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<document id="B7FBCB73256E8421E23846446D66E84D" ID-DOI="10.5281/zenodo.182997" ID-GBIF-Dataset="41df3ff4-efb7-4884-8acf-cf0ca7bb8f09" ID-ISSN="1175-5326" ID-Zenodo-Dep="182997" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459885852836" checkinUser="plazi" docAuthor="Bustamante, Martín R. &amp; Mendelson, Joseph R." docDate="2008" docId="038C1C586229D5164DF67B288F9BC01F" docLanguage="en" docName="zt01820p059.pdf" docOrigin="Zootaxa 1820" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Pristimantis gagliardoi Bustamante &amp; Mendelson, new species" docType="treatment" docVersion="8" lastPageNumber="57" masterDocId="FFB564206228D51E4D6178128D03C315" masterDocTitle="A new frog species (Strabomantidae: Pristimantis) from the High Andes of Southeastern Ecuador" masterLastPageNumber="59" masterPageNumber="49" pageNumber="50" updateTime="1698227758550" updateUser="plazi">
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<mods:title id="F9B301FC431E23D38D0844215C48F13A">A new frog species (Strabomantidae: Pristimantis) from the High Andes of Southeastern Ecuador</mods:title>
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<mods:namePart id="58CD546D46D276B84743EDF47259BDA8">Bustamante, Martín R.</mods:namePart>
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<mods:namePart id="AC0CF7BC07F5017A87F26AAE031F99D5">Mendelson, Joseph R.</mods:namePart>
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<mods:typeOfResource id="0BAEE6E52884C8467D1CF2ECC990DB39">text</mods:typeOfResource>
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<mods:date id="B7507CE6572CC4963EFB2B803500BABC">2008</mods:date>
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<mods:number id="159FC9FAAF3B7876F60CAD9C80034F65">1820</mods:number>
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<mods:identifier id="ED05D221FB5F8F7B8A9A178451FB0B73" type="DOI">10.5281/zenodo.182997</mods:identifier>
<mods:identifier id="8AF55174B433C86C63116A4AAA54466C" type="GBIF-Dataset">41df3ff4-efb7-4884-8acf-cf0ca7bb8f09</mods:identifier>
<mods:identifier id="7EF3FE8F42CED1C5652D92D8B0146943" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1E62D52EAE3468760FA0A72909FC9F4B" type="Zenodo-Dep">182997</mods:identifier>
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<treatment id="038C1C586229D5164DF67B288F9BC01F" ID-DOI="http://doi.org/10.5281/zenodo.5613998" ID-GBIF-Taxon="119371472" ID-Zenodo-Dep="5613998" LSID="urn:lsid:plazi:treatment:038C1C586229D5164DF67B288F9BC01F" httpUri="http://treatment.plazi.org/id/038C1C586229D5164DF67B288F9BC01F" lastPageId="8" lastPageNumber="57" pageId="1" pageNumber="50">
<subSubSection id="C33FFEC56229D51F4DF67B288C07C062" pageId="1" pageNumber="50" type="nomenclature">
<paragraph id="8B9AAD4E6229D51F4DF67B288E9BC041" blockId="1.[151,920,826,887]" box="[151,920,826,852]" pageId="1" pageNumber="50">
<heading id="D0D21A226229D51F4DF67B288E9BC041" bold="true" box="[151,920,826,852]" fontSize="11" level="1" pageId="1" pageNumber="50" reason="1">
<emphasis id="B951715C6229D51F4DF67B288E9BC041" bold="true" box="[151,920,826,852]" pageId="1" pageNumber="50">
<taxonomicName id="4C25D6CD6229D51F4DF67B288FF8C041" authority="Bustamante &amp; Mendelson" authorityName="Bustamante &amp; Mendelson" box="[151,763,826,852]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="50" phylum="Chordata" rank="species" species="gagliardoi" status="sp. nov.">
<emphasis id="B951715C6229D51F4DF67B288CAEC041" bold="true" box="[151,429,826,852]" italics="true" pageId="1" pageNumber="50">Pristimantis gagliardoi</emphasis>
Bustamante &amp; Mendelson
</taxonomicName>
,
<taxonomicNameLabel id="A262CC276229D51F4E697B288E9BC041" box="[776,920,826,852]" pageId="1" pageNumber="50" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9AAD4E6229D51F4DF67B4F8C07C062" blockId="1.[151,920,826,887]" box="[151,260,861,887]" pageId="1" pageNumber="50">
<figureCitation id="131EB1CB6229D51F4DF67B4F8C07C062" box="[151,260,861,887]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,255,1788,1812]" captionStartId-1="3.[151,255,2009,2033]" captionStartId-2="4.[151,255,1752,1776]" captionStartId-3="5.[151,255,1832,1856]" captionStartId-4="6.[151,255,1598,1622]" captionTargetBox-0="[151,1435,1077,1761]" captionTargetBox-1="[226,1360,1154,1985]" captionTargetBox-2="[226,1360,594,1728]" captionTargetBox-3="[415,1171,674,1808]" captionTargetBox-4="[498,1171,474,1554]" captionTargetId-0="figure@2.[151,1436,1077,1762]" captionTargetId-1="figure@3.[226,1360,1154,1985]" captionTargetId-2="figure@4.[226,1360,594,1728]" captionTargetId-3="figure@5.[415,1171,674,1808]" captionTargetId-4="figure@6.[415,1171,474,1574]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Dorsal (left) and ventral (right) views of Pristimantis gagliardoi in preservative; holotype, adult female (QCAZ 27103), SVL = 32.49 mm." captionText-1="FIGURE 2. Lateral view of the head holotype of Pristimantis gagliardoi, holotype, adult female (QCAZ 27103)." captionText-2="FIGURE 3. (A) Right hand of Pristimantis gagliardoi; (B) right foot of Pristimantis gagliardoi. Holotype, adult female (QCAZ 27103)." captionText-3="FIGURE 4. Dorsolateral (A) and ventral (B) views of Pristimantis gagliardoi in live; holotype, adult female (QCAZ 27103), SVL = 32.49 mm." captionText-4="FIGURE 5. Nuptial pad on Finger I of a Pristimantis gagliardoi male (QCAZ 27580)." httpUri-0="https://zenodo.org/record/182998/files/figure.png" httpUri-1="https://zenodo.org/record/182999/files/figure.png" httpUri-2="https://zenodo.org/record/183000/files/figure.png" httpUri-3="https://zenodo.org/record/183001/files/figure.png" httpUri-4="https://zenodo.org/record/183002/files/figure.png" pageId="1" pageNumber="50">Figs. 1–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33FFEC56229D51F4DF67BBF893AC79A" pageId="1" pageNumber="50" type="materials_examined">
<paragraph id="8B9AAD4E6229D51F4DF67BBF8982C0FA" blockId="1.[151,1437,941,1967]" pageId="1" pageNumber="50">
<emphasis id="B951715C6229D51F4DF67BBF8C13C0D2" bold="true" box="[151,272,941,967]" pageId="1" pageNumber="50">
<typeStatus id="549E13EC6229D51F4DF67BBF8C08C0D2" box="[151,267,941,967]" pageId="1" pageNumber="50" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED34358B6229D51F4C767BBF8C64C0D2" box="[279,359,941,967]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="1" pageNumber="50">QCAZ</collectionCode>
27103, an adult female (
<figureCitation id="131EB1CB6229D51F4FEB7BBF8FFBC0D2" box="[650,760,941,967]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,255,1788,1812]" captionStartId-1="3.[151,255,2009,2033]" captionStartId-2="4.[151,255,1752,1776]" captionStartId-3="5.[151,255,1832,1856]" captionTargetBox-0="[151,1435,1077,1761]" captionTargetBox-1="[226,1360,1154,1985]" captionTargetBox-2="[226,1360,594,1728]" captionTargetBox-3="[415,1171,674,1808]" captionTargetId-0="figure@2.[151,1436,1077,1762]" captionTargetId-1="figure@3.[226,1360,1154,1985]" captionTargetId-2="figure@4.[226,1360,594,1728]" captionTargetId-3="figure@5.[415,1171,674,1808]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Dorsal (left) and ventral (right) views of Pristimantis gagliardoi in preservative; holotype, adult female (QCAZ 27103), SVL = 32.49 mm." captionText-1="FIGURE 2. Lateral view of the head holotype of Pristimantis gagliardoi, holotype, adult female (QCAZ 27103)." captionText-2="FIGURE 3. (A) Right hand of Pristimantis gagliardoi; (B) right foot of Pristimantis gagliardoi. Holotype, adult female (QCAZ 27103)." captionText-3="FIGURE 4. Dorsolateral (A) and ventral (B) views of Pristimantis gagliardoi in live; holotype, adult female (QCAZ 27103), SVL = 32.49 mm." httpUri-0="https://zenodo.org/record/182998/files/figure.png" httpUri-1="https://zenodo.org/record/182999/files/figure.png" httpUri-2="https://zenodo.org/record/183000/files/figure.png" httpUri-3="https://zenodo.org/record/183001/files/figure.png" pageId="1" pageNumber="50">Figs. 1–4</figureCitation>
) obtained by MRB at La Libertad, Reserva Mazar, (
<geoCoordinate id="EE11CB896229D51F483D7BBF8DEBC0FA" direction="south" orientation="latitude" pageId="1" pageNumber="50" precision="15" value="-2.5463889">S 02° 32' 47''</geoCoordinate>
S,
<geoCoordinate id="EE11CB896229D51F4C6E7BC78CDAC0FA" box="[271,473,981,1007]" direction="west" orientation="longitude" pageId="1" pageNumber="50" precision="1" value="-78.698364">078° 41' 54.1'' W</geoCoordinate>
,
<quantity id="4CDD00AB6229D51F4C857BC78F38C0FA" box="[484,571,981,1007]" metricMagnitude="3" metricUnit="m" metricValue="2.895" pageId="1" pageNumber="50" unit="m" value="2895.0">2895m</quantity>
). Provincia Cañar,
<collectingCountry id="F332EDDE6229D51F4E787BC78E7EC0FA" box="[793,893,981,1007]" name="Ecuador" pageId="1" pageNumber="50">Ecuador</collectingCountry>
, on
<date id="FF9B8B8E6229D51F4ECC7BC7897FC0FA" box="[941,1148,981,1007]" pageId="1" pageNumber="50" value="2005-10-25">October 25, 2005</date>
.
