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<document id="EBC2E8FA060AE2292EDBD515F0A99B94" ID-DOI="10.11646/zootaxa.3964.4.2" ID-GBIF-Dataset="01eeeab3-a425-492d-9df3-19af964721f3" ID-ISSN="1175-5326" ID-Zenodo-Dep="232717" ID-ZooBank="02FA7DA4-4234-4F70-AD07-0536E7B4A6FA" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461077248415" checkinUser="plazi" docAuthor="Jepson, James E." docDate="2015" docId="CB2C2F09FFECFFECA8DFFB56FA64A150" docLanguage="en" docName="zt03964p432.pdf" docOrigin="Zootaxa 3964 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Karataumantispa Jepson, 2015, gen.nov." docType="treatment" docVersion="7" lastPageNumber="429" masterDocId="37155771FFEBFFE6A848FFB2FFF1A404" masterDocTitle="A review of the current state of knowledge of fossil Mantispidae (Insecta: Neuroptera)" masterLastPageNumber="432" masterPageNumber="419" pageNumber="426" updateTime="1698586685506" updateUser="plazi">
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<mods:title id="A2EB91EC6E9341DC9381417CD12D25CD">A review of the current state of knowledge of fossil Mantispidae (Insecta: Neuroptera)</mods:title>
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<mods:namePart id="D65529F9D3ADE49C97DB146979B87736">Jepson, James E.</mods:namePart>
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<mods:date id="BA5F2C5791EC9F4C5FB29A69A8B70850">2015</mods:date>
<mods:detail id="5221B4EDFE543FE4170D11604FA9A16D" type="volume">
<mods:number id="165C26D296E617B474D6E32BBF116993">3964</mods:number>
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<mods:identifier id="9D35E64E836AC9718265644A27D43741" type="DOI">10.11646/zootaxa.3964.4.2</mods:identifier>
<mods:identifier id="35B5C11FC8C5CA11DDB3E4F2EB045A02" type="GBIF-Dataset">01eeeab3-a425-492d-9df3-19af964721f3</mods:identifier>
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<treatment id="CB2C2F09FFECFFECA8DFFB56FA64A150" ID-DOI="http://doi.org/10.5281/zenodo.5661111" ID-GBIF-Taxon="119617821" ID-Zenodo-Dep="5661111" LSID="urn:lsid:plazi:treatment:CB2C2F09FFECFFECA8DFFB56FA64A150" httpUri="http://treatment.plazi.org/id/CB2C2F09FFECFFECA8DFFB56FA64A150" lastPageId="10" lastPageNumber="429" pageId="7" pageNumber="426">
<subSubSection id="0B9FCD94FFECFFE1A8DFFB56FDC6A0FA" box="[151,567,1251,1278]" pageId="7" pageNumber="426" type="nomenclature">
<paragraph id="433A9E1FFFECFFE1A8DFFB56FDC6A0FA" blockId="7.[151,567,1251,1278]" box="[151,567,1251,1278]" pageId="7" pageNumber="426">
<heading id="18722973FFECFFE1A8DFFB56FDC6A0FA" bold="true" box="[151,567,1251,1278]" fontSize="11" level="1" pageId="7" pageNumber="426" reason="1">
<emphasis id="71F1420DFFECFFE1A8DFFB56FDC6A0FA" bold="true" box="[151,567,1251,1278]" pageId="7" pageNumber="426">
Genus:
<taxonomicName id="8485E59CFFECFFE1A8B0FB51FE3BA0F9" box="[248,458,1251,1277]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="71F1420DFFECFFE1A8B0FB51FE3BA0F9" bold="true" box="[248,458,1251,1277]" italics="true" pageId="7" pageNumber="426">Karataumantispa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AC2FF76FFECFFE1A999FB56FDC6A0FA" box="[465,567,1252,1278]" pageId="7" pageNumber="426" rank="genus">gen.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0B9FCD94FFECFFE1A8DFFA98FB27A147" box="[151,1238,1322,1347]" pageId="7" pageNumber="426" type="etymology">
<paragraph id="433A9E1FFFECFFE1A8DFFA98FB27A147" blockId="7.[151,1238,1322,1347]" box="[151,1238,1322,1347]" pageId="7" pageNumber="426">
<emphasis id="71F1420DFFECFFE1A8DFFA98FEEFA147" bold="true" box="[151,286,1322,1347]" pageId="7" pageNumber="426">Etymology.</emphasis>
<emphasis id="71F1420DFFECFFE1A96CFA99FE70A146" box="[292,385,1323,1346]" italics="true" pageId="7" pageNumber="426">Karatau</emphasis>
after the fossil locality of the species and
<taxonomicName id="8485E59CFFECFFE1AB12FA99FC35A146" box="[858,964,1323,1346]" class="Insecta" family="Mantispidae" genus="Mantispa" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="71F1420DFFECFFE1AB12FA99FC35A146" box="[858,964,1323,1346]" italics="true" pageId="7" pageNumber="426">Mantispa</emphasis>
</taxonomicName>
a genus of
<taxonomicName id="8485E59CFFECFFE1AC0CFA98FB23A147" box="[1092,1234,1322,1347]" class="Insecta" family="Mantispidae" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="family">Mantispidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="0B9FCD94FFECFFE1A8DFFAC0FCEAA21F" pageId="7" pageNumber="426" type="materials_examined">
<paragraph id="433A9E1FFFECFFE1A8DFFAC0FD1CA1AB" blockId="7.[151,814,1394,1455]" pageId="7" pageNumber="426">
<emphasis id="71F1420DFFECFFE1A8DFFAC0FEC3A18F" bold="true" box="[151,306,1394,1419]" pageId="7" pageNumber="426">
<typeStatus id="9C3E20BDFFECFFE1A8DFFAC0FF23A18F" box="[151,210,1394,1419]" pageId="7" pageNumber="426">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="8485E59CFFECFFE1A971FAC1FEFDA1AA" authority="(Khramov, 2013) Mesithone" authorityName="(Khramov, 2013) Mesithone" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="carnaria">
<emphasis id="71F1420DFFECFFE1A971FAC1FD97A18E" box="[313,614,1395,1418]" italics="true" pageId="7" pageNumber="426">Karataumantispa carnaria</emphasis>
(
<bibRefCitation id="2714E3EEFFECFFE1AA3CFAC0FCD7A18F" author="Khramov" box="[628,806,1394,1419]" pageId="7" pageNumber="426" refString="Khramov, A. V. (2013) New mantidflies (Neuroptera: Mantispidae) from the Upper Jurassic of Kazakhstan. Insect Systematics &amp; Evolution, 44, 221 - 230. http: // dx. doi. org / 10.1163 / 1876312 X- 44032094" type="journal article" year="2013">Khramov, 2013</bibRefCitation>
)
<emphasis id="71F1420DFFECFFE1A8DFFA24FEFDA1AA" box="[151,268,1430,1454]" italics="true" pageId="7" pageNumber="426">Mesithone</emphasis>
</taxonomicName>
<taxonomicName id="8485E59CFFECFFE1A95BFA25FDB0A1AA" authority="Khramov, 2013" authorityName="Khramov" authorityYear="2013" box="[275,577,1430,1455]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="carnaria">
carnaria
<bibRefCitation id="2714E3EEFFECFFE1A934FA24FDB0A1AA" author="Khramov" box="[380,577,1430,1455]" pageId="7" pageNumber="426" refString="Khramov, A. V. (2013) New mantidflies (Neuroptera: Mantispidae) from the Upper Jurassic of Kazakhstan. Insect Systematics &amp; Evolution, 44, 221 - 230. http: // dx. doi. org / 10.1163 / 1876312 X- 44032094" type="journal article" year="2013" yearSuffix="p">Khramov, 2013 p</bibRefCitation>
</taxonomicName>
. 222, figs. 1–4.
</paragraph>
<paragraph id="433A9E1FFFECFFE1A8DFFA6CFCEAA21F" blockId="7.[151,859,1502,1563]" pageId="7" pageNumber="426">
<emphasis id="71F1420DFFECFFE1A8DFFA6CFECEA1F3" bold="true" box="[151,319,1502,1527]" pageId="7" pageNumber="426">Other species.</emphasis>
<taxonomicName id="8485E59CFFECFFE1A90EFA6DFEFDA21F" authority="(Khramov, 2013) Mesithone" authorityName="(Khramov, 2013) Mesithone" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="monstruosa">
<emphasis id="71F1420DFFECFFE1A90EFA6DFD65A1F2" box="[326,660,1503,1526]" italics="true" pageId="7" pageNumber="426">Karataumantispa monstruosa</emphasis>
(
<bibRefCitation id="2714E3EEFFECFFE1AAE9FA6CFCA2A1F3" author="Khramov" box="[673,851,1502,1527]" pageId="7" pageNumber="426" refString="Khramov, A. V. (2013) New mantidflies (Neuroptera: Mantispidae) from the Upper Jurassic of Kazakhstan. Insect Systematics &amp; Evolution, 44, 221 - 230. http: // dx. doi. org / 10.1163 / 1876312 X- 44032094" type="journal article" year="2013">Khramov, 2013</bibRefCitation>
)
<emphasis id="71F1420DFFECFFE1A8DFF9B1FEFDA21F" box="[151,268,1539,1563]" italics="true" pageId="7" pageNumber="426">Mesithone</emphasis>
</taxonomicName>
<taxonomicName id="8485E59CFFECFFE1A95BF9B1FD93A21E" authority="Khramov, 2013" authorityName="Khramov" authorityYear="2013" box="[275,610,1538,1563]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="monstruosa">
monstruosa
<bibRefCitation id="2714E3EEFFECFFE1A9D4F9B0FD93A21E" author="Khramov" box="[412,610,1538,1563]" pageId="7" pageNumber="426" refString="Khramov, A. V. (2013) New mantidflies (Neuroptera: Mantispidae) from the Upper Jurassic of Kazakhstan. Insect Systematics &amp; Evolution, 44, 221 - 230. http: // dx. doi. org / 10.1163 / 1876312 X- 44032094" type="journal article" year="2013" yearSuffix="p">Khramov, 2013 p</bibRefCitation>
</taxonomicName>
. 224, figs. 5–11.
</paragraph>
</subSubSection>
<subSubSection id="0B9FCD94FFECFFEEA8DFF9F8FD55A6F4" lastPageId="8" lastPageNumber="427" pageId="7" pageNumber="426" type="diagnosis">
<paragraph id="433A9E1FFFECFFE1A8DFF9F8FCBCA313" blockId="7.[151,1437,1610,2031]" pageId="7" pageNumber="426">
<emphasis id="71F1420DFFECFFE1A8DFF9F8FEE0A267" bold="true" box="[151,273,1610,1635]" pageId="7" pageNumber="426">Diagnosis.</emphasis>
The genus differs from other fossil mantispids based on the combination of the following characters: elongate forewings; subcostal space proximally very narrow, distally dilated; Rs with at least seven branches; MP deeply forked after the split of MA and MP; wing colour pattern minimal to absent, with small patches of colour on some crossveins; forelegs with two rows of spines on inner edge of the fore femur, one row with relatively smaller spines than the other; fore tibia with hook-like prostrate setae of similar size to the larger femora spines; combined length of the fore tibia and fore tarsus greater than fore femur.
</paragraph>
<paragraph id="433A9E1FFFECFFEEA88FF890FD55A6F4" blockId="7.[151,1437,1610,2031]" lastBlockId="8.[151,1436,151,1184]" lastPageId="8" lastPageNumber="427" pageId="7" pageNumber="426">
<emphasis id="71F1420DFFECFFE1A88FF890FE93A33F" bold="true" box="[199,354,1826,1851]" pageId="7" pageNumber="426">Comparison.</emphasis>
Elongate forewings are observed in all other fossil mantispids, except
<taxonomicName id="8485E59CFFECFFE1AC9CF891FA92A33E" box="[1236,1379,1827,1850]" class="Insecta" family="Mantispidae" genus="Promantispa" higherTaxonomySource="CoL" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="71F1420DFFECFFE1AC9CF891FA92A33E" box="[1236,1379,1827,1850]" italics="true" pageId="7" pageNumber="426">Promantispa</emphasis>
</taxonomicName>
and
<taxonomicName id="8485E59CFFECFFE1A8DFF8F4FEDDA35A" box="[151,300,1862,1886]" class="Insecta" family="Mantispidae" genus="Symphrasites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="71F1420DFFECFFE1A8DFF8F4FEDDA35A" box="[151,300,1862,1886]" italics="true" pageId="7" pageNumber="426">Symphrasites</emphasis>
</taxonomicName>
, which have a slightly broader shaped wing. The narrow subcostal space, which dilates distally, is similar to
<emphasis id="71F1420DFFECFFE1A942F8D9FE5FA386" box="[266,430,1899,1922]" italics="true" pageId="7" pageNumber="426">Mesomantispa</emphasis>
, however, in most other fossil genera the subcostal space is a uniform thickness for most of its length and slightly narrowed proximally (near wing base).
