155 lines
15 KiB
XML
155 lines
15 KiB
XML
<document id="9FAB6FC1FC4369DAA257537E8445743D" ID-CLB-Dataset="28054" ID-DOI="10.11646/zootaxa.4508.2.1" ID-GBIF-Dataset="da79b654-bd50-4715-8ed3-80fa7a00d469" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606934" ID-ZooBank="5E2BC894-1919-4F63-8EF5-BAAC91913388" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585631772" checkinUser="plazi" docAuthor="Tang, William, Skelley, Paul & Pérez-Farrera, Miguel Angel" docDate="2018" docId="F95B87D7FFB0951EFF49FA02D7175A1C" docLanguage="en" docName="zootaxa.4508.2.1.pdf" docOrigin="Zootaxa 4508 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceratophila (Ceratophila)" docType="treatment" docVersion="7" lastPageNumber="157" masterDocId="0562FFAFFFB59518FFDEFF8BD26C5B01" masterDocTitle="Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)" masterLastPageNumber="178" masterPageNumber="151" pageNumber="156" updateTime="1698712984586" updateUser="ExternalLinkService">
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<mods:mods id="0D2AA722D9EA1CCF672B69DF9A7AD0A2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B30BEBB1355B62F1975C72B3AEC296D6">
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<mods:title id="A3A266323AAA39048A64788CEB002DCD">Ceratophila, a new genus of erotylid beetles (Erotylidae: Pharaxonothinae) inhabiting male cones of the cycad Ceratozamia (Cycadales: Zamiaceae)</mods:title>
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</mods:titleInfo>
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<mods:name id="0F23FA7E9B5776C1BA153F5F02C9245C" type="personal">
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<mods:role id="6C67BBC6A77C8496BDD85DC992224BED">
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<mods:roleTerm id="7AAA61F58010F809F669D86CF15B49D9">Author</mods:roleTerm>
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<mods:namePart id="38820C88A30DBCB641D3D53BB52F2195">Tang, William</mods:namePart>
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</mods:name>
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<mods:name id="F9DC675C268E9473562DE36A45B47745" type="personal">
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<mods:roleTerm id="83F9976FE9B0AF7EF7B075D1B7CD519E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="26F353470367F54D5F4127B2F6B3CD05">Skelley, Paul</mods:namePart>
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</mods:name>
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<mods:name id="87B351EFBD4D4B9B6B8FEE8C89EFA476" type="personal">
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<mods:role id="3F0F8BC0B718EF8A3438A35CE6239DBD">
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<mods:roleTerm id="A1099B9A7FEDBA5167349838EC1906BA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="430D408A4E08DC6A9B07D332C177FBB8">Pérez-Farrera, Miguel Angel</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="C5A1977F6A15DC67611284F942348BC9">text</mods:typeOfResource>
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<mods:titleInfo id="6C8848C02828FB0CEB9154F6AA2419C3">
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<mods:title id="26349C3BEB2771748D71997F1464E44A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="76727DAF6BD7830BE3ADD8825EA93E27">
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<mods:date id="291C17A5990A01986286528568339B6E">2018</mods:date>
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<mods:detail id="DB680558022C2AFAC84BBEDBD41099A9" type="pubDate">
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<mods:number id="8C04BBB2F8E092FFB04D3EA3E0FDE6E9">2018-10-31</mods:number>
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</mods:detail>
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<mods:detail id="62EB9064BB274D72CF40C298ABC03C58" type="volume">
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<mods:number id="E2190A0974474310B2BE1AAC13D7102D">4508</mods:number>
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</mods:detail>
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<mods:detail id="EB8BDE6CAFE980518D1004B6DB938B01" type="issue">
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<mods:number id="6C3CC65B10247946ED7035B0E5FAFCE7">2</mods:number>
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<mods:start id="CD2FF023C509F6C9BAC529DBEC3CED27">151</mods:start>
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<mods:classification id="51A6F3530EBD51993A43D682DEE0594F">journal article</mods:classification>
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<mods:identifier id="1BC6AC7A676D66F16B17414D4317FE99" type="CLB-Dataset">28054</mods:identifier>
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<mods:identifier id="B15890D89FD0C438D32F2C4BDEB868C6" type="DOI">10.11646/zootaxa.4508.2.1</mods:identifier>
