treatments-xml/data/03/87/C2/0387C21220396974FF067304E867FA20.xml
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<document id="ADBCAD07D1E3D89674C3AF55B00026BD" ID-CLB-Dataset="294422" ID-DOI="10.11646/zootaxa.5437.3.1" ID-GBIF-Dataset="6c295c3c-1b9e-4ea6-9d17-4005b99c5a5b" ID-ISSN="1175-5326" ID-Zenodo-Dep="10984877" ID-ZooBank="F06B9D4F-B890-41E2-AC14-9C7DD717DFE8" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1713344302398" checkinUser="plazi" docAuthor="Özak, Argun Akif, Kurt, Tuba Terbiyik, Kamanli, Seyit Ali, Akbulut, Gürkan, Yanar, Alper &amp; Boxshall, Geoffrey A." docDate="2024" docId="0387C21220396974FF067304E867FA20" docLanguage="en" docName="zootaxa.5437.3.1.pdf" docOrigin="Zootaxa 5437 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5437.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Caligus sarosensis Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall 2024, sp. nov." docType="treatment" docUuid="C1548B11-522F-4D50-A2C0-0E6A65FAE685" docUuidSource="ZooBank" docVersion="2" lastPageNumber="325" masterDocId="FFBEBA6A2037696CFF91752BEA50FFBD" masterDocTitle="Two new species of planktonic Caligus O. F. Müller 1785 (Copepoda: Caligidae) from Türkiye with an updated review and checklist of planktonic caligids" masterLastPageNumber="335" masterPageNumber="301" pageNumber="315" updateTime="1713527568683" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="84824BAFFC229447A6E3ACB62A020C37">Two new species of planktonic Caligus O. F. Müller 1785 (Copepoda: Caligidae) from Türkiye with an updated review and checklist of planktonic caligids</mods:title>
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<mods:namePart id="CB2B33F7B1E212FA5DE7BBEC71887191">Özak, Argun Akif</mods:namePart>
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<mods:affiliation id="36055C1DD70EC3AF2794DBE32A5F1035">Department of Fish Diseases, Faculty of Fisheries, University of Çukurova, 01330 Sarıçam, Adana, Türkiye</mods:affiliation>
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<mods:namePart id="50DF9B192464F726FB7635A00E51F478">Kurt, Tuba Terbiyik</mods:namePart>
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<mods:affiliation id="B1AE5C529BFA2135A2C375989DB106C8">Department of Basic Sciences, Faculty of Fisheries, University of Çukurova, 01330 Sarıçam, Adana, Türkiye</mods:affiliation>
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<mods:affiliation id="0312749E23686DE269D453B53DFE8406">Department of Biology, Faculty of Art and Science, Burdur Mehmet Akif Ersoy University, Burdur 15030, Türkiye</mods:affiliation>
<mods:nameIdentifier id="BE5A7EE30521EA07C5EDA658D88E9A56" type="email">alikamanli06@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B08739408EB2DF87DCB79BA562901123">Akbulut, Gürkan</mods:namePart>
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<mods:affiliation id="AB68D9D95FBC9A1B3299D67A42344772">Department of Basic Sciences, Faculty of Fisheries, University of Çukurova, 01330 Sarıçam, Adana, Türkiye</mods:affiliation>
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<mods:namePart id="DC288E6A00099D9AF23D4057CD8C568F">Yanar, Alper</mods:namePart>
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<mods:affiliation id="D96896F27FBBEA14F1D58CC7888D992D">Department of Marine Sciences, Faculty of Marine Sciences and Technology, Iskenderun Technical University, Iskenderun-Hatay 31200, Türkiye</mods:affiliation>
<mods:nameIdentifier id="9CB26572BFC04E455E417A67DCB499AE" type="email">alperynr@gmail.com &amp; alper.yanar@iste.edu.tr</mods:nameIdentifier>
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<mods:namePart id="AD029503BF364AA2767435AE8C9C13DC">Boxshall, Geoffrey A.</mods:namePart>
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<mods:affiliation id="887B94B1E81B0FDF4CD7D4A4890F8E4F">Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K.</mods:affiliation>
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<emphasis id="B95AAF1620396962FF067304EB27F9F4" bold="true" box="[151,375,1583,1610]" italics="true" pageId="14" pageNumber="315">Caligus sarosensis</emphasis>
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LSIDurn:lsid:zoobank.org:act:
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material
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<materialsCitation id="3B46795920396962FF0673C8EB4DF8F8" collectingDate="2019-02-20" collectionCode="NHMUK, SABSW" country="Turkey" elevation="60" latitude="40.631252" location="Turkiye" longLatPrecision="1" longitude="26.727695" pageId="14" pageNumber="315" specimenCode="NHMUK 2022, SABSW1" specimenCount="1" typeStatus="holotype">
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stored in collections of the Natural History Museum, London (Reg, No,
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. 200). Type Locality: Stn.
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, Gulf of Saros,
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(
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): depth
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; date
<date id="FF9055C420396962FF067200EB49F8F8" box="[151,281,1835,1861]" pageId="14" pageNumber="315" value="2019-02-20">
<collectingDate id="EFD4AC2C20396962FF067200EB49F8F8" box="[151,281,1835,1861]" pageId="14" pageNumber="315" value="2019-02-20">20.02.2019</collectingDate>
</date>
.
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<emphasis id="B95AAF1620396962FF567264EB02F8D4" bold="true" box="[199,338,1871,1897]" pageId="14" pageNumber="315">Etymology:</emphasis>
The species name refers to the Gulf of Saros (
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, Ţrkiye) where the material was collected.
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<paragraph id="8B91730420396963FF567290EF6FFC84" blockId="14.[151,1437,1763,2042]" lastBlockId="15.[151,1437,151,2013]" lastPageId="15" lastPageNumber="316" pageId="14" pageNumber="315">
Adult female [
<figureCitation id="13156F8120396962FEF27290EB8FF868" box="[355,479,1979,2005]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStart-3="FIGURE 11" captionStart-4="FIGURE 12" captionStart-5="FIGURE 13" captionStart-6="FIGURE 14" captionStartId-0="16.[151,250,1865,1889]" captionStartId-1="17.[151,250,1856,1880]" captionStartId-2="18.[151,250,1909,1933]" captionStartId-3="19.[151,250,1891,1915]" captionStartId-4="20.[151,250,1909,1933]" captionStartId-5="21.[151,250,1943,1967]" captionStartId-6="22.[151,250,1876,1900]" captionTargetBox-0="[254,1334,278,1838]" captionTargetBox-1="[210,1377,181,1832]" captionTargetBox-2="[206,1382,186,1880]" captionTargetBox-3="[198,1390,181,1867]" captionTargetBox-4="[186,1401,186,1880]" captionTargetBox-5="[226,1361,186,1914]" captionTargetBox-6="[151,1435,181,1853]" captionTargetId-0="figure-66@16.[250,1338,273,1842]" captionTargetId-1="figure-19@17.[210,1377,181,1832]" captionTargetId-2="figure-27@18.[201,1386,181,1884]" captionTargetId-3="figure-19@19.[198,1390,181,1867]" captionTargetId-4="figure-27@20.[181,1406,181,1884]" captionTargetId-5="figure-19@21.[221,1366,181,1919]" captionTargetId-6="figure-27@22.[151,1435,181,1853]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="22" captionText-0="FIGURE 8. Caligus sarosensis sp. nov. Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment." captionText-1="FIGURE 9. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Habitus dorsal; B. Habitus ventral; C. Two subrectangular flap like processes (arrowheads) on either side of the medioventral surface of genital complex (arrowheads); D. Abdomen (ventral) with overlapping cuticular folds (white arrows) on anterolateral parts, digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) and 2 sensillae (long black arrows) on mid-anteroventral surface." captionText-2="FIGURE 10. Caligus sarosensis sp. nov. Adult female. A. Antenna, minute distal seta (arrowhead) on outer margin at midlength of terminal claw; B. Postantennal process; C. Maxillule; D. Mouth cone; E. Mandible; F. Maxilla; G. Maxilliped; H. Sternal furca and intercoxal sclerite of leg 1." captionText-3="FIGURE 11. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Antenna, minute distal seta (black arrowhead) on outer margin at mid-length of terminal claw; B. Postantennal process with accessory tine at base (white arrowhead); C. Maxillule; D. Maxilla; E. Distal segment of maxilla with spirally arranged strips of serrated membrane on calamus and bilaterally serrated membrane on canna; F. Maxilliped." captionText-4="FIGURE 12. Caligus sarosensis sp. nov. Adult female. A. Leg 1; B. Endopod; C. Terminal elements on distal segment of leg 1; D. Leg 2; E. Exopod of leg 2; F. Proximal outer spine on third exopodal segment of leg 2; G. Endopod of leg 2." captionText-5="FIGURE 13. Caligus sarosensis sp. nov. Adult female. A. Leg 3; B. Outer spines on third exopodal segment of leg 3; C. Leg 4; D. Exopodal spines on leg 4; E. Leg 5." captionText-6="FIGURE 14. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Distal segment of leg 1; B. Exopod of leg 2, proximal outer spine on third exopodal segment arrowed; C. Leg 4 exopodal spines surrounded with serrated hyaline membrane." figureDoi-0="http://doi.org/10.5281/zenodo.10984901" figureDoi-1="http://doi.org/10.5281/zenodo.10984903" figureDoi-2="http://doi.org/10.5281/zenodo.10984905" figureDoi-3="http://doi.org/10.5281/zenodo.10984911" figureDoi-4="http://doi.org/10.5281/zenodo.10984913" figureDoi-5="http://doi.org/10.5281/zenodo.10984915" figureDoi-6="http://doi.org/10.5281/zenodo.10984917" httpUri-0="https://zenodo.org/record/10984901/files/figure.png" httpUri-1="https://zenodo.org/record/10984903/files/figure.png" httpUri-2="https://zenodo.org/record/10984905/files/figure.png" httpUri-3="https://zenodo.org/record/10984911/files/figure.png" httpUri-4="https://zenodo.org/record/10984913/files/figure.png" httpUri-5="https://zenodo.org/record/10984915/files/figure.png" httpUri-6="https://zenodo.org/record/10984917/files/figure.png" pageId="14" pageNumber="315">Figs. 814</figureCitation>
]. Body (
<figureCitation id="13156F8120396962FDA97290E8CAF868" box="[568,666,1979,2005]" captionStart="FIGURE 8" captionStartId="16.[151,250,1865,1889]" captionTargetBox="[254,1334,278,1838]" captionTargetId="figure-66@16.[250,1338,273,1842]" captionTargetPageId="16" captionText="FIGURE 8. Caligus sarosensis sp. nov. Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment." figureDoi="http://doi.org/10.5281/zenodo.10984901" httpUri="https://zenodo.org/record/10984901/files/figure.png" pageId="14" pageNumber="315">Figs. 8A</figureCitation>
,
<figureCitation id="13156F8120396962FD327290E897F868" box="[675,711,1979,2005]" captionStart="FIGURE 9" captionStartId="17.[151,250,1856,1880]" captionTargetBox="[210,1377,181,1832]" captionTargetId="figure-19@17.[210,1377,181,1832]" captionTargetPageId="17" captionText="FIGURE 9. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Habitus dorsal; B. Habitus ventral; C. Two subrectangular flap like processes (arrowheads) on either side of the medioventral surface of genital complex (arrowheads); D. Abdomen (ventral) with overlapping cuticular folds (white arrows) on anterolateral parts, digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) and 2 sensillae (long black arrows) on mid-anteroventral surface." figureDoi="http://doi.org/10.5281/zenodo.10984903" httpUri="https://zenodo.org/record/10984903/files/figure.png" pageId="14" pageNumber="315">9A</figureCitation>
) comprising cephalothorax, free fourth pedigerous somite, genital complex, and 1-segmented abdomen.Total body length
<quantity id="4CD6DEE120396962FD6972F4E909F847" box="[760,857,2015,2042]" metricMagnitude="-3" metricUnit="m" metricValue="2.06" pageId="14" pageNumber="315" unit="mm" value="2.06">2.06 mm</quantity>
including caudal rami. Dorsal cephalothoracic shield longer than wide, 1.06 × 0.77, narrowing anteriorly towards narrow frontal plate bearing pair of large lunules; lateral margins convex, with broad hyaline membrane around lateral margins. Cephalothorax comprising approximately half of total body length. Thoracic zone of shield about as long as wide, 0.56 × 0.63, forming large posterior sinuses on either side; posterior margin extending beyond posterior end of lateral zones. Fourth pedigerous somite about 2.6 times wider than long (0.10 × 0.26). Genital complex (
<figureCitation id="13156F8120386963FC95740CE9DAFEFC" box="[772,906,295,321]" captionStart="FIGURE 8" captionStartId="16.[151,250,1865,1889]" captionTargetBox="[254,1334,278,1838]" captionTargetId="figure-66@16.[250,1338,273,1842]" captionTargetPageId="16" captionText="FIGURE 8. Caligus sarosensis sp. nov. Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment." figureDoi="http://doi.org/10.5281/zenodo.10984901" httpUri="https://zenodo.org/record/10984901/files/figure.png" pageId="15" pageNumber="316">Figs. 8A, B</figureCitation>
,