<taxonomicName id="8485E59CFFECFFE1ABD9F83DFC05A3A2" box="[913,1012,1935,1958]" class="Insecta" family="Mantispidae" genus="Feroseta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="71F1420DFFECFFE1ABD9F83DFC05A3A2" box="[913,1012,1935,1958]" italics="true" pageId="7" pageNumber="426">Feroseta</emphasis>
</taxonomicName>
differs in that its subcostal space is narrow proximally, dilated at midpoint, and narrow distally. The number of branches of Rs (with at least 7) differs from some of the other genera of mantispids:
<emphasis id="71F1420DFFECFFE1AAF0F865FCAAA3EA" box="[696,859,2007,2030]" italics="true" pageId="7" pageNumber="426">Mesomantispa</emphasis>
and
<taxonomicName id="8485E59CFFECFFE1ABD2F864FBDEA3EA" box="[922,1071,2006,2030]" class="Insecta" family="Mantispidae" genus="Symphrasites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="71F1420DFFECFFE1ABD2F864FBDEA3EA" box="[922,1071,2006,2030]" italics="true" pageId="7" pageNumber="426">Symphrasites</emphasis>
</taxonomicName>
11,
<emphasis id="71F1420DFFECFFE1AC20F864FA9FA3EA" box="[1128,1390,2006,2030]" italics="true" pageId="7" pageNumber="426">
<taxonomicName id="8485E59CFFECFFE1AC20F864FB1FA3EA" box="[1128,1262,2006,2030]" class="Insecta" family="Mantispidae" genus="Climaciella" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="genus">Climaciella</taxonomicName>
?
<taxonomicName id="8485E59CFFECFFE1AD4CF864FA9FA3EA" box="[1284,1390,2006,2030]" class="Insecta" family="Mantispidae" genus="Climaciella" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="henrotayi">henrotayi</taxonomicName>
</emphasis>
10,
<taxonomicName id="8485E59CFFE3FFEEA8DFFF2AFE5BA4AB" box="[151,426,152,176]" class="Insecta" family="Mantispidae" genus="Gerstaeckerella" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="427" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="71F1420DFFE3FFEEA8DFFF2AFE5BA4AB" box="[151,426,152,176]" italics="true" pageId="8" pageNumber="427">Gerstaeckerella asiatica</emphasis>
</taxonomicName>
and
<emphasis id="71F1420DFFE3FFEEA9AAFF2AFD45A4AB" box="[482,692,152,175]" italics="true" pageId="8" pageNumber="427">Sinomesomantispa</emphasis>
9,
<emphasis id="71F1420DFFE3FFEEAA91FF2AFC94A4B4" box="[729,869,152,176]" italics="true" pageId="8" pageNumber="427">Clavifemora</emphasis>
&gt;7, all other fossil genera 7 branches or less. The deeply forked MP (close to split of MA and MP) is similar to
<taxonomicName id="8485E59CFFE3FFEEAB04FF0EFC10A4D0" box="[844,993,188,212]" class="Insecta" family="Mantispidae" genus="Symphrasites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="71F1420DFFE3FFEEAB04FF0EFC10A4D0" box="[844,993,188,212]" italics="true" pageId="8" pageNumber="427">Symphrasites</emphasis>
</taxonomicName>
, but in all other fossil genera MP is not forked until the distal gradate series of crossveins near the posterior wing margin. A minimal colour pattern on the wings differs from
<emphasis id="71F1420DFFE3FFEEA931FEB7FDECA518" box="[377,541,261,284]" italics="true" pageId="8" pageNumber="427">Mesomantispa</emphasis>
and
<emphasis id="71F1420DFFE3FFEEAA12FEB6FD17A518" box="[602,742,260,284]" italics="true" pageId="8" pageNumber="427">Clavifemora</emphasis>
, which have spots,
<emphasis id="71F1420DFFE3FFEEAB9AFEB6FB5AA518" box="[978,1195,260,284]" italics="true" pageId="8" pageNumber="427">Archaeodrepanicus</emphasis>
which has 2–3 dark bands,
<emphasis id="71F1420DFFE3FFEEA8AFFE9AFE48A53B" box="[231,441,296,319]" italics="true" pageId="8" pageNumber="427">Sinomesomantispa</emphasis>
, which has a distinct thick dark band in distal part of wing, and
<emphasis id="71F1420DFFE3FFEEACD9FE9AFAB9A53B" box="[1169,1352,296,319]" italics="true" pageId="8" pageNumber="427">Doratomantispa</emphasis>
, which has small spots and colouration around some crossveins. The colour pattern is absent or not preserved in the rest of the mantispid fossil genera. The arrangement of the spines on the fore femur, in two rows, with one row of spines smaller than the other differs from the other genera. Two rows of spines are observed in the mesomantispine genera; however the spines are of a uniform size.
<emphasis id="71F1420DFFE3FFEEAA95FE0AFC76A5CB" box="[733,903,440,463]" italics="true" pageId="8" pageNumber="427">Micromantispa</emphasis>
has two rows with six large spines and many smaller spines.
<emphasis id="71F1420DFFE3FFEEA90DFE6FFDFBA5F0" box="[325,522,477,500]" italics="true" pageId="8" pageNumber="427">Doratomanatispa</emphasis>
has six large cuticular spines.
<taxonomicName id="8485E59CFFE3FFEEAB2DFE6FFBFDA5F0" box="[869,1036,477,500]" class="Insecta" family="Mantispidae" genus="Dicromantispa" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="71F1420DFFE3FFEEAB2DFE6FFBFDA5F0" box="[869,1036,477,500]" italics="true" pageId="8" pageNumber="427">Dicromantispa</emphasis>
</taxonomicName>
has a large basal spine, many short teeth-like spines and at least three intermediate sized spines.