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<mods:identifier id="F6CA684B8A5E50FB21268199433B2392" type="GBIF-Dataset">da79b654-bd50-4715-8ed3-80fa7a00d469</mods:identifier>
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<mods:identifier id="6E00400E66437AD3C04B9C80889DBA71" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="7F0F57D50ADFFBB162C476F62F4D96F5" type="Zenodo-Dep">2606934</mods:identifier>
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<mods:identifier id="35540A124E49C6F659D53C82143777E8" type="ZooBank">5E2BC894-1919-4F63-8EF5-BAAC91913388</mods:identifier>
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</mods:mods>
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<treatment id="F95B87D7FFB0951EFF49FA02D7175A1C" ID-DOI="http://doi.org/10.5281/zenodo.5958001" ID-GBIF-Taxon="154495719" ID-Zenodo-Dep="5958001" LSID="urn:lsid:plazi:treatment:F95B87D7FFB0951EFF49FA02D7175A1C" httpUri="http://treatment.plazi.org/id/F95B87D7FFB0951EFF49FA02D7175A1C" lastPageId="6" lastPageNumber="157" pageId="5" pageNumber="156">
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<subSubSection id="39E8654AFFB0951DFF49FA02D63A5EA2" box="[151,1110,1417,1443]" pageId="5" pageNumber="156" type="nomenclature">
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<paragraph id="714D36C1FFB0951DFF49FA02D63A5EA2" blockId="5.[151,1110,1417,1443]" box="[151,1110,1417,1443]" pageId="5" pageNumber="156">
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<heading id="2A0581ADFFB0951DFF49FA02D63A5EA2" bold="true" box="[151,1110,1417,1443]" fontSize="11" level="1" pageId="5" pageNumber="156" reason="1">
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<emphasis id="4386EAD3FFB0951DFF49FA02D63A5EA2" bold="true" box="[151,1110,1417,1443]" pageId="5" pageNumber="156">
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<taxonomicName id="B6F24D42FFB0951DFF49FA02D3BC5EA2" box="[151,464,1417,1443]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
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<emphasis id="4386EAD3FFB0951DFF49FA02D34B5EA2" bold="true" box="[151,295,1417,1443]" italics="true" pageId="5" pageNumber="156">Ceratophila</emphasis>
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(
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<emphasis id="4386EAD3FFB0951DFEE9FA02D3AA5EA2" bold="true" box="[311,454,1417,1443]" italics="true" pageId="5" pageNumber="156">Ceratophila</emphasis>
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)
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</taxonomicName>
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Tang, Skelley & Pérez-Farrera, nominate subgenus
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="39E8654AFFB0951DFF49FA5BD1F75EE7" box="[151,923,1488,1510]" pageId="5" pageNumber="156" type="materials_examined">
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<paragraph id="714D36C1FFB0951DFF49FA5BD1F75EE7" blockId="5.[151,923,1488,1510]" box="[151,923,1488,1510]" pageId="5" pageNumber="156">
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<emphasis id="4386EAD3FFB0951DFF49FA5BD34E5EE7" bold="true" box="[151,290,1488,1510]" pageId="5" pageNumber="156">
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<typeStatus id="AE498863FFB0951DFF49FA5BD2A05EE7" box="[151,204,1488,1510]" pageId="5" pageNumber="156">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="B6F24D42FFB0951DFEF7FA5BD0C85EE7" authority="Tang & Skelley & Pérez-Farrera, 2018" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[297,676,1488,1510]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="subGenus" status="sp. nov." subGenus="Ceratophila">
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<emphasis id="4386EAD3FFB0951DFEF7FA5BD3C85EE7" box="[297,420,1488,1510]" italics="true" pageId="5" pageNumber="156">Ceratophila</emphasis>
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(
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<emphasis id="4386EAD3FFB0951DFE6CFA5BD0415EE7" box="[434,557,1488,1510]" italics="true" pageId="5" pageNumber="156">Ceratophila</emphasis>
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)
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<emphasis id="4386EAD3FFB0951DFDE2FA5AD0C85EE7" box="[572,676,1489,1510]" italics="true" pageId="5" pageNumber="156">picipennis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="58B557A8FFB0951DFD6EFA5BD0855EE7" box="[688,745,1488,1510]" pageId="5" pageNumber="156" rank="species">
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<emphasis id="4386EAD3FFB0951DFD6EFA5BD0855EE7" bold="true" box="[688,745,1488,1510]" pageId="5" pageNumber="156">n. sp.</emphasis>
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</taxonomicNameLabel>
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, here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="39E8654AFFB0951DFF49F999D0D25D72" pageId="5" pageNumber="156" type="diagnosis">