<figureCitation id="13156F8120386963FC06740CE9EAFEFC" box="[919,954,295,321]" captionStart="FIGURE 9" captionStartId="17.[151,250,1856,1880]" captionTargetBox="[210,1377,181,1832]" captionTargetId="figure-19@17.[210,1377,181,1832]" captionTargetPageId="17" captionText="FIGURE 9. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Habitus dorsal; B. Habitus ventral; C. Two subrectangular flap like processes (arrowheads) on either side of the medioventral surface of genital complex (arrowheads); D. Abdomen (ventral) with overlapping cuticular folds (white arrows) on anterolateral parts, digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) and 2 sensillae (long black arrows) on mid-anteroventral surface." figureDoi="http://doi.org/10.5281/zenodo.10984903" httpUri="https://zenodo.org/record/10984903/files/figure.png" pageId="15" pageNumber="316">9A</figureCitation>
) 0.54 × 0.38, subrectangular, with parallel lateral margins and weakly lobate posterolateral corners, anterior part indented, forming folded narrow transition between fourth pedigerous somite and genital complex; lateral indentation and partial surface suture present on anteroventral surface of genital complex at about one third of complex length (
<figureCitation id="13156F8120386963FBB774B8EED8FE10" box="[1062,1160,403,429]" captionStart="FIGURE 8" captionStartId="16.[151,250,1865,1889]" captionTargetBox="[254,1334,278,1838]" captionTargetId="figure-66@16.[250,1338,273,1842]" captionTargetPageId="16" captionText="FIGURE 8. Caligus sarosensis sp. nov. Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment." figureDoi="http://doi.org/10.5281/zenodo.10984901" httpUri="https://zenodo.org/record/10984901/files/figure.png" pageId="15" pageNumber="316">Figs. 8B</figureCitation>
black arrows, 9B white arrows). Two subrectangular flap-like processes on medioventral surface either side of median line (
<figureCitation id="13156F8120386963FB75749CEF15FE6C" box="[1252,1349,439,465]" captionStart="FIGURE 8" captionStartId="16.[151,250,1865,1889]" captionTargetBox="[254,1334,278,1838]" captionTargetId="figure-66@16.[250,1338,273,1842]" captionTargetPageId="16" captionText="FIGURE 8. Caligus sarosensis sp. nov. Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment." figureDoi="http://doi.org/10.5281/zenodo.10984901" httpUri="https://zenodo.org/record/10984901/files/figure.png" pageId="15" pageNumber="316">Figs. 8B</figureCitation>
,
<figureCitation id="13156F8120386963FADE749CEFCCFE6C" box="[1359,1436,439,465]" captionStart="FIGURE 9" captionStartId="17.[151,250,1856,1880]" captionTargetBox="[210,1377,181,1832]" captionTargetId="figure-19@17.[210,1377,181,1832]" captionTargetPageId="17" captionText="FIGURE 9. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Habitus dorsal; B. Habitus ventral; C. Two subrectangular flap like processes (arrowheads) on either side of the medioventral surface of genital complex (arrowheads); D. Abdomen (ventral) with overlapping cuticular folds (white arrows) on anterolateral parts, digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) and 2 sensillae (long black arrows) on mid-anteroventral surface." figureDoi="http://doi.org/10.5281/zenodo.10984903" httpUri="https://zenodo.org/record/10984903/files/figure.png" pageId="15" pageNumber="316">9B, 9C</figureCitation>
arrowheads) (SV-SAR5), left and right processes subequal (41 × 74 µm, 45 × 74 µm, respectively). Posteroventral surface of genital complex swollen and produced into lobate posterolateral corners overlying oviduct openings (egg sac attachment area) (
<figureCitation id="13156F8120386963FE1B7708EBBCFD80" box="[394,492,547,573]" captionStart="FIGURE 8" captionStartId="16.[151,250,1865,1889]" captionTargetBox="[254,1334,278,1838]" captionTargetId="figure-66@16.[250,1338,273,1842]" captionTargetPageId="16" captionText="FIGURE 8. Caligus sarosensis sp. nov. Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment." figureDoi="http://doi.org/10.5281/zenodo.10984901" httpUri="https://zenodo.org/record/10984901/files/figure.png" pageId="15" pageNumber="316">Figs. 8C</figureCitation>
,
<figureCitation id="13156F8120386963FE697708E84EFD80" box="[504,542,547,573]" captionStart="FIGURE 9" captionStartId="17.[151,250,1856,1880]" captionTargetBox="[210,1377,181,1832]" captionTargetId="figure-19@17.[210,1377,181,1832]" captionTargetPageId="17" captionText="FIGURE 9. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Habitus dorsal; B. Habitus ventral; C. Two subrectangular flap like processes (arrowheads) on either side of the medioventral surface of genital complex (arrowheads); D. Abdomen (ventral) with overlapping cuticular folds (white arrows) on anterolateral parts, digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) and 2 sensillae (long black arrows) on mid-anteroventral surface." figureDoi="http://doi.org/10.5281/zenodo.10984903" httpUri="https://zenodo.org/record/10984903/files/figure.png" pageId="15" pageNumber="316">9D</figureCitation>
). Free abdomen 1-segmented, slender, 2.3 times longer than wide (0.32 × 0.14), genital complex and abdomen fused laterally, abdomen with numerous overlapping cuticular folds anterolaterally (
<figureCitation id="13156F8120386963FF0E7740EAADFD38" box="[159,253,619,645]" captionStart="FIGURE 8" captionStartId="16.[151,250,1865,1889]" captionTargetBox="[254,1334,278,1838]" captionTargetId="figure-66@16.[250,1338,273,1842]" captionTargetPageId="16" captionText="FIGURE 8. Caligus sarosensis sp. nov. Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment." figureDoi="http://doi.org/10.5281/zenodo.10984901" httpUri="https://zenodo.org/record/10984901/files/figure.png" pageId="15" pageNumber="316">Figs. 8C</figureCitation>
black arrows, 9D white arrows) (SV-SAR6): abdomen with paired swellings on anteroventral surface either side of mid-line, inner margins of each swelling connecting to another digitiform cuticular medial swelling carrying two tiny, triangular papilliform processes (
<figureCitation id="13156F8120386963FD137798E8B4FD70" box="[642,740,691,717]" captionStart="FIGURE 8" captionStartId="16.[151,250,1865,1889]" captionTargetBox="[254,1334,278,1838]" captionTargetId="figure-66@16.[250,1338,273,1842]" captionTargetPageId="16" captionText="FIGURE 8. Caligus sarosensis sp. nov. Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment." figureDoi="http://doi.org/10.5281/zenodo.10984901" httpUri="https://zenodo.org/record/10984901/files/figure.png" pageId="15" pageNumber="316">Figs. 8C</figureCitation>
arrowheads, 9D arrowheads) (SV-SAR6), mid-anteroventral surface ornamented with 2 sensilla (
<figureCitation id="13156F8120386963FDB677FCE8D6FD4C" box="[551,646,727,753]" captionStart="FIGURE 8" captionStartId="16.[151,250,1865,1889]" captionTargetBox="[254,1334,278,1838]" captionTargetId="figure-66@16.[250,1338,273,1842]" captionTargetPageId="16" captionText="FIGURE 8. Caligus sarosensis sp. nov. Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment." figureDoi="http://doi.org/10.5281/zenodo.10984901" httpUri="https://zenodo.org/record/10984901/files/figure.png" pageId="15" pageNumber="316">Figs. 8C</figureCitation>
long arrows, 9D long arrows) (SV-SAR6). Combined length of genital complex and abdomen about 84% of length of cephalothorax. Caudal ramus (
<figureCitation id="13156F8120386963FB9677D0EE3AFCA8" box="[1031,1130,763,789]" captionStart="FIGURE 8" captionStartId="16.[151,250,1865,1889]" captionTargetBox="[254,1334,278,1838]" captionTargetId="figure-66@16.[250,1338,273,1842]" captionTargetPageId="16" captionText="FIGURE 8. Caligus sarosensis sp. nov. Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment." figureDoi="http://doi.org/10.5281/zenodo.10984901" httpUri="https://zenodo.org/record/10984901/files/figure.png" pageId="15" pageNumber="316">Figs. 8C</figureCitation>
,
<figureCitation id="13156F8120386963FBE677D0EECAFCA8" box="[1143,1178,763,789]" captionStart="FIGURE 9" captionStartId="17.[151,250,1856,1880]" captionTargetBox="[210,1377,181,1832]" captionTargetId="figure-19@17.[210,1377,181,1832]" captionTargetPageId="17" captionText="FIGURE 9. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Habitus dorsal; B. Habitus ventral; C. Two subrectangular flap like processes (arrowheads) on either side of the medioventral surface of genital complex (arrowheads); D. Abdomen (ventral) with overlapping cuticular folds (white arrows) on anterolateral parts, digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) and 2 sensillae (long black arrows) on mid-anteroventral surface." figureDoi="http://doi.org/10.5281/zenodo.10984903" httpUri="https://zenodo.org/record/10984903/files/figure.png" pageId="15" pageNumber="316">9D</figureCitation>
) subrectangular, about 1.6 times longer than wide (0.08 × 0.05), bearing 6 plumose setae plus tuft of spinules along inner margin.
</paragraph>
<paragraph id="8B91730420386963FF567668EAA0F9B4" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="316">
Antennule (
<figureCitation id="13156F8120386963FEC17668EBFDFCE0" box="[336,429,835,861]" captionStart="FIGURE 8" captionStartId="16.[151,250,1865,1889]" captionTargetBox="[254,1334,278,1838]" captionTargetId="figure-66@16.[250,1338,273,1842]" captionTargetPageId="16" captionText="FIGURE 8. Caligus sarosensis sp. nov. Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment." figureDoi="http://doi.org/10.5281/zenodo.10984901" httpUri="https://zenodo.org/record/10984901/files/figure.png" pageId="15" pageNumber="316">Fig. 8D</figureCitation>
) 2-segmented; proximal segment about 1.7 times longer than slender distal segment; proximal segment armed with 25 plumose setae anteroventrally near anterior margin plus 2 naked dorsal setae (
<figureCitation id="13156F8120386963FF0E76A0EAA6FC18" box="[159,246,907,933]" captionStart="FIGURE 8" captionStartId="16.[151,250,1865,1889]" captionTargetBox="[254,1334,278,1838]" captionTargetId="figure-66@16.[250,1338,273,1842]" captionTargetPageId="16" captionText="FIGURE 8. Caligus sarosensis sp. nov. Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment." figureDoi="http://doi.org/10.5281/zenodo.10984901" httpUri="https://zenodo.org/record/10984901/files/figure.png" pageId="15" pageNumber="316">Fig. 8D</figureCitation>
arrowheads); slender distal segment carrying 1 subterminal seta on posterior margin and 11 terminal setae plus 2 aesthetascs (
<figureCitation id="13156F8120386963FEE37684EB9AFC74" box="[370,458,943,969]" captionStart="FIGURE 8" captionStartId="16.[151,250,1865,1889]" captionTargetBox="[254,1334,278,1838]" captionTargetId="figure-66@16.[250,1338,273,1842]" captionTargetPageId="16" captionText="FIGURE 8. Caligus sarosensis sp. nov. Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment." figureDoi="http://doi.org/10.5281/zenodo.10984901" httpUri="https://zenodo.org/record/10984901/files/figure.png" pageId="15" pageNumber="316">Fig. 8D</figureCitation>
arrows). Antenna (
<figureCitation id="13156F8120386963FD377684E949FC74" box="[678,793,943,969]" captionStart="FIGURE 10" captionStartId="18.[151,250,1909,1933]" captionTargetBox="[206,1382,186,1880]" captionTargetId="figure-27@18.[201,1386,181,1884]" captionTargetPageId="18" captionText="FIGURE 10. Caligus sarosensis sp. nov. Adult female. A. Antenna, minute distal seta (arrowhead) on outer margin at midlength of terminal claw; B. Postantennal process; C. Maxillule; D. Mouth cone; E. Mandible; F. Maxilla; G. Maxilliped; H. Sternal furca and intercoxal sclerite of leg 1." figureDoi="http://doi.org/10.5281/zenodo.10984905" httpUri="https://zenodo.org/record/10984905/files/figure.png" pageId="15" pageNumber="316">Figs. 10A</figureCitation>
,
<figureCitation id="13156F8120386963FCB67684E908FC74" box="[807,856,943,969]" captionStart="FIGURE 11" captionStartId="19.[151,250,1891,1915]" captionTargetBox="[198,1390,181,1867]" captionTargetId="figure-19@19.[198,1390,181,1867]" captionTargetPageId="19" captionText="FIGURE 11. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Antenna, minute distal seta (black arrowhead) on outer margin at mid-length of terminal claw; B. Postantennal process with accessory tine at base (white arrowhead); C. Maxillule; D. Maxilla; E. Distal segment of maxilla with spirally arranged strips of serrated membrane on calamus and bilaterally serrated membrane on canna; F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.10984911" httpUri="https://zenodo.org/record/10984911/files/figure.png" pageId="15" pageNumber="316">11A</figureCitation>
) 3-segmented; proximal segment without posterior process; middle segment subrectangular; distal segment forming acutely curved claw, with minute distal seta on outer margin at mid-length (
<figureCitation id="13156F8120386963FE4576DCE814FBAC" box="[468,580,1015,1041]" captionStart="FIGURE 10" captionStartId="18.[151,250,1909,1933]" captionTargetBox="[206,1382,186,1880]" captionTargetId="figure-27@18.[201,1386,181,1884]" captionTargetPageId="18" captionText="FIGURE 10. Caligus sarosensis sp. nov. Adult female. A. Antenna, minute distal seta (arrowhead) on outer margin at midlength of terminal claw; B. Postantennal process; C. Maxillule; D. Mouth cone; E. Mandible; F. Maxilla; G. Maxilliped; H. Sternal furca and intercoxal sclerite of leg 1." figureDoi="http://doi.org/10.5281/zenodo.10984905" httpUri="https://zenodo.org/record/10984905/files/figure.png" pageId="15" pageNumber="316">Figs. 10A</figureCitation>