</paragraph>
<paragraph id="8B9AAD4E6229D51F4DA77BEF893AC79A" blockId="1.[151,1437,941,1967]" pageId="1" pageNumber="50">
<emphasis id="B951715C6229D51F4DA77BEF8C81C702" bold="true" box="[198,386,1021,1047]" pageId="1" pageNumber="50">Paratopotypes.</emphasis>
<collectionCode id="ED34358B6229D51F4CED7BEF8CDFC702" box="[396,476,1021,1047]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="1" pageNumber="50">QCAZ</collectionCode>
27104, an adult female collected by MRB on
<date id="FF9B8B8E6229D51F49757BEF89E0C702" box="[1044,1251,1021,1047]" pageId="1" pageNumber="50" value="2005-10-22">22 October 2005</date>
;
<collectionCode id="ED34358B6229D51F49907BEF8842C702" box="[1265,1345,1021,1047]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="1" pageNumber="50">QCAZ</collectionCode>
27568, 27578, 27580–1, adult males, and
<collectionCode id="ED34358B6229D51F4F207C378F92C72A" box="[577,657,1061,1087]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="1" pageNumber="50">QCAZ</collectionCode>
27577, 27579, two adult females, collected by the authors and Michelle R.Cummer on
<date id="FF9B8B8E6229D51F4CD67C5F8FB9C772" box="[439,698,1101,1127]" pageId="1" pageNumber="50" value="2004-02-26" valueMax="2004-02-27" valueMin="2004-02-26">26–27 February 2004</date>
;
<collectionCode id="ED34358B6229D51F4FA67C5F8E14C772" box="[711,791,1101,1127]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="1" pageNumber="50">QCAZ</collectionCode>
29561, an adult male collected by MRB on
<date id="FF9B8B8E6229D51F484B7C5F8DD6C79A" pageId="1" pageNumber="50" value="2005-03-16">16 March 2005</date>
;
<collectionCode id="ED34358B6229D51F4D807C678C32C79A" box="[225,305,1141,1167]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="1" pageNumber="50">QCAZ</collectionCode>
32623, an adult female collected on
<date id="FF9B8B8E6229D51F4F837C678EC7C79A" box="[738,964,1141,1167]" pageId="1" pageNumber="50" value="2006-11-20">20 November 2006</date>
by MRB.
</paragraph>
</subSubSection>
<subSubSection id="C33FFEC56229D51C4DA77C8F8DD5C0EF" lastPageId="2" lastPageNumber="51" pageId="1" pageNumber="50" type="diagnosis">
<paragraph id="8B9AAD4E6229D51F4DA77C8F8CC7C5F2" blockId="1.[151,1437,941,1967]" pageId="1" pageNumber="50">
<emphasis id="B951715C6229D51F4DA77C8F8C45C7A2" bold="true" box="[198,326,1181,1207]" pageId="1" pageNumber="50">Diagnosis.</emphasis>
A member of
<taxonomicName id="4C25D6CD6229D51F4C927C8F8F9FC7A3" box="[499,668,1181,1206]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="50" phylum="Chordata" rank="species" species="unistrigatus">
<emphasis id="B951715C6229D51F4C927C8F8F9FC7A3" box="[499,668,1181,1206]" italics="true" pageId="1" pageNumber="50">P. unistrigatus</emphasis>
</taxonomicName>
(
<emphasis id="B951715C6229D51F4FCF7C8F8FECC7A3" box="[686,751,1181,1206]" italics="true" pageId="1" pageNumber="50">sensu</emphasis>
<bibRefCitation id="EFB4D0BF6229D51F4F967C8F8930C7A2" author="Lynch" box="[759,1075,1181,1207]" pageId="1" pageNumber="58" refString="Lynch, J. D., &amp; Duellman, W. E. (1997) Frogs of the genus Eleutherodactylus in western Ecuador. Systematics, Ecology, and biogeography. Special Publications, The University of Kansas, Natural History Museum, 23, 1 - 236." type="journal article" year="1997">Lynch and Duellman 1997</bibRefCitation>
, as modified by
<bibRefCitation id="EFB4D0BF6229D51F49977C8F8C3BC7CA" author="Duellman" pageId="1" pageNumber="58" refString="Duellman, W. E. &amp; Pramuk, J. B. (1999) Frogs of the genus Eleutherodactylus (Anura: Leptodactylidae) in the Andes of Northern Peru. Scientific Papers Natural History Museum, University of Kansas, 13, 1 - 78." type="journal article" year="1999">Duellman and Pramuk 1999</bibRefCitation>
,
<bibRefCitation id="EFB4D0BF6229D51F4C277CD78F2EC7CA" author="Hedges" box="[326,557,1221,1247]" pageId="1" pageNumber="58" refString="Hedges, S. B., Duellman, W. E. &amp; Heinicke M. P. (2008). New world direct-developing frogs (Anura: Terrarana): Molecular phylogeny, classification, biogeography and conservation. Zootaxa, 1737, 1 - 182." type="journal article" year="2008">
Hedges
<emphasis id="B951715C6229D51F4CC67CD78CDFC7CB" box="[423,476,1221,1246]" italics="true" pageId="1" pageNumber="50">et al</emphasis>
. 2008
</bibRefCitation>
,) diagnosed by having (1) skin texture of dorsum shagreen with scattered tubercles, those on venter areolate; (2) tympanic membrane and annulus evident, 29.7–40% of eye length on females and 25.2–32.3% on males, with supratympanic ridge obscuring upper and posterodorsal edges; (3) snout rounded in profile and dorsal views; (4) upper eyelid bearing one large, and several small scattered, conical tubercles; cranial crests absent; (5) dentigerous process of the vomer triangular and narrowly separated, each bearing 4–6 teeth; (6) vocal slits and vocal sac absent in males, nuptial pads present (
<figureCitation id="131EB1CB6229D51F49D57D9F89FFC6B2" box="[1204,1276,1421,1447]" captionStart="FIGURE 5" captionStartId="6.[151,255,1598,1622]" captionTargetBox="[498,1171,474,1554]" captionTargetId="figure@6.[415,1171,474,1574]" captionTargetPageId="6" captionText="FIGURE 5. Nuptial pad on Finger I of a Pristimantis gagliardoi male (QCAZ 27580)." httpUri="https://zenodo.org/record/183002/files/figure.png" pageId="1" pageNumber="50">Fig. 5</figureCitation>
); (7) first finger shorter than the second, fingers with expanded discs; (8) fingers with lateral fringes; (9) ulnar tubercles small; (10) heel with one enlarged conical tubercle; tarsus with 3–4 tubercles along the outer edge; (11) two metatarsal tubercles prominent, inner oval, about three times the size of the outer subconical tubercle; numerous supernumerary plantar tubercles; (12) toes with lateral fringes; (13) in ethanol, dorsum tan, gray or dark gray, with darker markings (sometimes chevrons or bars) outlined by white or black lines; venter cream to pale cream with darker spots; palmar and plantar surfaces gray, with Fingers I–II, and Toes I–III being the palest; (14) adults medium sized, in males 19.06–24.33 SVL (= 22.17 + 2.01, n = 6), females 26.83–33.64 SVL (= 30.63 + 3.10, n = 5).
</paragraph>
<paragraph id="8B9AAD4E6229D51F4DA47EE78F35C4BA" blockId="1.[151,1437,941,1967]" pageId="1" pageNumber="50">
We assign this species to the
<taxonomicName id="4C25D6CD6229D51F4F497EE78E4EC41B" box="[552,845,1781,1806]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="50" phylum="Chordata" rank="species" species="unistrigatus">
<emphasis id="B951715C6229D51F4F497EE78E4EC41B" box="[552,845,1781,1806]" italics="true" pageId="1" pageNumber="50">Pristimantis unistrigatus</emphasis>
</taxonomicName>
group based on the following characters: narrow head (head width 38.5–42.6% of SVL in females and 38.3–41.9% in males); absence of cranial crests; upper eyelid as wide as, or wider than, IOD (75.1–96.9% in females, 96.0–112.0% in males); skin on venter aerolate; nuptial pads present; tympanic membrane evident; discs on fingers and toes broad; Finger I shorter than Finger II and toes being unwebbed.
</paragraph>
<paragraph id="8B9AAD4E622AD51C4DA778858DD5C0EF" blockId="2.[151,1437,151,1018]" pageId="2" pageNumber="51">
<emphasis id="B951715C622AD51C4DA778858C77C3A4" bold="true" box="[198,372,151,177]" pageId="2" pageNumber="51">Comparisons.</emphasis>
<taxonomicName id="4C25D6CD622AD51C4C1A788A8F8FC3A4" box="[379,652,152,177]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C622AD51C4C1A788A8F8FC3A4" box="[379,652,152,177]" italics="true" pageId="2" pageNumber="51">Pristimantis gagliardoi</emphasis>
</taxonomicName>
may easily be identified by the presence of large, conical tubercles on the eyelids and heels. This species differs from other species in the group that have some degree of development of such tubercles as follows and by the presence/absece of other characters (condition of
<taxonomicName id="4C25D6CD622AD51C486478FA889FC214" box="[1285,1436,232,257]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C622AD51C486478FA889FC214" box="[1285,1436,232,257]" italics="true" pageId="2" pageNumber="51">P. gagliardoi</emphasis>
</taxonomicName>
in parentheses) (
<tableCitation id="C6A798F5622AD51C4C3679028CB3C23F" box="[343,432,272,298]" captionStart="TABLE 1" captionStartId="3.[151,239,593,617]" captionTargetBox="[159,1419,650,751]" captionTargetPageId="3" captionText="TABLE 1. Character states in similar Pristimantis species associated to the unistrigatus group." httpUri="http://table.plazi.org/id/DF5AFDC6622BD51D4DF67A43898FC17D" pageId="2" pageNumber="51" tableUuid="DF5AFDC6622BD51D4DF67A43898FC17D">Table 1</tableCitation>
):
<taxonomicName id="4C25D6CD622AD51C4CAA79028E9BC23F" authority="Lynch 1979" authorityName="Lynch" authorityYear="1979" box="[459,920,272,298]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="cryptomelas">
<emphasis id="B951715C622AD51C4CAA79028FF2C23C" box="[459,753,272,297]" italics="true" pageId="2" pageNumber="51">Pristimantis cryptomelas</emphasis>
(
<bibRefCitation id="EFB4D0BF622AD51C4E6079028E93C23F" author="Lynch" box="[769,912,272,298]" pageId="2" pageNumber="58" refString="Lynch, J. D. (1979) Leptodactylid frogs of the genus Eleutherodactylus from the Andes of southern Ecuador. Miscellaneous Publications. Museum of Natural History, University of Kansas, 66, 1 - 62." type="journal article" year="1979">Lynch 1979</bibRefCitation>
)
</taxonomicName>
is the species most similar to
<taxonomicName id="4C25D6CD622AD51C499D79028896C23C" box="[1276,1429,272,297]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C622AD51C499D79028896C23C" box="[1276,1429,272,297]" italics="true" pageId="2" pageNumber="51">P. gagliardoi</emphasis>
</taxonomicName>
, but differs by having a black coloration on the anterior and posterior sides of thighs, the posterior lower flanks, and the concealed portion of the shank (pale reddish), by having smooth skin (shagreen with scattered distinct tubercles), by lacking a conical tubercle over the eye (present), and by having larger pads on the digits of the hands and feet.
<taxonomicName id="4C25D6CD622AD51C4CF679A28EE0C2DF" authority="Lynch &amp; Duellman, 1980" authorityName="Lynch &amp; Duellman" authorityYear="1980" box="[407,995,432,458]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="inusitatus">
<emphasis id="B951715C622AD51C4CF679A28F9CC2DC" box="[407,671,432,457]" italics="true" pageId="2" pageNumber="51">Pristimantis inusitatus</emphasis>
(
<bibRefCitation id="EFB4D0BF622AD51C4FCC79A28ED8C2DF" author="Lynch" box="[685,987,432,458]" pageId="2" pageNumber="58" refString="Lynch, J. D. &amp; Duellman, W. E. (1980) The Eleutherodactylus of the Amazonian slopes of the Ecuadorian Andes. Miscellaneous Publications, Museum of Natural History, University of Kansas, 69, 1 - 86." type="journal article" year="1980">Lynch &amp; Duellman, 1980</bibRefCitation>
)
</taxonomicName>
differs by having a green dorsum and white venter (dorsum pale brown with variable dark brown markings, venter cream with variable dark brown reticulations or markings), conical tubercle on upper eyelid (present, but larger in
<taxonomicName id="4C25D6CD622AD51C490F7A12880BC10C" box="[1134,1288,512,538]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C622AD51C490F7A12897FC10C" box="[1134,1148,512,537]" italics="true" pageId="2" pageNumber="51">P</emphasis>
.