<taxonomicName id="8485E59CFFE3FFEEAB77FDB3FC53A61C" box="[831,930,513,536]" class="Insecta" family="Mantispidae" genus="Feroseta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="71F1420DFFE3FFEEAB77FDB3FC53A61C" box="[831,930,513,536]" italics="true" pageId="8" pageNumber="427">Feroseta</emphasis>
</taxonomicName>
also has one large basal spine, but with three minor and 7–8 smaller spines. The large hook-like prostrate setae are not observed in the other mantispid genera, the prostrate setae are less hook-like and smaller than the femora spines in
<emphasis id="71F1420DFFE3FFEEABAAFDFAFB4AA664" box="[994,1211,584,608]" italics="true" pageId="8" pageNumber="427">Archaeodrepanicus</emphasis>
,
<emphasis id="71F1420DFFE3FFEEAC82FDFAFA6DA65B" box="[1226,1436,584,607]" italics="true" pageId="8" pageNumber="427">Sinomesomantispa</emphasis>
and
<emphasis id="71F1420DFFE3FFEEA88EFDDEFEA3A680" box="[198,338,620,644]" italics="true" pageId="8" pageNumber="427">Clavifemora</emphasis>
. They are smaller and peg-like in
<emphasis id="71F1420DFFE3FFEEAA9AFDDFFC78A680" box="[722,905,621,644]" italics="true" pageId="8" pageNumber="427">Doratomantispa</emphasis>
, and absent in
<emphasis id="71F1420DFFE3FFEEAC7AFDDFFB2DA680" box="[1074,1244,621,644]" italics="true" pageId="8" pageNumber="427">Micromantispa</emphasis>
(which has three long spines on tibia) and
<emphasis id="71F1420DFFE3FFEEA9E6FD22FDACA6A3" box="[430,605,656,679]" italics="true" pageId="8" pageNumber="427">
<taxonomicName id="8485E59CFFE3FFEEA9E6FD22FDA8A6A3" box="[430,601,656,679]" class="Insecta" family="Mantispidae" genus="Dicromantispa" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="427" phylum="Arthropoda" rank="genus">Dicromantispa</taxonomicName>
.
</emphasis>
The combined length of the fore tibia and fore tarsus being longer than the fore femur; is similar in
<emphasis id="71F1420DFFE3FFEEA98FFD07FD80A6C8" box="[455,625,693,716]" italics="true" pageId="8" pageNumber="427">Micromantispa</emphasis>
, but shorter in
<emphasis id="71F1420DFFE3FFEEAB7DFD06FBFFA6C8" box="[821,1038,692,716]" italics="true" pageId="8" pageNumber="427">Archaeodrepanicus</emphasis>
,
<emphasis id="71F1420DFFE3FFEEAC6BFD06FB04A6CF" box="[1059,1269,692,715]" italics="true" pageId="8" pageNumber="427">Sinomesomantispa</emphasis>
,
<emphasis id="71F1420DFFE3FFEEAD41FD06FDFBA6EB" italics="true" pageId="8" pageNumber="427">
Clavifemora, Doratomantispa,
<taxonomicName id="8485E59CFFE3FFEEA915FD6AFDF6A6EB" box="[349,519,728,751]" class="Insecta" family="Mantispidae" genus="Dicromantispa" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="427" phylum="Arthropoda" rank="genus">Dicromantispa</taxonomicName>
,
</emphasis>
and
<emphasis id="71F1420DFFE3FFEEAA72FD6BFD55A6F4" box="[570,676,729,752]" italics="true" pageId="8" pageNumber="427">
<taxonomicName id="8485E59CFFE3FFEEAA72FD6BFD51A6F4" box="[570,672,729,752]" class="Insecta" family="Mantispidae" genus="Feroseta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="427" phylum="Arthropoda" rank="genus">Feroseta</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0B9FCD94FFE3FFECA88FFD4EFF27A7A1" lastPageId="10" lastPageNumber="429" pageId="8" pageNumber="427" type="discussion">
<paragraph id="433A9E1FFFE3FFEEA88FFD4EFAD7A0A4" blockId="8.[151,1436,151,1184]" pageId="8" pageNumber="427">
<emphasis id="71F1420DFFE3FFEEA88FFD4EFECAA711" bold="true" box="[199,315,764,789]" pageId="8" pageNumber="427">Remarks.</emphasis>
These species of mantispid were originally placed within the genus
<emphasis id="71F1420DFFE3FFEEAC01FD4EFB4FA710" box="[1097,1214,764,788]" italics="true" pageId="8" pageNumber="427">Mesithone</emphasis>
<bibRefCitation id="2714E3EEFFE3FFEEAC8EFD4EFA9AA710" author="Panfilov" box="[1222,1387,764,789]" pageId="8" pageNumber="427" refString="Panfilov, D. V. (1980) New representatives of lacewings (Neuroptera) from the Jurassic of Karatau. In: Dolin, V. G., Panfilov, D. V., Ponomarenko, A. G. &amp; Pritykina, L. N. (Eds), Fossil insects of the Mesozoic. Naukova Dumka, Kiev, pp. 82 - 111." type="book chapter" year="1980">Panfilov, 1980</bibRefCitation>
and the subfamily Mesithoninae
<bibRefCitation id="2714E3EEFFE3FFEEA9BFFCADFD54A73C" author="Panfilov" box="[503,677,799,824]" pageId="8" pageNumber="427" refString="Panfilov, D. V. (1980) New representatives of lacewings (Neuroptera) from the Jurassic of Karatau. In: Dolin, V. G., Panfilov, D. V., Ponomarenko, A. G. &amp; Pritykina, L. N. (Eds), Fossil insects of the Mesozoic. Naukova Dumka, Kiev, pp. 82 - 111." type="book chapter" year="1980">Panfilov, 1980</bibRefCitation>
(
<bibRefCitation id="2714E3EEFFE3FFEEAAF6FCADFC8FA73C" author="Khramov" box="[702,894,799,824]" pageId="8" pageNumber="427" refString="Khramov, A. V. (2013) New mantidflies (Neuroptera: Mantispidae) from the Upper Jurassic of Kazakhstan. Insect Systematics &amp; Evolution, 44, 221 - 230. http: // dx. doi. org / 10.1163 / 1876312 X- 44032094" type="journal article" year="2013">Khramov, 2013</bibRefCitation>
). The subfamily Mesomantispinae was also synonymized with Mesithoninae (
<bibRefCitation id="2714E3EEFFE3FFEEAA5DFCF6FD3AA758" author="Khramov" box="[533,715,836,861]" pageId="8" pageNumber="427" refString="Khramov, A. V. (2013) New mantidflies (Neuroptera: Mantispidae) from the Upper Jurassic of Kazakhstan. Insect Systematics &amp; Evolution, 44, 221 - 230. http: // dx. doi. org / 10.1163 / 1876312 X- 44032094" type="journal article" year="2013">Khramov, 2013</bibRefCitation>
). The specimens and other genera of Mesomantispinae do show some similarities with
<emphasis id="71F1420DFFE3FFEEA9D0FCDAFDFDA784" box="[408,524,872,896]" italics="true" pageId="8" pageNumber="427">Mesithone</emphasis>
: long recurrent humeral vein, Sc veinlets closely spaced with majority forked in forewing, general structure of Rs and M. However, they differ by having a combination of: Elongate forewing [broad and ovate in
<emphasis id="71F1420DFFE3FFEEA9CCFC02FE08A7CC" box="[388,505,944,968]" italics="true" pageId="8" pageNumber="427">Mesithone</emphasis>
]; the distal widening of the subcostal space [narrows in
<emphasis id="71F1420DFFE3FFEEACD8FC02FAF4A7CC" box="[1168,1285,944,968]" italics="true" pageId="8" pageNumber="427">Mesithone</emphasis>
]; Sc curving towards R1 [subcostal space narrows distally and Sc and R1 are fused in
<emphasis id="71F1420DFFE3FFEEAB81FC66FBCCA7E8" box="[969,1085,980,1004]" italics="true" pageId="8" pageNumber="427">Mesithone</emphasis>
]; R is fused with M basally for a long distance in the forewing [R and M not fused basally or fused for short distance in
<emphasis id="71F1420DFFE3FFEEACCDFC4AFB08A014" box="[1157,1273,1016,1040]" italics="true" pageId="8" pageNumber="427">Mesithone</emphasis>
—the drawing of the
<typeStatus id="9C3E20BDFFE3FFEEA896FBAEFEB2A031" box="[222,323,1052,1077]" pageId="8" pageNumber="427" type="holotype">holotype</typeStatus>
,
<taxonomicName id="8485E59CFFE3FFEEA906FBAFFE2AA030" box="[334,475,1052,1076]" class="Insecta" family="Mantispidae" genus="Mantispa" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="427" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="71F1420DFFE3FFEEA906FBAFFE2AA030" box="[334,475,1052,1076]" italics="true" pageId="8" pageNumber="427">M. maculata</emphasis>
</taxonomicName>
in
<bibRefCitation id="2714E3EEFFE3FFEEAA48FBAEFCFBA031" author="Panfilov" box="[512,778,1052,1077]" pageId="8" pageNumber="427" refString="Panfilov, D. V. (1980) New representatives of lacewings (Neuroptera) from the Jurassic of Karatau. In: Dolin, V. G., Panfilov, D. V., Ponomarenko, A. G. &amp; Pritykina, L. N. (Eds), Fossil insects of the Mesozoic. Naukova Dumka, Kiev, pp. 82 - 111." type="book chapter" year="1980">Panfilov (1980: fig. 86)</bibRefCitation>
, shows the fusing of M and R, this however is not the case from examining the photograph (
<figureCitation id="DBBE829AFFE3FFEEAA42FB8DFDBFA05C" box="[522,590,1087,1112]" captionStart="FIGURE 2" captionStartId="8.[151,250,1939,1961]" captionTargetBox="[154,1436,1243,1917]" captionTargetId="figure@8.[151,1436,1237,1917]" captionTargetPageId="8" captionText="FIGURE 2. Photograph of Mesithone maculata Panfilov 1980 (holotype) highlighting the differences in venation to Karataumantispa gen. nov. (Photograph: V. N. Makarkin)." httpUri="https://zenodo.org/record/232719/files/figure.png" pageId="8" pageNumber="427">Fig. 2</figureCitation>
) and from
<collectingCountry id="3B92DE8FFFE3FFEEAA83FBF2FCF4A053" box="[715,773,1088,1111]" name="Vietnam" pageId="8" pageNumber="427">V. N.</collectingCountry>
Makarkin (who has viewed the fossil (pers. comm))], M is always fused with R for some distance in the forewing of
<taxonomicName id="8485E59CFFE3FFEEAB7DFBD6FB27A078" authority="Makarkin et al., 2013" authorityName="Makarkin et al." authorityYear="2013" box="[821,1238,1124,1149]" class="Insecta" family="Mantispidae" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="427" phylum="Arthropoda" rank="family">
Mantispidae (Makarkin
<emphasis id="71F1420DFFE3FFEEAC0FFBD7FB70A078" box="[1095,1153,1124,1148]" italics="true" pageId="8" pageNumber="427">et al.</emphasis>
, 2013)
</taxonomicName>
; CuA pectinately branched in forewing [non-pectinate in
<emphasis id="71F1420DFFE3FFEEAA1BFB3AFD39A0A4" box="[595,712,1160,1184]" italics="true" pageId="8" pageNumber="427">Mesithone</emphasis>
]. These differences are highlighted in figures 2 and 3.
</paragraph>
<caption id="17FACE97FFE3FFEEA8DFF821FD15A3C3" httpUri="https://zenodo.org/record/232719/files/figure.png" pageId="8" pageNumber="427" targetBox="[154,1436,1243,1917]" targetPageId="8">
<paragraph id="433A9E1FFFE3FFEEA8DFF821FD15A3C3" blockId="8.[151,1436,1939,1991]" pageId="8" pageNumber="427">
<emphasis id="71F1420DFFE3FFEEA8DFF821FEEBA3AC" bold="true" box="[151,282,1939,1961]" pageId="8" pageNumber="427">FIGURE 2.</emphasis>
Photograph of
<emphasis id="71F1420DFFE3FFEEA986F821FDC6A3AC" box="[462,567,1939,1960]" italics="true" pageId="8" pageNumber="427">Mesithone</emphasis>
<taxonomicName id="8485E59CFFE3FFEEAA0DF821FCB9A3AD" authority="Panfilov 1980" authorityName="Panfilov" authorityYear="1980" box="[581,840,1939,1961]" class="Insecta" family="Mantispidae" genus="Mantispa" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="427" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="71F1420DFFE3FFEEAA0DF821FD52A3AC" box="[581,675,1939,1960]" italics="true" pageId="8" pageNumber="427">maculata</emphasis>
Panfilov 1980
</taxonomicName>
(holotype) highlighting the differences in venation to
<taxonomicName id="8485E59CFFE3FFEEA8DFF800FEB9A3C3" box="[151,328,1970,1991]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="427" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="71F1420DFFE3FFEEA8DFF800FEB9A3C3" box="[151,328,1970,1991]" italics="true" pageId="8" pageNumber="427">Karataumantispa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AC2FF76FFE3FFEEA906F800FE5CA3C3" box="[334,429,1970,1991]" pageId="8" pageNumber="427" rank="genus">
<emphasis id="71F1420DFFE3FFEEA906F800FE5CA3C3" bold="true" box="[334,429,1970,1991]" pageId="8" pageNumber="427">gen. nov.</emphasis>
</taxonomicNameLabel>
(Photograph: V.N. Makarkin).
</paragraph>
</caption>
<caption id="17FACE97FFE2FFEFA8DFF8F3FE4BA3D6" httpUri="https://zenodo.org/record/232720/files/figure.png" pageId="9" pageNumber="428" targetBox="[151,1436,199,1834]" targetPageId="9">
<paragraph id="433A9E1FFFE2FFEFA8DFF8F3FE4BA3D6" blockId="9.[151,1437,1857,2002]" pageId="9" pageNumber="428">
<emphasis id="71F1420DFFE2FFEFA8DFF8F3FEE5A352" bold="true" box="[151,276,1857,1879]" pageId="9" pageNumber="428">FIGURE 3.</emphasis>
A)
<taxonomicName id="8485E59CFFE2FFEFA975F8F0FCF7A353" authority="Khramov, 2013" authorityName="Khramov" authorityYear="2013" box="[317,774,1857,1879]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="carnaria">
<emphasis id="71F1420DFFE2FFEFA975F8F0FDBEA353" box="[317,591,1858,1879]" italics="true" pageId="9" pageNumber="428">Karataumantispa carnaria</emphasis>
(Khramov, 2013)
</taxonomicName>
(holotype) with CuA highlighted. B.
<taxonomicName id="8485E59CFFE2FFEFACC3F8F0FEB4A372" authority="Khramov, 2013" authorityName="Khramov" authorityYear="2013" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="carnaria">
<emphasis id="71F1420DFFE2FFEFACC3F8F0FA6DA353" box="[1163,1436,1858,1879]" italics="true" pageId="9" pageNumber="428">Karataumantispa carnaria</emphasis>
(Khramov, 2013)
</taxonomicName>
close up of forewing venation (holotype). C.
<taxonomicName id="8485E59CFFE2FFEFAB5AF8D3FB05A372" authority="Khramov, 2013" authorityName="Khramov" authorityYear="2013" box="[786,1268,1888,1910]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="monstruosa">
<emphasis id="71F1420DFFE2FFEFAB5AF8D3FBCEA372" box="[786,1087,1889,1910]" italics="true" pageId="9" pageNumber="428">Karataumantispa monstruosa</emphasis>
(Khramov, 2013)
</taxonomicName>
close up of hind wing venation (holotype). All wings with annotated wing venation, highlighting the major differences that exclude them from
<emphasis id="71F1420DFFE2FFEFA8DFF82FFEF0A3B6" box="[151,257,1949,1970]" italics="true" pageId="9" pageNumber="428">Mesithone</emphasis>
: the fusion of R and M, the distal widening of the subcostal space, Sc bending towards R1 space, and CuA pectinate. (Photographs: A. Khramov).
</paragraph>
</caption>
<paragraph id="433A9E1FFFE1FFECA88FFF25FD42A544" blockId="10.[151,1437,151,1365]" pageId="10" pageNumber="429">
The new placement of these species within Mesomantispinae is based upon the possession of the characters given in the diagnosis of the subfamily (
<bibRefCitation id="2714E3EEFFE1FFECAA20FF0EFCB4A4D0" author="Jepson" box="[616,837,188,212]" pageId="10" pageNumber="429" refString="Jepson, J. E., Heads, S. W., Makarkin, V. N. &amp; Ren, D. (2013) New fossil Mantidflies (Insecta: Neuroptera: Mantispidae) from the Mesozoic of North-Eastern China. Palaeontology, 56, 603 - 613. http: // dx. doi. org / 10.1111 / pala. 12005" type="journal article" year="2013">
Jepson
<emphasis id="71F1420DFFE1FFECAAF6FF0FFD08A4D0" box="[702,761,188,212]" italics="true" pageId="10" pageNumber="429">et al.</emphasis>
, 2013
</bibRefCitation>
), for example: short broad prothorax, femur without major spines, slightly arched tibia with prostrate setae, subcostal space dilated distally, humeral vein recurrent, Sc veinlets closely spaced with the majority forked, inconspicuous pterostigma, M forked distal to origin of Rs, MA and MP parallel, and CuA pectinately branched.
</paragraph>
<paragraph id="433A9E1FFFE1FFECA88FFEFEFF27A7A1" blockId="10.[151,1437,151,1365]" pageId="10" pageNumber="429">
They differ from the other genera of Mesomantispinae in the structure of the forelegs.
<emphasis id="71F1420DFFE1FFECACDBFEFEFA9DA560" box="[1171,1388,332,356]" italics="true" pageId="10" pageNumber="429">Archaeodrepanicus</emphasis>
and
<emphasis id="71F1420DFFE1FFECA8DFFEC2FE98A583" box="[151,361,368,391]" italics="true" pageId="10" pageNumber="429">Sinomesomantispa</emphasis>
have smaller more flattened non hook-like prostrate setae, and
<emphasis id="71F1420DFFE1FFECAC09FEC2FAE2A583" box="[1089,1299,368,391]" italics="true" pageId="10" pageNumber="429">Sinomesomantispa</emphasis>
has smaller spines. The combined length of the foretibia and foretarsus is greater than that of the femur in
<taxonomicName id="8485E59CFFE1FFECAC90FE27FA6DA5A8" box="[1240,1436,405,428]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="71F1420DFFE1FFECAC90FE27FA6DA5A8" box="[1240,1436,405,428]" italics="true" pageId="10" pageNumber="429">Karataumantispa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AC2FF76FFE1FFECA8DFFE05FEF2A5D4" box="[151,259,439,464]" pageId="10" pageNumber="429" rank="genus">
<emphasis id="71F1420DFFE1FFECA8DFFE05FEF2A5D4" bold="true" box="[151,259,439,464]" pageId="10" pageNumber="429">gen. nov.</emphasis>
</taxonomicNameLabel>
; this is shorter in the Chinese fossil mantispids.
<emphasis id="71F1420DFFE1FFECAB76FE0AFC3BA5D4" box="[830,970,440,464]" italics="true" pageId="10" pageNumber="429">Clavifemora</emphasis>
has a very different foreleg structure to
<taxonomicName id="8485E59CFFE1FFECA8DFFE6FFEAAA5F0" box="[151,347,477,500]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="71F1420DFFE1FFECA8DFFE6FFEAAA5F0" box="[151,347,477,500]" italics="true" pageId="10" pageNumber="429">Karataumantispa</emphasis>
</taxonomicName>
<emphasis id="71F1420DFFE1FFECA92CFE6EFE21A5F1" bold="true" box="[356,464,476,501]" pageId="10" pageNumber="429">
<taxonomicNameLabel id="6AC2FF76FFE1FFECA92CFE6EFE21A5F1" box="[356,464,476,501]" pageId="10" pageNumber="429" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, with its swollen club-like forelegs.
<taxonomicName id="8485E59CFFE1FFECAB31FE6FFBCCA5F0" box="[889,1085,477,500]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="71F1420DFFE1FFECAB31FE6FFBCCA5F0" box="[889,1085,477,500]" italics="true" pageId="10" pageNumber="429">Karataumantispa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AC2FF76FFE1FFECAC0EFE6EFB43A5F1" box="[1094,1202,476,501]" pageId="10" pageNumber="429" rank="genus">
<emphasis id="71F1420DFFE1FFECAC0EFE6EFB43A5F1" bold="true" box="[1094,1202,476,501]" pageId="10" pageNumber="429">gen. nov.</emphasis>
</taxonomicNameLabel>
also differs in wing venation to the other mesomantispines. The subcostal space is less strongly dilated distally in the other genera (except
<emphasis id="71F1420DFFE1FFECA8BBFD97FE67A638" box="[243,406,549,572]" italics="true" pageId="10" pageNumber="429">Mesomantispa</emphasis>
). The radial sector is less densely branched in
<emphasis id="71F1420DFFE1FFECABFFFD96FB61A638" box="[951,1168,548,572]" italics="true" pageId="10" pageNumber="429">Archaeodrepanicus</emphasis>
(6 branches) and more branched in
<emphasis id="71F1420DFFE1FFECA96BFDFAFE04A65B" box="[291,501,584,607]" italics="true" pageId="10" pageNumber="429">Sinomesomantispa</emphasis>
(9) and
<emphasis id="71F1420DFFE1FFECAA1FFDFAFD0AA65B" box="[599,763,584,607]" italics="true" pageId="10" pageNumber="429">Mesomantispa</emphasis>
(11) (this is a tentative character because the exact number of Rs branches is not known); R1 and Rs curve distally towards apex of wing in
<taxonomicName id="8485E59CFFE1FFECAC5AFDDFFB27A680" box="[1042,1238,621,644]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="71F1420DFFE1FFECAC5AFDDFFB27A680" box="[1042,1238,621,644]" italics="true" pageId="10" pageNumber="429">Karataumantispa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AC2FF76FFE1FFECAC94FDDEFAB4A681" box="[1244,1349,620,645]" pageId="10" pageNumber="429" rank="genus">
<emphasis id="71F1420DFFE1FFECAC94FDDEFAB4A681" bold="true" box="[1244,1349,620,645]" pageId="10" pageNumber="429">gen. nov.</emphasis>
</taxonomicNameLabel>
and the Chinese genera, these are straight and horizontal in
<emphasis id="71F1420DFFE1FFECAB58FD22FC42A6A3" box="[784,947,656,679]" italics="true" pageId="10" pageNumber="429">Mesomantispa</emphasis>
.
<emphasis id="71F1420DFFE1FFECAB80FD22FB9DA6A3" box="[968,1132,656,679]" italics="true" pageId="10" pageNumber="429">Mesomantispa</emphasis>
has a wider cubital area (approximately half the width of the wing);
<taxonomicName id="8485E59CFFE1FFECAAC1FD07FCBCA6C8" box="[649,845,693,716]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="71F1420DFFE1FFECAAC1FD07FCBCA6C8" box="[649,845,693,716]" italics="true" pageId="10" pageNumber="429">Karataumantispa</emphasis>
</taxonomicName>
<emphasis id="71F1420DFFE1FFECAB1DFD06FC4EA6C9" bold="true" box="[853,959,692,717]" pageId="10" pageNumber="429">
<taxonomicNameLabel id="6AC2FF76FFE1FFECAB1DFD06FC4EA6C9" box="[853,959,692,717]" pageId="10" pageNumber="429" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
has a similar cubital width to the Chinese genera (approximately one third of wing width). MP forks just after the split of M into MA and MP in
<taxonomicName id="8485E59CFFE1FFECA8DFFD4FFEAAA710" box="[151,347,765,788]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="71F1420DFFE1FFECA8DFFD4FFEAAA710" box="[151,347,765,788]" italics="true" pageId="10" pageNumber="429">Karataumantispa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AC2FF76FFE1FFECA92AFD4EFE3DA711" box="[354,460,764,789]" pageId="10" pageNumber="429" rank="genus">
<emphasis id="71F1420DFFE1FFECA92AFD4EFE3DA711" bold="true" box="[354,460,764,789]" pageId="10" pageNumber="429">gen. nov.</emphasis>
</taxonomicNameLabel>
and much closer to the posterior wing margin in all other genera. The colour pattern is not as well developed (this is a tentative character because it may be taphonomic) in
<taxonomicName id="8485E59CFFE1FFECAC38FC92FAC5A733" box="[1136,1332,800,823]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="71F1420DFFE1FFECAC38FC92FAC5A733" box="[1136,1332,800,823]" italics="true" pageId="10" pageNumber="429">Karataumantispa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AC2FF76FFE1FFECAD75FC92FA6DA73C" box="[1341,1436,800,824]" pageId="10" pageNumber="429" rank="genus">gen.nov.</taxonomicNameLabel>
<emphasis id="71F1420DFFE1FFECA8DFFCF6FE81A758" box="[151,368,836,860]" italics="true" pageId="10" pageNumber="429">Archaeodrepanicus</emphasis>
has two-three dark bands across the wing;
<emphasis id="71F1420DFFE1FFECAB1FFCF6FBD8A75F" box="[855,1065,836,859]" italics="true" pageId="10" pageNumber="429">Sinomesomantispa</emphasis>
has a distinct dark band in distal part of the wing;
<emphasis id="71F1420DFFE1FFECA92CFCDAFE01A784" box="[356,496,872,896]" italics="true" pageId="10" pageNumber="429">Clavifemora</emphasis>
has three dark spots similar to
<emphasis id="71F1420DFFE1FFECAB29FCDAFBF5A77B" box="[865,1028,872,895]" italics="true" pageId="10" pageNumber="429">Mesomantispa</emphasis>
, which has large dark spots on the wing.
</paragraph>
</subSubSection>
<subSubSection id="0B9FCD94FFE1FFECA88FFC02FA64A150" pageId="10" pageNumber="429" type="reference_group">
<paragraph id="433A9E1FFFE1FFECA88FFC02FA64A150" blockId="10.[151,1437,151,1365]" pageId="10" pageNumber="429">
<taxonomicName id="8485E59CFFE1FFECA88FFC02FDE8A7CC" box="[199,537,944,968]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="species" species="monstruosa">
<emphasis id="71F1420DFFE1FFECA88FFC02FDE8A7CC" box="[199,537,944,968]" italics="true" pageId="10" pageNumber="429">Karataumantispa monstruosa</emphasis>
</taxonomicName>
is less well-preserved than
<taxonomicName id="8485E59CFFE1FFECAB2AFC03FC19A7C3" box="[866,1000,944,968]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="species" species="carnaria">
<emphasis id="71F1420DFFE1FFECAB2AFC03FC19A7C3" box="[866,1000,944,968]" italics="true" pageId="10" pageNumber="429">K. carnaria</emphasis>
</taxonomicName>
and therefore possesses fewer of the aforementioned wing characters that place it within Mesomantispinae.
<taxonomicName id="8485E59CFFE1FFECAB94FC67FAC1A7E8" box="[988,1328,981,1004]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="species" species="monstruosa">
<emphasis id="71F1420DFFE1FFECAB94FC67FAC1A7E8" box="[988,1328,981,1004]" italics="true" pageId="10" pageNumber="429">Karataumantispa monstruosa</emphasis>
</taxonomicName>
however shares some similarities with other mesomantispines,
<emphasis id="71F1420DFFE1FFECAABDFC4AFC68A00B" box="[757,921,1016,1039]" italics="true" pageId="10" pageNumber="429">Mesomantispa</emphasis>
and
<emphasis id="71F1420DFFE1FFECAB98FC4AFB58A014" box="[976,1193,1016,1040]" italics="true" pageId="10" pageNumber="429">Archaeodrepanicus</emphasis>
, with the presence of 3r1-rs and the position of 2r1-rs. On the venation drawings in the original description of
<taxonomicName id="8485E59CFFE1FFECAC31FBAFFAECA030" box="[1145,1309,1053,1076]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="species" species="monstruosa">
<emphasis id="71F1420DFFE1FFECAC31FBAFFAECA030" box="[1145,1309,1053,1076]" italics="true" pageId="10" pageNumber="429">K. monstruosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="2714E3EEFFE1FFECAD63FBAEFEA7A05C" author="Khramov" pageId="10" pageNumber="429" refString="Khramov, A. V. (2013) New mantidflies (Neuroptera: Mantispidae) from the Upper Jurassic of Kazakhstan. Insect Systematics &amp; Evolution, 44, 221 - 230. http: // dx. doi. org / 10.1163 / 1876312 X- 44032094" type="journal article" year="2013">Khramov, 2013: Figs. 9–11</bibRefCitation>
) the fusion of Sc and R is shown in the forewing, this however has not been confirmed from the study of high resolution images of the specimen, however the subcostal space is distally dilated, and the hind wing shows that Sc curves towards R1 (
<figureCitation id="DBBE829AFFE1FFECAA51FB35FDA4A0A4" box="[537,597,1159,1184]" captionStart="FIGURE 3. A" captionStartId="9.[151,250,1857,1879]" captionTargetBox="[151,1436,199,1834]" captionTargetId="figure@9.[151,1436,193,1836]" captionTargetPageId="9" captionText="FIGURE 3. A) Karataumantispa carnaria (Khramov, 2013) (holotype) with CuA highlighted. B. Karataumantispa carnaria (Khramov, 2013) close up of forewing venation (holotype). C. Karataumantispa monstruosa (Khramov, 2013) close up of hind wing venation (holotype). All wings with annotated wing venation, highlighting the major differences that exclude them from Mesithone: the fusion of R and M, the distal widening of the subcostal space, Sc bending towards R 1 space, and CuA pectinate. (Photographs: A. Khramov)." httpUri="https://zenodo.org/record/232720/files/figure.png" pageId="10" pageNumber="429">Fig.3</figureCitation>
C). The cubital area is also poorly preserved. It was originally described as the CuA being dichotomously branched and CuP deeply forked, however from the photograph and drawings (
<bibRefCitation id="2714E3EEFFE1FFECA8D6FB7DFE22A0EC" author="Khramov" box="[158,467,1231,1256]" pageId="10" pageNumber="429" refString="Khramov, A. V. (2013) New mantidflies (Neuroptera: Mantispidae) from the Upper Jurassic of Kazakhstan. Insect Systematics &amp; Evolution, 44, 221 - 230. http: // dx. doi. org / 10.1163 / 1876312 X- 44032094" type="journal article" year="2013">Khramov, 2013: Figs. 9–11</bibRefCitation>
) the structure of the cubital area cannot be interpreted with confidence.
<taxonomicName id="8485E59CFFE1FFECACB1FB63FA6DA0EC" box="[1273,1436,1233,1256]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="species" species="monstruosa">
<emphasis id="71F1420DFFE1FFECACB1FB63FA6DA0EC" box="[1273,1436,1233,1256]" italics="true" pageId="10" pageNumber="429">K. monstruosa</emphasis>
</taxonomicName>
is therefore tentatively placed within
<taxonomicName id="8485E59CFFE1FFECAA1FFB47FCEAA108" box="[599,795,1269,1292]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="71F1420DFFE1FFECAA1FFB47FCEAA108" box="[599,795,1269,1292]" italics="true" pageId="10" pageNumber="429">Karataumantispa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AC2FF76FFE1FFECAB60FB46FC66A109" box="[808,919,1268,1293]" pageId="10" pageNumber="429" rank="genus">
<emphasis id="71F1420DFFE1FFECAB60FB46FC66A109" bold="true" box="[808,919,1268,1293]" pageId="10" pageNumber="429">gen. nov.</emphasis>
</taxonomicNameLabel>
(pending the discovery of better preserved specimens) due to it differing slightly in foreleg structure, it has a much broader tibia and femur, covered with numerous small hairs and smaller spines (with respect to femur), and the pronotum is also stouter than
<taxonomicName id="8485E59CFFE1FFECAD44FA8FFA7FA150" box="[1292,1422,1341,1364]" class="Insecta" family="Berothidae" genus="Karataumantispa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="species" species="carnaria">
<emphasis id="71F1420DFFE1FFECAD44FA8FFA7FA150" box="[1292,1422,1341,1364]" italics="true" pageId="10" pageNumber="429">K. carnaria</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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