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<paragraph id="714D36C1FFB0951DFF49F999D0D25D72" blockId="5.[151,1437,1554,2011]" pageId="5" pageNumber="156">
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<emphasis id="4386EAD3FFB0951DFF49F999D3355D2A" bold="true" box="[151,345,1554,1579]" pageId="5" pageNumber="156">Adult diagnosis.</emphasis>
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The subgenus
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<emphasis id="4386EAD3FFB0951DFDDBF998D0745D2B" box="[517,536,1555,1578]" italics="true" pageId="5" pageNumber="156">C</emphasis>
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. (
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<taxonomicName id="B6F24D42FFB0951DFDEFF998D0D45D2A" box="[561,696,1555,1579]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
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<emphasis id="4386EAD3FFB0951DFDEFF998D0D45D2A" box="[561,696,1555,1579]" italics="true" pageId="5" pageNumber="156">Ceratophila</emphasis>
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</taxonomicName>
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) is distinguished from the subgenus
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<emphasis id="4386EAD3FFB0951DFB82F998D6035D2B" box="[1116,1135,1555,1578]" italics="true" pageId="5" pageNumber="156">C</emphasis>
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. (
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<taxonomicName id="B6F24D42FFB0951DFB59F998D6845D2A" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1159,1256,1555,1579]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="172" phylum="Tracheophyta" rank="subGenus" subGenus="Vovidesa">
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<emphasis id="4386EAD3FFB0951DFB59F998D6845D2A" box="[1159,1256,1555,1579]" italics="true" pageId="5" pageNumber="156">Vovidesa</emphasis>
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</taxonomicName>
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) by the convex pronotal surface near lateral carina; lateral carina distinctly thicker anteriorly; metatibia triangularly dilated toward apex; and apparently lacking sexual dimorphism.
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</paragraph>
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</subSubSection>
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<subSubSection id="39E8654AFFB0951DFF19F9F5D7F05CDB" pageId="5" pageNumber="156" type="description">
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<paragraph id="714D36C1FFB0951DFF19F9F5D7F05CDB" blockId="5.[151,1437,1554,2011]" pageId="5" pageNumber="156">
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<emphasis id="4386EAD3FFB0951DFF19F9F5D3CD5D96" bold="true" box="[199,417,1662,1687]" pageId="5" pageNumber="156">Adult description.</emphasis>
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Length
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<quantity id="B60A9B24FFB0951DFDDFF9F5D0CB5D97" box="[513,679,1662,1687]" metricMagnitude="-3" metricUnit="m" metricValue="4.07" metricValueMax="5.11" metricValueMin="3.03" pageId="5" pageNumber="156" unit="mm" value="4.07" valueMax="5.11" valueMin="3.03">3.03–5.11 mm</quantity>
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, width
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<quantity id="B60A9B24FFB0951DFD23F9F5D1C85D97" box="[765,932,1662,1687]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="2.04" metricValueMin="1.26" pageId="5" pageNumber="156" unit="mm" value="1.65" valueMax="2.04" valueMin="1.26">1.26–2.04 mm</quantity>
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. Mouthparts: submentum without long erect or reclining setae. Pronotum: nearly quadrate, length/width = 0.78–0.90; anterior margin near eyes weakly emarginate, anterior angles slightly projecting, anterior angles rounded; surface near lateral carinae not explanate, carinae in dorsal view evenly arcuate; in lateral view thickening anteriorly, 2× thicker than posterior edge; pronotal disc lacking longitudinal groove extending anteriorly from lateral basal pore in margin. Elytra: in most populations often laterally with dark coloration that may extend to disc, maculation merges gradually with lighter ground color anteriad and posteriad; lacking visible setae on surface. Legs: protibia with distal lateral tooth strongly developed in both sexes, apex of tooth with cluster of 3 short stout spinules; meso- and metatibiae moderately angularly dilated at apex which is fringed with short stout spinules. Male genitalia: median lobe and tegmen compressed laterally, penile struts 1.15–1.90× longer than median lobe. Sexual dimorphism absent, except in
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<emphasis id="4386EAD3FFB0951DFB0EF848D68F5CDB" box="[1232,1251,1987,2010]" italics="true" pageId="5" pageNumber="156">C</emphasis>
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.
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<taxonomicName id="B6F24D42FFB0951DFB2FF848D7F85CDA" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[1265,1428,1987,2011]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="156" phylum="Tracheophyta" rank="species" species="chemnicki" subGenus="Ceratophila">
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(
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<emphasis id="4386EAD3FFB0951DFB24F848D7615CDB" box="[1274,1293,1987,2010]" italics="true" pageId="5" pageNumber="156">C</emphasis>
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.)
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<emphasis id="4386EAD3FFB0951DFAFDF848D7F85CDA" box="[1315,1428,1987,2011]" italics="true" pageId="5" pageNumber="156">chemnicki</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="39E8654AFFB3951EFF19FF1CD7175A1C" pageId="6" pageNumber="157" type="discussion">
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<paragraph id="714D36C1FFB3951EFF19FF1CD7175A1C" blockId="6.[151,1436,151,285]" pageId="6" pageNumber="157">
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<emphasis id="4386EAD3FFB3951EFF19FF1CD3585BB1" bold="true" box="[199,308,151,176]" pageId="6" pageNumber="157">Remarks</emphasis>
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.
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<taxonomicName id="B6F24D42FFB3951EFE9AFF13D0035BB1" box="[324,623,152,176]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="6" pageNumber="155" phylum="Tracheophyta" rank="subGenus" subGenus="Ceratophila">
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<emphasis id="4386EAD3FFB3951EFE9AFF13D3A05BB1" box="[324,460,152,176]" italics="true" pageId="6" pageNumber="157">Ceratophila</emphasis>
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(
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<emphasis id="4386EAD3FFB3951EFE00FF13D0095BB1" box="[478,613,152,176]" italics="true" pageId="6" pageNumber="157">Ceratophila</emphasis>
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)
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</taxonomicName>
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is notably different from
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<emphasis id="4386EAD3FFB3951EFC44FF13D1C15BAE" box="[922,941,152,175]" italics="true" pageId="6" pageNumber="157">C</emphasis>
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. (
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<taxonomicName id="B6F24D42FFB3951EFC18FF13D64B5BB1" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[966,1063,152,176]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="6" pageNumber="172" phylum="Tracheophyta" rank="subGenus" subGenus="Vovidesa">
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<emphasis id="4386EAD3FFB3951EFC18FF13D64B5BB1" box="[966,1063,152,176]" italics="true" pageId="6" pageNumber="157">Vovidesa</emphasis>
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</taxonomicName>
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) in an apparent lack of external sexual dimorphism (exception, see remarks under
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<taxonomicName id="B6F24D42FFB3951EFD3AFF37D1165BD5" authorityName="Tang & Skelley & Pérez-Farrera" authorityYear="2018" box="[740,890,188,212]" class="Cycadopsida" family="Zamiaceae" genus="Ceratophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="6" pageNumber="157" phylum="Tracheophyta" rank="species" species="chemnicki">
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<emphasis id="4386EAD3FFB3951EFD3AFF37D1165BD5" box="[740,890,188,212]" italics="true" pageId="6" pageNumber="157">C. chemnicki</emphasis>
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</taxonomicName>
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) and in other noted external characters. Many populations of
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<taxonomicName id="B6F24D42FFB3951EFE9BFF6BD3BA5BF6" box="[325,470,224,247]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="6" pageNumber="157" phylum="Tracheophyta" rank="genus">
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<emphasis id="4386EAD3FFB3951EFE9BFF6BD3BA5BF6" box="[325,470,224,247]" italics="true" pageId="6" pageNumber="157">Ceratozamia</emphasis>
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</taxonomicName>
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appear to have a member of each subgenus on the male cones. The morphological differences between these beetles imply they occupy different niches on the cones, fulfilling different functions.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |