437 lines
48 KiB
XML
437 lines
48 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.889.39090" ID-GBIF-Dataset="f50f6008-500c-492f-84b8-295ba991ec7b" ID-PMC="PMC6872603" ID-Pensoft-Pub="1313-2970-889-81" ID-Pensoft-UUID="0C546DE0BA425539879C1421D8C25A63" ID-PubMed="31777439" ID-ZooBank="7E7A3D196D7040398C087B248A27EB33" ModsDocID="1313-2970-889-81" checkinTime="1574358895821" checkinUser="pensoft" docAuthor="Hendrich, Lars, Shaverdo, Helena, Hajek, Jiri & Balke, Michael" docDate="2019" docId="DD105FE9178258F581598A383B009095" docLanguage="en" docName="ZooKeys 889: 81-152" docOrigin="ZooKeys 889" docSource="http://dx.doi.org/10.3897/zookeys.889.39090" docTitle="Copelatus irregularis W. J. Macleay 1871" docType="treatment" docVersion="4" id="0C546DE0BA425539879C1421D8C25A63" lastPageNumber="81" masterDocId="0C546DE0BA425539879C1421D8C25A63" masterDocTitle="Taxonomic revision of Australian Copelatus Erichson, 1832 (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="152" masterPageNumber="81" pageNumber="81" updateTime="1668168070667" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of Australian Copelatus Erichson, 1832 (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hendrich, Lars</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hajek, Jiri</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>889</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>81</mods:start>
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<mods:end>152</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.889.39090</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.889.39090</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-889-81</mods:identifier>
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<mods:identifier type="ZooBank">7E7A3D196D7040398C087B248A27EB33</mods:identifier>
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<mods:identifier type="Pensoft-UUID">0C546DE0BA425539879C1421D8C25A63</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="160511454" LSID="urn:lsid:plazi:treatment:DD105FE9178258F581598A383B009095" httpUri="http://treatment.plazi.org/id/DD105FE9178258F581598A383B009095" lastPageNumber="81" pageId="0" pageNumber="81">
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<subSubSection pageId="0" pageNumber="81" type="nomenclature">
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<paragraph pageId="0" pageNumber="81">
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<taxonomicName LSID="DD105FE9-1782-58F5-8159-8A383B009095" authority="W. J. Macleay, 1871" authorityName="W. J. Macleay" authorityYear="1871" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus irregularis" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="irregularis">Copelatus irregularis W.J. Macleay, 1871</taxonomicName>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus and colouration of Copelatus irregularis, male. Total length 7.7 mm." figureDoi="10.3897/zookeys.889.39090.figure5" httpUri="https://binary.pensoft.net/fig/356474" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure5">Figures 5</figureCitation>
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,
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Copelatus irregularis A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure14" httpUri="https://binary.pensoft.net/fig/356483" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure14">14</figureCitation>
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,
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<figureCitation captionStart="Figure 23" captionStartId="F22" captionText="Figure 23. Distribution of Copelatus irregularis." figureDoi="10.3897/zookeys.889.39090.figure23" httpUri="https://binary.pensoft.net/fig/356491" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure23">23</figureCitation>
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,
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<figureCitation captionStart="Figure 31" captionStartId="F29" captionText="Figure 31. Habitat of Copelatus irregularis and C. clarki A residual pool of the stream bed of Oakly Creek, Queensland (Loc. 28) B only of C. clarki, Stradbroke Island, temporary sedge swamp near Brown Lake, Queensland. Photographs by L. Hendrich." figureDoi="10.3897/zookeys.889.39090.figure31" httpUri="https://binary.pensoft.net/fig/356498" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure31">31</figureCitation>
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,
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<figureCitation captionStart="Figure 34" captionStartId="F32" captionText="Figure 34. Habitat of Copelatus irregularis A Queensland, Paluma Road 4 km west of Bruce Highway, Maiden Hair Fern Creek, 270 m, shady and rocky residual pools with rotten leaves B of C. irregularis and C. nigrolineatus, Queensland, Winfield, Winfield Road, muddy forest pool, rich in rotten leaves, near the street, 21 m. Photographs by L. Hendrich." figureDoi="10.3897/zookeys.889.39090.figure34" httpUri="https://binary.pensoft.net/fig/356501" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure34">34</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="reference_group">
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<paragraph pageId="0" pageNumber="81">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus irregularis" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="irregularis">Copelatus irregularis</taxonomicName>
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W.J.