arrowhead, 11A arrowhead) plus proximal seta. Postantennal process (
<figureCitation id="13156F8120386963FAF576DCEA99FB88" captionStart="FIGURE 10" captionStartId="18.[151,250,1909,1933]" captionTargetBox="[206,1382,186,1880]" captionTargetId="figure-27@18.[201,1386,181,1884]" captionTargetPageId="18" captionText="FIGURE 10. Caligus sarosensis sp. nov. Adult female. A. Antenna, minute distal seta (arrowhead) on outer margin at midlength of terminal claw; B. Postantennal process; C. Maxillule; D. Mouth cone; E. Mandible; F. Maxilla; G. Maxilliped; H. Sternal furca and intercoxal sclerite of leg 1." figureDoi="http://doi.org/10.5281/zenodo.10984905" httpUri="https://zenodo.org/record/10984905/files/figure.png" pageId="15" pageNumber="316">Figs. 10B</figureCitation>
,
<figureCitation id="13156F8120386963FF427130EB52FB88" box="[211,258,1051,1077]" captionStart="FIGURE 11" captionStartId="19.[151,250,1891,1915]" captionTargetBox="[198,1390,181,1867]" captionTargetId="figure-19@19.[198,1390,181,1867]" captionTargetPageId="19" captionText="FIGURE 11. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Antenna, minute distal seta (black arrowhead) on outer margin at mid-length of terminal claw; B. Postantennal process with accessory tine at base (white arrowhead); C. Maxillule; D. Maxilla; E. Distal segment of maxilla with spirally arranged strips of serrated membrane on calamus and bilaterally serrated membrane on canna; F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.10984911" httpUri="https://zenodo.org/record/10984911/files/figure.png" pageId="15" pageNumber="316">11B</figureCitation>
) weakly curved, carrying accessory tine proximally (
<figureCitation id="13156F8120386963FCC57130E9E6FB88" box="[852,950,1051,1077]" captionStart="FIGURE 11" captionStartId="19.[151,250,1891,1915]" captionTargetBox="[198,1390,181,1867]" captionTargetId="figure-19@19.[198,1390,181,1867]" captionTargetPageId="19" captionText="FIGURE 11. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Antenna, minute distal seta (black arrowhead) on outer margin at mid-length of terminal claw; B. Postantennal process with accessory tine at base (white arrowhead); C. Maxillule; D. Maxilla; E. Distal segment of maxilla with spirally arranged strips of serrated membrane on calamus and bilaterally serrated membrane on canna; F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.10984911" httpUri="https://zenodo.org/record/10984911/files/figure.png" pageId="15" pageNumber="316">Fig. 11B</figureCitation>
arrowhead) plus two papillae on basal part, each with 6 sensilla; similar papilla with 6 sensilla present on adjacent ventral surface. Maxillule (
<figureCitation id="13156F8120386963FB767114EF08FBE4" box="[1255,1368,1087,1113]" captionStart="FIGURE 10" captionStartId="18.[151,250,1909,1933]" captionTargetBox="[206,1382,186,1880]" captionTargetId="figure-27@18.[201,1386,181,1884]" captionTargetPageId="18" captionText="FIGURE 10. Caligus sarosensis sp. nov. Adult female. A. Antenna, minute distal seta (arrowhead) on outer margin at midlength of terminal claw; B. Postantennal process; C. Maxillule; D. Mouth cone; E. Mandible; F. Maxilla; G. Maxilliped; H. Sternal furca and intercoxal sclerite of leg 1." figureDoi="http://doi.org/10.5281/zenodo.10984905" httpUri="https://zenodo.org/record/10984905/files/figure.png" pageId="15" pageNumber="316">Figs. 10C</figureCitation>
,
<figureCitation id="13156F8120386963FAF57114EFC4FBE4" box="[1380,1428,1087,1113]" captionStart="FIGURE 11" captionStartId="19.[151,250,1891,1915]" captionTargetBox="[198,1390,181,1867]" captionTargetId="figure-19@19.[198,1390,181,1867]" captionTargetPageId="19" captionText="FIGURE 11. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Antenna, minute distal seta (black arrowhead) on outer margin at mid-length of terminal claw; B. Postantennal process with accessory tine at base (white arrowhead); C. Maxillule; D. Maxilla; E. Distal segment of maxilla with spirally arranged strips of serrated membrane on calamus and bilaterally serrated membrane on canna; F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.10984911" httpUri="https://zenodo.org/record/10984911/files/figure.png" pageId="15" pageNumber="316">11C</figureCitation>
) comprising slightly curved dentiform posterior process, anterior papilla bearing 1 long and 2 small setae; posterior part of dentiform process ornamented with narrow marginal flange bilaterally. Mouth tube (
<figureCitation id="13156F8120386963FB1871ACEEBDFB1C" box="[1161,1261,1159,1185]" captionStart="FIGURE 10" captionStartId="18.[151,250,1909,1933]" captionTargetBox="[206,1382,186,1880]" captionTargetId="figure-27@18.[201,1386,181,1884]" captionTargetPageId="18" captionText="FIGURE 10. Caligus sarosensis sp. nov. Adult female. A. Antenna, minute distal seta (arrowhead) on outer margin at midlength of terminal claw; B. Postantennal process; C. Maxillule; D. Mouth cone; E. Mandible; F. Maxilla; G. Maxilliped; H. Sternal furca and intercoxal sclerite of leg 1." figureDoi="http://doi.org/10.5281/zenodo.10984905" httpUri="https://zenodo.org/record/10984905/files/figure.png" pageId="15" pageNumber="316">Fig. 10D</figureCitation>
) approximately 1.1 times longer than wide. Distal margins of labium and labrum fringed with hyaline membrane. Labrum with submarginal row of minute denticles. Mandible (
<figureCitation id="13156F8120386963FD5971E4E97EFB54" box="[712,814,1231,1257]" captionStart="FIGURE 10" captionStartId="18.[151,250,1909,1933]" captionTargetBox="[206,1382,186,1880]" captionTargetId="figure-27@18.[201,1386,181,1884]" captionTargetPageId="18" captionText="FIGURE 10. Caligus sarosensis sp. nov. Adult female. A. Antenna, minute distal seta (arrowhead) on outer margin at midlength of terminal claw; B. Postantennal process; C. Maxillule; D. Mouth cone; E. Mandible; F. Maxilla; G. Maxilliped; H. Sternal furca and intercoxal sclerite of leg 1." figureDoi="http://doi.org/10.5281/zenodo.10984905" httpUri="https://zenodo.org/record/10984905/files/figure.png" pageId="15" pageNumber="316">Fig. 10E</figureCitation>
) curved distally and bearing 12 inner teeth near apex. Maxilla (
<figureCitation id="13156F8120386963FF6D71D8EB0EFAB0" box="[252,350,1267,1293]" captionStart="FIGURE 10" captionStartId="18.[151,250,1909,1933]" captionTargetBox="[206,1382,186,1880]" captionTargetId="figure-27@18.[201,1386,181,1884]" captionTargetPageId="18" captionText="FIGURE 10. Caligus sarosensis sp. nov. Adult female. A. Antenna, minute distal seta (arrowhead) on outer margin at midlength of terminal claw; B. Postantennal process; C. Maxillule; D. Mouth cone; E. Mandible; F. Maxilla; G. Maxilliped; H. Sternal furca and intercoxal sclerite of leg 1." figureDoi="http://doi.org/10.5281/zenodo.10984905" httpUri="https://zenodo.org/record/10984905/files/figure.png" pageId="15" pageNumber="316">Fig. 10F</figureCitation>
,
<figureCitation id="13156F8120386963FEF971D8EBC9FAB0" box="[360,409,1267,1293]" captionStart="FIGURE 11" captionStartId="19.[151,250,1891,1915]" captionTargetBox="[198,1390,181,1867]" captionTargetId="figure-19@19.[198,1390,181,1867]" captionTargetPageId="19" captionText="FIGURE 11. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Antenna, minute distal seta (black arrowhead) on outer margin at mid-length of terminal claw; B. Postantennal process with accessory tine at base (white arrowhead); C. Maxillule; D. Maxilla; E. Distal segment of maxilla with spirally arranged strips of serrated membrane on calamus and bilaterally serrated membrane on canna; F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.10984911" httpUri="https://zenodo.org/record/10984911/files/figure.png" pageId="15" pageNumber="316">11D</figureCitation>
) 2-segmented, brachiform; proximal segment (lacertus) unarmed; distal segment (brachium) armed with small subdistal outer hyaline membrane (flabellum) plus 2 elements at apex (short canna and long calamus) (
<figureCitation id="13156F8120386963FE9A7010EB39FAE8" box="[267,361,1339,1365]" captionStart="FIGURE 11" captionStartId="19.[151,250,1891,1915]" captionTargetBox="[198,1390,181,1867]" captionTargetId="figure-19@19.[198,1390,181,1867]" captionTargetPageId="19" captionText="FIGURE 11. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Antenna, minute distal seta (black arrowhead) on outer margin at mid-length of terminal claw; B. Postantennal process with accessory tine at base (white arrowhead); C. Maxillule; D. Maxilla; E. Distal segment of maxilla with spirally arranged strips of serrated membrane on calamus and bilaterally serrated membrane on canna; F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.10984911" httpUri="https://zenodo.org/record/10984911/files/figure.png" pageId="15" pageNumber="316">Fig 11E</figureCitation>
), canna with bilaterally serrated hyaline membrane, calamus longer than canna and ornamented with spirally arranged strips of serrated membrane (
<figureCitation id="13156F8120386963FD4C7074E91BFAC4" box="[733,843,1375,1401]" captionStart="FIGURE 10" captionStartId="18.[151,250,1909,1933]" captionTargetBox="[206,1382,186,1880]" captionTargetId="figure-27@18.[201,1386,181,1884]" captionTargetPageId="18" captionText="FIGURE 10. Caligus sarosensis sp. nov. Adult female. A. Antenna, minute distal seta (arrowhead) on outer margin at midlength of terminal claw; B. Postantennal process; C. Maxillule; D. Mouth cone; E. Mandible; F. Maxilla; G. Maxilliped; H. Sternal furca and intercoxal sclerite of leg 1." figureDoi="http://doi.org/10.5281/zenodo.10984905" httpUri="https://zenodo.org/record/10984905/files/figure.png" pageId="15" pageNumber="316">Figs. 10F</figureCitation>
,
<figureCitation id="13156F8120386963FCC77074E9D7FAC4" box="[854,903,1375,1401]" captionStart="FIGURE 11" captionStartId="19.[151,250,1891,1915]" captionTargetBox="[198,1390,181,1867]" captionTargetId="figure-19@19.[198,1390,181,1867]" captionTargetPageId="19" captionText="FIGURE 11. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Antenna, minute distal seta (black arrowhead) on outer margin at mid-length of terminal claw; B. Postantennal process with accessory tine at base (white arrowhead); C. Maxillule; D. Maxilla; E. Distal segment of maxilla with spirally arranged strips of serrated membrane on calamus and bilaterally serrated membrane on canna; F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.10984911" httpUri="https://zenodo.org/record/10984911/files/figure.png" pageId="15" pageNumber="316">11E</figureCitation>
). Maxilliped (
<figureCitation id="13156F8120386963FBB47074EEC7FAC4" box="[1061,1175,1375,1401]" captionStart="FIGURE 10" captionStartId="18.[151,250,1909,1933]" captionTargetBox="[206,1382,186,1880]" captionTargetId="figure-27@18.[201,1386,181,1884]" captionTargetPageId="18" captionText="FIGURE 10. Caligus sarosensis sp. nov. Adult female. A. Antenna, minute distal seta (arrowhead) on outer margin at midlength of terminal claw; B. Postantennal process; C. Maxillule; D. Mouth cone; E. Mandible; F. Maxilla; G. Maxilliped; H. Sternal furca and intercoxal sclerite of leg 1." figureDoi="http://doi.org/10.5281/zenodo.10984905" httpUri="https://zenodo.org/record/10984905/files/figure.png" pageId="15" pageNumber="316">Figs. 10G</figureCitation>
,
<figureCitation id="13156F8120386963FB327074EE9FFAC4" box="[1187,1231,1375,1401]" captionStart="FIGURE 11" captionStartId="19.[151,250,1891,1915]" captionTargetBox="[198,1390,181,1867]" captionTargetId="figure-19@19.[198,1390,181,1867]" captionTargetPageId="19" captionText="FIGURE 11. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Antenna, minute distal seta (black arrowhead) on outer margin at mid-length of terminal claw; B. Postantennal process with accessory tine at base (white arrowhead); C. Maxillule; D. Maxilla; E. Distal segment of maxilla with spirally arranged strips of serrated membrane on calamus and bilaterally serrated membrane on canna; F. Maxilliped." figureDoi="http://doi.org/10.5281/zenodo.10984911" httpUri="https://zenodo.org/record/10984911/files/figure.png" pageId="15" pageNumber="316">11F</figureCitation>
) comprising large proximal segment (corpus) and distal subchela representing fused endopodal segments plus curved terminal claw; subchela armed with small seta at base of claw, tip of claw extending almost to middle of corpus; myxal area smooth. Sternal furca (
<figureCitation id="13156F8120386963FE0E70E0E857FA58" box="[415,519,1483,1509]" captionStart="FIGURE 10" captionStartId="18.[151,250,1909,1933]" captionTargetBox="[206,1382,186,1880]" captionTargetId="figure-27@18.[201,1386,181,1884]" captionTargetPageId="18" captionText="FIGURE 10. Caligus sarosensis sp. nov. Adult female. A. Antenna, minute distal seta (arrowhead) on outer margin at midlength of terminal claw; B. Postantennal process; C. Maxillule; D. Mouth cone; E. Mandible; F. Maxilla; G. Maxilliped; H. Sternal furca and intercoxal sclerite of leg 1." figureDoi="http://doi.org/10.5281/zenodo.10984905" httpUri="https://zenodo.org/record/10984905/files/figure.png" pageId="15" pageNumber="316">Fig. 10H</figureCitation>
) with short, straight, weakly divergent tines rounded at tip, tines with marginal flanges.
</paragraph>
<paragraph id="8B91730420386963FF567338EE73F8B8" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="316">
Leg 1 (
<figureCitation id="13156F8120386963FE867338EB2CF990" box="[279,380,1555,1581]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-27@20.[181,1406,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Caligus sarosensis sp. nov. Adult female. A. Leg 1; B. Endopod; C. Terminal elements on distal segment of leg 1; D. Leg 2; E. Exopod of leg 2; F. Proximal outer spine on third exopodal segment of leg 2; G. Endopod of leg 2." figureDoi="http://doi.org/10.5281/zenodo.10984913" httpUri="https://zenodo.org/record/10984913/files/figure.png" pageId="15" pageNumber="316">Fig. 12A</figureCitation>
) biramous, with 2-segmented exopod and lobate vestigial endopod carrying minute denticle at apex (
<figureCitation id="13156F8120386963FF4B731CEB10F9EC" box="[218,320,1591,1617]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-27@20.[181,1406,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Caligus sarosensis sp. nov. Adult female. A. Leg 1; B. Endopod; C. Terminal elements on distal segment of leg 1; D. Leg 2; E. Exopod of leg 2; F. Proximal outer spine on third exopodal segment of leg 2; G. Endopod of leg 2." figureDoi="http://doi.org/10.5281/zenodo.10984913" httpUri="https://zenodo.org/record/10984913/files/figure.png" pageId="15" pageNumber="316">Fig. 12B</figureCitation>
). Sympod armed with lateral plumose seta and inner seta. First exopodal segment (
<figureCitation id="13156F8120386963FB44731CEF6AF9EC" box="[1237,1338,1591,1617]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-27@20.[181,1406,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Caligus sarosensis sp. nov. Adult female. A. Leg 1; B. Endopod; C. Terminal elements on distal segment of leg 1; D. Leg 2; E. Exopod of leg 2; F. Proximal outer spine on third exopodal segment of leg 2; G. Endopod of leg 2." figureDoi="http://doi.org/10.5281/zenodo.10984913" httpUri="https://zenodo.org/record/10984913/files/figure.png" pageId="15" pageNumber="316">Fig. 12A</figureCitation>
) bearing small spine at outer distal corner but lacking row of setules along free posterior margin (
<figureCitation id="13156F8120386963FB1E7370EF53F9C8" box="[1167,1283,1627,1653]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-27@20.[181,1406,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Caligus sarosensis sp. nov. Adult female. A. Leg 1; B. Endopod; C. Terminal elements on distal segment of leg 1; D. Leg 2; E. Exopod of leg 2; F. Proximal outer spine on third exopodal segment of leg 2; G. Endopod of leg 2." figureDoi="http://doi.org/10.5281/zenodo.10984913" httpUri="https://zenodo.org/record/10984913/files/figure.png" pageId="15" pageNumber="316">Figs. 12A</figureCitation>
,
<figureCitation id="13156F8120386963FA817370EF14F9C8" box="[1296,1348,1627,1653]" captionStart="FIGURE 14" captionStartId="22.[151,250,1876,1900]" captionTargetBox="[151,1435,181,1853]" captionTargetId="figure-27@22.[151,1435,181,1853]" captionTargetPageId="22" captionText="FIGURE 14. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Distal segment of leg 1; B. Exopod of leg 2, proximal outer spine on third exopodal segment arrowed; C. Leg 4 exopodal spines surrounded with serrated hyaline membrane." figureDoi="http://doi.org/10.5281/zenodo.10984917" httpUri="https://zenodo.org/record/10984917/files/figure.png" pageId="15" pageNumber="316">14A</figureCitation>
). Distal exopodal segment (
<figureCitation id="13156F8120386963FEE17354EBB0F924" box="[368,480,1663,1689]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-27@20.[181,1406,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Caligus sarosensis sp. nov. Adult female. A. Leg 1; B. Endopod; C. Terminal elements on distal segment of leg 1; D. Leg 2; E. Exopod of leg 2; F. Proximal outer spine on third exopodal segment of leg 2; G. Endopod of leg 2." figureDoi="http://doi.org/10.5281/zenodo.10984913" httpUri="https://zenodo.org/record/10984913/files/figure.png" pageId="15" pageNumber="316">Figs. 12C</figureCitation>
,
<figureCitation id="13156F8120386963FE7A7354E84CF924" box="[491,540,1663,1689]" captionStart="FIGURE 14" captionStartId="22.[151,250,1876,1900]" captionTargetBox="[151,1435,181,1853]" captionTargetId="figure-27@22.[151,1435,181,1853]" captionTargetPageId="22" captionText="FIGURE 14. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Distal segment of leg 1; B. Exopod of leg 2, proximal outer spine on third exopodal segment arrowed; C. Leg 4 exopodal spines surrounded with serrated hyaline membrane." figureDoi="http://doi.org/10.5281/zenodo.10984917" httpUri="https://zenodo.org/record/10984917/files/figure.png" pageId="15" pageNumber="316">14A</figureCitation>
) with 4 terminal elements (spines 13 and inner seta 4) on distal margin; spine 1 (outermost) shortest, middle 2 spines almost equal in length and each bearing single accessory process (
<figureCitation id="13156F8120386963FAB97388EFDEF900" box="[1320,1422,1699,1725]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-27@20.[181,1406,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Caligus sarosensis sp. nov. Adult female. A. Leg 1; B. Endopod; C. Terminal elements on distal segment of leg 1; D. Leg 2; E. Exopod of leg 2; F. Proximal outer spine on third exopodal segment of leg 2; G. Endopod of leg 2." figureDoi="http://doi.org/10.5281/zenodo.10984913" httpUri="https://zenodo.org/record/10984913/files/figure.png" pageId="15" pageNumber="316">Fig. 12C</figureCitation>
), seta 4 (innermost) about 4 times longer than middle two spines 2 and 3 and slightly longer than segment; posterior margin with three plumose setae (
<figureCitation id="13156F8120386963FD8373C0E82AF8B8" box="[530,634,1771,1797]" captionStart="FIGURE 14" captionStartId="22.[151,250,1876,1900]" captionTargetBox="[151,1435,181,1853]" captionTargetId="figure-27@22.[151,1435,181,1853]" captionTargetPageId="22" captionText="FIGURE 14. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Distal segment of leg 1; B. Exopod of leg 2, proximal outer spine on third exopodal segment arrowed; C. Leg 4 exopodal spines surrounded with serrated hyaline membrane." figureDoi="http://doi.org/10.5281/zenodo.10984917" httpUri="https://zenodo.org/record/10984917/files/figure.png" pageId="15" pageNumber="316">Fig. 14A</figureCitation>
), each distinctly longer than segment.
</paragraph>
<paragraph id="8B9173042038697CFF567224EB17FF44" blockId="15.[151,1437,151,2013]" lastBlockId="16.[151,1436,151,249]" lastPageId="16" lastPageNumber="317" pageId="15" pageNumber="316">
Leg 2 (
<figureCitation id="13156F8120386963FE887224EBD0F894" box="[281,384,1807,1833]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-27@20.[181,1406,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Caligus sarosensis sp. nov. Adult female. A. Leg 1; B. Endopod; C. Terminal elements on distal segment of leg 1; D. Leg 2; E. Exopod of leg 2; F. Proximal outer spine on third exopodal segment of leg 2; G. Endopod of leg 2." figureDoi="http://doi.org/10.5281/zenodo.10984913" httpUri="https://zenodo.org/record/10984913/files/figure.png" pageId="15" pageNumber="316">Fig. 12D</figureCitation>
) biramous with 3-segmented rami. First exopodal segment with pinnate seta on inner margin and long spine at outer distal corner reflexed obliquely back across surface of second exopodal segment; second segment lacking outer spine, armed only with pinnate seta on inner margin (
<figureCitation id="13156F8120386963FC4A727CEE18F8CC" box="[987,1096,1879,1905]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-27@20.[181,1406,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Caligus sarosensis sp. nov. Adult female. A. Leg 1; B. Endopod; C. Terminal elements on distal segment of leg 1; D. Leg 2; E. Exopod of leg 2; F. Proximal outer spine on third exopodal segment of leg 2; G. Endopod of leg 2." figureDoi="http://doi.org/10.5281/zenodo.10984913" httpUri="https://zenodo.org/record/10984913/files/figure.png" pageId="15" pageNumber="316">Figs. 12E</figureCitation>
,
<figureCitation id="13156F8120386963FBC2727CEED5F8CC" box="[1107,1157,1879,1905]" captionStart="FIGURE 14" captionStartId="22.[151,250,1876,1900]" captionTargetBox="[151,1435,181,1853]" captionTargetId="figure-27@22.[151,1435,181,1853]" captionTargetPageId="22" captionText="FIGURE 14. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Distal segment of leg 1; B. Exopod of leg 2, proximal outer spine on third exopodal segment arrowed; C. Leg 4 exopodal spines surrounded with serrated hyaline membrane." figureDoi="http://doi.org/10.5281/zenodo.10984917" httpUri="https://zenodo.org/record/10984917/files/figure.png" pageId="15" pageNumber="316">14B</figureCitation>
): third exopodal segment with 5 inner plumose setae plus 3 spines; proximal outer spine (smallest) naked and slender (
<figureCitation id="13156F8120386963FB477250EF0FF828" box="[1238,1375,1915,1941]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-27@20.[181,1406,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Caligus sarosensis sp. nov. Adult female. A. Leg 1; B. Endopod; C. Terminal elements on distal segment of leg 1; D. Leg 2; E. Exopod of leg 2; F. Proximal outer spine on third exopodal segment of leg 2; G. Endopod of leg 2." figureDoi="http://doi.org/10.5281/zenodo.10984913" httpUri="https://zenodo.org/record/10984913/files/figure.png" pageId="15" pageNumber="316">Figs. 12E,F</figureCitation>
,
<figureCitation id="13156F8120386963FAFF7250EFCDF828" box="[1390,1437,1915,1941]" captionStart="FIGURE 14" captionStartId="22.[151,250,1876,1900]" captionTargetBox="[151,1435,181,1853]" captionTargetId="figure-27@22.[151,1435,181,1853]" captionTargetPageId="22" captionText="FIGURE 14. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Distal segment of leg 1; B. Exopod of leg 2, proximal outer spine on third exopodal segment arrowed; C. Leg 4 exopodal spines surrounded with serrated hyaline membrane." figureDoi="http://doi.org/10.5281/zenodo.10984917" httpUri="https://zenodo.org/record/10984917/files/figure.png" pageId="15" pageNumber="316">14B</figureCitation>
arrow), middle spine with hyaline membrane bilaterally; terminal spine ornamented with hyaline membrane along outer margin and row of setules along inner margin (
<figureCitation id="13156F8120386963FD7472E8E922F860" box="[741,882,1987,2013]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-27@20.[181,1406,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Caligus sarosensis sp. nov. Adult female. A. Leg 1; B. Endopod; C. Terminal elements on distal segment of leg 1; D. Leg 2; E. Exopod of leg 2; F. Proximal outer spine on third exopodal segment of leg 2; G. Endopod of leg 2." figureDoi="http://doi.org/10.5281/zenodo.10984913" httpUri="https://zenodo.org/record/10984913/files/figure.png" pageId="15" pageNumber="316">Figs. 12E, F</figureCitation>
,
<figureCitation id="13156F8120386963FCEC72E8E9FFF860" box="[893,943,1987,2013]" captionStart="FIGURE 14" captionStartId="22.[151,250,1876,1900]" captionTargetBox="[151,1435,181,1853]" captionTargetId="figure-27@22.[151,1435,181,1853]" captionTargetPageId="22" captionText="FIGURE 14. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Distal segment of leg 1; B. Exopod of leg 2, proximal outer spine on third exopodal segment arrowed; C. Leg 4 exopodal spines surrounded with serrated hyaline membrane." figureDoi="http://doi.org/10.5281/zenodo.10984917" httpUri="https://zenodo.org/record/10984917/files/figure.png" pageId="15" pageNumber="316">14B</figureCitation>
). First endopodal segment armed with inner plumose seta; second endopodal segment armed with 2 inner plumose setae and ornamented with rows of setules along outer and inner margins; third segment with 6 plumose setae and bearing tuft of setules proximal to outermost seta (
<figureCitation id="13156F812027697CFF4375F4EB6BFF44" box="[210,315,223,249]" captionStart="FIGURE 12" captionStartId="20.[151,250,1909,1933]" captionTargetBox="[186,1401,186,1880]" captionTargetId="figure-27@20.[181,1406,181,1884]" captionTargetPageId="20" captionText="FIGURE 12. Caligus sarosensis sp. nov. Adult female. A. Leg 1; B. Endopod; C. Terminal elements on distal segment of leg 1; D. Leg 2; E. Exopod of leg 2; F. Proximal outer spine on third exopodal segment of leg 2; G. Endopod of leg 2." figureDoi="http://doi.org/10.5281/zenodo.10984913" httpUri="https://zenodo.org/record/10984913/files/figure.png" pageId="16" pageNumber="317">Fig. 12G</figureCitation>
).
</paragraph>
<caption id="DF51238C2027697CFF067262EE9FF84F" ID-DOI="http://doi.org/10.5281/zenodo.10984901" ID-Zenodo-Dep="10984901" httpUri="https://zenodo.org/record/10984901/files/figure.png" pageId="16" pageNumber="317" startId="16.[151,250,1865,1889]" targetBox="[254,1334,278,1838]" targetPageId="16" targetType="figure">
<paragraph id="8B9173042027697CFF067262EE9FF84F" blockId="16.[151,1437,1865,2034]" pageId="16" pageNumber="317">
<emphasis id="B95AAF162027697CFF067262EB42F8DC" bold="true" box="[151,274,1865,1889]" pageId="16" pageNumber="317">FIGURE 8.</emphasis>
<taxonomicName id="4C2E08872027697CFE867262EB85F8DC" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[279,469,1865,1889]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF162027697CFE867262EB85F8DC" box="[279,469,1865,1889]" italics="true" pageId="16" pageNumber="317">Caligus sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2027697CFE4B7261E879F8DC" box="[474,553,1866,1889]" pageId="16" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
Adult female. A. Habitus dorsal; B. Genitoabdomen (ventral) with partial surface suture on anteroventral surface of genital complex (arrows) and two subrectangular flap like processes (arrowheads); C. Abdomen (ventral) with overlapping cuticular folds (arrows) on anterolateral parts and digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) plus 2 sensillae (long arrows) on mid-anteroventral surface; D. Antennule with 2 naked dorsal setae (arrowheads) on proximal segment and 2 aesthetascs (arrows) on distal segment.
</paragraph>
</caption>
<caption id="DF51238C2026697DFF06726BE804F854" ID-DOI="http://doi.org/10.5281/zenodo.10984903" ID-Zenodo-Dep="10984903" httpUri="https://zenodo.org/record/10984903/files/figure.png" pageId="17" pageNumber="318" startId="17.[151,250,1856,1880]" targetBox="[210,1377,181,1832]" targetPageId="17" targetType="figure">
<paragraph id="8B9173042026697DFF06726BE804F854" blockId="17.[151,1437,1856,2025]" pageId="17" pageNumber="318">
<emphasis id="B95AAF162026697DFF06726BEB43F8E5" bold="true" box="[151,275,1856,1880]" pageId="17" pageNumber="318">FIGURE 9.</emphasis>
<taxonomicName id="4C2E08872026697DFE8B726AEB89F8E5" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[282,473,1857,1881]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF162026697DFE8B726AEB89F8E5" box="[282,473,1857,1881]" italics="true" pageId="17" pageNumber="318">Caligus sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2026697DFE71726AE860F8E5" box="[480,560,1857,1880]" pageId="17" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Habitus dorsal; B. Habitus ventral; C. Two subrectangular flap like processes (arrowheads) on either side of the medioventral surface of genital complex (arrowheads); D. Abdomen (ventral) with overlapping cuticular folds (white arrows) on anterolateral parts, digitiform cuticular medial swelling carrying two tiny triangular papilliform processes (arrowheads) and 2 sensillae (long black arrows) on mid-anteroventral surface.
</paragraph>
</caption>
<caption id="DF51238C2025697EFF06725EE809F868" ID-DOI="http://doi.org/10.5281/zenodo.10984905" ID-Zenodo-Dep="10984905" httpUri="https://zenodo.org/record/10984905/files/figure.png" pageId="18" pageNumber="319" startId="18.[151,250,1909,1933]" targetBox="[206,1382,186,1880]" targetPageId="18" targetType="figure">
<paragraph id="8B9173042025697EFF06725EE809F868" blockId="18.[151,1437,1909,2005]" pageId="18" pageNumber="319">
<emphasis id="B95AAF162025697EFF06725EEB4BF830" bold="true" box="[151,283,1909,1933]" pageId="18" pageNumber="319">FIGURE 10</emphasis>
.
<taxonomicName id="4C2E08872025697EFEB8725EEBBAF830" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[297,490,1909,1933]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF162025697EFEB8725EEBBAF830" box="[297,490,1909,1933]" italics="true" pageId="18" pageNumber="319">Caligus sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2025697EFE63725EE814F831" box="[498,580,1909,1932]" pageId="18" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
Adult female. A. Antenna, minute distal seta (arrowhead) on outer margin at midlength of terminal claw; B. Postantennal process; C. Maxillule; D. Mouth cone; E. Mandible; F. Maxilla; G. Maxilliped; H. Sternal furca and intercoxal sclerite of leg 1.
</paragraph>
</caption>
<caption id="DF51238C2024697FFF067248EE61F85A" ID-DOI="http://doi.org/10.5281/zenodo.10984911" ID-Zenodo-Dep="10984911" httpUri="https://zenodo.org/record/10984911/files/figure.png" pageId="19" pageNumber="320" startId="19.[151,250,1891,1915]" targetBox="[198,1390,181,1867]" targetPageId="19" targetType="figure">
<paragraph id="8B9173042024697FFF067248EE61F85A" blockId="19.[151,1437,1891,2023]" pageId="19" pageNumber="320">
<emphasis id="B95AAF162024697FFF067248EB4FF8C6" bold="true" box="[151,287,1891,1915]" pageId="19" pageNumber="320">FIGURE 11.</emphasis>
<taxonomicName id="4C2E08872024697FFEB47248EBB4F8C6" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[293,484,1891,1915]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF162024697FFEB47248EBB4F8C6" box="[293,484,1891,1915]" italics="true" pageId="19" pageNumber="320">Caligus sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2024697FFE78724FE869F8C6" box="[489,569,1892,1915]" pageId="19" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Antenna, minute distal seta (black arrowhead) on outer margin at mid-length of terminal claw; B. Postantennal process with accessory tine at base (white arrowhead); C. Maxillule; D. Maxilla; E. Distal segment of maxilla with spirally arranged strips of serrated membrane on calamus and bilaterally serrated membrane on canna; F. Maxilliped.
</paragraph>
</caption>
<caption id="DF51238C20236978FF06725EEF4CF80C" ID-DOI="http://doi.org/10.5281/zenodo.10984913" ID-Zenodo-Dep="10984913" httpUri="https://zenodo.org/record/10984913/files/figure.png" pageId="20" pageNumber="321" startId="20.[151,250,1909,1933]" targetBox="[186,1401,186,1880]" targetPageId="20" targetType="figure">
<paragraph id="8B91730420236978FF06725EEF4CF80C" blockId="20.[151,1436,1909,1969]" pageId="20" pageNumber="321">
<emphasis id="B95AAF1620236978FF06725EEB70F830" bold="true" box="[151,288,1909,1933]" pageId="20" pageNumber="321">FIGURE 12.</emphasis>
<taxonomicName id="4C2E088720236978FEB7725EEBB6F830" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[294,486,1909,1933]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF1620236978FEB7725EEBB6F830" box="[294,486,1909,1933]" italics="true" pageId="20" pageNumber="321">Caligus sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D20236978FE7D725EE86CF831" box="[492,572,1909,1932]" pageId="20" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
Adult female. A. Leg 1; B. Endopod; C. Terminal elements on distal segment of leg 1; D. Leg 2; E. Exopod of leg 2; F. Proximal outer spine on third exopodal segment of leg 2; G. Endopod of leg 2.
</paragraph>
</caption>
<caption id="DF51238C20226979FF0672BCE869F86E" ID-DOI="http://doi.org/10.5281/zenodo.10984915" ID-Zenodo-Dep="10984915" httpUri="https://zenodo.org/record/10984915/files/figure.png" pageId="21" pageNumber="322" startId="21.[151,250,1943,1967]" targetBox="[226,1361,186,1914]" targetPageId="21" targetType="figure">
<paragraph id="8B91730420226979FF0672BCE869F86E" blockId="21.[151,1436,1943,2004]" pageId="21" pageNumber="322">
<emphasis id="B95AAF1620226979FF0672BCEB70F812" bold="true" box="[151,288,1943,1967]" pageId="21" pageNumber="322">FIGURE 13.</emphasis>
<taxonomicName id="4C2E088720226979FEB672B3EBB7F812" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[295,487,1944,1968]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF1620226979FEB672B3EBB7F812" box="[295,487,1944,1968]" italics="true" pageId="21" pageNumber="322">Caligus sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D20226979FE7C72B3E86EF812" box="[493,574,1944,1967]" pageId="21" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
Adult female. A. Leg 3; B. Outer spines on third exopodal segment of leg 3; C. Leg 4; D. Exopodal spines on leg 4; E. Leg 5.
</paragraph>
</caption>
<caption id="DF51238C2021697AFF06727FE8F2F808" ID-DOI="http://doi.org/10.5281/zenodo.10984917" ID-Zenodo-Dep="10984917" httpUri="https://zenodo.org/record/10984917/files/figure.png" pageId="22" pageNumber="323" startId="22.[151,250,1876,1900]" targetBox="[151,1435,181,1853]" targetPageId="22" targetType="figure">
<paragraph id="8B9173042021697AFF06727FE8F2F808" blockId="22.[151,1437,1876,1973]" pageId="22" pageNumber="323">
<emphasis id="B95AAF162021697AFF06727FEB4FF8D0" bold="true" box="[151,287,1876,1901]" pageId="22" pageNumber="323">FIGURE 14.</emphasis>
<taxonomicName id="4C2E08872021697AFEB4727EEBB4F8D1" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[293,484,1877,1901]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF162021697AFEB4727EEBB4F8D1" box="[293,484,1877,1901]" italics="true" pageId="22" pageNumber="323">Caligus sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2021697AFE78727EE869F8D1" box="[489,569,1877,1900]" pageId="22" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Distal segment of leg 1; B. Exopod of leg 2, proximal outer spine on third exopodal segment arrowed; C. Leg 4 exopodal spines surrounded with serrated hyaline membrane.
</paragraph>
</caption>
<paragraph id="8B9173042020697BFF5675BCEB24FE48" blockId="23.[151,1437,151,681]" pageId="23" pageNumber="324">
Leg 3 (
<figureCitation id="13156F812020697BFEB175BCEBD9FF0C" box="[288,393,151,177]" captionStart="FIGURE 13" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[226,1361,186,1914]" captionTargetId="figure-19@21.[221,1366,181,1919]" captionTargetPageId="21" captionText="FIGURE 13. Caligus sarosensis sp. nov. Adult female. A. Leg 3; B. Outer spines on third exopodal segment of leg 3; C. Leg 4; D. Exopodal spines on leg 4; E. Leg 5." figureDoi="http://doi.org/10.5281/zenodo.10984915" httpUri="https://zenodo.org/record/10984915/files/figure.png" pageId="23" pageNumber="324">Fig. 13A</figureCitation>
) with narrow intercoxal sclerite, coxa and basis fused into flattened apron-like protopod ornamented with extended strips of hyaline membrane along lateral and posterior margins. Inner coxal seta and outer basal seta both pinnate. Exopod 3-segmented, first segment with outer spine extending beyond middle of second segment, orientated parallel with longitudinal axis of ramus, base of spine with pecten-like strip of with hyaline membrane; second segment with outer spine and inner plumose seta, plus setules along outer margin; third segment with outer row of setules and 3 outer spines (
<figureCitation id="13156F812020697BFD787460E91CFED8" box="[745,844,331,357]" captionStart="FIGURE 13" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[226,1361,186,1914]" captionTargetId="figure-19@21.[221,1366,181,1919]" captionTargetPageId="21" captionText="FIGURE 13. Caligus sarosensis sp. nov. Adult female. A. Leg 3; B. Outer spines on third exopodal segment of leg 3; C. Leg 4; D. Exopodal spines on leg 4; E. Leg 5." figureDoi="http://doi.org/10.5281/zenodo.10984915" httpUri="https://zenodo.org/record/10984915/files/figure.png" pageId="23" pageNumber="324">Fig. 13B</figureCitation>
) (first and second spines almost equal in length, third spine slightly shorter than first 2) plus 4 pinnate setae (
<figureCitation id="13156F812020697BFC957444E93DFE34" box="[772,877,367,393]" captionStart="FIGURE 13" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[226,1361,186,1914]" captionTargetId="figure-19@21.[221,1366,181,1919]" captionTargetPageId="21" captionText="FIGURE 13. Caligus sarosensis sp. nov. Adult female. A. Leg 3; B. Outer spines on third exopodal segment of leg 3; C. Leg 4; D. Exopodal spines on leg 4; E. Leg 5." figureDoi="http://doi.org/10.5281/zenodo.10984915" httpUri="https://zenodo.org/record/10984915/files/figure.png" pageId="23" pageNumber="324">Fig. 13A</figureCitation>
). Endopod (
<figureCitation id="13156F812020697BFC677444EE0BFE34" box="[1014,1115,367,393]" captionStart="FIGURE 13" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[226,1361,186,1914]" captionTargetId="figure-19@21.[221,1366,181,1919]" captionTargetPageId="21" captionText="FIGURE 13. Caligus sarosensis sp. nov. Adult female. A. Leg 3; B. Outer spines on third exopodal segment of leg 3; C. Leg 4; D. Exopodal spines on leg 4; E. Leg 5." figureDoi="http://doi.org/10.5281/zenodo.10984915" httpUri="https://zenodo.org/record/10984915/files/figure.png" pageId="23" pageNumber="324">Fig. 13A</figureCitation>
) 2-segmented; first segment armed with long, inner pinnate seta and forming flap-like velum extending to base of exopod, ornamented with row of fine setules along free posterior margin; second segment bearing 6 pinnate setae and bearing row of long setules along outer margin.
</paragraph>
<paragraph id="8B9173042020697BFF5674D4E817FD14" blockId="23.[151,1437,151,681]" pageId="23" pageNumber="324">
Leg 4 (
<figureCitation id="13156F812020697BFE8474D4EB28FDA4" box="[277,376,511,537]" captionStart="FIGURE 13" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[226,1361,186,1914]" captionTargetId="figure-19@21.[221,1366,181,1919]" captionTargetPageId="21" captionText="FIGURE 13. Caligus sarosensis sp. nov. Adult female. A. Leg 3; B. Outer spines on third exopodal segment of leg 3; C. Leg 4; D. Exopodal spines on leg 4; E. Leg 5." figureDoi="http://doi.org/10.5281/zenodo.10984915" httpUri="https://zenodo.org/record/10984915/files/figure.png" pageId="23" pageNumber="324">Fig. 13C</figureCitation>
) uniramous. Protopodal segment with outer seta derived from basis. Exopod 2-segmented; first segment armed with long slender, bilaterally flanged, outer distal spine; compound distal segment with 1 lateral spine and 3 apical spines along oblique distal margin; inner apical spine longest. Inner and middle apical spines each with pecten at base (
<figureCitation id="13156F812020697BFEEF7740EBA1FD38" box="[382,497,619,645]" captionStart="FIGURE 13" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[226,1361,186,1914]" captionTargetId="figure-19@21.[221,1366,181,1919]" captionTargetPageId="21" captionText="FIGURE 13. Caligus sarosensis sp. nov. Adult female. A. Leg 3; B. Outer spines on third exopodal segment of leg 3; C. Leg 4; D. Exopodal spines on leg 4; E. Leg 5." figureDoi="http://doi.org/10.5281/zenodo.10984915" httpUri="https://zenodo.org/record/10984915/files/figure.png" pageId="23" pageNumber="324">Figs. 13D</figureCitation>
,
<figureCitation id="13156F812020697BFE6C7740E87FFD38" box="[509,559,619,645]" captionStart="FIGURE 14" captionStartId="22.[151,250,1876,1900]" captionTargetBox="[151,1435,181,1853]" captionTargetId="figure-27@22.[151,1435,181,1853]" captionTargetPageId="22" captionText="FIGURE 14. Caligus sarosensis sp. nov. Adult female. Drishti applied confocal laser scanning microscope images (CLSM-D). A. Distal segment of leg 1; B. Exopod of leg 2, proximal outer spine on third exopodal segment arrowed; C. Leg 4 exopodal spines surrounded with serrated hyaline membrane." figureDoi="http://doi.org/10.5281/zenodo.10984917" httpUri="https://zenodo.org/record/10984917/files/figure.png" pageId="23" pageNumber="324">14C</figureCitation>
). Armature of rami of legs 14 as follows (Roman numerals indicating spines and Arabic numerals indicating setae).
</paragraph>
<paragraph id="8B9173042020697BFDB677F7E9B4FC23" blockId="23.[551,1023,732,927]" pageId="23" pageNumber="324">
Exopod Endopod Leg 1 I-0;
<collectionCode id="ED3FEBC12020697BFD7C762DE95CFC9D" box="[749,780,774,800]" pageId="23" pageNumber="324">III</collectionCode>
,1,3 vestigial Leg 2 I-1; 0-1; II, I, 5 01; 02; 6 Leg 3 I-0; I-1;
<collectionCode id="ED3FEBC12020697BFC9D7670E97BFCC8" box="[780,811,859,885]" pageId="23" pageNumber="324">III</collectionCode>
, 4 01; 6 Leg 4 I-0; I,
<collectionCode id="ED3FEBC12020697BFC9F76AEE97AFC22" box="[782,810,901,927]" pageId="23" pageNumber="324">III</collectionCode>
absent
</paragraph>
<paragraph id="8B9173042020697BFF5676E5E8ADFB92" blockId="23.[151,1436,973,1072]" pageId="23" pageNumber="324">
Leg 5 (
<figureCitation id="13156F812020697BFE8276E6EB24FC5A" box="[275,372,973,999]" captionStart="FIGURE 13" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[226,1361,186,1914]" captionTargetId="figure-19@21.[221,1366,181,1919]" captionTargetPageId="21" captionText="FIGURE 13. Caligus sarosensis sp. nov. Adult female. A. Leg 3; B. Outer spines on third exopodal segment of leg 3; C. Leg 4; D. Exopodal spines on leg 4; E. Leg 5." figureDoi="http://doi.org/10.5281/zenodo.10984915" httpUri="https://zenodo.org/record/10984915/files/figure.png" pageId="23" pageNumber="324">Fig. 13E</figureCitation>
) located on posterolateral ventral surface of genital complex and represented by 2 small papillae; outer (protopodal) papilla with single long pinnate seta; inner (exopodal) papilla carrying 2 shorter unequal pinnate setae; sensilla located on surface close to inner papilla.
</paragraph>
</subSubSection>
<subSubSection id="C334208F20206974FF0671AAE867FA20" lastPageId="24" lastPageNumber="325" pageId="23" pageNumber="324" type="discussion">
<paragraph id="8B9173042020697BFF0671AAEB59FB26" blockId="23.[151,265,1153,1179]" box="[151,265,1153,1179]" pageId="23" pageNumber="324">
<heading id="D0D9C4682020697BFF0671AAEB59FB26" bold="true" box="[151,265,1153,1179]" fontSize="11" level="1" pageId="23" pageNumber="324" reason="1">
<emphasis id="B95AAF162020697BFF0671AAEB59FB26" bold="true" box="[151,265,1153,1179]" pageId="23" pageNumber="324">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9173042020697BFF0671E2EED7FA2A" blockId="23.[151,1437,1225,2043]" pageId="23" pageNumber="324">
The new species,
<taxonomicName id="4C2E08872020697BFECE71E1EBA5FB5E" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[351,501,1226,1252]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF162020697BFECE71E1EBA5FB5E" box="[351,501,1226,1252]" italics="true" pageId="23" pageNumber="324">C. sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2020697BFE6A71E1E804FB59" box="[507,596,1226,1252]" pageId="23" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
exhibits numerous detailed similarities in the structure and setation of legs 1 to 4 with
<taxonomicName id="4C2E08872020697BFEB371C5EBEFFABA" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[290,447,1262,1288]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="izmiriensis" status="sp. nov.">
<emphasis id="B95AAF162020697BFEB371C5EBEFFABA" box="[290,447,1262,1288]" italics="true" pageId="23" pageNumber="324">C. izmiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2020697BFE5871C5E874FAB5" box="[457,548,1262,1288]" pageId="23" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
(described above) and
<taxonomicName id="4C2E08872020697BFCA571C6EE09FABA" authorityName="Suarez-Morales, Kim &amp; Escamilla" authorityYear="2012" box="[820,1113,1261,1287]" class="Copepoda" family="Caligidae" genus="Metacaligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="yucatanensis">
<emphasis id="B95AAF162020697BFCA571C6EE09FABA" box="[820,1113,1261,1287]" italics="true" pageId="23" pageNumber="324">Metacaligus yucatanensis</emphasis>
</taxonomicName>
is very similar to both new species, having an unusual leg 1 with a rounded distal margin on the second exopodal segment and spines 1 to 3 are small and clustered close to the base of seta 4. In addition, spines 2 and 3 each bear an accessory process and seta 4 is longer than the segment. This configuration differs from that of
<taxonomicName id="4C2E08872020697BFC0A7071EE02FACE" authorityName="Suarez-Morales, Kim &amp; Escamilla" authorityYear="2012" box="[923,1106,1370,1395]" class="Copepoda" family="Caligidae" genus="Metacaligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="yucatanensis">
<emphasis id="B95AAF162020697BFC0A7071EE02FACE" box="[923,1106,1370,1395]" italics="true" pageId="23" pageNumber="324">M. yucatanensis</emphasis>
</taxonomicName>
only in the lack of accessory processes on spines 2 and 3, which are absent according to
<bibRefCitation id="EFBF0EF52020697BFCBE7056EED3FA2A" author="Suarez-Morales, E. &amp; Kim, I. - H. &amp; Escamilla, B. J." box="[815,1155,1405,1431]" pageId="23" pageNumber="324" pagination="804 - 818" refId="ref19296" refString="Suarez-Morales, E., Kim, I. - H. &amp; Escamilla, B. J. (2012 b) On some Caligids (Copepoda: Caligidae) from plankton of a coastal lagoon in the Gulf of Mexico with a description of a new species of Metacaligus. Zoological Studies, 51, 804 - 818." type="journal article" year="2012">
Suárez-Morales
<emphasis id="B95AAF162020697BFC767055EE70FA2A" box="[999,1056,1405,1431]" italics="true" pageId="23" pageNumber="324">et al.</emphasis>
(2012b)
</bibRefCitation>
.
</paragraph>
<paragraph id="8B9173042020697BFF567089E81BF899" blockId="23.[151,1437,1225,2043]" pageId="23" pageNumber="324">
In
<taxonomicName id="4C2E08872020697BFF777089EBD0FA06" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[230,384,1442,1468]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="izmiriensis" status="sp. nov.">
<emphasis id="B95AAF162020697BFF777089EBD0FA06" box="[230,384,1442,1468]" italics="true" pageId="23" pageNumber="324">C. izmiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2020697BFE197089EBB1FA01" box="[392,481,1442,1468]" pageId="23" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
the second exopodal segment of leg 2 carries a tiny vestige of an outer margin spine but both
<taxonomicName id="4C2E08872020697BFE9070EDEBC9FA62" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[257,409,1478,1504]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF162020697BFE9070EDEBC9FA62" box="[257,409,1478,1504]" italics="true" pageId="23" pageNumber="324">C.sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2020697BFE3270EDEBAEFA5D" box="[419,510,1478,1504]" pageId="23" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C2E08872020697BFDAB70EDE8A4FA62" authorityName="Suarez-Morales, Kim &amp; Escamilla" authorityYear="2012" box="[570,756,1478,1503]" class="Copepoda" family="Caligidae" genus="Metacaligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="yucatanensis">
<emphasis id="B95AAF162020697BFDAB70EDE8A4FA62" box="[570,756,1478,1503]" italics="true" pageId="23" pageNumber="324">M. yucatanensis</emphasis>
</taxonomicName>
lack any vestige of this spine. The loss of this spine is an extremely rare character state in
<taxonomicName id="4C2E08872020697BFD9370C2E809F9BE" authorityName="O. F. Muller" authorityYear="1785" box="[514,601,1513,1539]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="B95AAF162020697BFD9370C2E809F9BE" box="[514,601,1513,1539]" italics="true" pageId="23" pageNumber="324">Caligus</emphasis>
</taxonomicName>
. In leg 3 of
<taxonomicName id="4C2E08872020697BFD4E70C1E928F9BE" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[735,888,1514,1540]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="izmiriensis" status="sp. nov.">
<emphasis id="B95AAF162020697BFD4E70C1E928F9BE" box="[735,888,1514,1540]" italics="true" pageId="23" pageNumber="324">C. izmiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2020697BFCEE70C1E987F9B9" box="[895,983,1514,1540]" pageId="23" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
the gap between the rami is smaller than in the other two species, so the velum is not quite so broad, but all three share an elongate second exopodal segment and the outer spine on the first exopodal segment is straight and reaches only about two thirds of the distance along the second segment. There is also a pecten-like, rounded strip of hyaline membrane positioned across the base of the outer spine on the first exopodal segment in all these species. The segmentation and armature of leg 4 is similar in these three species and all have the same arrangement of pectens. Despite these detailed similarities, these species can be readily distinguished by the sternal furca: in
<taxonomicName id="4C2E08872020697BFCBD73E9E995F966" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[812,965,1730,1756]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="izmiriensis" status="sp. nov.">
<emphasis id="B95AAF162020697BFCBD73E9E995F966" box="[812,965,1730,1756]" italics="true" pageId="23" pageNumber="324">C. izmiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2020697BFC5A73E9EE73F961" box="[971,1059,1730,1756]" pageId="23" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
the sternal furca has long, closely adpressed tines, compared with short, widely separated tines in
<taxonomicName id="4C2E08872020697BFCCD73CDE9A1F942" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[860,1009,1766,1792]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF162020697BFCCD73CDE9A1F942" box="[860,1009,1766,1792]" italics="true" pageId="23" pageNumber="324">C. sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2020697BFC6673CDEE01F8BD" box="[1015,1105,1766,1792]" pageId="23" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
, and by the complete absence of the sternal furca in
<taxonomicName id="4C2E08872020697BFE1F7221E814F89E" authorityName="Suarez-Morales, Kim &amp; Escamilla" authorityYear="2012" box="[398,580,1802,1827]" class="Copepoda" family="Caligidae" genus="Metacaligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="yucatanensis">
<emphasis id="B95AAF162020697BFE1F7221E814F89E" box="[398,580,1802,1827]" italics="true" pageId="23" pageNumber="324">M. yucatanensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9173042020697BFF567206EA99F846" blockId="23.[151,1437,1225,2043]" pageId="23" pageNumber="324">
The paired cephalothoracic appendages of the two new species are rather generic and lack any distinctive characteristics. However, the sensory papillae associated with the postantennal process are multisensillate in both species and the presence of 6 or 7 sensilla per papilla is unusually high as there are rarely more than
<quantity id="4CD6DEE12020697BFA99725EEF63F832" box="[1288,1331,1909,1935]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="23" pageNumber="324" unit="in" value="4.0">4 in</quantity>
the great majority of
<taxonomicName id="4C2E08872020697BFEB072B2EB82F80E" box="[289,466,1945,1971]" pageId="23" pageNumber="324">
<emphasis id="B95AAF162020697BFEB072B2EB28F80E" box="[289,376,1945,1971]" italics="true" pageId="23" pageNumber="324">Caligus</emphasis>
species
</taxonomicName>
where the number is known. The presence of an accessory tine on the postantennal process of
<taxonomicName id="4C2E08872020697BFE817295EBCFF86A" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[272,415,1982,2008]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF162020697BFE817295EBCFF86A" box="[272,415,1982,2008]" italics="true" pageId="23" pageNumber="324">C sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2020697BFE347295EBAFF865" box="[421,511,1982,2008]" pageId="23" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
, serves to distinguish it from
<taxonomicName id="4C2E08872020697BFCD87295E9B3F86A" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[841,995,1982,2008]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="izmiriensis" status="sp. nov.">
<emphasis id="B95AAF162020697BFCD87295E9B3F86A" box="[841,995,1982,2008]" italics="true" pageId="23" pageNumber="324">C. izmiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D2020697BFC7B7295EE12F865" box="[1002,1090,1982,2008]" pageId="23" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
which lacks such an accessory tine.
</paragraph>
<paragraph id="8B917304202F6974FF5675BCEE32FC50" blockId="24.[151,1437,151,1437]" pageId="24" pageNumber="325">
The new species from the Gulf of Saros exhibits a slender dorsal cephalothoracic shield which narrows anteriorly towards the frontal plate which bears the paired lunules, and the lateral margins of the shield are provided with unusually broad marginal membrane. This general morphotype is typical of so-called “planktonic” caligids, such as
<taxonomicName id="4C2E0887202F6974FF657428E8DEFEA0" authority="Shen &amp; Li, 1959" authorityName="Shen &amp; Li" authorityYear="1959" box="[244,654,258,285]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B95AAF16202F6974FF657428EB92FEA0" box="[244,450,259,285]" italics="true" pageId="24" pageNumber="325">Caligus undulatus</emphasis>
<bibRefCitation id="EFBF0EF5202F6974FE5A7428E8DEFEA0" author="Shen, C. J. &amp; Li, H. L." box="[459,654,258,285]" pageId="24" pageNumber="325" pagination="12 - 20" refId="ref18806" refString="Shen, C. J. &amp; Li, H. L. (1959) Parasitic copepods from fishes of China. IV. Caligoida, Caligidae (3). Acta Zoologica Sinica, 11, 12 - 20." type="journal article" year="1959">Shen &amp; Li, 1959</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="EFBF0EF5202F6974FD0A7428E9DFFEA0" author="Ohtsuka, S. &amp; Nawata, M. &amp; Nishida, Y. &amp; Nitta, M. &amp; Hirano, K. &amp; Adachi, K. &amp; Kondo, Y. &amp; Venmathi Maran, B. A. &amp; Suarez-Morales, E." box="[667,911,259,285]" pageId="24" pageNumber="325" pagination="52271" refId="ref18258" refString="Ohtsuka, S., Nawata, M., Nishida, Y., Nitta, M., Hirano, K., Adachi, K., Kondo, Y., Venmathi Maran, B. A. &amp; Suarez-Morales, E. (2020) Discovery of the fish host of the &quot; planktonic &quot; caligid Caligus undulatus Shen &amp; Li, 1959 (Crustacea: Copepoda: Siphonostomatoida). Biodiversity Data Journal, 8, e 52271. https: // doi. org / 10.3897 / BDJ. 8. e 52271" type="journal article" year="2020">
Ohtsuka
<emphasis id="B95AAF16202F6974FC937428E96CFEA0" box="[770,828,259,285]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
(2020)
</bibRefCitation>
listed the narrow frontal plate as a diagnostic characteristic of their newly proposed
<taxonomicName id="4C2E0887202F6974FDCB740CE97AFEFC" authorityName="Shen &amp; Li" authorityYear="1959" box="[602,810,295,321]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B95AAF16202F6974FDCB740CE97AFEFC" box="[602,810,295,321]" italics="true" pageId="24" pageNumber="325">Caligus undulatus</emphasis>
</taxonomicName>
-group of species which comprised five, possibly six, species, each of which had originally been described from plankton samples rather than from material obtained from a host fish. As defined by
<bibRefCitation id="EFBF0EF5202F6974FE267444E8F4FE34" author="Ohtsuka, S. &amp; Nawata, M. &amp; Nishida, Y. &amp; Nitta, M. &amp; Hirano, K. &amp; Adachi, K. &amp; Kondo, Y. &amp; Venmathi Maran, B. A. &amp; Suarez-Morales, E." box="[439,676,367,393]" pageId="24" pageNumber="325" pagination="52271" refId="ref18258" refString="Ohtsuka, S., Nawata, M., Nishida, Y., Nitta, M., Hirano, K., Adachi, K., Kondo, Y., Venmathi Maran, B. A. &amp; Suarez-Morales, E. (2020) Discovery of the fish host of the &quot; planktonic &quot; caligid Caligus undulatus Shen &amp; Li, 1959 (Crustacea: Copepoda: Siphonostomatoida). Biodiversity Data Journal, 8, e 52271. https: // doi. org / 10.3897 / BDJ. 8. e 52271" type="journal article" year="2020">
Ohtsuka
<emphasis id="B95AAF16202F6974FD8B7444E801FE34" box="[538,593,367,393]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
(2020)
</bibRefCitation>
, the
<taxonomicName id="4C2E0887202F6974FD467444E935FE34" authorityName="Shen &amp; Li" authorityYear="1959" box="[727,869,367,393]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B95AAF16202F6974FD467444E935FE34" box="[727,869,367,393]" italics="true" pageId="24" pageNumber="325">C. undulatus</emphasis>
</taxonomicName>
-group is characterized by: 1), leg 4 is 3-segmented with IV spines on compound distal exopodal segment; 2), three plumose setae are present on posterior margin of second exopodal segment of leg 1; 3), outer spines on distal exopodal segment of leg 2 are small or reduced; 4), antenna with weak or well-developed process on proximal segment; 5), body with relatively narrow frontal plate; 6), female genital complex longer than wide, sometimes with outer margin undulated; and 7), male urosome slender, with 2-segmented abdomen. In addition to
<taxonomicName id="4C2E0887202F6974FD147708EE4BFD80" authority=", Ohtsuka et al. (2020)" authorityName="Ohtsuka" authorityYear="2020" box="[645,1051,547,573]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B95AAF16202F6974FD147708E946FD80" box="[645,790,547,573]" italics="true" pageId="24" pageNumber="325">C. undulatus</emphasis>
,
<bibRefCitation id="EFBF0EF5202F6974FCB67708EE4BFD80" author="Ohtsuka, S. &amp; Nawata, M. &amp; Nishida, Y. &amp; Nitta, M. &amp; Hirano, K. &amp; Adachi, K. &amp; Kondo, Y. &amp; Venmathi Maran, B. A. &amp; Suarez-Morales, E." box="[807,1051,547,573]" pageId="24" pageNumber="325" pagination="52271" refId="ref18258" refString="Ohtsuka, S., Nawata, M., Nishida, Y., Nitta, M., Hirano, K., Adachi, K., Kondo, Y., Venmathi Maran, B. A. &amp; Suarez-Morales, E. (2020) Discovery of the fish host of the &quot; planktonic &quot; caligid Caligus undulatus Shen &amp; Li, 1959 (Crustacea: Copepoda: Siphonostomatoida). Biodiversity Data Journal, 8, e 52271. https: // doi. org / 10.3897 / BDJ. 8. e 52271" type="journal article" year="2020">
Ohtsuka
<emphasis id="B95AAF16202F6974FC1C7708E997FD80" box="[909,967,547,573]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
(2020)
</bibRefCitation>
</taxonomicName>
included in this species group
<taxonomicName id="4C2E0887202F6974FA127708E8BCFDDC" authority="Suarez-Morales, Camisotti &amp; Martin, 2012" authorityName="Suarez-Morales, Camisotti &amp; Martin" authorityYear="2012" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="evelynae">
<emphasis id="B95AAF16202F6974FA127708EAA9FDDC" italics="true" pageId="24" pageNumber="325">C. evelynae</emphasis>
<bibRefCitation id="EFBF0EF5202F6974FE90776CE8BCFDDC" author="Suarez-Morales, E. &amp; Camisotti, H. &amp; Martin, A." box="[257,748,583,609]" pageId="24" pageNumber="325" pagination="59 - 71" refId="ref19233" refString="Suarez-Morales, E., Camisotti, H. &amp; Martin, A. (2012 a) A new species of Caligus (Copepoda, Siphonostomatoida) from the plankton of the Caribbean coast of Venezuela with a key to species. ZooKeys, 201, 59 - 71. https: // doi. org / 10.3897 / zookeys. 201.3099" type="journal article" year="2012">Suárez-Morales, Camisotti &amp; Martin, 2012</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2E0887202F6974FD69776CEFC8FDDC" authority="Venmathi Maran, Ohtsuka &amp; Jitchum, 2012" authorityName="Venmathi Maran, Ohtsuka &amp; Jitchum" authorityYear="2012" box="[760,1432,583,609]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="longiramus">
<emphasis id="B95AAF16202F6974FD69776CE9CAFDDC" box="[760,922,583,609]" italics="true" pageId="24" pageNumber="325">C. longiramus</emphasis>
<bibRefCitation id="EFBF0EF5202F6974FC33776CEFC8FDDC" author="Venmathi Maran, B. A. &amp; Ohtsuka, S. &amp; Jitchum, P." box="[930,1432,583,609]" pageId="24" pageNumber="325" pagination="87 - 95" refId="ref19556" refString="Venmathi Maran, B. A., Ohtsuka, S. &amp; Jitchum, P. (2012 a) Occurrence of caligid copepods (Crustacea) in plankton samples collected from Japan and Thailand, with the description of a new Species. Species Diversity, 17, 87 - 95. https: // doi. org / 10.12782 / sd. 17.1.087" type="journal article" year="2012">Venmathi Maran, Ohtsuka &amp; Jitchum, 2012</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2E0887202F6974FF067740E95AFD38" authority="Venmathi Maran, Ohtsuka &amp; Shang, 2012" authorityName="Venmathi Maran, Ohtsuka &amp; Shang" authorityYear="2012" box="[151,778,619,645]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="ogawai">
<emphasis id="B95AAF16202F6974FF067740EB5EFD39" box="[151,270,619,645]" italics="true" pageId="24" pageNumber="325">C. ogawai</emphasis>
<bibRefCitation id="EFBF0EF5202F6974FE887740E95AFD38" author="Venmathi Maran, B. A. &amp; Ohtsuka, S. &amp; Shang, X." box="[281,778,619,645]" pageId="24" pageNumber="325" pagination="201 - 219" refId="ref19622" refString="Venmathi Maran, B. A., Ohtsuka, S. &amp; Shang, X. (2012 b) Records of adult caligiform copepods (Crustacea: Copepoda: Siphonostomatoida) in marine plankton from East Asia, including descriptions of two new species of Caligus (Caligidae). Species Diversity, 17, 201 - 219. https: // doi. org / 10.12782 / sd. 17.2.201" type="journal article" year="2012">Venmathi Maran, Ohtsuka &amp; Shang, 2012</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2E0887202F6974FC887740EE3CFD38" authority="Heegaard, 1972" authorityName="Heegaard" authorityYear="1972" box="[793,1132,619,645]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="tripedalis">
<emphasis id="B95AAF16202F6974FC887740E9F9FD38" box="[793,937,619,645]" italics="true" pageId="24" pageNumber="325">C. tripedalis</emphasis>
<bibRefCitation id="EFBF0EF5202F6974FC257740EE3CFD38" author="Heegaard, P. E." box="[948,1132,619,645]" pageId="24" pageNumber="325" pagination="295 - 317" refId="ref16692" refString="Heegaard, P. E. (1972) Caliginae and Euryphorinae of the Dana Expedition (Crustacea Copepoda, Caligidae). Steenstrupia, 2, 295 - 317." type="journal article" year="1972">Heegaard, 1972</bibRefCitation>
</taxonomicName>
, and possibly
<taxonomicName id="4C2E0887202F6974FA867740EB1BFD14" authority="Heegaard, 1966" authorityName="Heegaard" authorityYear="1966" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="hyalinae">
<emphasis id="B95AAF16202F6974FA867740EFCCFD38" box="[1303,1436,619,645]" italics="true" pageId="24" pageNumber="325">C. hyalinae</emphasis>
<bibRefCitation id="EFBF0EF5202F6974FF0677A4EB1BFD14" author="Heegaard, P. E." box="[151,331,655,681]" pageId="24" pageNumber="325" pagination="187 - 197" refId="ref16663" refString="Heegaard, P. E. (1966) Parasitic copepods from Texas. Videnskabelige Meddelelser fra Dansk Naturhistoriske Forening, Copenhagen, 129, 187 - 197." type="journal article" year="1966">Heegaard, 1966</bibRefCitation>
</taxonomicName>
. Subsequently,
<bibRefCitation id="EFBF0EF5202F6974FE6A77A4E971FD14" author="Boxshall, G. A. &amp; Bernot, J. P." box="[507,801,655,681]" pageId="24" pageNumber="325" pagination="545 - 567" refId="ref15523" refString="Boxshall, G. A. &amp; Bernot, J. P. (2023) Resolving taxonomic and nomenclatural problems in the genus Caligus O. F. M ¸ ller, 1785 (Copepoda: Caligidae). Zootaxa, 5360 (4), 545 - 567. https: // doi. org / 10.11646 / zootaxa. 5360.4.5" type="journal article" year="2023">Boxshall &amp; Bernot (2023)</bibRefCitation>
considered that
<taxonomicName id="4C2E0887202F6974FC4877A4EE0AFD14" authorityName="Heegaard" authorityYear="1966" box="[985,1114,655,681]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="hyalinae">
<emphasis id="B95AAF16202F6974FC4877A4EE0AFD14" box="[985,1114,655,681]" italics="true" pageId="24" pageNumber="325">C. hyalinae</emphasis>
</taxonomicName>
should be treated as a junior subjective synonym of
<taxonomicName id="4C2E0887202F6974FE337798E8EFFD70" authority="Wilson, 1905" authorityName="Wilson" authorityYear="1905" box="[418,703,690,717]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B95AAF16202F6974FE337798E84AFD70" box="[418,538,691,717]" italics="true" pageId="24" pageNumber="325">C. chelifer</emphasis>
<bibRefCitation id="EFBF0EF5202F6974FDB27798E8EFFD70" author="Wilson, C. B." box="[547,703,690,717]" pageId="24" pageNumber="325" pagination="479 - 672" refId="ref19836" refString="Wilson, C. B. (1905) North American parasitic copepods belonging to the family Caligidae, Part I. - The Caliginae. Proceedings of the United States National Museum, 28, 479 - 672. https: // doi. org / 10.5479 / si. 00963801.28 - 1404.479" type="journal article" year="1905">Wilson, 1905</bibRefCitation>
</taxonomicName>
, and they confirmed that this species does also belong in the
<taxonomicName id="4C2E0887202F6974FA127798EB55FD4C" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B95AAF16202F6974FA127798EB55FD4C" italics="true" pageId="24" pageNumber="325">C. undulatus</emphasis>
</taxonomicName>
-group. Both new species share
<taxonomicName id="4C2E0887202F6974FDF777FCE8A4FD4C" box="[614,756,727,753]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B95AAF16202F6974FDF777FCE8A4FD4C" box="[614,756,727,753]" italics="true" pageId="24" pageNumber="325">C. undulatus</emphasis>
</taxonomicName>
-group characteristics 1, 2, 4, 5, and 6.
<taxonomicName id="4C2E0887202F6974FB3177FCEF26FD4D" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[1184,1398,727,753]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="izmiriensis" status="sp. nov.">
<emphasis id="B95AAF16202F6974FB3177FCEF26FD4D" box="[1184,1398,727,753]" italics="true" pageId="24" pageNumber="325">Caligus izmiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D202F6974FAED77FCEA99FCA8" pageId="24" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
also shares character 7 but the male (and, therefore, the state of characteristic 7) is unknown in
<taxonomicName id="4C2E0887202F6974FA9777D0EFCDFCA9" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[1286,1437,763,789]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF16202F6974FA9777D0EFCDFCA9" box="[1286,1437,763,789]" italics="true" pageId="24" pageNumber="325">C. sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D202F6974FF067634EAA0FC84" box="[151,240,799,825]" pageId="24" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
Characteristic 4 is not robust since it incorporates the two extremes (and presumably all states in between) of a well-developed process on the proximal segment of the antenna versus proximal process lacking. Characteristic 3 is also variably expressed within the species group, ranging from minute in
<taxonomicName id="4C2E0887202F6974FB82764CEAB8FC18" authority="(Venmathi Maran et al. 2012 a)" baseAuthorityName="Venmathi Maran" baseAuthorityYear="2012" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="ogawai">
<emphasis id="B95AAF16202F6974FB82764CEED7FC3D" box="[1043,1159,871,897]" italics="true" pageId="24" pageNumber="325">C. ogawai</emphasis>
(
<bibRefCitation id="EFBF0EF5202F6974FB09764CEAB0FC18" author="Venmathi Maran, B. A. &amp; Ohtsuka, S. &amp; Jitchum, P." pageId="24" pageNumber="325" pagination="87 - 95" refId="ref19556" refString="Venmathi Maran, B. A., Ohtsuka, S. &amp; Jitchum, P. (2012 a) Occurrence of caligid copepods (Crustacea) in plankton samples collected from Japan and Thailand, with the description of a new Species. Species Diversity, 17, 87 - 95. https: // doi. org / 10.12782 / sd. 17.1.087" type="journal article" year="2012">
Venmathi Maran
<emphasis id="B95AAF16202F6974FAF2764CEFCDFC3C" box="[1379,1437,871,897]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
2012a
</bibRefCitation>
)
</taxonomicName>
, via slender and setiform in
<taxonomicName id="4C2E0887202F6974FDB876A0EE58FC18" authority="(Suarez-Morales et al. 2012 a)" baseAuthorityName="Suarez-Morales" baseAuthorityYear="2012" box="[553,1032,907,933]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="evelynae">
<emphasis id="B95AAF16202F6974FDB876A0E8FDFC18" box="[553,685,907,933]" italics="true" pageId="24" pageNumber="325">C. evelynae</emphasis>
(
<bibRefCitation id="EFBF0EF5202F6974FD2D76A0EE51FC18" author="Suarez-Morales, E. &amp; Camisotti, H. &amp; Martin, A." box="[700,1025,907,933]" pageId="24" pageNumber="325" pagination="59 - 71" refId="ref19233" refString="Suarez-Morales, E., Camisotti, H. &amp; Martin, A. (2012 a) A new species of Caligus (Copepoda, Siphonostomatoida) from the plankton of the Caribbean coast of Venezuela with a key to species. ZooKeys, 201, 59 - 71. https: // doi. org / 10.3897 / zookeys. 201.3099" type="journal article" year="2012">
Suárez-Morales
<emphasis id="B95AAF16202F6974FCE676A0E9E0FC18" box="[887,944,907,933]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
2012a
</bibRefCitation>
)
</taxonomicName>
, to about the same size as the distal outer spine in
<taxonomicName id="4C2E0887202F6974FEA97684E911FC74" authority="(Venmathi Maran et al. 2012 b)" baseAuthorityName="Venmathi Maran" baseAuthorityYear="2012" box="[312,833,943,969]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="longiramus">
<emphasis id="B95AAF16202F6974FEA97684EB88FC74" box="[312,472,943,969]" italics="true" pageId="24" pageNumber="325">C. longiramus</emphasis>
(
<bibRefCitation id="EFBF0EF5202F6974FE767684E969FC74" author="Venmathi Maran, B. A. &amp; Ohtsuka, S. &amp; Shang, X." box="[487,825,943,969]" pageId="24" pageNumber="325" pagination="201 - 219" refId="ref19622" refString="Venmathi Maran, B. A., Ohtsuka, S. &amp; Shang, X. (2012 b) Records of adult caligiform copepods (Crustacea: Copepoda: Siphonostomatoida) in marine plankton from East Asia, including descriptions of two new species of Caligus (Caligidae). Species Diversity, 17, 201 - 219. https: // doi. org / 10.12782 / sd. 17.2.201" type="journal article" year="2012">
Venmathi Maran
<emphasis id="B95AAF16202F6974FD3E7684E8B7FC74" box="[687,743,943,969]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
2012b
</bibRefCitation>
)
</taxonomicName>
. In
<taxonomicName id="4C2E0887202F6974FCFA7684EE51FC75" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[875,1025,943,969]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF16202F6974FCFA7684EE51FC75" box="[875,1025,943,969]" italics="true" pageId="24" pageNumber="325">C. sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D202F6974FB997684EE30FC74" box="[1032,1120,943,969]" pageId="24" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
the proximal outer spine on the distal exopodal segment of leg 2 is minute and in
<taxonomicName id="4C2E0887202F6974FD7C76F8E9D7FC51" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[749,903,979,1005]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="izmiriensis" status="sp. nov.">
<emphasis id="B95AAF16202F6974FD7C76F8E9D7FC51" box="[749,903,979,1005]" italics="true" pageId="24" pageNumber="325">C. izmiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D202F6974FC1F76F8E9B7FC50" box="[910,999,979,1005]" pageId="24" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
it is small.
</paragraph>
<paragraph id="8B917304202F6974FF5676DCE867FA20" blockId="24.[151,1437,151,1437]" pageId="24" pageNumber="325">
<taxonomicName id="4C2E0887202F6974FF5676DCEBCFFBAD" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[199,415,1015,1041]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="izmiriensis" status="sp. nov.">
<emphasis id="B95AAF16202F6974FF5676DCEBCFFBAD" box="[199,415,1015,1041]" italics="true" pageId="24" pageNumber="325">Caligus izmiriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D202F6974FE3676DCE850FBAC" box="[423,512,1015,1041]" pageId="24" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C2E0887202F6974FDA676DCE89DFBAD" authority="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall, 2024" authorityName="Özak &amp; Kurt &amp; Kamanli &amp; Akbulut &amp; Yanar &amp; Boxshall" authorityYear="2024" box="[567,717,1015,1041]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="sarosensis" status="sp. nov.">
<emphasis id="B95AAF16202F6974FDA676DCE89DFBAD" box="[567,717,1015,1041]" italics="true" pageId="24" pageNumber="325">C. sarosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269126D202F6974FD4576DCE97DFBAC" box="[724,813,1015,1041]" pageId="24" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
both exhibit the same general morphotype as members of the
<taxonomicName id="4C2E0887202F6974FF717130EB3FFB88" box="[224,367,1051,1077]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B95AAF16202F6974FF717130EB3FFB88" box="[224,367,1051,1077]" italics="true" pageId="24" pageNumber="325">C. undulatus</emphasis>
</taxonomicName>
-group, especially the narrow frontal plate and the broad marginal membrane around the dorsal cephalothoracic shield. We propose to place them in the
<taxonomicName id="4C2E0887202F6974FC907114E9DEFBE4" box="[769,910,1087,1113]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B95AAF16202F6974FC907114E9DEFBE4" box="[769,910,1087,1113]" italics="true" pageId="24" pageNumber="325">C. undulatus</emphasis>
</taxonomicName>
- group together with
<taxonomicName id="4C2E0887202F6974FBEA7114EF58FBE4" box="[1147,1288,1087,1113]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B95AAF16202F6974FBEA7114EF58FBE4" box="[1147,1288,1087,1113]" italics="true" pageId="24" pageNumber="325">C. undulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2E0887202F6974FA857114EFC5FBE4" box="[1300,1429,1087,1113]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="evelynae">
<emphasis id="B95AAF16202F6974FA857114EFC5FBE4" box="[1300,1429,1087,1113]" italics="true" pageId="24" pageNumber="325">C. evelynae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2E0887202F6974FF067148EB6CFBC0" box="[151,316,1123,1149]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="longiramus">
<emphasis id="B95AAF16202F6974FF067148EB6CFBC0" box="[151,316,1123,1149]" italics="true" pageId="24" pageNumber="325">C. longiramus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2E0887202F6974FEDC7148EB94FBC1" box="[333,452,1123,1149]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="ogawai">
<emphasis id="B95AAF16202F6974FEDC7148EB94FBC1" box="[333,452,1123,1149]" italics="true" pageId="24" pageNumber="325">C. ogawai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2E0887202F6974FE477148E836FBC0" authorityName="Heegaard" authorityYear="1972" box="[470,614,1123,1149]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="tripedalis">
<emphasis id="B95AAF16202F6974FE477148E836FBC0" box="[470,614,1123,1149]" italics="true" pageId="24" pageNumber="325">C. tripedalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2E0887202F6974FD357148E94FFBC0" authorityName="Wilson" authorityYear="1905" box="[676,799,1123,1149]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="B95AAF16202F6974FD357148E94FFBC0" box="[676,799,1123,1149]" italics="true" pageId="24" pageNumber="325">C. chelifer</emphasis>
</taxonomicName>
. Interestingly
<taxonomicName id="4C2E0887202F6974FC587148EEBFFBC1" authorityName="Suarez-Morales, Kim &amp; Escamilla" authorityYear="2012" box="[969,1263,1123,1149]" class="Copepoda" family="Caligidae" genus="Metacaligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="yucatanensis">
<emphasis id="B95AAF16202F6974FC587148EEBFFBC1" box="[969,1263,1123,1149]" italics="true" pageId="24" pageNumber="325">Metacaligus yucatanensis</emphasis>
</taxonomicName>
also shares
<taxonomicName id="4C2E0887202F6974FA127148EB55FB1C" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B95AAF16202F6974FA127148EB55FB1C" italics="true" pageId="24" pageNumber="325">C. undulatus</emphasis>
</taxonomicName>
-group characteristics 1, 2, 4, 5, 6 and 7. Regarding characteristic 3, the proximal outer spine on exopodal segment 3 of leg
<quantity id="4CD6DEE1202F6974FEF47180EBC3FB78" box="[357,403,1195,1221]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="24" pageNumber="325" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C2E0887202F6974FE0C7180E807FB79" authorityName="Suarez-Morales, Kim &amp; Escamilla" authorityYear="2012" box="[413,599,1195,1220]" class="Copepoda" family="Caligidae" genus="Metacaligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="yucatanensis">
<emphasis id="B95AAF16202F6974FE0C7180E807FB79" box="[413,599,1195,1220]" italics="true" pageId="24" pageNumber="325">M. yucatanensis</emphasis>
</taxonomicName>
is long although extremely slender and setiform in appearance (
<bibRefCitation id="EFBF0EF5202F6974FAD47180EBD5FB54" author="Suarez-Morales, E. &amp; Kim, I. - H. &amp; Escamilla, B. J." pageId="24" pageNumber="325" pagination="804 - 818" refId="ref19296" refString="Suarez-Morales, E., Kim, I. - H. &amp; Escamilla, B. J. (2012 b) On some Caligids (Copepoda: Caligidae) from plankton of a coastal lagoon in the Gulf of Mexico with a description of a new species of Metacaligus. Zoological Studies, 51, 804 - 818." type="journal article" year="2012">
Suárez-Morales
<emphasis id="B95AAF16202F6974FF6A71E4EB64FB54" box="[251,308,1231,1257]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
2012b
</bibRefCitation>
), somewhat like that of
<taxonomicName id="4C2E0887202F6974FD0771E4E949FB54" box="[662,793,1231,1257]" class="Copepoda" family="Caligidae" genus="Caligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="evelynae">
<emphasis id="B95AAF16202F6974FD0771E4E949FB54" box="[662,793,1231,1257]" italics="true" pageId="24" pageNumber="325">C. evelynae</emphasis>
</taxonomicName>
. Despite this impressive array of shared character states, the two new species can be readily distinguished from
<taxonomicName id="4C2E0887202F6974FC9871D8E991FAB1" authorityName="Suarez-Morales, Kim &amp; Escamilla" authorityYear="2012" box="[777,961,1267,1292]" class="Copepoda" family="Caligidae" genus="Metacaligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="yucatanensis">
<emphasis id="B95AAF16202F6974FC9871D8E991FAB1" box="[777,961,1267,1292]" italics="true" pageId="24" pageNumber="325">M. yucatanensis</emphasis>
</taxonomicName>
by the presence of a sternal furca in both new species. All species currently placed in
<taxonomicName id="4C2E0887202F6974FD17703CE940FA8C" authorityName="Thomsen" authorityYear="1949" box="[646,784,1303,1329]" class="Copepoda" family="Caligidae" genus="Metacaligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="B95AAF16202F6974FD17703CE940FA8C" box="[646,784,1303,1329]" italics="true" pageId="24" pageNumber="325">Metacaligus</emphasis>
</taxonomicName>
lack a sternal furca (
<bibRefCitation id="EFBF0EF5202F6974FC69703CEE94FA8C" author="Dojiri, M. &amp; Ho, J. - S." box="[1016,1220,1303,1329]" pageId="24" pageNumber="325" refId="ref16258" refString="Dojiri, M. &amp; Ho, J. - S. (2013) Systematics of the Caligidae, copepods parasitic on marine fishes. Crustaceana Monographs 18. Brill, Leiden and Boston, 448 pp. https: // doi. org / 10.1163 / 9789004204256" type="book" year="2013">Dojiri &amp; Ho 2013</bibRefCitation>
;
<bibRefCitation id="EFBF0EF5202F6974FB41703CEB53FAE8" author="Suarez-Morales, E. &amp; Kim, I. - H. &amp; Escamilla, B. J." pageId="24" pageNumber="325" pagination="804 - 818" refId="ref19296" refString="Suarez-Morales, E., Kim, I. - H. &amp; Escamilla, B. J. (2012 b) On some Caligids (Copepoda: Caligidae) from plankton of a coastal lagoon in the Gulf of Mexico with a description of a new species of Metacaligus. Zoological Studies, 51, 804 - 818." type="journal article" year="2012">
Suárez-Morales
<emphasis id="B95AAF16202F6974FA19703CEAE4FAE8" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
2012b
</bibRefCitation>
) and the presence of a sternal furca prevents the placement of the new species in
<taxonomicName id="4C2E0887202F6974FB077010EF70FAE8" authorityName="Thomsen" authorityYear="1949" box="[1174,1312,1339,1365]" class="Copepoda" family="Caligidae" genus="Metacaligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="B95AAF16202F6974FB077010EF70FAE8" box="[1174,1312,1339,1365]" italics="true" pageId="24" pageNumber="325">Metacaligus</emphasis>
</taxonomicName>
. However, in view of the close similarity between the new species and
<taxonomicName id="4C2E0887202F6974FCBF7074E9B3FAC5" authorityName="Suarez-Morales, Kim &amp; Escamilla" authorityYear="2012" box="[814,995,1375,1400]" class="Copepoda" family="Caligidae" genus="Metacaligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="yucatanensis">
<emphasis id="B95AAF16202F6974FCBF7074E9B3FAC5" box="[814,995,1375,1400]" italics="true" pageId="24" pageNumber="325">M. yucatanensis</emphasis>
</taxonomicName>
, it is necessary to review the validity of
<taxonomicName id="4C2E0887202F6974FF0670A8EB71FA20" authorityName="Thomsen" authorityYear="1949" box="[151,289,1411,1437]" class="Copepoda" family="Caligidae" genus="Metacaligus" kingdom="Animalia" order="Siphonostomatoida" pageId="24" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="B95AAF16202F6974FF0670A8EB71FA20" box="[151,289,1411,1437]" italics="true" pageId="24" pageNumber="325">Metacaligus</emphasis>
</taxonomicName>
as a generic level taxon.
</paragraph>
</subSubSection>
</treatment>
</document>