<emphasis id="B951715C622AD51C49EC7A12880BC10C" box="[1165,1288,512,537]" italics="true" pageId="2" pageNumber="51">gagliardoi</emphasis>
</taxonomicName>
), snout protruding in profile and subcauminate in dorsal view (rounded in both views).
<taxonomicName id="4C25D6CD622AD51C494F7A3A8C63C17F" authority="Lynch &amp; Duellman, 1980" authorityName="Lynch &amp; Duellman" authorityYear="1980" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="eriphus">
<emphasis id="B951715C622AD51C494F7A3A881CC154" box="[1070,1311,552,577]" italics="true" pageId="2" pageNumber="51">Pristimantis eriphus</emphasis>
(
<bibRefCitation id="EFB4D0BF622AD51C48517A3A8C5BC17F" author="Lynch" pageId="2" pageNumber="58" refString="Lynch, J. D. &amp; Duellman, W. E. (1980) The Eleutherodactylus of the Amazonian slopes of the Ecuadorian Andes. Miscellaneous Publications, Museum of Natural History, University of Kansas, 69, 1 - 86." type="journal article" year="1980">Lynch &amp; Duellman, 1980</bibRefCitation>
)
</taxonomicName>
has a straight canthus rostralis (weakly angular), adult females have red iris (bronze) and flanks with white areas and slightly oblique white bars on posterior flanks and hidden surfaces of hind limbs (bars on flanks).
<taxonomicName id="4C25D6CD622AD51C4C3E7AB28E9CC1AF" authority="Lynch &amp; Duellman, 1980" authorityName="Lynch &amp; Duellman" authorityYear="1980" box="[351,927,672,698]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="prolatus">
<emphasis id="B951715C622AD51C4C3E7AB28F54C1AC" box="[351,599,672,697]" italics="true" pageId="2" pageNumber="51">Pristimantis prolatus</emphasis>
(
<bibRefCitation id="EFB4D0BF622AD51C4F077AB28E94C1AF" author="Lynch" box="[614,919,672,698]" pageId="2" pageNumber="58" refString="Lynch, J. D. &amp; Duellman, W. E. (1980) The Eleutherodactylus of the Amazonian slopes of the Ecuadorian Andes. Miscellaneous Publications, Museum of Natural History, University of Kansas, 69, 1 - 86." type="journal article" year="1980">Lynch &amp; Duellman, 1980</bibRefCitation>
)
</taxonomicName>
has no ulnar tubercles (few conical), vocal slits and vocal sac present in males (absent), Iris is gray brown (bronze), snout in profile nearly truncate and accuimnate in dorsal view (rounded in both views).
<taxonomicName id="4C25D6CD622AD51C4E627AE2889FC01F" authority="Duellman and Pramuk, 1999" authorityName="Duellman and Pramuk" authorityYear="1999" box="[771,1436,752,778]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="nephophilus">
<emphasis id="B951715C622AD51C4E627AE28929C01C" box="[771,1066,752,777]" italics="true" pageId="2" pageNumber="51">Pristimantis nephophilus</emphasis>
(
<bibRefCitation id="EFB4D0BF622AD51C495A7AE28897C01F" author="Duellman" box="[1083,1428,752,778]" pageId="2" pageNumber="58" refString="Duellman, W. E. &amp; Pramuk, J. B. (1999) Frogs of the genus Eleutherodactylus (Anura: Leptodactylidae) in the Andes of Northern Peru. Scientific Papers Natural History Museum, University of Kansas, 13, 1 - 78." type="journal article" year="1999">Duellman and Pramuk, 1999</bibRefCitation>
)
</taxonomicName>
has a few, small tubercles on the posterior eyelid (small tubercles scattered over surface of eyelid, and a single, large, central, conical tubercle), dark brown coloration with cream spots on the posterior thighs (pale reddish), brown coloration on the venter (cream with variable markings), all tubercles on the feet and hands weakly developed (strongly developed), smoother skin, lacking pronounced tubercles (shagreen with scattered distinct tubercles), and discs on the hind digits being slightly smaller than those of fingers (equal in size).
</paragraph>
</subSubSection>
<caption id="DF5AFDC6622AD51C4DF67EEE8F09C42E" httpUri="https://zenodo.org/record/182998/files/figure.png" pageId="2" pageNumber="51" targetBox="[151,1435,1077,1761]" targetPageId="2">
<paragraph id="8B9AAD4E622AD51C4DF67EEE8F09C42E" blockId="2.[151,1436,1788,1851]" pageId="2" pageNumber="51">
<emphasis id="B951715C622AD51C4DF67EEE8C16C401" bold="true" box="[151,277,1788,1812]" pageId="2" pageNumber="51">FIGURE 1</emphasis>
. Dorsal (left) and ventral (right) views of
<taxonomicName id="4C25D6CD622AD51C4F8E7EEF8EE8C401" box="[751,1003,1789,1812]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C622AD51C4F8E7EEF8EE8C401" box="[751,1003,1789,1812]" italics="true" pageId="2" pageNumber="51">Pristimantis gagliardoi</emphasis>
</taxonomicName>
in preservative; holotype, adult female (QCAZ 27103), SVL = 32.49 mm.
</paragraph>
</caption>
<subSubSection id="C33FFEC5622AD5194DA77F6389C6C2B7" lastPageId="7" lastPageNumber="56" pageId="2" pageNumber="51" type="description">
<paragraph id="8B9AAD4E622AD51D4DA77F638CD7C10F" blockId="2.[151,1436,1905,2012]" lastBlockId="3.[151,1437,152,538]" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51">
<emphasis id="B951715C622AD51C4DA77F638F2AC49E" bold="true" box="[198,553,1905,1931]" pageId="2" pageNumber="51">
Description of the
<typeStatus id="549E13EC622AD51C4CD67F638F27C49E" box="[439,548,1905,1931]" pageId="2" pageNumber="51" type="holotype">holotype</typeStatus>
.
</emphasis>
Head slightly wider than long, head length 85% of head width (
<figureCitation id="131EB1CB622AD51C48207F60888AC499" box="[1345,1417,1906,1932]" captionStart="FIGURE 1" captionStartId="2.[151,255,1788,1812]" captionTargetBox="[151,1435,1077,1761]" captionTargetId="figure@2.[151,1436,1077,1762]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal (left) and ventral (right) views of Pristimantis gagliardoi in preservative; holotype, adult female (QCAZ 27103), SVL = 32.49 mm." httpUri="https://zenodo.org/record/182998/files/figure.png" pageId="2" pageNumber="51">Fig. 1</figureCitation>
); snout relatively short (snout to eye distance 15.1% of SVL), rounded in dorsal view and in profile (
<figureCitation id="131EB1CB622AD51C487D7F88888EC4A1" box="[1308,1421,1946,1972]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,255,1788,1812]" captionStartId-1="3.[151,255,2009,2033]" captionTargetBox-0="[151,1435,1077,1761]" captionTargetBox-1="[226,1360,1154,1985]" captionTargetId-0="figure@2.[151,1436,1077,1762]" captionTargetId-1="figure@3.[226,1360,1154,1985]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Dorsal (left) and ventral (right) views of Pristimantis gagliardoi in preservative; holotype, adult female (QCAZ 27103), SVL = 32.49 mm." captionText-1="FIGURE 2. Lateral view of the head holotype of Pristimantis gagliardoi, holotype, adult female (QCAZ 27103)." httpUri-0="https://zenodo.org/record/182998/files/figure.png" httpUri-1="https://zenodo.org/record/182999/files/figure.png" pageId="2" pageNumber="51">Figs. 1–2</figureCitation>
); eye to nostril 92.7% length of eye; nostrils not protuberant, directed anterolaterally; canthus rostralis weakly angular; loreal region concave; lips not flared; interorbital area flat, broader than upper eyelid (upper eyelid width 85.9% of interorbital distance); upper eyelid with distinct conical fleshy tubercle situated centrally on outer margin of eyelid; cranial crests absent; supratympanic fold weak, but partially obscuring upper margin of tympanic annulus and tympanum; side of head nearly vertical; tympanic annulus distinct; tympanic membrane weakly pustular, not thickened; length of tympanic annulus 40% length of eye; postrictal tubercles large, subconical, situated ventral and posteroventral to tympanic annulus; skin on head finely granular; choanae very small, oval, not concealed by palatal shelf of maxillary; vomerine odontophores oblique, posteromedial to choanae, oval in outline, about 3–
<date id="FF9B8B8E622BD51D4F1879A28F8BC2DF" box="[633,648,432,458]" pageId="3" pageNumber="52">4</date>
x size of choana, separated medially by distance less than width of odontophore, each bearing 4 teeth; tongue wider than long, its posterior border bilobbed, posterior half not adherent to floor of mouth.
</paragraph>
<caption id="DF5AFDC6622BD51D4DF67A43898FC17D" ID-Table-UUID="DF5AFDC6622BD51D4DF67A43898FC17D" box="[151,1164,593,617]" httpUri="http://table.plazi.org/id/DF5AFDC6622BD51D4DF67A43898FC17D" pageId="3" pageNumber="52" targetBox="[159,1419,650,751]" targetIsTable="true" targetPageId="3">
<paragraph id="8B9AAD4E622BD51D4DF67A43898FC17D" blockId="3.[151,1164,593,617]" box="[151,1164,593,617]" pageId="3" pageNumber="52">
<emphasis id="B951715C622BD51D4DF67A438C0BC17C" bold="true" box="[151,264,593,617]" pageId="3" pageNumber="52">TABLE 1.</emphasis>
Character states in similar
<taxonomicName id="4C25D6CD622BD51D4F4C7A408FACC17C" box="[557,687,594,617]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="52" phylum="Chordata" rank="genus">
<emphasis id="B951715C622BD51D4F4C7A408FACC17C" box="[557,687,594,617]" italics="true" pageId="3" pageNumber="52">Pristimantis</emphasis>
</taxonomicName>
species associated to the
<taxonomicName id="4C25D6CD622BD51D4EA17A40893CC17C" box="[960,1087,594,617]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="unistrigatus">
<emphasis id="B951715C622BD51D4EA17A40893CC17C" box="[960,1087,594,617]" italics="true" pageId="3" pageNumber="52">unistrigatus</emphasis>
</taxonomicName>
group.