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<bibRefCitation author="Macleay, WJ" journalOrPublisher="Transactions of the Entomological Society of New South Wales" pageId="0" pageNumber="81" pagination="79 - 205" refId="B22" refString="Macleay, WJ, 1871. Notes on a collection of insects from Gayndah. Transactions of the Entomological Society of New South Wales 2: 79 - 205" title="Notes on a collection of insects from Gayndah." volume="2" year="1871">Macleay 1871</bibRefCitation>
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: 126 (original description);
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<bibRefCitation DOI="https://doi.org/10.1071/AJZS057" author="Watts, CHS" journalOrPublisher="Australian Journal of Zoology Supplement Series" pageId="0" pageNumber="81" pagination="1 - 166" refId="B36" refString="Watts, CHS, 1978. A revision of the Australian Dytiscidae (Coleoptera). Australian Journal of Zoology Supplement Series 57: 1 - 166" title="A revision of the Australian Dytiscidae (Coleoptera)." url="https://doi.org/10.1071/AJZS057" volume="57" year="1978">Watts 1978</bibRefCitation>
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: 126 (description);
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<bibRefCitation author="Watts, CHS" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="81" pagination="22 - 28" refId="B37" refString="Watts, CHS, 1985. A faunal assessment of Australian Hydradephaga. Proceedings of the Academy of Natural Sciences of Philadelphia 137: 22 - 28" title="A faunal assessment of Australian Hydradephaga." volume="137" year="1985">Watts 1985</bibRefCitation>
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: 26 (general distribution);
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<bibRefCitation author="Larson, DJ" journalOrPublisher="Proceedings of the Royal Society of Queensland" pageId="0" pageNumber="81" pagination="47 - 63" refId="B20" refString="Larson, DJ, 1993. Ecology of tropical Australian Hydradephaga (Insecta: Coleoptera). Part 1. Natural history and distribution of northern Queensland species. Proceedings of the Royal Society of Queensland 103: 47 - 63" title="Ecology of tropical Australian Hydradephaga (Insecta: Coleoptera). Part 1. Natural history and distribution of northern Queensland species." volume="103" year="1993">Larson 1993</bibRefCitation>
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: 52 (habitat information);
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<bibRefCitation DOI="https://doi.org/10.1111/j.1440-6055.1997.tb01469.x" author="Larson, DJ" journalOrPublisher="Australian Journal of Entomology" pageId="0" pageNumber="81" pagination="269 - 285" refId="B21" refString="Larson, DJ, 1997. Habitat and community patterns of tropical Australian Hydradephagan water beetles (Coleoptera: Dytiscidae, Gyrinidae, Noteridae). Australian Journal of Entomology 36: 269 - 285" title="Habitat and community patterns of tropical Australian Hydradephagan water beetles (Coleoptera: Dytiscidae, Gyrinidae, Noteridae)." url="https://doi.org/10.1111/j.1440-6055.1997.tb01469.x" volume="36" year="1997">Larson 1997</bibRefCitation>
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: 275 (habitat information);
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<bibRefCitation author="Hendrich, L" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="81" pagination="43 - 58" refId="B15" refString="Hendrich, L, 2003. Austrodytesplateni sp. nov., and a faunal analysis of the Hydradephaga of the Pilbara region, Western Australia (Coleoptera: Dytiscidae, Gyrinidae, Haliplidae). Koleopterologische Rundschau 73: 43 - 58" title="Austrodytesplateni sp. nov., and a faunal analysis of the Hydradephaga of the Pilbara region, Western Australia (Coleoptera: Dytiscidae, Gyrinidae, Haliplidae)." volume="73" year="2003">Hendrich 2003</bibRefCitation>
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: 51 (new records);
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<bibRefCitation author="Watts, CHS" journalOrPublisher="Identification Ecology Guide" pageId="0" pageNumber="81" pagination="1 - 110" refId="B38" refString="Watts, CHS, 2002. Checklist and guides to the identification, to genus, of adults and larval Australian water beetles of the families Dytiscidae, Noteridae, Hygrobiidae, Haliplidae, Gyrinidae, Hydraenidae and the superfamily Hydrophiloidea (Insecta-Coleoptera). Cooperative Research Centre for Freshwater Ecology (Australia). Identification Ecology Guide 43: 1 - 110" title="Checklist and guides to the identification, to genus, of adults and larval Australian water beetles of the families Dytiscidae, Noteridae, Hygrobiidae, Haliplidae, Gyrinidae, Hydraenidae and the superfamily Hydrophiloidea (Insecta-Coleoptera). Cooperative Research Centre for Freshwater Ecology (Australia)." volume="43" year="2002">Watts 2002</bibRefCitation>
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: 42 (checklist);
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<bibRefCitation author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="81" publicationUrl="http://www.waterbeetles.eu" refId="B28" refString="Nilsson, AN, Hajek, J, 2019. .http://www.waterbeetles.eu" url="http://www.waterbeetles.eu" year="2019">
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Nilsson and
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<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
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2019
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</bibRefCitation>
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: 62 (catalogue).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="type locality">
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<paragraph pageId="0" pageNumber="81">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="81">"[Australia, Queensland] Lizard Island".</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="type material">
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<paragraph pageId="0" pageNumber="81">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="81">ANIC, studied but the exact data not noted.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="materials_examined">
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<paragraph pageId="0" pageNumber="81">Material studied (221 specimens).</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Western Australia:</emphasis>
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8 exs., "WA, E-Pilbara, Weeli Wolli Springs, NW of Newman [555 m],
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<geoCoordinate degrees="22" direction="south" minutes="54.886" orientation="latitude" precision="1" value="-22.914766">22°54.886'S</geoCoordinate>
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,
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<geoCoordinate degrees="119" direction="east" minutes="12.661" orientation="longitude" precision="1" value="119.21101">119°12.661'E</geoCoordinate>
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, 2005, C.H.S. Watts leg.", "DNA Balke 2655", "DNA Balke 2656" [green printed labels] (ZSM); 3 exs., "AUSTRALIA/WA/Shire of Wyndham - East Kimberley, Gibb River Raod, Barnett River Gorge,
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<geoCoordinate degrees="16" direction="south" minutes="32" orientation="latitude" precision="1" seconds="15.17" value="-16.537546">16°32'15.17"S</geoCoordinate>
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,
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<geoCoordinate degrees="126" direction="east" minutes="7" orientation="longitude" precision="1" seconds="30.43" value="126.12512">126°7'30.43"E</geoCoordinate>
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, 450 m, 19.6.1999, Hendrich leg. Loc. 17/117" (CLH); 7 exs., "WA Palm Springs 9/09/2001, JSC" (DPAW); 4 exs., "WA Warrie Springs 8/09/2001 MDS" (DPAW); 4 exs., "AUSTRALIA WA, Mt Fanny 08/07/1969, Giles 50 mi 180,
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<geoCoordinate degrees="25" direction="south" minutes="47" orientation="latitude" precision="925" value="-25.783333">25°47'S</geoCoordinate>
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,
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<geoCoordinate degrees="128" direction="east" minutes="35" orientation="longitude" precision="925" value="128.58333">128°35'E</geoCoordinate>
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, Giuliani D.D. leg." (WAM); 3 exs., "AUSTRALIA WA, Synnot Range 01/09/1969,
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<geoCoordinate degrees="16" direction="south" minutes="25" orientation="latitude" precision="925" value="-16.416666">16°25'S</geoCoordinate>
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,
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<geoCoordinate degrees="125" direction="east" minutes="28" orientation="longitude" precision="925" value="125.46667">125°28'E</geoCoordinate>
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, Giuliani D.D. leg." (WAM); 1 ex., "AUSTRALIA/WA, Gill Pinnacle, 10/07/1969,
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<geoCoordinate degrees="24" direction="south" minutes="54" orientation="latitude" precision="925" value="-24.9">24°54'S</geoCoordinate>
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,
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<geoCoordinate degrees="128" direction="east" minutes="47" orientation="longitude" precision="925" value="128.78334">128°47'E</geoCoordinate>
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, Giuliani D.D. leg." (WAM); 2 exs., "AUSTRALIA/WA, Charnley River, 25 mi E Beverley Springs, 09/09/1969,
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<geoCoordinate degrees="16" direction="south" minutes="15" orientation="latitude" precision="925" value="-16.25">16°15'S</geoCoordinate>
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,
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<geoCoordinate degrees="125" direction="east" minutes="26" orientation="longitude" precision="925" value="125.433334">125°26'E</geoCoordinate>
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, Giuliani D.D. leg." (WAM); 4 exs., "AUSTRALIA/ WA, Nullagine [
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<geoCoordinate degrees="21.8833" direction="south" orientation="latitude" precision="5" value="-21.8833">-21.8833</geoCoordinate>
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,
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<geoCoordinate degrees="120.1166" direction="east" orientation="longitude" precision="5" value="120.1166">120.1166</geoCoordinate>
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] 19.-20.1.1974, Douglas A.M. & M.J. leg." (WAM); 2 exs., "AUSTRALIA/WA, Marandoo Camp, at light,
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<geoCoordinate degrees="22" direction="south" minutes="38" orientation="latitude" precision="925" value="-22.633333">22°38'S</geoCoordinate>
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,
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<geoCoordinate degrees="118" direction="east" minutes="06" orientation="longitude" precision="925" value="118.1">118°06'E</geoCoordinate>
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, 5.-19.5.1980, Houston, T.F. et al. leg." (WAM); 1 ex., "AUSTRALIA/WA, Lyons River, Mt Augustus, at light,
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<geoCoordinate degrees="24" direction="south" minutes="19" orientation="latitude" precision="15" seconds="12" value="-24.320002">24°19'12"S</geoCoordinate>
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,
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<geoCoordinate degrees="116" direction="east" minutes="49" orientation="longitude" precision="15" seconds="48" value="116.83">116°49'48"E</geoCoordinate>
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, 2.-3-IX.1980, Howard, C.A. & Houston, T.F. leg." (WAM); 1 ex., "Western Australia, Pilbara, Ashburton (S), Millstream, Deep Reach [
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<geoCoordinate degrees="21.58333" direction="south" orientation="latitude" precision="1" value="-21.58333">-21.58333</geoCoordinate>
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,
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<geoCoordinate degrees="117.0667" direction="east" orientation="longitude" precision="5" value="117.0667">117.0667</geoCoordinate>
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], 8.11.1970, Britton, E.B." (ANIC); 1 ex., "Australia, WA, Pilbara, Hamersley Range, 17 km S Auski Roadhouse, Fig. Tree Crossing 400 m, 28.8.2002,
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<geoCoordinate degrees="22" direction="south" minutes="32" orientation="latitude" precision="925" value="-22.533333">22°32'S</geoCoordinate>
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,
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<geoCoordinate degrees="118" direction="east" minutes="44" orientation="longitude" precision="925" value="118.73333">118°44'E</geoCoordinate>
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, Hendrich leg. (Loc. WA 9/173)" (CLH); 2 exs., "Australia, WA, Pilbara, Hamersley Range, Karijini N.P., Dales Gorge [Fortescue Falls and Circular Pool], 400 m, 28.8.2002,
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<geoCoordinate degrees="22" direction="south" minutes="29" orientation="latitude" precision="925" value="-22.483334">22°29'S</geoCoordinate>
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,
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<geoCoordinate degrees="118" direction="east" minutes="35" orientation="longitude" precision="925" value="118.583336">118°35'E</geoCoordinate>
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, Hendrich leg. (Loc. WA 10/174)" (CLH); 2 exs., "Australia, WA, Pilbara, Hamersley Range, Karijini N.P., Kalamina Gorge, 450 m, 29.8.2002,
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<geoCoordinate degrees="22" direction="south" minutes="25" orientation="latitude" precision="925" value="-22.416666">22°25'S</geoCoordinate>
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,
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<geoCoordinate degrees="118" direction="east" minutes="23" orientation="longitude" precision="925" value="118.38333">118°23'E</geoCoordinate>
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, Hendrich leg. (Loc. WA 11/175)" (CLH); 1 ex., "Australia, WA, Pilbara, Hamersley Range, Karijini N.P., Knox Gorge, 450 m, 29.8.2002,
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<geoCoordinate degrees="22" direction="south" minutes="21" orientation="latitude" precision="925" value="-22.35">22°21'S</geoCoordinate>
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,
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<geoCoordinate degrees="118" direction="east" minutes="18" orientation="longitude" precision="925" value="118.3">118°18'E</geoCoordinate>
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, Hendrich leg. (Loc. WA 12/176)" (CLH). 1 ex., "W AUSTRALIA, E-Pilbara, Weeli Wolli Springs,
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<geoCoordinate degrees="22" direction="south" minutes="54.886" orientation="latitude" precision="1" value="-22.914766">22°54.886'S</geoCoordinate>
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,
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<geoCoordinate degrees="119" direction="east" minutes="12.661" orientation="longitude" precision="1" value="119.21101">119°12.661'E</geoCoordinate>
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, 555 m, xi.2001, M.
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<normalizedToken originalValue="Häckel">Haeckel</normalizedToken>
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leg." (NMPC).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Northern Territory</emphasis>
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: 12 exs., "Australia NT, Trephina Gorge, East MacDonnell Ranges [
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<geoCoordinate degrees="23.529" direction="south" orientation="latitude" precision="55" value="-23.529">-23.529</geoCoordinate>
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,
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<geoCoordinate degrees="134.375" direction="east" orientation="longitude" precision="55" value="134.375">134.375</geoCoordinate>
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], October 1972, leg. M. Baehr" (CLH, ZSM); 1 ex., "Australia, NT, Litchfield NP, Florence Falls,
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<geoCoordinate degrees="12" direction="south" minutes="51" orientation="latitude" precision="15" seconds="15" value="-12.854167">12°51'15"S</geoCoordinate>
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,
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<geoCoordinate degrees="132" direction="east" minutes="45" orientation="longitude" precision="15" seconds="16" value="132.75444">132°45'16"E</geoCoordinate>
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, 63 m, at light, 12.V.2006, leg. Berger & Dostal (5/06)" (CGW); 2 exs., "Australia, NT, 80 km W Roper Bar,
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<geoCoordinate degrees="14" direction="south" minutes="54" orientation="latitude" precision="15" seconds="19" value="-14.905277">14°54'19"S</geoCoordinate>
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,
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<geoCoordinate degrees="133" direction="east" minutes="57" orientation="longitude" precision="15" seconds="28" value="133.95778">133°57'28"E</geoCoordinate>
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, 78 m, at light, 14.V.2006, leg. Berger & Dostal (7/06)" (CGW); 1 ex., "Australia, NT, 40 km v. Port Roper,
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<geoCoordinate degrees="14" direction="south" minutes="54" orientation="latitude" precision="1" seconds="04.8" value="-14.901333">14°54'04.8"S</geoCoordinate>
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,
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<geoCoordinate degrees="135" direction="east" minutes="03" orientation="longitude" precision="1" seconds="24.7" value="135.05687">135°03'24.7"E</geoCoordinate>
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, 134 m, 15.V.2006, leg. Berger & Dostal" (CGW); 1 male, "Australia, N.T./Kakadu N.P., Jim Jim District, Barramundie Gorge, Maguk, 50 m, 31.10.1996,
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<geoCoordinate degrees="13" direction="south" minutes="18.823" orientation="latitude" precision="1" value="-13.313717">13°18.823'S</geoCoordinate>
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,
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<geoCoordinate degrees="132" direction="east" minutes="26.198" orientation="longitude" precision="1" value="132.43663">132°26.198'E</geoCoordinate>
|
||
. L. Hendrich leg./ Lok. 9" (NMW); 8 males, 4 females, "Australia, N.T./Kakadu N.P., Gunlom Camp. Area, pool in Monsoonal Forest, ca. 50 m, 3.11.1996,
|
||
<geoCoordinate degrees="13" direction="south" minutes="26.082" orientation="latitude" precision="1" value="-13.4347">13°26.082'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="132" direction="east" minutes="24.929" orientation="longitude" precision="1" value="132.41548">132°24.929'E</geoCoordinate>
|
||
L. Hendrich leg./Lok. 15" (NMW); 1 male "Thorey 1867" [partly hw], "Nov: Holl: bor:" [hw], "Coll. Mus. Vindob." (NHW). 2 exs., "Australia: NT, Litchfield NP, TJAYNERA FALLS,
|
||
<geoCoordinate degrees="13" direction="south" minutes="15" orientation="latitude" precision="925" value="-13.25">13°15'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="130" direction="east" minutes="44" orientation="longitude" precision="925" value="130.73334">130°44'E</geoCoordinate>
|
||
, 63 m, S.
|
||
<normalizedToken originalValue="Jákl">Jakl</normalizedToken>
|
||
leg., 20
|
||
<normalizedToken originalValue="–27.XI.2008”">-27.XI.2008"</normalizedToken>
|
||
(NMPC); 1 ex., "Australia: NT, Victoria Highway, 110 km to KUNUNURRA,
|
||
<geoCoordinate degrees="15" direction="south" minutes="57" orientation="latitude" precision="925" value="-15.95">15°57'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="129" direction="east" minutes="33" orientation="longitude" precision="925" value="129.55">129°33'E</geoCoordinate>
|
||
, 76 m, S.
|
||
<normalizedToken originalValue="Jákl">Jakl</normalizedToken>
|
||
leg., 30.XI.2008" (NMPC); 1 ex., "Australia: NT, Nitmiluk NP, Edit Falls,
|
||
<geoCoordinate degrees="14" direction="south" minutes="10" orientation="latitude" precision="925" value="-14.166667">14°10'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="132" direction="east" minutes="06" orientation="longitude" precision="925" value="132.1">132°06'E</geoCoordinate>
|
||
, 37 m, S.
|
||
<normalizedToken originalValue="Jákl">Jakl</normalizedToken>
|
||
leg., 3.XII.2008" (NMPC); 1 ex., "Australia: NT, 70 km SW of Mataranka,
|
||
<geoCoordinate degrees="15" direction="south" minutes="1" orientation="latitude" precision="925" value="-15.016666">15°1'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="132" direction="east" minutes="50" orientation="longitude" precision="925" value="132.83333">132°50'E</geoCoordinate>
|
||
, 190 m, S.
|
||
<normalizedToken originalValue="Jákl">Jakl</normalizedToken>
|
||
leg., 22.
|
||
<normalizedToken originalValue="–23.XII.2008”">-23.XII.2008"</normalizedToken>
|
||
(NMPC); 2 exs., "Australia NT, 25 km S of Katherine, 168 m,
|
||
<geoCoordinate degrees="14" direction="south" minutes="31" orientation="latitude" precision="925" value="-14.516666">14°31'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="132" direction="east" minutes="25" orientation="longitude" precision="925" value="132.41667">132°25'E</geoCoordinate>
|
||
, 16.-17.1.2009, Sv.
