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<document id="5AF05C9721794E49557E118143B2DF96" ID="10.11646/zootaxa.4236.1.2SLASH10251" ID-CLB-Dataset="36479" ID-DOI="10.11646/zootaxa.4236.1.2" ID-GBIF-Dataset="0f39b5c0-0af5-476f-8448-a02339cc1dfd" ID-ISSN="1175-5326" ID-Zenodo-Dep="322014" ID-ZooBank="23412386-CB17-49CA-9C47-BD71DD9C5372" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487752177037" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N. &amp; Yushkova, Olga V." docDate="2017" docId="4F3A5135FFB3DC084FE3FEBC6D99F89C" docLanguage="en" docName="zootaxa.4236.1.2SLASH10251.pdf" docOrigin="Zootaxa 4236 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhabdoblattella euptera Anisyutkin, sp.nov." docType="treatment" docVersion="7" lastPageNumber="52" masterDocId="B303294DFFB8DC034F74FFCC6F0CFF9F" masterDocTitle="New data on cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from India and Sri Lanka, with descriptions of new species and the genital complex of Aptera fusca (Thunberg, 1784)" masterLastPageNumber="64" masterPageNumber="41" pageNumber="52" updateTime="1698437163562" updateUser="ExternalLinkService">
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<mods:title id="A9BDE9DC52E76BEB2BD9D924D393ED04">New data on cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from India and Sri Lanka, with descriptions of new species and the genital complex of Aptera fusca (Thunberg, 1784)</mods:title>
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<mods:namePart id="CCAA00667A12C79283C0FF6A552A0A7A">Anisyutkin, Leonid N.</mods:namePart>
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<mods:namePart id="55BC9F1C311DC6018406675C4E7C8353">Yushkova, Olga V.</mods:namePart>
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<mods:title id="075008EB2ADECC343B27AB939D4E6E5D">Zootaxa</mods:title>
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<mods:date id="6BBAA35F717953DA6EAD9E12803A4DFE">2017</mods:date>
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<mods:identifier id="CD927327F5BD1EA95B2948F1F5A98B2C" type="DOI">10.11646/zootaxa.4236.1.2</mods:identifier>
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<treatment id="4F3A5135FFB3DC084FE3FEBC6D99F89C" ID-DOI="http://doi.org/10.5281/zenodo.6015705" ID-GBIF-Taxon="127697123" ID-Zenodo-Dep="6015705" LSID="urn:lsid:plazi:treatment:4F3A5135FFB3DC084FE3FEBC6D99F89C" httpUri="http://treatment.plazi.org/id/4F3A5135FFB3DC084FE3FEBC6D99F89C" lastPageNumber="52" pageId="11" pageNumber="52">
<subSubSection id="8F89B3A8FFB3DC084FE3FEBC6EBCFE34" pageId="11" pageNumber="52" type="nomenclature">
<paragraph id="C72CE023FFB3DC084FE3FEBC6DBCFE15" blockId="11.[151,688,368,427]" box="[151,688,368,394]" pageId="11" pageNumber="52">
<heading id="9C64574FFFB3DC084FE3FEBC6DBCFE15" bold="true" box="[151,688,368,394]" fontSize="11" level="1" pageId="11" pageNumber="52" reason="1">
<emphasis id="F5E73C31FFB3DC084FE3FEBC6DBCFE15" bold="true" box="[151,688,368,394]" pageId="11" pageNumber="52">
<taxonomicName id="00939BA0FFB3DC084FE3FEBC6D42FE15" authority="Anisyutkin" authorityName="Anisyutkin" box="[151,590,368,394]" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="euptera" status="sp. nov.">
<emphasis id="F5E73C31FFB3DC084FE3FEBC6EB5FE15" bold="true" box="[151,441,368,394]" italics="true" pageId="11" pageNumber="52">Rhabdoblattella euptera</emphasis>
Anisyutkin
</taxonomicName>
,
<taxonomicNameLabel id="EED4814AFFB3DC084D2EFEBC6DBCFE15" box="[602,688,368,394]" pageId="11" pageNumber="52" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C72CE023FFB3DC084FE3FE5E6EBCFE34" blockId="11.[151,688,368,427]" box="[151,432,402,427]" pageId="11" pageNumber="52">
(
<figureCitation id="5FA8FCA6FFB3DC084FEBFE5E6E0FFE34" box="[159,259,402,427]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="11" pageNumber="52">Figs. 4D</figureCitation>
, E, 6AP, 11K)
</paragraph>
</subSubSection>
<subSubSection id="8F89B3A8FFB3DC084FE3FE166BFFFD88" pageId="11" pageNumber="52" type="materials_examined">
<paragraph id="C72CE023FFB3DC084FE3FE166BFFFD88" blockId="11.[151,1437,474,1795]" pageId="11" pageNumber="52">
<materialsCitation id="77FBEA7EFFB3DC084FE3FE166BFFFD88" ID-GBIF-Occurrence="1434656613" collectingDate="1972-11-25" collectionCode="SOUTH, MHNG" collectorName="C. Besuchet &amp; I. Lobl" country="India" elevation="475" location="Valara Fall" municipality="Cardamon Hills" pageId="11" pageNumber="52" specimenCount="1" stateProvince="Kerala">
<emphasis id="F5E73C31FFB3DC084FE3FE166E0AFE6C" bold="true" box="[151,262,474,499]" pageId="11" pageNumber="52">Material.</emphasis>
Holotype—male,
<collectionCode id="A18278E6FFB3DC084EA8FE176D35FE6C" box="[476,569,475,499]" pageId="11" pageNumber="52">SOUTH</collectionCode>
<collectingCountry id="BF84A0B3FFB3DC084D30FE176D9AFE6C" box="[580,662,475,499]" name="India" pageId="11" pageNumber="52">INDIA</collectingCountry>
, state
<collectingRegion id="05572EC1FFB3DC084D96FE166C23FE6C" box="[738,815,474,499]" country="India" name="Kerala" pageId="11" pageNumber="52">Kerala</collectingRegion>
,
<collectingMunicipality id="27487A59FFB3DC084C4AFE166CFAFE6C" box="[830,1014,474,499]" pageId="11" pageNumber="52">Cardamon Hills</collectingMunicipality>
,
<location id="C24CB6F8FFB3DC084B71FE166B89FE6C" LSID="urn:lsid:plazi:treatment:4F3A5135FFB3DC084FE3FEBC6D99F89C:C24CB6F8FFB3DC084B71FE166B89FE6C" box="[1029,1157,474,499]" country="India" municipality="Cardamon Hills" name="Valara Fall" pageId="11" pageNumber="52" stateProvince="Kerala">Valara Fall</location>
,
<locationDeviation id="D69EF901FFB3DC084BE7FE166A94FE6D" box="[1171,1432,474,499]" pageId="11" pageNumber="52">
<quantity id="006B4DC6FFB3DC084BE7FE166BD2FE6C" box="[1171,1246,474,499]" metricMagnitude="4" metricUnit="m" metricValue="4.6" pageId="11" pageNumber="52" unit="km" value="46.0">46 km</quantity>
SW of Munnar
</locationDeviation>
,
<elevation id="4CBE0710FFB3DC084FE3FE326E15FD89" box="[151,281,510,535]" metricMagnitude="2" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="11" pageNumber="52" unit="m" value="475.0" valueMax="500.0" valueMin="450.0">
<quantity id="006B4DC6FFB3DC084FE3FE326E15FD89" box="[151,281,510,535]" metricMagnitude="2" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="11" pageNumber="52" unit="m" value="475.0" valueMax="500.0" valueMin="450.0">450500 m</quantity>
</elevation>
; sieving in the forest near river,
<collectingDate id="A3693F0BFFB3DC084DF6FE326C51FD88" box="[642,861,510,535]" pageId="11" pageNumber="52" value="1972-11-25">
<date id="B32DC6E3FFB3DC084DF6FE326C51FD88" box="[642,861,510,535]" pageId="11" pageNumber="52" value="1972-11-25">25 November 1972</date>
</collectingDate>
, coll.
<collectorName id="6A6685F5FFB3DC084CD4FE336B27FD88" box="[928,1067,510,535]" pageId="11" pageNumber="52">C. Besuchet</collectorName>
/
<collectorName id="6A6685F5FFB3DC084B40FE326B70FD88" box="[1076,1148,510,535]" pageId="11" pageNumber="52">I. Löbl</collectorName>
(
<collectionCode id="A18278E6FFB3DC084BFEFE336BEAFD88" box="[1162,1254,511,535]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="11" pageNumber="52">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8F89B3A8FFB3DC084FB3FDEF6D63FDC0" pageId="11" pageNumber="52" type="etymology">
<paragraph id="C72CE023FFB3DC084FB3FDEF6D63FDC0" blockId="11.[151,1437,474,1795]" pageId="11" pageNumber="52">
<emphasis id="F5E73C31FFB3DC084FB3FDEF6E42FDA4" bold="true" box="[199,334,547,571]" pageId="11" pageNumber="52">Etymology.</emphasis>
The name of species derived from the Greek words &quot;eu&quot;—well and &quot;ptero&quot;—wing, i.e. a cockroach with the well developed wings.
</paragraph>
</subSubSection>
<subSubSection id="8F89B3A8FFB3DC084FB3FDA66D99F89C" pageId="11" pageNumber="52" type="description">
<paragraph id="C72CE023FFB3DC084FB3FDA66C0DFACC" blockId="11.[151,1437,474,1795]" pageId="11" pageNumber="52">
<emphasis id="F5E73C31FFB3DC084FB3FDA66E55FD1C" bold="true" box="[199,345,618,643]" pageId="11" pageNumber="52">Description.</emphasis>
Male (the
<typeStatus id="18285E81FFB3DC084EAEFDA66D4DFD1C" box="[474,577,618,643]" pageId="11" pageNumber="52" type="holotype">holotype</typeStatus>
). General colour yellowish brown (
<figureCitation id="5FA8FCA6FFB3DC084CADFDA66B39FD1C" box="[985,1077,618,643]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="11" pageNumber="52">Fig. 4D</figureCitation>
, E); facial part of head yellow, with brown vertex; eyes black; pronotum, tegmina and abdomen densely speckled with small dark maculae; legs yellowish. Surfaces smooth and lustrous; antennae with lustrous proximal 1213 segments, the rest of segments dull. Head convex (
<figureCitation id="5FA8FCA6FFB3DC084E08FD1A6EC1FD70" box="[380,461,726,751]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="11" pageNumber="52">Figs. 4</figureCitation>
E, 6A); epicranial sutures indistinct; ocellar spots distinct; a weak transverse hollow located between antennal sockets; distance between eyes 0.8 times eye length; distance between antennal sockets about twice of the scape length (about
<quantity id="006B4DC6FFB3DC084D24FCD36DA0FCA8" box="[592,684,799,823]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="52" unit="mm" value="0.8">0.8 mm</quantity>
); approximate length ratio of 3rd5th segments of maxillary palps 1.0: 1.0: 1.2. Pronotum as in
<figureCitation id="5FA8FCA6FFB3DC084E9FFC8E6D42FCC4" box="[491,590,834,859]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="11" pageNumber="52">Figs. 4D</figureCitation>
,
<figureCitation id="5FA8FCA6FFB3DC084D2DFC8E6D64FCC4" box="[601,616,834,859]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">6</figureCitation>
B. Tegmina and wings completely developed, surpassed abdominal apex, rounded apically (
<figureCitation id="5FA8FCA6FFB3DC084E12FCAA6EC8FCE0" box="[358,452,870,895]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="11" pageNumber="52">Fig. 4D</figureCitation>
). Tegmina weakly sclerotized; venation distinct;
<emphasis id="F5E73C31FFB3DC084C83FCAB6B1EFCE1" box="[1015,1042,871,894]" italics="true" pageId="11" pageNumber="52">Sc</emphasis>
thickened (well visible on ventral side of tegmen);
<emphasis id="F5E73C31FFB3DC084E2CFC406E65FC3C" box="[344,361,908,931]" italics="true" pageId="11" pageNumber="52">R</emphasis>
,
<emphasis id="F5E73C31FFB3DC084E0DFC406E9CFC3C" box="[377,400,908,931]" italics="true" pageId="11" pageNumber="52">M</emphasis>
and
<emphasis id="F5E73C31FFB3DC084EBEFC476EF0FC3D" box="[458,508,907,930]" italics="true" pageId="11" pageNumber="52">CuA</emphasis>
stems fused at base;
<emphasis id="F5E73C31FFB3DC084D85FC476C2FFC3D" box="[753,803,907,930]" italics="true" pageId="11" pageNumber="52">CuP</emphasis>
distinct. Anterior margin of fore femur armed as in the
<typeStatus id="18285E81FFB3DC084FE3FC636FCBFC58" box="[151,199,943,967]" pageId="11" pageNumber="52">type</typeStatus>
B, with 67 spines, including 2 apical ones. Fore tibiae not thickened distally. Structure of hind tarsi as described for the genus (see above); arolium about half of the claw length. Abdominal tergites 6th and 7th specialized (
<figureCitation id="5FA8FCA6FFB3DC084E57FC3A6E6BFB90" box="[291,359,1014,1039]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
C, D). 6th tergite with three median membranous areas, medial and paired laterals, median area rounded; lateral ones located along with caudal margin of tergite, with small membranous tubercles (
<figureCitation id="5FA8FCA6FFB3DC084A7FFBD66A43FBAC" box="[1291,1359,1050,1075]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
C). 7th tergite with a larger membranous area, as compared to 6th tergite, the membranous area separated by two sclerotized arms connected in the middle of tergite; a small tripartite membranous lobe situated at the point of contact of two sclerotized arms (
<figureCitation id="5FA8FCA6FFB3DC084D7DFB4A6D41FB00" box="[521,589,1158,1183]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
D). Posterolateral angles of tergites obtuse (
<figureCitation id="5FA8FCA6FFB3DC084B49FB4A6B8EFB00" box="[1085,1154,1158,1183]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
C, D). Anal plate (tergite X) elongated, caudal margin with distinct median incision (
<figureCitation id="5FA8FCA6FFB3DC084C3BFB666C9AFB5C" box="[847,918,1194,1219]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
E). Cerci longer, as compared to
<taxonomicName id="00939BA0FFB3DC084A69FB606A90FB5C" box="[1309,1436,1194,1219]" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="alexeevi" status="sp. nov.">
<emphasis id="F5E73C31FFB3DC084A69FB606A90FB5C" box="[1309,1436,1194,1219]" italics="true" pageId="11" pageNumber="52">R. alexeevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EED4814AFFB3DC084FE3FB036FE5FB78" box="[151,233,1231,1255]" pageId="11" pageNumber="52" rank="species">
<emphasis id="F5E73C31FFB3DC084FE3FB036FE5FB78" bold="true" box="[151,233,1231,1255]" pageId="11" pageNumber="52">sp.nov.</emphasis>
</taxonomicNameLabel>
, flatten, with segments distinctly separated (
<figureCitation id="5FA8FCA6FFB3DC084D8AFB026C4AFB78" box="[766,838,1230,1255]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
E). Paraprocts of blaberid-type, left paraproct with caudally directed sclerotized lamellate outgrowth (
<figureCitation id="5FA8FCA6FFB3DC084D96FB3E6C2BFA94" box="[738,807,1266,1291]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
F, G,
<emphasis id="F5E73C31FFB3DC084C1EFB3E6CA8FA94" box="[874,932,1266,1291]" italics="true" pageId="11" pageNumber="52">l.out.</emphasis>
). Hypandrium nearly symmetrical, with its caudal margin rounded and partly membranous (
<figureCitation id="5FA8FCA6FFB3DC084D91FADA6C3CFAB0" box="[741,816,1302,1327]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
H); a median tooth present (
<figureCitation id="5FA8FCA6FFB3DC084BFBFADA6BD5FAB0" box="[1167,1241,1302,1327]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
H, I,
<emphasis id="F5E73C31FFB3DC084A56FADB6A40FAB1" box="[1314,1356,1303,1326]" italics="true" pageId="11" pageNumber="52">m.t.</emphasis>
); styli symmetrical and fusiform, bent at the base (
<figureCitation id="5FA8FCA6FFB3DC084DF7FAF66DC9FACC" box="[643,709,1338,1363]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
H, I).
</paragraph>
<paragraph id="C72CE023FFB3DC084FB3FA926E58F998" blockId="11.[151,1437,474,1795]" pageId="11" pageNumber="52">
Male genitalia (
<figureCitation id="5FA8FCA6FFB3DC084E0FFA926EC6FAE8" box="[379,458,1374,1399]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Figs. 6</figureCitation>
JP, 11K). Similar to those of
<taxonomicName id="00939BA0FFB3DC084C5CFA936CA9FAE8" box="[808,933,1374,1399]" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="alexeevi" status="sp. nov.">
<emphasis id="F5E73C31FFB3DC084C5CFA936C35FAE9" box="[808,825,1375,1398]" italics="true" pageId="11" pageNumber="52">R</emphasis>
.
<emphasis id="F5E73C31FFB3DC084C3DFA926CA9FAE8" box="[841,933,1374,1399]" italics="true" pageId="11" pageNumber="52">alexeevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EED4814AFFB3DC084CDAFA936B0CFAE8" box="[942,1024,1375,1399]" pageId="11" pageNumber="52" rank="species">
<emphasis id="F5E73C31FFB3DC084CDAFA936B0CFAE8" bold="true" box="[942,1024,1375,1399]" pageId="11" pageNumber="52">sp.nov.</emphasis>
</taxonomicNameLabel>
, but the caudal part of sclerite R1T with cleft (
<figureCitation id="5FA8FCA6FFB3DC084E65FA4E6E58FA04" box="[273,340,1410,1435]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
J,
<emphasis id="F5E73C31FFB3DC084E18FA4E6E84FA04" box="[364,392,1410,1435]" italics="true" pageId="11" pageNumber="52">cl.</emphasis>
), R3 with branches of about equal length, R5 with striations, apical part of sclerite L2D (L1) absent, basal part with distinct &quot;curved upward subsclerite&quot; (
<figureCitation id="5FA8FCA6FFB3DC084C38FA6A6C9CFA20" box="[844,912,1446,1471]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
K, L
<emphasis id="F5E73C31FFB3DC084CB9FA6B6CF8FA21" box="[973,1012,1447,1470]" italics="true" pageId="11" pageNumber="52">u.s.</emphasis>
), large “chaeta-bearing area” located near L2D (
<figureCitation id="5FA8FCA6FFB3DC084E7BFA066E5DFA7C" box="[271,337,1482,1507]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
K,
<emphasis id="F5E73C31FFB3DC084E07FA066EA5FA7C" box="[371,425,1482,1507]" italics="true" pageId="11" pageNumber="52">ch.a.</emphasis>
), sclerite L3 (L2d) without basal subsclerite, &quot;folded structure&quot; distinct, with small bristles (
<figureCitation id="5FA8FCA6FFB3DC084FEBFA226FEDF998" box="[159,225,1518,1543]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
MP,
<emphasis id="F5E73C31FFB3DC084E57FA236E48F998" box="[291,324,1519,1543]" italics="true" pageId="11" pageNumber="52">f.s.</emphasis>
).
</paragraph>
<paragraph id="C72CE023FFB3DC084FB3F9DE6E9CF9B4" blockId="11.[151,1437,474,1795]" box="[199,400,1554,1579]" pageId="11" pageNumber="52">Female unknown.</paragraph>
<paragraph id="C72CE023FFB3DC084FB3F9FB6FCDF9EC" blockId="11.[151,1437,474,1795]" pageId="11" pageNumber="52">
<emphasis id="F5E73C31FFB3DC084FB3F9FB6E7AF9D0" bold="true" box="[199,374,1591,1615]" pageId="11" pageNumber="52">Measurements</emphasis>
(mm). Head length: 2.9, width 3.1, pronotum length 4.4, width 5.5, tegmina length 15.0, width 4.8.
</paragraph>
<paragraph id="C72CE023FFB3DC084FB3F9B26D99F89C" blockId="11.[151,1437,474,1795]" pageId="11" pageNumber="52">
<emphasis id="F5E73C31FFB3DC084FB3F9B26E6DF908" bold="true" box="[199,353,1662,1687]" pageId="11" pageNumber="52">Comparison.</emphasis>
The new species readily differs from all its congeners, excep for
<taxonomicName id="00939BA0FFB3DC084B18F9B36BE6F908" box="[1132,1258,1662,1687]" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="alexeevi" status="sp. nov.">
<emphasis id="F5E73C31FFB3DC084B18F9B36BE6F908" box="[1132,1258,1662,1687]" italics="true" pageId="11" pageNumber="52">R. alexeevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EED4814AFFB3DC084B81F9B36A4BF908" box="[1269,1351,1663,1687]" pageId="11" pageNumber="52" rank="species">
<emphasis id="F5E73C31FFB3DC084B81F9B36A4BF908" bold="true" box="[1269,1351,1663,1687]" pageId="11" pageNumber="52">sp.nov.</emphasis>
</taxonomicNameLabel>
, in the presence of abdominal tergal glands (
<figureCitation id="5FA8FCA6FFB3DC084D4BF96E6D8FF924" box="[575,643,1698,1723]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI VIII— 6 th 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="11" pageNumber="52">Fig. 6</figureCitation>
C, D). From the latter species,
<taxonomicName id="00939BA0FFB3DC084C94F9686B59F924" box="[992,1109,1700,1723]" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="euptera" status="sp. nov.">
<emphasis id="F5E73C31FFB3DC084C94F9686B59F924" box="[992,1109,1700,1723]" italics="true" pageId="11" pageNumber="52">R. euptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EED4814AFFB3DC084B29F96F6BBCF924" box="[1117,1200,1699,1723]" pageId="11" pageNumber="52" rank="species">
<emphasis id="F5E73C31FFB3DC084B29F96F6BBCF924" bold="true" box="[1117,1200,1699,1723]" pageId="11" pageNumber="52">sp.nov.</emphasis>
</taxonomicNameLabel>
differs in having the completely developed tegmina and wings surpassing the abdominal apex, the structure of tergal glands (see under the description above) and the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>