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<mods:title id="9344C4451525DF27E31EA2944E0AFFCF">The taxonomy of Indian gorgonians: an assessment of the descriptive records of gorgonians (Anthozoa: Octocorallia: Alcyonacea) recorded as occurring in the territorial waters of India, along with neighbouring regions and the Andaman and Nicobar Islands, and the highlighting of perceived unethical practice</mods:title>
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<mods:namePart id="27145CEF2F982B8C82B2AACE0BA39E00">Ramvilas, Ghosh</mods:namePart>
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<mods:namePart id="3A98E992EAD2B9E53631451227D6432C">Alderslade, Philip</mods:namePart>
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<treatment id="0388B6417B72FFC8FF56FCE7FBF8F930" ID-DOI="http://doi.org/10.5281/zenodo.7640997" ID-GBIF-Taxon="209290939" ID-Zenodo-Dep="7640997" LSID="urn:lsid:plazi:treatment:0388B6417B72FFC8FF56FCE7FBF8F930" httpUri="http://treatment.plazi.org/id/0388B6417B72FFC8FF56FCE7FBF8F930" lastPageNumber="101" pageId="100" pageNumber="101">
<subSubSection id="C33B54DC7B72FFC8FF56FCE7FDA7FCF7" box="[151,601,798,825]" pageId="100" pageNumber="101" type="nomenclature">
<paragraph id="8B9E07577B72FFC8FF56FCE7FDA7FCF7" blockId="100.[151,601,798,825]" box="[151,601,798,825]" pageId="100" pageNumber="101">
<heading id="D0D6B03B7B72FFC8FF56FCE7FDA7FCF7" bold="true" box="[151,601,798,825]" fontSize="11" level="2" pageId="100" pageNumber="101" reason="4">
<taxonomicName id="4C217CD47B72FFC8FF56FCE7FDA7FCF7" ID-CoL="5B8KB" authority="Grassoff, 2000" authorityName="Grasshoff" authorityYear="2000" box="[151,601,798,825]" class="Anthozoa" family="Ellisellidae" genus="Verrucella" kingdom="Animalia" order="Alcyonacea" pageId="100" pageNumber="101" phylum="Cnidaria" rank="species" species="gubalensis">
<emphasis id="B955DB457B72FFC8FF56FCE7FDA7FCF7" bold="true" box="[151,601,798,825]" pageId="100" pageNumber="101">
<emphasis id="B955DB457B72FFC8FF56FCE7FE63FCF6" bold="true" box="[151,413,798,825]" italics="true" pageId="100" pageNumber="101">Verrucella gubalensis</emphasis>
Grassoff, 2000
</emphasis>
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<subSubSection id="C33B54DC7B72FFC8FF56FC9AFAA0FC28" pageId="100" pageNumber="101" type="reference_group">
<paragraph id="8B9E07577B72FFC8FF56FC9AFC7AFCB4" blockId="100.[151,900,867,891]" box="[151,900,867,891]" pageId="100" pageNumber="101">
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<treatmentCitation id="0A8021467B72FFC8FF56FC9AFDAFFCB4" author="Grasshoff, M." box="[151,593,867,891]" page="114" pageId="100" pageNumber="101" year="2000">
<taxonomicName id="4C217CD47B72FFC8FF56FC9AFDAFFCB4" ID-CoL="5B8KB" authority="Grasshoff, 2000: 114" authorityName="Grasshoff" authorityPageNumber="114" authorityYear="2000" box="[151,593,867,891]" class="Anthozoa" family="Ellisellidae" genus="Verrucella" kingdom="Animalia" order="Alcyonacea" pageId="100" pageNumber="101" phylum="Cnidaria" rank="species" species="gubalensis">
<emphasis id="B955DB457B72FFC8FF56FC9AFE8DFCB4" box="[151,371,867,891]" italics="true" pageId="100" pageNumber="101">Verrucella gubalensis</emphasis>
<bibRefCitation id="EFB07AA67B72FFC8FEBBFC9AFDAFFCB4" author="Grasshoff, M." box="[378,593,867,891]" pageId="100" pageNumber="101" pagination="1 - 125" refId="ref101466" refString="Grasshoff, M. (2000) The gorgonians of the Sinai coast and the Strait of Gubal, Red Sea (Coelenterata, Octocorallia). Courier Forschungsinstitut Senckenberg, 224, 1 - 125." type="journal article" year="2000">Grasshoff, 2000: 114</bibRefCitation>
</taxonomicName>
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, fig. 204&amp;205 (
<collectingRegion id="49E5C9B57B72FFC8FD30FC9AFCDFFCB4" box="[753,801,867,891]" country="Papua New Guinea" name="Gulf" pageId="100" pageNumber="101">Gulf</collectingRegion>
of
<collectingRegion id="49E5C9B57B72FFC8FC82FC9AFC86FCB4" box="[835,888,867,891]" country="Egypt" name="As Suways" pageId="100" pageNumber="101">Suez</collectingRegion>
).
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<paragraph id="8B9E07577B72FFC8FF56FC50FAA0FC28" blockId="100.[151,1436,937,1000]" pageId="100" pageNumber="101">
Opinion: There is not enough evidence that this species occurs in the region, and the Indian material is probably the same as at least one of the other
<taxonomicName id="4C217CD47B72FFC8FDC2FC34FD2EFC28" ID-CoL="84VMY" authority="Milne Edwards &amp; Haime, 1857" box="[515,720,973,999]" family="Ellisellidae" kingdom="Animalia" order="Alcyonacea" pageId="100" pageNumber="101" rank="genus">
<emphasis id="B955DB457B72FFC8FDC2FC34FD86FC28" box="[515,632,973,999]" italics="true" pageId="100" pageNumber="101">Verrucella</emphasis>
species
</taxonomicName>
described by
<bibRefCitation id="EFB07AA67B72FFC8FCADFC34FBD8FC28" author="Fernando, S. A." box="[876,1062,973,999]" pageId="100" pageNumber="101" refId="ref100707" refString="Fernando, S. A. (2011) Monograph on Gorgonids (Sea fans) of India. Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Parangipettai, 145 pp." type="book" year="2011">Fernando (2011)</bibRefCitation>
and
<bibRefCitation id="EFB07AA67B72FFC8FB9CFC34FAA4FC28" author="Fernando, S. &amp; Venkataraman, K. &amp; Raghunathan, C." box="[1117,1370,973,1000]" pageId="100" pageNumber="101" refId="ref100744" refString="Fernando, S. A, Venkataraman, K. &amp; Raghunathan, C. (2017) Gorgonians of Indian sea. Director, Zoological Survey of India, Kolkata, 330 pp. Available from: https: // www. nhbs. com / (accessed 21 March 2022)" type="book" year="2017">
Fernando
<emphasis id="B955DB457B72FFC8FB0CFC37FB01FC28" box="[1229,1279,973,999]" italics="true" pageId="100" pageNumber="101">et al</emphasis>
. (2017)
</bibRefCitation>
.
</paragraph>
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<subSubSection id="C33B54DC7B72FFC8FF56FBECFBF8F930" pageId="100" pageNumber="101" type="description">
<paragraph id="8B9E07577B72FFC8FF56FBECFED7FBE0" blockId="100.[151,1436,1045,1252]" box="[151,297,1045,1071]" pageId="100" pageNumber="101">Justification:</paragraph>
<paragraph id="8B9E07577B72FFC8FF06FBC0FEEDFB54" blockId="100.[151,1436,1045,1252]" pageId="100" pageNumber="101">
<emphasis id="B955DB457B72FFC8FF06FBC0FC57FB9C" box="[199,937,1081,1107]" italics="true" pageId="100" pageNumber="101">These Indian records seem to be either invalid or unconfirmable</emphasis>
:
<bibRefCitation id="EFB07AA67B72FFC8FC7BFBC0FB26FB9C" author="Fernando, S. A." box="[954,1240,1081,1107]" pageId="100" pageNumber="101" refId="ref100707" refString="Fernando, S. A. (2011) Monograph on Gorgonids (Sea fans) of India. Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Parangipettai, 145 pp." type="book" year="2011">Fernando 2011: 118119</bibRefCitation>
, pl. 78, fig. 11c (SE coast);
<bibRefCitation id="EFB07AA67B72FFC8FEDCFBA7FDD6FBB8" author="Kumar, J. S. Y. &amp; Raghunathan, C. &amp; Raghuraman, R. &amp; Sreeraj, C. R. &amp; Venkataraman, K." box="[285,552,1117,1144]" pageId="100" pageNumber="101" refId="ref103651" refString="Kumar, J. S. Y., Raghunathan, C., Raghuraman, R., Sreeraj, C. R. &amp; Venkataraman, K. (2014 a) Handbook on Gorgonians (Octocorallia) of Andaman and Nicobar Islands. Director, Zoological Survey of India, Kolkata, 119 pp." type="book" year="2014">
Kumar
<emphasis id="B955DB457B72FFC8FEB2FBA7FE5BFBB8" box="[371,421,1117,1143]" italics="true" pageId="100" pageNumber="101">et al</emphasis>
. 2014a: 40
</bibRefCitation>
, pl. 17, fig. AD (Andaman Is.);
<bibRefCitation id="EFB07AA67B72FFC8FC65FBA4FB33FBB8" author="Fernando, S. &amp; Venkataraman, K. &amp; Raghunathan, C." box="[932,1229,1117,1144]" pageId="100" pageNumber="101" refId="ref100744" refString="Fernando, S. A, Venkataraman, K. &amp; Raghunathan, C. (2017) Gorgonians of Indian sea. Director, Zoological Survey of India, Kolkata, 330 pp. Available from: https: // www. nhbs. com / (accessed 21 March 2022)" type="book" year="2017">
Fernando
<emphasis id="B955DB457B72FFC8FBD7FBA7FBB6FBB8" box="[1046,1096,1117,1143]" italics="true" pageId="100" pageNumber="101">et al</emphasis>
. 2017: 270
</bibRefCitation>
, pl. 126, fig. 11c (SE coast).
</paragraph>
<paragraph id="8B9E07577B72FFC8FF06FB5CFCFFFB2C" blockId="100.[151,1436,1045,1252]" pageId="100" pageNumber="101">
<emphasis id="B955DB457B72FFC8FF06FB5CFC8BFB70" box="[199,885,1189,1215]" italics="true" pageId="100" pageNumber="101">These Indian records appear to represent an identical species</emphasis>
:
<bibRefCitation id="EFB07AA67B72FFC8FC45FB5CFB9BFB70" author="Fernando, S. A." box="[900,1125,1189,1215]" pageId="100" pageNumber="101" refId="ref100707" refString="Fernando, S. A. (2011) Monograph on Gorgonids (Sea fans) of India. Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Parangipettai, 145 pp." type="book" year="2011">Fernando 2011: 120</bibRefCitation>
, pl. 80, fig. 11c (Pamban);
<bibRefCitation id="EFB07AA67B72FFC8FF56FB30FE45FB2C" author="Fernando, S. &amp; Venkataraman, K. &amp; Raghunathan, C." box="[151,443,1225,1252]" pageId="100" pageNumber="101" refId="ref100744" refString="Fernando, S. A, Venkataraman, K. &amp; Raghunathan, C. (2017) Gorgonians of Indian sea. Director, Zoological Survey of India, Kolkata, 330 pp. Available from: https: // www. nhbs. com / (accessed 21 March 2022)" type="book" year="2017">
Fernando
<emphasis id="B955DB457B72FFC8FEC9FB33FEC7FB2C" box="[264,313,1225,1251]" italics="true" pageId="100" pageNumber="101">et al</emphasis>
. 2017: 276
</bibRefCitation>
, pl. 129, fig. 11c (Pamban).
</paragraph>
<paragraph id="8B9E07577B72FFC8FF06FAEBFED3F9E8" blockId="100.[151,1437,1297,1792]" pageId="100" pageNumber="101">
<emphasis id="B955DB457B72FFC8FF06FAEBFE63FAE4" box="[199,413,1297,1323]" italics="true" pageId="100" pageNumber="101">Literature analysis</emphasis>
: The descriptions of the Indian material assigned to this species by
<bibRefCitation id="EFB07AA67B72FFC8FB6FFAE8FA95FAE4" author="Fernando, S. A." box="[1198,1387,1297,1323]" pageId="100" pageNumber="101" refId="ref100707" refString="Fernando, S. A. (2011) Monograph on Gorgonids (Sea fans) of India. Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Parangipettai, 145 pp." type="book" year="2011">Fernando (2011)</bibRefCitation>
and
<bibRefCitation id="EFB07AA67B72FFC8FF56FACCFE6DFA80" author="Fernando, S. &amp; Venkataraman, K. &amp; Raghunathan, C." box="[151,403,1333,1360]" pageId="100" pageNumber="101" refId="ref100744" refString="Fernando, S. A, Venkataraman, K. &amp; Raghunathan, C. (2017) Gorgonians of Indian sea. Director, Zoological Survey of India, Kolkata, 330 pp. Available from: https: // www. nhbs. com / (accessed 21 March 2022)" type="book" year="2017">
Fernando
<emphasis id="B955DB457B72FFC8FEC9FACFFEC4FA80" box="[264,314,1333,1359]" italics="true" pageId="100" pageNumber="101">et al</emphasis>
. (2017)
</bibRefCitation>
are identical, but the sclerite images are so small that it is impossible to compare them with those figured in the description of the
<typeStatus id="549AB9F57B72FFC8FD8DFAA0FD4CFABC" box="[588,690,1369,1395]" pageId="100" pageNumber="101" type="holotype">holotype</typeStatus>
.
<bibRefCitation id="EFB07AA67B72FFC8FD7FFAA0FC81FABC" author="Grasshoff, M." box="[702,895,1369,1395]" pageId="100" pageNumber="101" pagination="1 - 121" refId="ref101429" refString="Grasshoff, M. (1999) The shallow water gorgonians of New Caledonia and adjacent islands (Coelenterata: Octocorallia). Senckenbergiana biologica, 78 (1 / 2), 1 - 121." type="journal article" year="1999">Grasshoff (1999)</bibRefCitation>
reports the species has double heads
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long and slightly larger than the
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long spindles in the polyps.
<bibRefCitation id="EFB07AA67B72FFC8FBE5FA84FB21FA58" author="Fernando, S. A." box="[1060,1247,1405,1431]" pageId="100" pageNumber="101" refId="ref100707" refString="Fernando, S. A. (2011) Monograph on Gorgonids (Sea fans) of India. Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Parangipettai, 145 pp." type="book" year="2011">Fernando (2011)</bibRefCitation>
and
<bibRefCitation id="EFB07AA67B72FFC8FAD6FA84FEF8FA74" author="Fernando, S. &amp; Venkataraman, K. &amp; Raghunathan, C." pageId="100" pageNumber="101" refId="ref100744" refString="Fernando, S. A, Venkataraman, K. &amp; Raghunathan, C. (2017) Gorgonians of Indian sea. Director, Zoological Survey of India, Kolkata, 330 pp. Available from: https: // www. nhbs. com / (accessed 21 March 2022)" type="book" year="2017">
Fernando
<emphasis id="B955DB457B72FFC8FA49FA87FF53FA74" italics="true" pageId="100" pageNumber="101">et al</emphasis>
. (2017)
</bibRefCitation>
report double heads of exactly the same size as in the
<typeStatus id="549AB9F57B72FFC8FCA9FA58FC33FA74" box="[872,973,1441,1467]" pageId="100" pageNumber="101" type="holotype">holotype</typeStatus>
, but the spindles are up to
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long. On the information given, the specimen appears to be the same as
<taxonomicName id="4C217CD47B72FFC8FC56FA3CFA9AFA10" authority="Fernando, 2011" authorityName="Fernando" authorityYear="2011" box="[919,1380,1477,1503]" class="Anthozoa" family="Ellisellidae" genus="Verrucella" kingdom="Animalia" order="Alcyonacea" pageId="100" pageNumber="101" phylum="Cnidaria" rank="species" species="pambanensis">
<emphasis id="B955DB457B72FFC8FC56FA3CFB59FA10" box="[919,1191,1477,1503]" italics="true" pageId="100" pageNumber="101">Verrucella pambanensis</emphasis>
<bibRefCitation id="EFB07AA67B72FFC8FB71FA3CFA9AFA10" author="Fernando, S. A." box="[1200,1380,1477,1503]" pageId="100" pageNumber="101" refId="ref100707" refString="Fernando, S. A. (2011) Monograph on Gorgonids (Sea fans) of India. Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Parangipettai, 145 pp." type="book" year="2011">Fernando, 2011</bibRefCitation>
</taxonomicName>
. The description of that species says both the double heads and the spindles are up to
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, but a figure shows a spindle
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long.
</paragraph>
<paragraph id="8B9E07577B72FFC8FF06F9CBFBF8F930" blockId="100.[151,1437,1297,1792]" pageId="100" pageNumber="101">
A completely different looking specimen is assigned to this species in
<bibRefCitation id="EFB07AA67B72FFC8FBCBF9CBFAFAF984" author="Kumar, J. S. Y. &amp; Raghunathan, C. &amp; Raghuraman, R. &amp; Sreeraj, C. R. &amp; Venkataraman, K." box="[1034,1284,1585,1612]" pageId="100" pageNumber="101" refId="ref103651" refString="Kumar, J. S. Y., Raghunathan, C., Raghuraman, R., Sreeraj, C. R. &amp; Venkataraman, K. (2014 a) Handbook on Gorgonians (Octocorallia) of Andaman and Nicobar Islands. Director, Zoological Survey of India, Kolkata, 119 pp." type="book" year="2014">
Kumar
<emphasis id="B955DB457B72FFC8FBA2F9CBFB67F984" box="[1123,1177,1585,1611]" italics="true" pageId="100" pageNumber="101">et al</emphasis>
. (2014a)
</bibRefCitation>
, but the true characteristics of the material are unknown because much of the description is taken word-for-word from Grasshoffs original text, including even the same sclerite measurements, and some of the figured sclerites can also be found in modified form in Fernando
<emphasis id="B955DB457B72FFC8FE2BF967FDE2F978" box="[490,540,1693,1719]" italics="true" pageId="100" pageNumber="101">et al</emphasis>
.s (2017) description of
<taxonomicName id="4C217CD47B72FFC8FCEEF964FC1AF978" authorityName="Grasshoff" authorityYear="1999" box="[815,996,1693,1719]" class="Anthozoa" family="Ellisellidae" genus="Ellisella" kingdom="Animalia" order="Alcyonacea" pageId="100" pageNumber="101" phylum="Cnidaria" rank="species" species="eustala">
<emphasis id="B955DB457B72FFC8FCEEF964FC1AF978" box="[815,996,1693,1719]" italics="true" pageId="100" pageNumber="101">Ellisella eustala</emphasis>
</taxonomicName>
and some in the
<bibRefCitation id="EFB07AA67B72FFC8FB63F964FA62F978" author="Fernando, S. &amp; Venkataraman, K. &amp; Raghunathan, C." box="[1186,1436,1693,1720]" pageId="100" pageNumber="101" refId="ref100744" refString="Fernando, S. A, Venkataraman, K. &amp; Raghunathan, C. (2017) Gorgonians of Indian sea. Director, Zoological Survey of India, Kolkata, 330 pp. Available from: https: // www. nhbs. com / (accessed 21 March 2022)" type="book" year="2017">
Fernando
<emphasis id="B955DB457B72FFC8FAD2F967FABAF978" box="[1299,1348,1693,1719]" italics="true" pageId="100" pageNumber="101">et al</emphasis>
. (2017)
</bibRefCitation>
description of
<emphasis id="B955DB457B72FFC8FEF8F938FE1DF914" box="[313,483,1729,1755]" italics="true" pageId="100" pageNumber="101">
<taxonomicName id="4C217CD47B72FFC8FEF8F938FE1EF914" authorityName="Grasshoff" authorityYear="1999" box="[313,480,1729,1755]" class="Anthozoa" family="Ellisellidae" genus="Ellisella" kingdom="Animalia" order="Alcyonacea" pageId="100" pageNumber="101" phylum="Cnidaria" rank="species" species="azilia">Ellisella azilia</taxonomicName>
.
</emphasis>
<bibRefCitation id="EFB07AA67B72FFC8FE2BF93BFD37F913" author="Kumar, J. S. Y. &amp; Raghunathan, C. &amp; Venkataraman, K." box="[490,713,1729,1756]" pageId="100" pageNumber="101" pagination="15 - 33" refId="ref103709" refString="Kumar, J. S. Y., Raghunathan, C. &amp; Venkataraman, K. (2015) Abundance of shallow water octocorals in the Andaman and Nicobar Archipelago, India. In: Venkataraman, K. &amp; Sivaperuman, C. (Eds.), Marine faunal diversity in India: taxonomy, ecology and conservation. London Academic Press, London, pp. 15 - 33. https: // doi. org / 10.1016 / B 978 - 0 - 12 - 801948 - 1.00002 - 1" type="book chapter" year="2015">
Kumar
<emphasis id="B955DB457B72FFC8FDFEF93BFD8EF914" box="[575,624,1729,1755]" italics="true" pageId="100" pageNumber="101">et al</emphasis>
. (2015)
</bibRefCitation>
lists the species and provide a colony figure, while
<bibRefCitation id="EFB07AA67B72FFC8FACEF93BFF10F930" author="Kumar, J. S. Y. &amp; Geetha, S. &amp; Raghunathan, C. &amp; Sornaraj, R." pageId="100" pageNumber="101" pagination="96 - 102" refId="ref103354" refString="Kumar, J. S. Y., Geetha, S., Raghunathan, C. &amp; Sornaraj, R. (2018 a) Diversity and distribution of gorgonians (Octocorallia) in the Long Island and it's adjacent areas in Middle Andaman, India. Indian Journal of Geo-Marine Sciences, 47, 96 - 102." type="journal article" year="2018">
Kumar
<emphasis id="B955DB457B72FFC8FAA5F93BFA6BF914" box="[1380,1429,1729,1755]" italics="true" pageId="100" pageNumber="101">et al</emphasis>
. (2018a)
</bibRefCitation>
just lists the species. This species is probably endemic to the
<collectingRegion id="49E5C9B57B72FFC8FC63F91CFBFCF930" box="[930,1026,1765,1791]" country="Egypt" name="Al Bahr al Ahmar" pageId="100" pageNumber="101">Red Sea</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>