|
||
<normalizedToken originalValue="Bílý">Bily</normalizedToken>
|
||
leg." (NMPC).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Queensland</emphasis>
|
||
: 2 exs., "AUSTRALIA, QLD. Mulgrave R, nr The Fisheries Nov. 9/90 Larson" (ZSM); 3 exs., "AUSTRALIA QLD. McLeod R nr base Windsor Tableland Nov. 12/90 Larson" (ZSM); 4 exs., "Australia S QLD, N Stradbroke Island, creek @Trans-island Rd. 1 km SE Brown Lake, 18.X.2014,
|
||
<geoCoordinate degrees="27" direction="south" minutes="30" orientation="latitude" precision="1" seconds="8.43" value="-27.502342">27°30'8.43"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="153" direction="east" minutes="26" orientation="longitude" precision="1" seconds="10.78" value="153.43633">153°26'10.78"E</geoCoordinate>
|
||
, Lars Hendrich leg. (QLD2/14)" (NMW); 1 ex., "QL, 35-47 km SWW Mt. Gamet, 31.I.-4.II.2012, leg. S. Prebsl" (CGW); 11 exs., "Australia, QL, Ravenshoe, 90 km W Innisfail, 900 m, 20.I.1993, leg. G. Wewalka (16)" (CGW); 18 exs., "Australia, QL, Greenvale, 150 km NW Charters Towers, 600 m, 19.I.1993, leg. G. Wewalka (12
|
||
<normalizedToken originalValue="–14)”">-14)"</normalizedToken>
|
||
(CGW); 6 exs., "Australia QLD/30 km NNW Mareeba, near Mitchell Lake, 9.11. 1996 Hendrich leg./Lok. 20" (CLH); 6 exs., "Australia: S QLD, 40 km E Bundaberg, Tusky Creek, 9 m, 26.IX.2006, 24.39.139S 152.01.477E, L. & E. Hendrich leg. (QLD 52)" (ZSM); 6 exs., "Australia: C QLD, 18 km S Calen, Mt. Charlton-Calen Road, Boulder Creek, 42 m, 23.IX.2006, 21.00.365S 148.43.231E, L. & E. Hendrich leg. (QLD 45)" (ZSM); 3 exs., "Australia: C QLD, 10 km S Mizani, Lake Kinchant, seapage, 48 m, 24.IX.2006, 21.11.580S 148.53.522E, L. & E. Hendrich leg. (QLD 46)" (ZSM); 11 exs., "Australia: C QLD, Paluma Road 4 km W Bruce Hwy, Maiden Hair Fern Crk., 270 m, 20.IX.2006, 19.00.162S 146.17.070E, L. & E. Hendrich leg. (QLD 40)", 4 specimens with "DNA Balke 1863", "DNA Balke 1864", "DNA Balke 2140", "DNA Balke 2141" [green printed labels] (CLH, ZSM); 2 exs., "Australia: S QLD, Winfield, Winfield Road, forest pool, 21 m, 26.IX.2006, 24.34.084S 152.00.513E, L. & E. Hendrich leg. (QLD 54)" (CLH); 1 ex., "Queensland, Pt. Denison [Port Denison, now Bowen -20.05, 148.25]" (ANIC); 1 ex., "Queensland, North Burnett (R) Gayndah, [-25.63333333, 151.6]" (ANIC); 1 ex., "Queensland Tablelands, N side of Lake Tinaroo, 9.11.1966 [-17.16667, 145.55], Britton, E.B." (ANIC). 1 ex., "Queensland, Tablelands, Mt. Lewis, tin working site, 3.12.1968 [
|
||
<geoCoordinate degrees="16.58333015" direction="south" orientation="latitude" precision="1" value="-16.58333">-16.58333015</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="145.2832947" direction="east" orientation="longitude" precision="1" value="145.2833">145.2832947</geoCoordinate>
|
||
], Britton, E.B. & Misko, S." (ANIC); 1 ex., "Queensland, Townsville (C), Mt. Spec, 5.1.1965, -18.95, 146.1833333, Brooks, J.G." (ANIC); 1 ex., "Queensland Rockhampton (R) Pistol Gap, Byfield, 10.1.1970 [
|
||
<geoCoordinate degrees="22.83333333" direction="south" orientation="latitude" precision="1" value="-22.833334">-22.83333333</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="150.6666667" direction="east" orientation="longitude" precision="1" value="150.66667">150.6666667</geoCoordinate>
|
||
], Britton, Holloway & Misko" (ANIC); 1 ex., "Queensland, 9 miles W of Paluma, Charters Towers (R), 15.4.1969 [-19.1, 146.0667], Common, I.F.B. & Upton, M.S." (ANIC); 1 ex., "North Queensland, Magnetic Island [
|
||
<geoCoordinate degrees="19.13333" direction="south" orientation="latitude" precision="1" value="-19.13333">-19.13333</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="146.8333" direction="east" orientation="longitude" precision="5" value="146.8333">146.8333</geoCoordinate>
|
||
], Lea, A.M." (ANIC); 1 ex., "Queensland Tablelands (R) Mary Creek [-16.55, 145.2], 4.12.1968 Britton, E.B. & Misko, S." (ANIC); 1 ex., "Queensland, Townsville (general), 26.3.1968 [
|
||
<geoCoordinate degrees="19.26667023" direction="south" orientation="latitude" precision="1" value="-19.26667">-19.26667023</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="146.8166656" direction="east" orientation="longitude" precision="1" value="146.81667">146.8166656</geoCoordinate>
|
||
] Ferrar, P." (ANIC); 1 ex., "Queensland, Cardstone, 16.12.1965 [
|
||
<geoCoordinate degrees="17.76666667" direction="south" orientation="latitude" precision="1" value="-17.766666">-17.76666667</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="145.5833333" direction="east" orientation="longitude" precision="1" value="145.58333">145.5833333</geoCoordinate>
|
||
] Hyde, K." (ANIC); 1 ex., "Queensland, Townsville (general) [
|
||
<geoCoordinate degrees="19.26667023" direction="south" orientation="latitude" precision="1" value="-19.26667">-19.26667023</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="146.8166656" direction="east" orientation="longitude" precision="1" value="146.81667">146.8166656</geoCoordinate>
|
||
], 21.12.1967, Ferrar, P." (ANIC); 1 ex., "Queensland Tablelands (R), Mary Creek, 5.12.1968 [-16.55, 145.2] Britton, E.B. & Misko, S." (ANIC); 1 ex., "QLD, Magnetic Island [-19.13, 146.83], A.M. Lea",
|
||
<normalizedToken originalValue="“K.215158”">"K.215158"</normalizedToken>
|
||
(AMS); 1 ex., "Queensland [
|
||
<geoCoordinate degrees="22.82407" direction="south" orientation="latitude" precision="1" value="-22.82407">-22.82407</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="147.63635" direction="east" orientation="longitude" precision="1" value="147.63635">147.63635</geoCoordinate>
|
||
] K.K. Spence",
|
||
<normalizedToken originalValue="“K.215159”">"K.215159"</normalizedToken>
|
||
(AMS); 1 ex., "Queensland [
|
||
<geoCoordinate degrees="23.37805" direction="south" orientation="latitude" precision="1" value="-23.37805">-23.37805</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="150.51361" direction="east" orientation="longitude" precision="1" value="150.51361">150.51361</geoCoordinate>
|
||
] T.G. Sloane",
|
||
<normalizedToken originalValue="“K.215160”">"K.215160"</normalizedToken>
|
||
(AMS); 1 ex., "Queensland Magnetic Island [-19.13, 146.83] A.M. Lea",
|
||
<normalizedToken originalValue="“K.215161”">"K.215161"</normalizedToken>
|
||
(AMS); 1 ex., "Queensland Hayman Island [
|
||
<geoCoordinate degrees="20.0508" direction="south" orientation="latitude" precision="5" value="-20.0508">-20.0508</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="148.88783" direction="east" orientation="longitude" precision="1" value="148.88783">148.88783</geoCoordinate>
|
||
] Jan 1935, F.A. McNeil",
|
||
<normalizedToken originalValue="“K.215162”">"K.215162"</normalizedToken>
|
||
(AMS); 1 ex., "Australia Queensland [
|
||
<geoCoordinate degrees="22.82407" direction="south" orientation="latitude" precision="1" value="-22.82407">-22.82407</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="147.63635" direction="east" orientation="longitude" precision="1" value="147.63635">147.63635</geoCoordinate>
|
||
],
|
||
<normalizedToken originalValue="“K.215163”">"K.215163"</normalizedToken>
|
||
(AMS); 1 ex., "Australia Queensland, The Rock Pool, Carnarvon NP [
|
||
<geoCoordinate degrees="25.06756" direction="south" orientation="latitude" precision="1" value="-25.06756">-25.06756</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="148.246259" direction="east" orientation="longitude" precision="1" value="148.24626">148.246259</geoCoordinate>
|
||
] 26 Apr 2006, at MV lamp, D.R. Britton, J.R. Weiner,
|
||
<normalizedToken originalValue="“K.363072”">"K.363072"</normalizedToken>
|
||
(AMS); 1 ex., "Australia, Queensland, Tewah Creek via Tin Can Bay, 17. October 1970, T. Weir" (QM); 1 ex., "Australia, Queensland, Baldy Mt. Road, 6 miles SW Atherton, 1100 m, 27. December 1972, B.K.Cantrell" (QM); 1 ex., "Australia, Queensland, Bald Hills Station 4 km N of Isabella Falls, 15°15' 145°00', 29. December 1984, mv lamp, G. & A. Daniels" (QM); 2 exs., "Australia, Queensland, Kenilworth State Forest, via Kenilworth, 20. October 1972, B.K. & J.A. Cantrall" (QM); 4 exs. "Australia, Queensland, Davies Creek via Mareeba, 27. Nov. 1981, R.I. Storey" (QDPIB); 2 exs., "Australia Queensland, Walkamin, 20. March 1984, light trap, J.D. Brown" (QDPIB); 1 ex., "Australia, Queensland Danbulla S.F. via Yungaburra, 13. February 1992, at light, Storey, De Faveri & Huwer" (QDPIB); 1 ex., "Australia, Queensland, 10. Dec. 1985, 15 km WNW of South Johnstone, light trap, Fay & Halfpapp" (QDPIB); 1 ex., "Australia, Queensland, 9. IX. 1985, 15 km WNW of South Johnstone, light trap, Fay & Halfpapp" (QDPIB); 4 exs., "Australia, Queensland, 23 km E of Mareeba, 29. January 1989, at light, R.I. Storey" (QDPIB); 2 exs., "Australia, Queensland, Tolga, 10. December 1982, light trap, J.D. Brown" (QDPIB); 1 ex., "Australia, Queensland", Paluma, 9.-13. January 1989, light trap, rainforest, R.I. Storey (QDPIB); 1 ex., "Australia, Queensland, Windsor Tableland, 16. December 1984, at light, R.I. Storey (QDPIB); 1 ex., "Australia, Queensland, Garradunga, 17. December 1992, J. Hasenpusch" (QDPIB); 1 ex., "Australia, Queensland, Mt. Mulligen Plateau, 700 m, 15.-19. IV. 1985, at light, K.H. Halfpapp" (QDPIB); 1 ex., "Australia, Queensland, Yam Island 24-27 March 1988, at light, Fay, Halfpapp" (QDPIB); 1 ex., "Australia, Queensland, Pinnarendi Station 60 km W of Mt. Garnet, 7. February 1989, D. Heiner" (QDPIB); 3 exs., "N Queensland, 15.1.2000, Davis Creek, Sv.
|
||
<normalizedToken originalValue="Bílý">Bily</normalizedToken>
|
||
leg." (NMPC); 1 ex., "N Queensland, 29.1.-3.2.2000, The Lynd Junction, Sv.
|
||
<normalizedToken originalValue="Bílý">Bily</normalizedToken>
|
||
leg." (NMPC); 6 exs., "N Queensland, 5.2.2000, Ravenshoe, Sv.
|
||
<normalizedToken originalValue="Bílý">Bily</normalizedToken>
|
||
leg." (NMPC); 3 exs., "N Queensland, 13.2.2000, Undara, Sv.
|
||
<normalizedToken originalValue="Bílý">Bily</normalizedToken>
|
||
leg." (NMPC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="doubtful record">
|
||
<paragraph pageId="0" pageNumber="81">Doubtful record.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">1 ex., "Tasmania, Gladstone, Dorset, A.M. Lea" (SAMA), probably mislabelled.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="description">
|
||
<paragraph pageId="0" pageNumber="81">Description of male.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Body shape:</emphasis>
|
||
In dorsal view, oblong oval, broadest at midlength of elytra, moderately convex. Body outline almost continuous, with slight discontinuity between pronotum and elytra. Head relatively broad; anterior margin of clypeus not bordered. Pronotum broadest just before posterior angles, lateral margins moderately curved. Base of elytra as broad as pronotal base; lateral margins of elytra moderately curved (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus and colouration of Copelatus irregularis, male. Total length 7.7 mm." figureDoi="10.3897/zookeys.889.39090.figure5" httpUri="https://binary.pensoft.net/fig/356474" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure5">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.889.39090.figure5" httpUri="https://binary.pensoft.net/fig/356474" pageId="0" pageNumber="81" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" pageId="0" pageNumber="81">Figure 5.</emphasis>
|
||
Habitus and colouration of
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus irregularis" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="irregularis">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">Copelatus irregularis</emphasis>
|
||
</taxonomicName>
|
||
, male. Total length 7.7 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Colouration:</emphasis>
|
||
Body black, most of clypeus, sides of pronotum, base and tip of elytra, appendages and much of ventral surface testaceous (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus and colouration of Copelatus irregularis, male. Total length 7.7 mm." figureDoi="10.3897/zookeys.889.39090.figure5" httpUri="https://binary.pensoft.net/fig/356474" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure5">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Dorsal surface sculpture:</emphasis>
|
||
Whole surface shiny (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus and colouration of Copelatus irregularis, male. Total length 7.7 mm." figureDoi="10.3897/zookeys.889.39090.figure5" httpUri="https://binary.pensoft.net/fig/356474" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure5">Fig. 5</figureCitation>
|
||
). Head uniformly microreticulated, reticulation composed of moderately deeply impressed isodiametric very small meshes. Punctation composed of very small punctures sparsely spread on surface; rows of deep and coarse punctures present around inner margin of eyes and in small depression anterolaterally of eyes. Pronotum with some moderately strong, short striae, more dense and weak laterally; lateral beading of pronotum very thin and indistinct. Microreticulation and punctation similar to that of head; row of coarse setigerous punctures present along anterior margin, basal margin (except basomedially), and laterally close to sides. Elytra with microreticulation similar to that of head and pronotum, but less impressed. Punctation consisting of very fine sparse punctures. Each elytron with 11 strongly impressed discal and one submarginal longitudinal striae, intervals subequal, alternate striae tending to be shorter apically; submarginal stria reaching from little behind middle of elytron almost to end of stria 10. Serial punctures on elytron untraceable.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Antennae and legs:</emphasis>
|
||
Antenna with antennomeres long and slender. Protibia modified, distinctly broadened anteriorly (2/3rd) and strongly narrowed basally (1/3rd). Pro- and mesotarsomeres 1-3 distinctly broadened, with adhesive discs on their ventral side; claws simple.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Ventral part:</emphasis>
|
||
Finely microreticulated, with intermixed, sparsely distributed, very small punctures. Prosternal process rather flat, distinctly bordered laterally, weakly and bluntly pointed. Lateral parts of metaventrite tongue-shaped, very slender. Metacoxal lines close, well-marked and moderately divergent anteriorly. Metacoxae with some short and deep striae, abdominal ventrites 1-3 with numerous longitudinal striae.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Male genitalia:</emphasis>
|
||
Median lobe consisting of a few sclerites, well separated apically (
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Copelatus irregularis A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure14" httpUri="https://binary.pensoft.net/fig/356483" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure14">Fig. 14A, B, C</figureCitation>
|
||
). Shape of paramere narrowly triangular, with very dense, strong, long setae along dorsal margin (
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Copelatus irregularis A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure14" httpUri="https://binary.pensoft.net/fig/356483" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure14">Fig. 14D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="female">
|
||
<paragraph pageId="0" pageNumber="81">Female.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">Similar to male in habitus. Pro- and mesotarsomeres not broadened, without adhesive setae. Pronotal striae fine and dense.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="description">
|
||
<paragraph pageId="0" pageNumber="81">Measurements.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">TL = 6.8-7.8 mm; TL-H = 6.0-6.9 mm; MW = 3.2-3.4 mm.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="variability">
|
||
<paragraph pageId="0" pageNumber="81">Variability.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">All specimens studied are rather uniform in shape and size but vary in the extension of the testaceous elytral markings.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="differential diagnosis">
|
||
<paragraph pageId="0" pageNumber="81">Differential diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
The species is close to
|
||
<taxonomicName lsidName="C. bakewelli" pageId="0" pageNumber="81" rank="species" species="bakewelli">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. bakewelli</emphasis>
|
||
</taxonomicName>
|
||
but can be easily separated by the larger size, the dorsal colouration, and the form of the median lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="distribution">
|
||
<paragraph pageId="0" pageNumber="81">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
The species is widely distributed in the northern half of Australia. Records are from the Northern Territory (inland to the East MacDonnell Ranges), north-western Australia (Kimberley region and the Pilbara), and Queensland south to Brisbane (
|
||
<figureCitation captionStart="Figure 23" captionStartId="F22" captionText="Figure 23. Distribution of Copelatus irregularis." figureDoi="10.3897/zookeys.889.39090.figure23" httpUri="https://binary.pensoft.net/fig/356491" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure23">Fig. 23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
According to the literature,
|
||
<taxonomicName lsidName="C. irregularis" pageId="0" pageNumber="81" rank="species" species="irregularis">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. irregularis</emphasis>
|
||
</taxonomicName>
|
||
occurs also in New Guinea, cited in
|
||
<bibRefCitation author="Guignot, F" journalOrPublisher="Annales Historico- Naturales Musei Nationalis Hungarici (Series Nova)" pageId="0" pageNumber="81" pagination="51 - 60" refId="B11" refString="Guignot, F, 1956. Dytiscides recoltes par le Dr. L. Biro en Nouvelle Guinee et dans l'Ile de Java (Coleoptera). Annales Historico- Naturales Musei Nationalis Hungarici (Series Nova) 7: 51 - 60" title="Dytiscides recoltes par le Dr. L. Biro en Nouvelle Guinee et dans l'Ile de Java (Coleoptera)." volume="7" year="1956">Guignot (1956</bibRefCitation>
|
||
: 55) and
|
||
<bibRefCitation author="Zimmermann, A" journalOrPublisher="Junk, Berlin" pageId="0" pageNumber="81" refId="B40" refString="Zimmermann, A, 1920. Pars 71. Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: Schenkling S (Ed.) Coleopterorum Catalogus, Vol. IV. Junk, Berlin" title="Pars 71. Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: Schenkling S (Ed.) Coleopterorum Catalogus, Vol. IV." year="1920">Zimmermann (1920</bibRefCitation>
|
||
: 140); Papua in
|
||
<bibRefCitation author="Gueorguiev, VB" journalOrPublisher="Izvestija na Zoologitjeskija Institut s Musei Sofia" pageId="0" pageNumber="81" pagination="5 - 45" refId="B9" refString="Gueorguiev, VB, 1968. Essai de classification des coleopteres Dytiscidae. I. Tribus Copelatini (Colymbetinae). Izvestija na Zoologitjeskija Institut s Musei Sofia 28: 5 - 45" title="Essai de classification des coleopteres Dytiscidae. I. Tribus Copelatini (Colymbetinae)." volume="28" year="1968">
|
||
<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
|
||
(1968
|
||
</bibRefCitation>
|
||
: 9); Papua New Guinea: Central Province: Redscar Bay in
|
||
<bibRefCitation author="Regimbart, M" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="81" pagination="186 - 367" refId="B32" refString="Regimbart, M, 1899. Revision des Dytiscidae de la region Indo-Sino-Malaise. Annales de la Societe Entomologique de France 68: 186 - 367" title="Revision des Dytiscidae de la region Indo-Sino-Malaise." volume="68" year="1899">
|
||
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
|
||
(1899
|
||
</bibRefCitation>
|
||
: 306) and
|
||
<bibRefCitation author="Gueorguiev, VB" journalOrPublisher="Frustula Entomologica (Nuova Serie)" pageId="0" pageNumber="81" pagination="147 - 166" refId="B10" refString="Gueorguiev, VB, Rocchi, S, 1993. Contributo alla conoscenza dei Dytiscidae della Nuova Guinea (Coleoptera). Frustula Entomologica (Nuova Serie) 15: 147 - 166" title="Contributo alla conoscenza dei Dytiscidae della Nuova Guinea (Coleoptera)." volume="15" year="1993">
|
||
<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
|
||
and Rocchi (1993
|
||
</bibRefCitation>
|
||
: 158). However, all these records are in need of confirmation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="habitat">
|
||
<paragraph pageId="0" pageNumber="81">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
This is a widely distributed species and one of the most common
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">Copelatus</emphasis>
|
||
</taxonomicName>
|
||
in tropical northern Australia. It can be found in almost all habitat types, preferably in forested areas. Most specimens were obtained from shallow water, amongst leaves and plant debris, of isolated pools of seasonal creeks and streams, and pools adjacent to streams in Eucalypt or tropical woodland. Also, a few specimens were taken from slow flowing spring-fed streams. In the Pilbara (Hendrich 2002),
|
||
<taxonomicName lsidName="C. irregularis" pageId="0" pageNumber="81" rank="species" species="irregularis">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. irregularis</emphasis>
|
||
</taxonomicName>
|
||
was found in different isolated and half-shaded rocky and sandy pools (10-20 m², up to 1.5 m depth) of an intermittent stream. The bottom of most habitats consisted of sand and stones, with a thin layer of mud and plant debris (
|
||
<figureCitation captionStart="Figure 31" captionStartId="F29" captionText="Figure 31. Habitat of Copelatus irregularis and C. clarki A residual pool of the stream bed of Oakly Creek, Queensland (Loc. 28) B only of C. clarki, Stradbroke Island, temporary sedge swamp near Brown Lake, Queensland. Photographs by L. Hendrich." figureDoi="10.3897/zookeys.889.39090.figure31" httpUri="https://binary.pensoft.net/fig/356498" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure31">Figs 31A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 34" captionStartId="F32" captionText="Figure 34. Habitat of Copelatus irregularis A Queensland, Paluma Road 4 km west of Bruce Highway, Maiden Hair Fern Creek, 270 m, shady and rocky residual pools with rotten leaves B of C. irregularis and C. nigrolineatus, Queensland, Winfield, Winfield Road, muddy forest pool, rich in rotten leaves, near the street, 21 m. Photographs by L. Hendrich." figureDoi="10.3897/zookeys.889.39090.figure34" httpUri="https://binary.pensoft.net/fig/356501" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure34">34A</figureCitation>
|
||
).
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus irregularis" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="irregularis">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">Copelatus irregularis</emphasis>
|
||
</taxonomicName>
|
||
is also attracted to light.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |