFA8FF92FE78C961B151AF48" box="[441,519,769,791]" name="Georgia" pageId="9" pageNumber="306">Georgia</collectingCountry>
close to the border with
<collectingCountry id="F330C7C0FFA8FF92FF7CC97BB271AF6E" box="[189,295,795,817]" name="Azerbaijan" pageId="9" pageNumber="306">Azerbaijan</collectingCountry>
(
<figureCitation id="131C9BD5FFA8FF92FEF4C97BB229AF6F" box="[309,383,795,817]" captionStart="Fig" captionStartId="13.[226,257,1865,1884]" captionTargetBox="[189,1399,1145,1829]" captionTargetId="figure-51@13.[189,1399,1145,1829]" captionTargetPageId="13" captionText="Fig. 32. Records of Allotrichoma tskarishdidae sp.n. Рис. 32. Распространение Allotrichoma tskarishdidae sp.n." figureDoi="http://doi.org/10.5281/zenodo.13178010" httpUri="https://zenodo.org/record/13178010/files/figure.png" pageId="9" pageNumber="306">Fig. 32</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33DD4DBFFA8FF92FF23C956B2E8AF38" pageId="9" pageNumber="306" type="biology_ecology">
<paragraph id="8B988750FFA8FF92FF23C956B2E8AF38" blockId="9.[189,775,315,871]" pageId="9" pageNumber="306">ECOLOGY: All specimens were collected at the sparsely vegetated shores of rivers.</paragraph>
</subSubSection>
</treatment>
</document>