</paragraph>
</caption>
<paragraph id="8B9AAD4E622BD51D4DFE7A988856C1FA" pageId="3" pageNumber="52">
<table id="F9255FEE622B2AE14DFE7A988888C1FA" box="[159,1419,650,751]" gridcols="5" gridrows="3" pageId="3" pageNumber="52">
<tr id="3515AF0C622B2AE14DFE7A988888C18A" box="[159,1419,650,671]" gridrow="0" pageId="3" pageNumber="52">
<th id="76C4C670622B2AE14DFE7A988C1FC18A" box="[159,284,650,671]" gridcol="0" gridrow="0" pageId="3" pageNumber="52">Character</th>
<th id="76C4C670622B2AE14CE57A988F96C18A" box="[388,661,650,671]" gridcol="1" gridrow="0" pageId="3" pageNumber="52">
<emphasis id="B951715C622BD51D4CE57A988F97C18A" box="[388,660,650,671]" italics="true" pageId="3" pageNumber="52">
<taxonomicName id="4C25D6CD622BD51D4CE57A988F12C18A" box="[388,529,650,671]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="cryptomelas">P. cryptomelas</taxonomicName>
<taxonomicName id="4C25D6CD622BD51D4F557A988F97C18A" box="[564,660,650,671]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="eriphus">P. eriphus</taxonomicName>
</emphasis>
</th>
<th id="76C4C670622B2AE14F857A988E5AC18A" box="[740,857,650,671]" gridcol="2" gridrow="0" pageId="3" pageNumber="52">
<taxonomicName id="4C25D6CD622BD51D4F857A988E5AC18A" box="[740,857,650,671]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="inusitatus">
<emphasis id="B951715C622BD51D4F857A988E5AC18A" box="[740,857,650,671]" italics="true" pageId="3" pageNumber="52">P. inusitatus</emphasis>
</taxonomicName>
</th>
<th id="76C4C670622B2AE14EF57A9889A8C18A" box="[916,1195,650,671]" gridcol="3" gridrow="0" pageId="3" pageNumber="52">
<emphasis id="B951715C622BD51D4EF57A9889A8C18A" box="[916,1195,650,671]" italics="true" pageId="3" pageNumber="52">
<taxonomicName id="4C25D6CD622BD51D4EF57A988922C18A" box="[916,1057,650,671]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="nephophilus">P. nephophilus</taxonomicName>
<taxonomicName id="4C25D6CD622BD51D49257A9889A8C18A" box="[1092,1195,650,671]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="prolatus">P. prolatus</taxonomicName>
</emphasis>
</th>
<th id="76C4C670622B2AE149947A988888C18A" box="[1269,1419,650,671]" gridcol="4" gridrow="0" pageId="3" pageNumber="52">
<taxonomicName id="4C25D6CD622BD51D49947A988873C18A" box="[1269,1392,650,671]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C622BD51D49947A988873C18A" box="[1269,1392,650,671]" italics="true" pageId="3" pageNumber="52">P. gagliardoi</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3515AF0C622B2AE14DFE7AA08888C1D2" box="[159,1419,690,711]" gridrow="1" pageId="3" pageNumber="52">
<th id="76C4C670622B2AE14DFE7AA08C1FC1D2" box="[159,284,690,711]" gridcol="0" gridrow="1" pageId="3" pageNumber="52">SVL males</th>
<td id="76C4C670622B2AE14CE57AA08F96C1D2" box="[388,661,690,711]" gridcol="1" gridrow="1" pageId="3" pageNumber="52">28.2–30.2 18.1–25.2</td>
<td id="76C4C670622B2AE14F857AA08E5AC1D2" box="[740,857,690,711]" gridcol="2" gridrow="1" pageId="3" pageNumber="52">-</td>
<td id="76C4C670622B2AE14EF57AA089A8C1D2" box="[916,1195,690,711]" gridcol="3" gridrow="1" pageId="3" pageNumber="52">- 13.7–18.4</td>
<td id="76C4C670622B2AE149947AA08888C1D2" box="[1269,1419,690,711]" gridcol="4" gridrow="1" pageId="3" pageNumber="52">19.1–24.3</td>
</tr>
<tr id="3515AF0C622B2AE14DFE7AC88888C1FA" box="[159,1419,730,751]" gridrow="2" pageId="3" pageNumber="52">
<th id="76C4C670622B2AE14DFE7AC88C1FC1FA" box="[159,284,730,751]" gridcol="0" gridrow="2" pageId="3" pageNumber="52">SVL females</th>
<td id="76C4C670622B2AE14CE57AC88F96C1FA" box="[388,661,730,751]" gridcol="1" gridrow="2" pageId="3" pageNumber="52">38.6 25.8–29.0</td>
<td id="76C4C670622B2AE14F857AC88E5AC1FA" box="[740,857,730,751]" gridcol="2" gridrow="2" pageId="3" pageNumber="52">24.0–24.4</td>
<td id="76C4C670622B2AE14EF57AC889A8C1FA" box="[916,1195,730,751]" gridcol="3" gridrow="2" pageId="3" pageNumber="52">24.6–34.0 20.8–24.1</td>
<td id="76C4C670622B2AE149947AC88888C1FA" box="[1269,1419,730,751]" gridcol="4" gridrow="2" pageId="3" pageNumber="52">26.8–33.6</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9AAD4E622BD51D4DFE7B108888C002" blockId="3.[159,1419,770,860]" box="[159,1419,770,791]" pageId="3" pageNumber="52">Canthus rostralis Rounded Straight (weakl Sharp Weakly angular Moderately Weakly angular</paragraph>
<paragraph id="8B9AAD4E622BD51D4F557B0D897BC021" blockId="3.[159,1419,770,860]" box="[564,1144,799,820]" pageId="3" pageNumber="52">concave) (barely concave) sharp</paragraph>
<paragraph id="8B9AAD4E622BD51A4DFE7B558F6AC10F" blockId="3.[159,1419,770,860]" lastBlockId="4.[151,1437,152,538]" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52">
Snout in profile Rounded Round Protruding Rounded Nearly truncate Rounded Dorsum of head, body, and limbs with scattered subconical tubercles becoming larger and more abundant on lateral surfaces; dermal ridge from posteromedial edge of eyelid to scapular region; dorsolateral folds absent; flanks granular; cloacal sheath absent; enlarged tubercles in cloacal region present, moderately developed; skin on throat granular; skin on venter areolate; discoidal fold not evident; forearm slender; radio-ulna length 25% of SVL; ulnar tubercles few, round to subconical; thenar tubercle ovoid, about
<date id="FF9B8B8E622CD51A4985792A8802C244" box="[1252,1281,310,338]" pageId="4" pageNumber="53">3x</date>
size of bifid subpalmar tubercle; supernumerary tubercles few, large subconical; subarticular prominent, subconical; hand length longer than radioulna length 33.1% of SVL; Fingers II–IV bearing narrow lateral fringes; relative lengths of fingers I &lt;II &lt;IV &lt;III; disc on thumb slightly expanded; discs on Fingers II–IV extensively expanded, truncate, less than twice width of digits; Finger I having ventral pad defined by circumferential groove, absent on other fingers (
<figureCitation id="131EB1CB622CD51A4F737A128F5AC10F" box="[530,601,512,538]" captionStart="FIGURE 3" captionStartId="4.[151,255,1752,1776]" captionTargetBox="[226,1360,594,1728]" captionTargetId="figure@4.[226,1360,594,1728]" captionTargetPageId="4" captionText="FIGURE 3. (A) Right hand of Pristimantis gagliardoi; (B) right foot of Pristimantis gagliardoi. Holotype, adult female (QCAZ 27103)." httpUri="https://zenodo.org/record/183000/files/figure.png" pageId="4" pageNumber="53">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8B9AAD4E622BD51D4DFE7B7D886AC71A" pageId="3" pageNumber="52">
<table id="F9255FEE622B2AE14DFE7B7D8888C739" box="[159,1419,879,1068]" gridcols="6" gridrows="3" pageId="3" pageNumber="52">
<tr id="3515AF0C622B2AE14DFE7B7D8888C091" box="[159,1419,879,900]" gridrow="0" pageId="3" pageNumber="52">
<th id="76C4C670622B2AE14DFE7B7D8C67C091" box="[159,356,879,900]" gridcol="0" gridrow="0" pageId="3" pageNumber="52">Snout in dorsal view</th>
<th id="76C4C670622B2AE14CE57B7D8F11C091" box="[388,530,879,900]" gridcol="1" gridrow="0" pageId="3" pageNumber="52">Sub acuminate</th>
<th id="76C4C670622B2AE14F557B7D8FCEC091" box="[564,717,879,900]" gridcol="2" gridrow="0" pageId="3" pageNumber="52">Round</th>
<th id="76C4C670622B2AE14F857B7D8E7BC091" box="[740,888,879,900]" gridcol="3" gridrow="0" pageId="3" pageNumber="52">Sub acuminate</th>
<th id="76C4C670622B2AE14EF57B7D892FC091" box="[916,1068,879,900]" gridcol="4" gridrow="0" pageId="3" pageNumber="52">Rounded</th>
<th id="76C4C670622B2AE149257B7D8888C091" box="[1092,1419,879,900]" gridcol="5" gridrow="0" pageId="3" pageNumber="52">Acuminate Rounded</th>
</tr>
<tr id="3515AF0C622B2AE14DFE7B858888C0F2" box="[159,1419,919,999]" gridrow="1" pageId="3" pageNumber="52">
<th id="76C4C670622B2AE14DFE7B858C67C0F2" box="[159,356,919,999]" gridcol="0" gridrow="1" pageId="3" pageNumber="52">Eyelid</th>
<td id="76C4C670622B2AE14CE57B858F11C0F2" box="[388,530,919,999]" gridcol="1" gridrow="1" pageId="3" pageNumber="52">Bearing tuber- cles</td>
<td id="76C4C670622B2AE14F557B858FCEC0F2" box="[564,717,919,999]" gridcol="2" gridrow="1" pageId="3" pageNumber="52">Bearing one conical tubercle</td>
<td id="76C4C670622B2AE14F857B858E7BC0F2" box="[740,888,919,999]" gridcol="3" gridrow="1" pageId="3" pageNumber="52">Bearing a coni- cal tubercle</td>
<td id="76C4C670622B2AE14EF57B85892FC0F2" box="[916,1068,919,999]" gridcol="4" gridrow="1" pageId="3" pageNumber="52">With small tubercles poste- riorly</td>
<td id="76C4C670622B2AE149257B858888C0F2" box="[1092,1419,919,999]" gridcol="5" gridrow="1" pageId="3" pageNumber="52">Bearing one Distinct conipungent tubercle cal, fleshy</td>
</tr>
<tr id="3515AF0C622B2AE14DFE7BE88888C739" box="[159,1419,1018,1068]" gridrow="2" pageId="3" pageNumber="52">
<th id="76C4C670622B2AE14DFE7BE88C67C739" box="[159,356,1018,1068]" gridcol="0" gridrow="2" pageId="3" pageNumber="52">Ulnar tubercles</th>
<td id="76C4C670622B2AE14CE57BE88F11C739" box="[388,530,1018,1068]" gridcol="1" gridrow="2" pageId="3" pageNumber="52">Prominent</td>
<td id="76C4C670622B2AE14F557BE88FCEC739" box="[564,717,1018,1068]" gridcol="2" gridrow="2" pageId="3" pageNumber="52">Conical to sub- conical</td>
<td id="76C4C670622B2AE14F857BE88E7BC739" box="[740,888,1018,1068]" gridcol="3" gridrow="2" pageId="3" pageNumber="52">Conical</td>
<td id="76C4C670622B2AE14EF57BE8892FC739" box="[916,1068,1018,1068]" gridcol="4" gridrow="2" pageId="3" pageNumber="52">Few, round to subconical</td>
<td id="76C4C670622B2AE149257BE88888C739" box="[1092,1419,1018,1068]" gridcol="5" gridrow="2" pageId="3" pageNumber="52">Not present Few conical</td>
</tr>
</table>
</paragraph>
<caption id="DF5AFDC6622BD51D4DF67FCB8852C4E4" box="[151,1361,2009,2033]" httpUri="https://zenodo.org/record/182999/files/figure.png" pageId="3" pageNumber="52" targetBox="[226,1360,1154,1985]" targetPageId="3">
<paragraph id="8B9AAD4E622BD51D4DF67FCB8852C4E4" blockId="3.[151,1361,2009,2033]" box="[151,1361,2009,2033]" pageId="3" pageNumber="52">
<emphasis id="B951715C622BD51D4DF67FCB8C11C4E4" bold="true" box="[151,274,2009,2033]" pageId="3" pageNumber="52">FIGURE 2</emphasis>
. Lateral view of the head holotype of
<taxonomicName id="4C25D6CD622BD51D4FCB7FC88EA0C4E4" box="[682,931,2010,2033]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C622BD51D4FCB7FC88EA0C4E4" box="[682,931,2010,2033]" italics="true" pageId="3" pageNumber="52">Pristimantis gagliardoi</emphasis>
</taxonomicName>
, holotype, adult female (QCAZ 27103).
</paragraph>
</caption>
<caption id="DF5AFDC6622CD51A4DF67ECA8C41C407" httpUri="https://zenodo.org/record/183000/files/figure.png" pageId="4" pageNumber="53" targetBox="[226,1360,594,1728]" targetPageId="4">
<paragraph id="8B9AAD4E622CD51A4DF67ECA8C41C407" blockId="4.[151,1436,1752,1810]" pageId="4" pageNumber="53">
<emphasis id="B951715C622CD51A4DF67ECA8C11C5E5" bold="true" box="[151,274,1752,1776]" pageId="4" pageNumber="53">FIGURE 3</emphasis>
. (A) Right hand of
<taxonomicName id="4C25D6CD622CD51A4C877ECB8FDCC5E5" box="[486,735,1753,1776]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C622CD51A4C877ECB8FDCC5E5" box="[486,735,1753,1776]" italics="true" pageId="4" pageNumber="53">Pristimantis gagliardoi</emphasis>
</taxonomicName>
; (B) right foot of
<taxonomicName id="4C25D6CD622CD51A4EC17ECB899BC5E5" box="[928,1176,1753,1776]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C622CD51A4EC17ECB899BC5E5" box="[928,1176,1753,1776]" italics="true" pageId="4" pageNumber="53">Pristimantis gagliardoi</emphasis>
</taxonomicName>
. Holotype, adult female (QCAZ 27103).
</paragraph>
</caption>
<paragraph id="8B9AAD4E622CD51B4DA47F548FAFC217" blockId="4.[151,1437,1862,2008]" lastBlockId="5.[151,1437,152,618]" lastPageId="5" lastPageNumber="54" pageId="4" pageNumber="53">
<collectingCountry id="F332EDDE622CD51A4DA47F548C03C475" box="[197,256,1862,1888]" name="India" pageId="4" pageNumber="53">Hind</collectingCountry>
limbs relatively slender; tibia length 52.9% of SVL; foot length 95.3% of tibia length; heel bearing small, subconical tubercle; outer edge of tarsus bearing three large subconical tubercles; inner metatarsal tubercle flat, ovoid, about
<date id="FF9B8B8E622CD51A4CB97F848CF7C4BA" box="[472,500,1940,1968]" pageId="4" pageNumber="53">3x</date>
subconical outer metatarsal tubercle; supernumerary plantar tubercles prominent, round or subconical; subarticular tubercles round, not conical; toes bearing narrow lateral fringes, most developed on Toes IV–V; relative length of toes I &lt;II &lt;III &lt;V &lt;IV; webbing absent; discs on toes equal in size to discs on fingers; tip of Toe V extending to distal edge of distal subarticular tubercle on Toe IV; tip of Toe III not extending to that tubercle (
<figureCitation id="131EB1CB622DD51B4F3478FA8F9FC217" box="[597,668,232,258]" captionStart="FIGURE 3" captionStartId="4.[151,255,1752,1776]" captionTargetBox="[226,1360,594,1728]" captionTargetId="figure@4.[226,1360,594,1728]" captionTargetPageId="4" captionText="FIGURE 3. (A) Right hand of Pristimantis gagliardoi; (B) right foot of Pristimantis gagliardoi. Holotype, adult female (QCAZ 27103)." httpUri="https://zenodo.org/record/183000/files/figure.png" pageId="5" pageNumber="54">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8B9AAD4E622DD51B4DA4791D8F13C17F" blockId="5.[151,1437,152,618]" pageId="5" pageNumber="54">
<emphasis id="B951715C622DD51B4DA4791D8CEAC23C" bold="true" box="[197,489,271,297]" pageId="5" pageNumber="54">
Coloration of
<typeStatus id="549E13EC622DD51B4C17791D8CE7C23C" box="[374,484,271,297]" pageId="5" pageNumber="54" type="holotype">holotype</typeStatus>
.
</emphasis>
In life: Dorsum tan with brown bars outlined by thin black lines, suprascapular region with a dark brown W mark corresponding to suprascapular dermal ridges; dark brown interorbital bar; brown cloacal region. Dorsal surfaces of limbs tan with brown and dark brown bars. Dorsum of hands and feet with black blotches, more abundant towards distal portions of fingers and toes. Flanks tan with black blotches. Axial region and groin pink with brown blotches. Hidden parts of venter and forelimbs pink with brown blotches. Ventral surfaces of tarsus and outer toes dark brown. Yellow belly with dark brown spots, yellowish throat with brown blotches. Two dark brown blotches in ventral scapular region. Venter of hands dark brown, inner fingers (I and II) pale brown; black bars originating below eye and extending towards the lip. Iris bronze with black reticulations (
<figureCitation id="131EB1CB622DD51B4CDB7A428F00C17F" box="[442,515,592,618]" captionStart="FIGURE 4" captionStartId="5.[151,255,1832,1856]" captionTargetBox="[415,1171,674,1808]" captionTargetId="figure@5.[415,1171,674,1808]" captionTargetPageId="5" captionText="FIGURE 4. Dorsolateral (A) and ventral (B) views of Pristimantis gagliardoi in live; holotype, adult female (QCAZ 27103), SVL = 32.49 mm." httpUri="https://zenodo.org/record/183001/files/figure.png" pageId="5" pageNumber="54">Fig. 4</figureCitation>
).
</paragraph>
<caption id="DF5AFDC6622DD51B4DF67F3A8CB1C477" httpUri="https://zenodo.org/record/183001/files/figure.png" pageId="5" pageNumber="54" targetBox="[415,1171,674,1808]" targetPageId="5">
<paragraph id="8B9AAD4E622DD51B4DF67F3A8CB1C477" blockId="5.[151,1435,1832,1890]" pageId="5" pageNumber="54">
<emphasis id="B951715C622DD51B4DF67F3A8C17C455" bold="true" box="[151,276,1832,1856]" pageId="5" pageNumber="54">FIGURE 4</emphasis>
. Dorsolateral (A) and ventral (B) views of
<taxonomicName id="4C25D6CD622DD51B4F957F3B8EEDC455" box="[756,1006,1833,1856]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="54" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C622DD51B4F957F3B8EEDC455" box="[756,1006,1833,1856]" italics="true" pageId="5" pageNumber="54">Pristimantis gagliardoi</emphasis>
</taxonomicName>
in live; holotype, adult female (QCAZ 27103), SVL = 32.49 mm.
</paragraph>
</caption>
<paragraph id="8B9AAD4E622DD5184DA47F848C6AC2B7" blockId="5.[151,1436,1940,2008]" lastBlockId="6.[151,1436,152,418]" lastPageId="6" lastPageNumber="55" pageId="5" pageNumber="54">
In preservative: Dorsum gray with darker interorbital bar and a W over scapular region; dorsal surfaces of proximal forelimbs dull cream, distal forelimbs dull gray with darker markings; dorsal surfaces of Fingers I and II pale cream, Fingers III and IV pale gray with dark gray markings and paler discs; dorsal surfaces of thighs pinkish gray with darker blotches; shanks gray with darker bars, tarsus gray with darker markings, Toes IV and V same color, Toes I–III cream. Dorsum of tympanum with black markings; gray loreal, canthal, and upper labial regions with black markings. Venter tan with dark gray spots, pale cream throat with gray markings; venter of proximal forelimbs pinkish becoming darker on ulna and outer fingers, inner fingers cream; ventral hindlimbs pinkish with brown markings, on tarsus and Toes IV and V becoming dark grey, and Toes I– III cream (
<figureCitation id="131EB1CB622ED5184C72799A8C5BC2B7" box="[275,344,392,418]" captionStart="FIGURE 1" captionStartId="2.[151,255,1788,1812]" captionTargetBox="[151,1435,1077,1761]" captionTargetId="figure@2.[151,1436,1077,1762]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal (left) and ventral (right) views of Pristimantis gagliardoi in preservative; holotype, adult female (QCAZ 27103), SVL = 32.49 mm." httpUri="https://zenodo.org/record/182998/files/figure.png" pageId="6" pageNumber="55">Fig. 1</figureCitation>
).
</paragraph>
<caption id="DF5AFDC6622ED5184DF67E2C8936C540" box="[151,1077,1598,1622]" httpUri="https://zenodo.org/record/183002/files/figure.png" pageId="6" pageNumber="55" targetBox="[498,1171,474,1554]" targetPageId="6">
<paragraph id="8B9AAD4E622ED5184DF67E2C8936C540" blockId="6.[151,1077,1598,1622]" box="[151,1077,1598,1622]" pageId="6" pageNumber="55">
<emphasis id="B951715C622ED5184DF67E2C8C11C543" bold="true" box="[151,274,1598,1622]" pageId="6" pageNumber="55">FIGURE 5</emphasis>
. Nuptial pad on Finger I of a
<taxonomicName id="4C25D6CD622ED5184F317E2D8E4AC543" box="[592,841,1599,1622]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C622ED5184F317E2D8E4AC543" box="[592,841,1599,1622]" italics="true" pageId="6" pageNumber="55">Pristimantis gagliardoi</emphasis>
</taxonomicName>
male (QCAZ 27580).
</paragraph>
</caption>
<paragraph id="8B9AAD4E622ED5184DA77E948831C40C" blockId="6.[151,1436,1670,2017]" pageId="6" pageNumber="55">
<emphasis id="B951715C622ED5184DA77E948F14C5B5" bold="true" box="[198,535,1670,1696]" pageId="6" pageNumber="55">
Measurements of
<typeStatus id="549E13EC622ED5184CC57E948F11C5B5" box="[420,530,1670,1696]" pageId="6" pageNumber="55" type="holotype">holotype</typeStatus>
.
</emphasis>
SVL = 32.49; tibia length = 17.19; foot length = 16.38; head length = 10.84; head width = 12.75; interorbital distance = 4.03; upper eyelid width = 3.46; internarial distance = 2.79; eye to nostril distance = 3.57; snout to eye distance = 5.06; eye diameter = 3.85; tympanum diameter = 1.54; eye to tympanum distance = 1.40; radioulna length = 8.12; hand length = 10.78; and Finger I length = 5.13.
</paragraph>
<paragraph id="8B9AAD4E622ED5194DA47F3489C6C2B7" blockId="6.[151,1436,1670,2017]" lastBlockId="7.[151,1436,152,418]" lastPageId="7" lastPageNumber="56" pageId="6" pageNumber="55">
<emphasis id="B951715C622ED5184DA47F348C3CC455" bold="true" box="[197,319,1830,1856]" pageId="6" pageNumber="55">Variation.</emphasis>
Variation in measurements and proportions among additional specimens are presented in
<tableCitation id="C6A798F5622ED5184DF67F5D8DFBC47C" box="[151,248,1871,1897]" captionStart="TABLE 2" captionStartId="7.[151,239,471,495]" captionTargetBox="[159,1355,562,1271]" captionTargetPageId="7" captionText="TABLE 2. Measurements (in mm) of adults of Pristimantis gagliardoi. Range, mean SD are given. SVL = snout – vent length." httpUri="http://table.plazi.org/id/DF5AFDC6622FD5194DF679C58DE2C104" pageId="6" pageNumber="55" tableUuid="DF5AFDC6622FD5194DF679C58DE2C104">Tables 2</tableCitation>
and
<tableCitation id="C6A798F5622ED5184C517F5D8C42C47C" box="[304,321,1871,1897]" captionStart="TABLE 3" captionStartId="7.[151,239,1351,1375]" captionTargetBox="[159,1232,1442,1935]" captionTargetPageId="7" captionText="TABLE 3. Ranges of morphological proportions (in percentages) of adults of Pristimantis gagliardoi. SVL = snout – vent length." httpUri="http://table.plazi.org/id/DF5AFDC6622FD5194DF67D558DE2C694" pageId="6" pageNumber="55" tableUuid="DF5AFDC6622FD5194DF67D558DE2C694">3</tableCitation>
. Variation of coloration in life: QCAZ 27570: Dorsum with cream and pale brown bars outlined by thin black lines; flanks cream with bars originating on the dorsum; dorsum of limbs showing the same pattern as the body; groin and hidden surfaces of thighs colorless; venter cream; iris bronze with a brown medial stripe; upper lip with vertical brown and cream bars; QCAZ 27104: Dorsum tan with brown blotches outlined by thin black lines; flanks brown with diagonal bars becoming greenish toward the dorsum; dorsum of forelimbs with brown and greenish bars; tarsus tan with brown bars; dorsum of arms tan with brown blotches; dorsum of hands and feet with black blotches present on outer fingers; groin, armpit, and hidden parts of thighs pale pink with brown blotching; throat tan with brown blotching; venter of limbs tan with dark brown blotching with outer fingers and toes darker; black bars originating below eye and extending towards the lip. Iris bronze with black reticulations; QCAZ 27434: Same pattern as QCAZ 27104 but having diagonal green bars extending from the dorsum onto the hidden surfaces of thighs; groin and armpit are orange.
</paragraph>
</subSubSection>
<caption id="DF5AFDC6622FD5194DF679C58DE2C104" ID-Table-UUID="DF5AFDC6622FD5194DF679C58DE2C104" httpUri="http://table.plazi.org/id/DF5AFDC6622FD5194DF679C58DE2C104" pageId="7" pageNumber="56" targetBox="[159,1355,562,1271]" targetIsTable="true" targetPageId="7">
<paragraph id="8B9AAD4E622FD5194DF679C58DE2C104" blockId="7.[151,1436,471,529]" pageId="7" pageNumber="56">
<emphasis id="B951715C622FD5194DF679C58C09C2FA" bold="true" box="[151,266,471,495]" pageId="7" pageNumber="56">TABLE 2.</emphasis>
Measurements (in mm) of adults of
<taxonomicName id="4C25D6CD622FD5194FFA79CA8E96C2FA" box="[667,917,472,495]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="56" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C622FD5194FFA79CA8E96C2FA" box="[667,917,472,495]" italics="true" pageId="7" pageNumber="56">Pristimantis gagliardoi</emphasis>
</taxonomicName>
. Range, mean SD are given. SVL = snout–vent length.
</paragraph>
</caption>
<paragraph id="8B9AAD4E622FD5194DFE7A208821C7E2" pageId="7" pageNumber="56">
<table id="F9255FEE622F2AE14DFE7A208848C7E2" box="[159,1355,562,1271]" gridcols="3" gridrows="17" pageId="7" pageNumber="56">
<tr id="3515AF0C622F2AE14DFE7A208848C15C" box="[159,1355,562,585]" gridrow="0" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7A208CB9C15C" box="[159,442,562,585]" gridcol="0" gridrow="0" pageId="7" pageNumber="56">Character</th>
<th id="76C4C670622F2AE14FC97A208ECFC15C" box="[680,972,562,585]" gridcol="1" gridrow="0" pageId="7" pageNumber="56">
females
<emphasis id="B951715C622FD5194E607A208E0DC15C" box="[769,782,562,585]" italics="true" pageId="7" pageNumber="56">n</emphasis>
= 5
</th>
<th id="76C4C670622F2AE149477A208848C15C" box="[1062,1355,562,585]" gridcol="2" gridrow="0" pageId="7" pageNumber="56">
males
<emphasis id="B951715C622FD519490A7A20897BC15C" box="[1131,1144,562,585]" italics="true" pageId="7" pageNumber="56">n</emphasis>
= 5
</th>
</tr>
<tr id="3515AF0C622F2AE14DFE7A4F8848C161" box="[159,1355,605,628]" gridrow="1" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7A4F8CB9C161" box="[159,442,605,628]" gridcol="0" gridrow="1" pageId="7" pageNumber="56">SVL</th>
<td id="76C4C670622F2AE14FC97A4F8ECFC161" box="[680,972,605,628]" gridcol="1" gridrow="1" pageId="7" pageNumber="56">26.83–33.64 (30.63 ± 3.10)</td>
<td id="76C4C670622F2AE149477A4F8848C161" box="[1062,1355,605,628]" gridcol="2" gridrow="1" pageId="7" pageNumber="56">19.06–24.33 (22.17 ± 2.01)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7A9A8848C18A" box="[159,1355,648,671]" gridrow="2" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7A9A8CB9C18A" box="[159,442,648,671]" gridcol="0" gridrow="2" pageId="7" pageNumber="56">Tibia length</th>
<td id="76C4C670622F2AE14FC97A9A8ECFC18A" box="[680,972,648,671]" gridcol="1" gridrow="2" pageId="7" pageNumber="56">15.23–18.17 (16.68 ± 1.29)</td>
<td id="76C4C670622F2AE149477A9A8848C18A" box="[1062,1355,648,671]" gridcol="2" gridrow="2" pageId="7" pageNumber="56">10.98–14.97 (12.70 ± 1.50)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7AA08848C1DC" box="[159,1355,690,713]" gridrow="3" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7AA08CB9C1DC" box="[159,442,690,713]" gridcol="0" gridrow="3" pageId="7" pageNumber="56">Foot length</th>
<td id="76C4C670622F2AE14FC97AA08ECFC1DC" box="[680,972,690,713]" gridcol="1" gridrow="3" pageId="7" pageNumber="56">13.21–17.05 (15.23 ± 1.86)</td>
<td id="76C4C670622F2AE149477AA08848C1DC" box="[1062,1355,690,713]" gridcol="2" gridrow="3" pageId="7" pageNumber="56">9.22–13.04 (11.16 ± 1.46)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7ACF8848C1E1" box="[159,1355,733,756]" gridrow="4" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7ACF8CB9C1E1" box="[159,442,733,756]" gridcol="0" gridrow="4" pageId="7" pageNumber="56">Head length</th>
<td id="76C4C670622F2AE14FC97ACF8ECFC1E1" box="[680,972,733,756]" gridcol="1" gridrow="4" pageId="7" pageNumber="56">9.82–11.43 (10.70 ± 0.60)</td>
<td id="76C4C670622F2AE149477ACF8848C1E1" box="[1062,1355,733,756]" gridcol="2" gridrow="4" pageId="7" pageNumber="56">6.76–9.05 (7.87 ± 0.85)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7B1A8848C00A" box="[159,1355,776,799]" gridrow="5" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7B1A8CB9C00A" box="[159,442,776,799]" gridcol="0" gridrow="5" pageId="7" pageNumber="56">Head width</th>
<td id="76C4C670622F2AE14FC97B1A8ECFC00A" box="[680,972,776,799]" gridcol="1" gridrow="5" pageId="7" pageNumber="56">10.71–13.34 (12.22 ± 1.00)</td>
<td id="76C4C670622F2AE149477B1A8848C00A" box="[1062,1355,776,799]" gridcol="2" gridrow="5" pageId="7" pageNumber="56">7.73–10.19 (9.02 ± 0.94)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7B208848C05C" box="[159,1355,818,841]" gridrow="6" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7B208CB9C05C" box="[159,442,818,841]" gridcol="0" gridrow="6" pageId="7" pageNumber="56">Interorbital distance</th>
<td id="76C4C670622F2AE14FC97B208ECFC05C" box="[680,972,818,841]" gridcol="1" gridrow="6" pageId="7" pageNumber="56">3.20–4.03 (3.51 ± 0.33)</td>
<td id="76C4C670622F2AE149477B208848C05C" box="[1062,1355,818,841]" gridcol="2" gridrow="6" pageId="7" pageNumber="56">2.18–2.85 (2.60 ± 0.28)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7B4F8848C061" box="[159,1355,861,884]" gridrow="7" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7B4F8CB9C061" box="[159,442,861,884]" gridcol="0" gridrow="7" pageId="7" pageNumber="56">Upper eyelid width</th>
<td id="76C4C670622F2AE14FC97B4F8ECFC061" box="[680,972,861,884]" gridcol="1" gridrow="7" pageId="7" pageNumber="56">2.46–3.46 (3.25 ± 0.39)</td>
<td id="76C4C670622F2AE149477B4F8848C061" box="[1062,1355,861,884]" gridcol="2" gridrow="7" pageId="7" pageNumber="56">2.32–2.83 (2.65 ± 0.22)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7B9A8848C0B7" box="[159,1355,904,930]" gridrow="8" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7B9A8CB9C0B7" box="[159,442,904,930]" gridcol="0" gridrow="8" pageId="7" pageNumber="56">Radioulna length</th>
<td id="76C4C670622F2AE14FC97B9A8ECFC0B7" box="[680,972,904,930]" gridcol="1" gridrow="8" pageId="7" pageNumber="56">2.58–2.91 (2.81 ± 0.14)</td>
<td id="76C4C670622F2AE149477B9A8848C0B7" box="[1062,1355,904,930]" gridcol="2" gridrow="8" pageId="7" pageNumber="56">1.83–2.64 (2.23 ± 0.29)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7BA78848C0D9" box="[159,1355,949,972]" gridrow="9" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7BA78CB9C0D9" box="[159,442,949,972]" gridcol="0" gridrow="9" pageId="7" pageNumber="56">Eye to nostril distance</th>
<td id="76C4C670622F2AE14FC97BA78ECFC0D9" box="[680,972,949,972]" gridcol="1" gridrow="9" pageId="7" pageNumber="56">2.43–3.58 (3.23 ± 0.34)</td>
<td id="76C4C670622F2AE149477BA78848C0D9" box="[1062,1355,949,972]" gridcol="2" gridrow="9" pageId="7" pageNumber="56">2.01–2.83 (2.33 ± 0.31)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7BF28848C0E2" box="[159,1355,992,1015]" gridrow="10" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7BF28CB9C0E2" box="[159,442,992,1015]" gridcol="0" gridrow="10" pageId="7" pageNumber="56">Snout to eye distance</th>
<td id="76C4C670622F2AE14FC97BF28ECFC0E2" box="[680,972,992,1015]" gridcol="1" gridrow="10" pageId="7" pageNumber="56">4.52–5.75 (5.10±0.51)</td>
<td id="76C4C670622F2AE149477BF28848C0E2" box="[1062,1355,992,1015]" gridcol="2" gridrow="10" pageId="7" pageNumber="56">3.31–4.55 (3.85 ± 0.45)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7C188848C734" box="[159,1355,1034,1057]" gridrow="11" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7C188CB9C734" box="[159,442,1034,1057]" gridcol="0" gridrow="11" pageId="7" pageNumber="56">Eye diameter</th>
<td id="76C4C670622F2AE14FC97C188ECFC734" box="[680,972,1034,1057]" gridcol="1" gridrow="11" pageId="7" pageNumber="56">3.45–4.12 (3.78 ± 0.27)</td>
<td id="76C4C670622F2AE149477C188848C734" box="[1062,1355,1034,1057]" gridcol="2" gridrow="11" pageId="7" pageNumber="56">2.40–3.00 (2.83 ± 0.24)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7C278848C759" box="[159,1355,1077,1100]" gridrow="12" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7C278CB9C759" box="[159,442,1077,1100]" gridcol="0" gridrow="12" pageId="7" pageNumber="56">Tympanum diameter</th>
<td id="76C4C670622F2AE14FC97C278ECFC759" box="[680,972,1077,1100]" gridcol="1" gridrow="12" pageId="7" pageNumber="56">1.04–1.54 (1.25 ± 0.19)</td>
<td id="76C4C670622F2AE149477C278848C759" box="[1062,1355,1077,1100]" gridcol="2" gridrow="12" pageId="7" pageNumber="56">0.70–0.80 (0.76 ± 0.03)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7C728848C762" box="[159,1355,1120,1143]" gridrow="13" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7C728CB9C762" box="[159,442,1120,1143]" gridcol="0" gridrow="13" pageId="7" pageNumber="56">Eye to tympanum distance</th>
<td id="76C4C670622F2AE14FC97C728ECFC762" box="[680,972,1120,1143]" gridcol="1" gridrow="13" pageId="7" pageNumber="56">1.32–1.75 (1.51± 0.17)</td>
<td id="76C4C670622F2AE149477C728848C762" box="[1062,1355,1120,1143]" gridcol="2" gridrow="13" pageId="7" pageNumber="56">1.10–1.35 (1.18 ± 0.10)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7C988848C7B4" box="[159,1355,1162,1185]" gridrow="14" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7C988CB9C7B4" box="[159,442,1162,1185]" gridcol="0" gridrow="14" pageId="7" pageNumber="56">Radio-ulna length</th>
<td id="76C4C670622F2AE14FC97C988ECFC7B4" box="[680,972,1162,1185]" gridcol="1" gridrow="14" pageId="7" pageNumber="56">6.92–8.50 (7.83 ± 0.65)</td>
<td id="76C4C670622F2AE149477C988848C7B4" box="[1062,1355,1162,1185]" gridcol="2" gridrow="14" pageId="7" pageNumber="56">5.26–6.94 (5.85 ± 0.66)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7CA78848C7D9" box="[159,1355,1205,1228]" gridrow="15" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7CA78CB9C7D9" box="[159,442,1205,1228]" gridcol="0" gridrow="15" pageId="7" pageNumber="56">Hand length</th>
<td id="76C4C670622F2AE14FC97CA78ECFC7D9" box="[680,972,1205,1228]" gridcol="1" gridrow="15" pageId="7" pageNumber="56">8.43–10.78 (9.38 ± 1.00)</td>
<td id="76C4C670622F2AE149477CA78848C7D9" box="[1062,1355,1205,1228]" gridcol="2" gridrow="15" pageId="7" pageNumber="56">5.92–7.48 (6.56 ± 0.68)</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7CF28848C7E2" box="[159,1355,1248,1271]" gridrow="16" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7CF28CB9C7E2" box="[159,442,1248,1271]" gridcol="0" gridrow="16" pageId="7" pageNumber="56">Finger I length</th>
<td id="76C4C670622F2AE14FC97CF28ECFC7E2" box="[680,972,1248,1271]" gridcol="1" gridrow="16" pageId="7" pageNumber="56">3.95–5.13 (4.73 ± 0.54)</td>
<td id="76C4C670622F2AE149477CF28848C7E2" box="[1062,1355,1248,1271]" gridcol="2" gridrow="16" pageId="7" pageNumber="56">2.91–3.80 (3.27 ± 0.34)</td>
</tr>
</table>
</paragraph>
<caption id="DF5AFDC6622FD5194DF67D558DE2C694" ID-Table-UUID="DF5AFDC6622FD5194DF67D558DE2C694" httpUri="http://table.plazi.org/id/DF5AFDC6622FD5194DF67D558DE2C694" pageId="7" pageNumber="56" targetBox="[159,1232,1442,1935]" targetIsTable="true" targetPageId="7">
<paragraph id="8B9AAD4E622FD5194DF67D558DE2C694" blockId="7.[151,1436,1351,1409]" pageId="7" pageNumber="56">
<emphasis id="B951715C622FD5194DF67D558C0BC64A" bold="true" box="[151,264,1351,1375]" pageId="7" pageNumber="56">TABLE 3.</emphasis>
Ranges of morphological proportions (in percentages) of adults of
<emphasis id="B951715C622FD5194EB57D5A89D0C64A" box="[980,1235,1352,1375]" italics="true" pageId="7" pageNumber="56">
<taxonomicName id="4C25D6CD622FD5194EB57D5A89CCC64A" box="[980,1231,1352,1375]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="56" phylum="Chordata" rank="species" species="gagliardoi">Pristimantis gagliardoi</taxonomicName>
.
</emphasis>
SVL = snout–vent length.
</paragraph>
</caption>
<paragraph id="8B9AAD4E622FD5194DFE7DB089B6C49A" pageId="7" pageNumber="56">
<table id="F9255FEE622F2AE14DFE7DB089D3C49A" box="[159,1232,1442,1935]" gridcols="3" gridrows="12" pageId="7" pageNumber="56">
<tr id="3515AF0C622F2AE14DFE7DB089D3C6AC" box="[159,1232,1442,1465]" gridrow="0" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7DB08F28C6AC" box="[159,555,1442,1465]" gridcol="0" gridrow="0" pageId="7" pageNumber="56">Character</th>
<th id="76C4C670622F2AE14F8C7DB08E8CC6AC" box="[749,911,1442,1465]" gridcol="1" gridrow="0" pageId="7" pageNumber="56">females (n = 5)</th>
<th id="76C4C670622F2AE1492B7DB089D3C6AC" box="[1098,1232,1442,1465]" gridcol="2" gridrow="0" pageId="7" pageNumber="56">males (n =5)</th>
</tr>
<tr id="3515AF0C622F2AE14DFE7DDF89D3C6F1" box="[159,1232,1485,1508]" gridrow="1" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7DDF8F28C6F1" box="[159,555,1485,1508]" gridcol="0" gridrow="1" pageId="7" pageNumber="56">Tibia length/ SVL</th>
<td id="76C4C670622F2AE14F8C7DDF8E8CC6F1" box="[749,911,1485,1508]" gridcol="1" gridrow="1" pageId="7" pageNumber="56">51.7–57.5</td>
<td id="76C4C670622F2AE1492B7DDF89D3C6F1" box="[1098,1232,1485,1508]" gridcol="2" gridrow="1" pageId="7" pageNumber="56">53.8–61.5</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7DEA89D3C51A" box="[159,1232,1528,1551]" gridrow="2" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7DEA8F28C51A" box="[159,555,1528,1551]" gridcol="0" gridrow="2" pageId="7" pageNumber="56">Foot length/SVL</th>
<td id="76C4C670622F2AE14F8C7DEA8E8CC51A" box="[749,911,1528,1551]" gridcol="1" gridrow="2" pageId="7" pageNumber="56">47.5–50.4</td>
<td id="76C4C670622F2AE1492B7DEA89D3C51A" box="[1098,1232,1528,1551]" gridcol="2" gridrow="2" pageId="7" pageNumber="56">47.7–51.4</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7E3089D3C52C" box="[159,1232,1570,1593]" gridrow="3" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7E308F28C52C" box="[159,555,1570,1593]" gridcol="0" gridrow="3" pageId="7" pageNumber="56">Foot length/tibia length</th>
<td id="76C4C670622F2AE14F8C7E308E8CC52C" box="[749,911,1570,1593]" gridcol="1" gridrow="3" pageId="7" pageNumber="56">85.6–95.3</td>
<td id="76C4C670622F2AE1492B7E3089D3C52C" box="[1098,1232,1570,1593]" gridcol="2" gridrow="3" pageId="7" pageNumber="56">84.0–92.5</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7E5F89D3C571" box="[159,1232,1613,1636]" gridrow="4" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7E5F8F28C571" box="[159,555,1613,1636]" gridcol="0" gridrow="4" pageId="7" pageNumber="56">Head width/SVL</th>
<td id="76C4C670622F2AE14F8C7E5F8E8CC571" box="[749,911,1613,1636]" gridcol="1" gridrow="4" pageId="7" pageNumber="56">38.5–42.6</td>
<td id="76C4C670622F2AE1492B7E5F89D3C571" box="[1098,1232,1613,1636]" gridcol="2" gridrow="4" pageId="7" pageNumber="56">38.3–41.9</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7E6A89D3C59A" box="[159,1232,1656,1679]" gridrow="5" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7E6A8F28C59A" box="[159,555,1656,1679]" gridcol="0" gridrow="5" pageId="7" pageNumber="56">Head length /SVL</th>
<td id="76C4C670622F2AE14F8C7E6A8E8CC59A" box="[749,911,1656,1679]" gridcol="1" gridrow="5" pageId="7" pageNumber="56">33.4–37.8</td>
<td id="76C4C670622F2AE1492B7E6A89D3C59A" box="[1098,1232,1656,1679]" gridcol="2" gridrow="5" pageId="7" pageNumber="56">34.6–37.2</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7EB089D3C5AC" box="[159,1232,1698,1721]" gridrow="6" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7EB08F28C5AC" box="[159,555,1698,1721]" gridcol="0" gridrow="6" pageId="7" pageNumber="56">Head length/ head width</th>
<td id="76C4C670622F2AE14F8C7EB08E8CC5AC" box="[749,911,1698,1721]" gridcol="1" gridrow="6" pageId="7" pageNumber="56">85.0–91.7</td>
<td id="76C4C670622F2AE1492B7EB089D3C5AC" box="[1098,1232,1698,1721]" gridcol="2" gridrow="6" pageId="7" pageNumber="56">84.0–92.1</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7EDF89D3C5F1" box="[159,1232,1741,1764]" gridrow="7" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7EDF8F28C5F1" box="[159,555,1741,1764]" gridcol="0" gridrow="7" pageId="7" pageNumber="56">Eye–to–nostril distance/eye diameter</th>
<td id="76C4C670622F2AE14F8C7EDF8E8CC5F1" box="[749,911,1741,1764]" gridcol="1" gridrow="7" pageId="7" pageNumber="56">79.5–92.7</td>
<td id="76C4C670622F2AE1492B7EDF89D3C5F1" box="[1098,1232,1741,1764]" gridcol="2" gridrow="7" pageId="7" pageNumber="56">72.0–97.9</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7EEA89D3C41A" box="[159,1232,1784,1807]" gridrow="8" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7EEA8F28C41A" box="[159,555,1784,1807]" gridcol="0" gridrow="8" pageId="7" pageNumber="56">Upper eyelid width/IOD</th>
<td id="76C4C670622F2AE14F8C7EEA8E8CC41A" box="[749,911,1784,1807]" gridcol="1" gridrow="8" pageId="7" pageNumber="56">75.1–96.9</td>
<td id="76C4C670622F2AE1492B7EEA89D3C41A" box="[1098,1232,1784,1807]" gridcol="2" gridrow="8" pageId="7" pageNumber="56">96.0–112.0</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7F3089D3C42C" box="[159,1232,1826,1849]" gridrow="9" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7F308F28C42C" box="[159,555,1826,1849]" gridcol="0" gridrow="9" pageId="7" pageNumber="56">Radioulna length/SVL</th>
<td id="76C4C670622F2AE14F8C7F308E8CC42C" box="[749,911,1826,1849]" gridcol="1" gridrow="9" pageId="7" pageNumber="56">24.4–26.6</td>
<td id="76C4C670622F2AE1492B7F3089D3C42C" box="[1098,1232,1826,1849]" gridcol="2" gridrow="9" pageId="7" pageNumber="56">25.1–28.5</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7F5F89D3C471" box="[159,1232,1869,1892]" gridrow="10" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7F5F8F28C471" box="[159,555,1869,1892]" gridcol="0" gridrow="10" pageId="7" pageNumber="56">Hand length/radioulna length</th>
<td id="76C4C670622F2AE14F8C7F5F8E8CC471" box="[749,911,1869,1892]" gridcol="1" gridrow="10" pageId="7" pageNumber="56">103.2–132.7</td>
<td id="76C4C670622F2AE1492B7F5F89D3C471" box="[1098,1232,1869,1892]" gridcol="2" gridrow="10" pageId="7" pageNumber="56">101.5–125.9</td>
</tr>
<tr id="3515AF0C622F2AE14DFE7F6A89D3C49A" box="[159,1232,1912,1935]" gridrow="11" pageId="7" pageNumber="56">
<th id="76C4C670622F2AE14DFE7F6A8F28C49A" box="[159,555,1912,1935]" gridcol="0" gridrow="11" pageId="7" pageNumber="56">Finger I Length/hand length</th>
<td id="76C4C670622F2AE14F8C7F6A8E8CC49A" box="[749,911,1912,1935]" gridcol="1" gridrow="11" pageId="7" pageNumber="56">46.9–58.2</td>
<td id="76C4C670622F2AE1492B7F6A89D3C49A" box="[1098,1232,1912,1935]" gridcol="2" gridrow="11" pageId="7" pageNumber="56">41.8–66.0</td>
</tr>
</table>
</paragraph>
<subSubSection id="C33FFEC56220D5164DA478858973C23F" pageId="8" pageNumber="57" type="etymology">
<paragraph id="8B9AAD4E6220D5164DA478858973C23F" blockId="8.[151,1437,151,778]" pageId="8" pageNumber="57">
<emphasis id="B951715C6220D5164DA478858C51C3A4" bold="true" box="[197,338,151,177]" pageId="8" pageNumber="57">Etymology.</emphasis>
The specific name is a noun in the genitive case and is a patronym for Ron Gagliardo, dedicated collaborator in both research and conservation programs involving Neotropical amphibians. Gagliardo has been important in promoting amphibian research, environmental education, but especially for supporting the establishment of captive breeding programs for the threatened Ecuadorian frogs.
</paragraph>
</subSubSection>
<subSubSection id="C33FFEC56220D5164DA479258F9BC01F" pageId="8" pageNumber="57" type="distribution">
<paragraph id="8B9AAD4E6220D5164DA479258F9BC01F" blockId="8.[151,1437,151,778]" pageId="8" pageNumber="57">
<emphasis id="B951715C6220D5164DA479258CFCC244" bold="true" box="[197,511,311,337]" pageId="8" pageNumber="57">Distribution and ecology.</emphasis>
<taxonomicName id="4C25D6CD6220D5164F67792A8E1BC244" box="[518,792,312,337]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C6220D5164F67792A8E1BC244" box="[518,792,312,337]" italics="true" pageId="8" pageNumber="57">Pristimantis gagliardoi</emphasis>
</taxonomicName>
is known only from La Libertad, Reserva Mazar (
<geoCoordinate id="EE11CB896220D5164806792A8DB4C26F" direction="south" orientation="latitude" pageId="8" pageNumber="57" precision="15" value="-0.042222224">S 0 2 32</geoCoordinate>
' 47'' S, 0 78 41' 54.1'' W,
<quantity id="4CDD00AB6220D5164CBA79728F33C26F" box="[475,560,352,378]" metricMagnitude="3" metricUnit="m" metricValue="2.895" pageId="8" pageNumber="57" unit="m" value="2895.0">2895m</quantity>
), Provincia Cañar,
<collectingCountry id="F332EDDE6220D5164E7179728E77C26F" box="[784,884,352,378]" name="Ecuador" pageId="8" pageNumber="57">Ecuador</collectingCountry>
. This site, a small farm, is located on the western versant of the Ecuadorian Andes in Bosque de neblina montano (Montane cloud forest) and Bosque siempreverde montano alto (Evergreen montane forest) according to
<bibRefCitation id="EFB4D0BF6220D5164EF879A28998C2DF" box="[921,1179,432,458]" pageId="8" pageNumber="58" refString="Valencia, R., Cern, C., Palacios, W. &amp; Sierra, R. (1999) Las formaciones naturales de la Sierra del Ecuador. In Sierra, R. (Ed.) Propuesta Preliminar de Clasificacin de Vegetacin para el Ecuador Continental. Proyecto INEFAN / GEF - BIRF and Ecociencia, Quito, Ecuador, pp. 79 - 108." type="book chapter">
Valencia
<emphasis id="B951715C6220D516496679A2893EC2DC" box="[1031,1085,432,457]" italics="true" pageId="8" pageNumber="57">et al</emphasis>
. (1999)
</bibRefCitation>
(
<figureCitation id="131EB1CB6220D51649CF79A289FBC2DF" box="[1198,1272,432,458]" captionStart="FIGURE 6" captionStartId="8.[151,255,1718,1742]" captionTargetBox="[257,1292,834,1694]" captionTargetId="figure@8.[217,1369,834,1694]" captionTargetPageId="8" captionText="FIGURE 6. Map of Ecuador showing the type locality of Pristimantis gagliardoi." httpUri="https://zenodo.org/record/183003/files/figure.png" pageId="8" pageNumber="57">Fig. 6</figureCitation>
). The species has been found on leaves and branches at heights varying from
<quantity id="4CDD00AB6220D5164EFA79CA8932C2E7" box="[923,1073,472,498]" metricMagnitude="-1" metricUnit="m" metricValue="9.5" metricValueMax="15.0" metricValueMin="4.0" pageId="8" pageNumber="57" unit="m" value="0.95" valueMax="1.5" valueMin="0.4">0.40–1.50 m</quantity>
above ground (=
<quantity id="4CDD00AB6220D516484979CA888DC2E7" box="[1320,1422,472,498]" metricMagnitude="-1" metricUnit="m" metricValue="7.75" pageId="8" pageNumber="57" unit="m" value="0.775">0.775 m</quantity>
), mainly in secondary forests (12 of 16 individuals), and on vegetation over very small stream (2 of 16) and in primary forest (1 of 16). During our visits (six field trips since
<date id="FF9B8B8E6220D5164EE87A3A8934C157" box="[905,1079,552,578]" pageId="8" pageNumber="57" value="2004-02">February 2004</date>
) to La Libertad,
<taxonomicName id="4C25D6CD6220D51648627A3A889FC154" box="[1283,1436,552,577]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C6220D51648627A3A889FC154" box="[1283,1436,552,577]" italics="true" pageId="8" pageNumber="57">P. gagliardoi</emphasis>
</taxonomicName>
showed seasonal activity. During the wettest months we found it in primary and secondary forest as well as along small streams. In drier months (October and November) we found it along streams. An adult female (QCAZ 32623) obtained on
<date id="FF9B8B8E6220D5164C827AB28F85C1AF" box="[483,646,672,698]" pageId="8" pageNumber="57" value="1920-11">20 November</date>
, 2006, contained mature eggs in the oviducts.
<taxonomicName id="4C25D6CD6220D51649C47AB28DBEC1F4" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="gagliardoi">
<emphasis id="B951715C6220D51649C47AB28DBEC1F4" italics="true" pageId="8" pageNumber="57">Pristimantis gagliardoi</emphasis>
</taxonomicName>
occurs sympatrically with
<taxonomicName id="4C25D6CD6220D5164C9D7ADA8FF6C1F4" box="[508,757,712,737]" class="Amphibia" family="Hemiphractidae" genus="Gastrotheca" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="pseustes">
<emphasis id="B951715C6220D5164C9D7ADA8FF6C1F4" box="[508,757,712,737]" italics="true" pageId="8" pageNumber="57">Gastrotheca pseustes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C25D6CD6220D5164E647ADA8900C1F4" box="[773,1027,712,737]" class="Amphibia" family="Dendrobatidae" genus="Hyloxalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="B951715C6220D5164E647ADA8900C1F4" box="[773,1027,712,737]" italics="true" pageId="8" pageNumber="57">Hyloxalus vertebralis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C25D6CD6220D51649737ADA8982C1F4" box="[1042,1153,712,738]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="orestes">
<emphasis id="B951715C6220D51649737ADA8922C1F4" box="[1042,1057,712,737]" italics="true" pageId="8" pageNumber="57">P</emphasis>
.
<emphasis id="B951715C6220D516494E7ADA8982C1F4" box="[1071,1153,712,737]" italics="true" pageId="8" pageNumber="57">orestes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C25D6CD6220D51649F07ADA8846C1F4" box="[1169,1349,712,738]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="pycnodermis">
<emphasis id="B951715C6220D51649F07ADA89A3C1F4" box="[1169,1184,712,737]" italics="true" pageId="8" pageNumber="57">P</emphasis>
.
<emphasis id="B951715C6220D51649CE7ADA8846C1F4" box="[1199,1349,712,737]" italics="true" pageId="8" pageNumber="57">pycnodermis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C25D6CD6220D51648357ADA8DB7C01C" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="species" species="riveti">
<emphasis id="B951715C6220D51648357ADA8860C1F4" box="[1364,1379,712,737]" italics="true" pageId="8" pageNumber="57">P</emphasis>
.
<emphasis id="B951715C6220D51648137ADA8DB7C01C" italics="true" pageId="8" pageNumber="57">riveti</emphasis>
</taxonomicName>
and an undescribed
<taxonomicName id="4C25D6CD6220D5164CC47AE28F37C01C" box="[421,564,752,777]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="57" phylum="Chordata" rank="genus">
<emphasis id="B951715C6220D5164CC47AE28F37C01C" box="[421,564,752,777]" italics="true" pageId="8" pageNumber="57">Pristimantis</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>
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