317 lines
36 KiB
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317 lines
36 KiB
XML
<document id="5E7756293706F1EDBEE4352933BA6F6B" ID-DOI="10.5252/g2011n3a2" ID-ISSN="1638-9395" ID-Zenodo-Dep="5378171" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630284327775" checkinUser="marcus" docAuthor="Ivanov, Martin & Böhme, Madelaine" docDate="2011" docId="CA65878DFFF0FFA84566FA76FEE8EEE6" docLanguage="en" docName="Geodiversitas.2011.33.3.411-449.pdf" docOrigin="Geodiversitas 33 (3)" docSource="http://dx.doi.org/10.5252/g2011n3a2" docStyle="DocumentStyle:DF8BFA78045876A77BB343B212438C9A.3:Geodiversitas.2011-2012.journal_article.type1" docStyleId="DF8BFA78045876A77BB343B212438C9A" docStyleName="Geodiversitas.2011-2012.journal_article.type1" docStyleVersion="3" docTitle="Bavarioboa hermi Szyndlar & Schleich 1993" docType="treatment" docVersion="3" lastPageNumber="419" masterDocId="365CFFF5FFF5FFA04463FF9CFF9EEA6C" masterDocTitle="Snakes from Griesbeckerzell (Langhian, Early Badenian), North Alpine Foreland Basin (Germany), with comments on the evolution of snake faunas in Central Europe during the Miocene Climatic Optimum" masterLastPageNumber="449" masterPageNumber="411" pageNumber="416" updateTime="1699214033724" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="7E873EE18FECDED1558F87F29B5A8ECB">
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<mods:title id="894654A8E192FCD6E35C865D77947BDA">Snakes from Griesbeckerzell (Langhian, Early Badenian), North Alpine Foreland Basin (Germany), with comments on the evolution of snake faunas in Central Europe during the Miocene Climatic Optimum</mods:title>
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<mods:name id="3BF778D44503E209610053B8C8CA87C4" type="personal">
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<mods:namePart id="EEC99CAA1015325C88F48C89BDCCE5E5">Ivanov, Martin</mods:namePart>
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<mods:name id="F8D9A7837F51D78E84B10380FA548890" type="personal">
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<mods:roleTerm id="7AE26016CB49036524126B933B620E1A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3B03A66C8F89E46A4233249DDE1D2E80">Böhme, Madelaine</mods:namePart>
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<mods:typeOfResource id="DF81F4DD66B101B65F12EDA2FD0EBE9F">text</mods:typeOfResource>
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<mods:titleInfo id="CE771C7EB8C9F50C1276A78F64DDB3D9">
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<mods:title id="A53627D2598DB7112A0C6B0267FFDBC9">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part id="EBA3DAF0B9E367BB0D89ADABDFB1A7A2">
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<mods:date id="AE725CB6F7FF48495EDF9072C159096B">2011</mods:date>
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<mods:detail id="3C8C8129E6383030F6546819CB8FDC88" type="pubDate">
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<mods:number id="4B69C553151E5F0CACE94B129AC64E7F">2011-09-30</mods:number>
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<mods:detail id="3F7940DECE7032D4E4A6406866629A49" type="volume">
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<mods:number id="56BCA8144AC3A964F6CFE37142932B44">33</mods:number>
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</mods:detail>
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<mods:detail id="41245D371AE0B29A10F1387C26A9FF33" type="issue">
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<mods:number id="E75BBC6B1525FE7A53491EAE82C7E8E3">3</mods:number>
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<mods:start id="0876E3F08D819CFD4418BE20FA209FF9">411</mods:start>
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<mods:end id="DC01074C16649FEF74637B5D6E4DEAFC">449</mods:end>
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<mods:location id="8880AF0A63FC0AA7E15A7ECAF3884E9A">
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<mods:url id="4A2733785370A21CFEF887548B23B412">http://dx.doi.org/10.5252/g2011n3a2</mods:url>
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<mods:classification id="CC3E96144DB025ECB628118740FC9EA7">journal article</mods:classification>
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<mods:identifier id="125FB5BE0A08623DA6E9D40C5F6F2DB9" type="DOI">10.5252/g2011n3a2</mods:identifier>
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<mods:identifier id="D2FB100B5BF93AC84B91C505DEE73D91" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="2B24711FF5FE1AC32AC26EBAC957CCCA" type="Zenodo-Dep">5378171</mods:identifier>
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<treatment id="CA65878DFFF0FFA84566FA76FEE8EEE6" LSID="urn:lsid:plazi:treatment:CA65878DFFF0FFA84566FA76FEE8EEE6" httpUri="http://treatment.plazi.org/id/CA65878DFFF0FFA84566FA76FEE8EEE6" lastPageId="8" lastPageNumber="419" pageId="5" pageNumber="416">
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<subSubSection id="0AD66510FFF0FFA54566FA76FD93EC49" pageId="5" pageNumber="416" type="nomenclature">
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<paragraph id="4273369BFFF0FFA54566FA76FD93EC49" blockId="5.[235,525,1514,1573]" pageId="5" pageNumber="416">
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<heading id="193B81F7FFF0FFA54566FA76FD93EC49" centered="true" fontSize="11" level="2" pageId="5" pageNumber="416" reason="2">
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<taxonomicName id="85CC4D18FFF0FFA54566FA76FD93EC49" authority="Szyndlar & Schleich, 1993" authorityName="Szyndlar & Schleich" authorityYear="1993" class="Reptilia" family="Boidae" genus="Bavarioboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="416" phylum="Chordata" rank="species" species="hermi">
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<emphasis id="70B8EA89FFF0FFA54566FA76FE1CEC69" bold="true" box="[261,386,1514,1541]" italics="true" pageId="5" pageNumber="416">Bavarioboa</emphasis>
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aff.
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<emphasis id="70B8EA89FFF0FFA545D1FA76FE6CEC69" bold="true" box="[434,498,1514,1541]" italics="true" pageId="5" pageNumber="416">hermi</emphasis>
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<bibRefCitation id="265D4B6AFFF0FFA54488F996FD93EC49" author="SZYNDLAR Z. & SCHLEICH H. H." box="[235,525,1546,1573]" pageId="5" pageNumber="416" pagination="1 - 47" refId="ref23308" refString="SZYNDLAR Z. & SCHLEICH H. H. 1993. - Description of Miocene snakes from Petersbuch 2 with comments on the Lower and Middle Miocene ophidian faunas of Southern Germany. Stuttgarter Beitrage zur Naturkunde B 192: 1 - 47, 10 figs." type="journal article" year="1993">Szyndlar & Schleich, 1993</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0AD66510FFF0FFA54412F9CBFC78EA83" pageId="5" pageNumber="416" type="materials_examined">
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<paragraph id="4273369BFFF0FFA54412F9CBFF46ECCF" blockId="5.[113,648,1622,1699]" pageId="5" pageNumber="416">
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MATERIAL EXAMINED. — 2 trunk vertebrae (
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<collectionCode id="24DDAE5EFFF0FFA54626F9CAFD18EC02" box="[581,646,1622,1646]" country="0" httpUri="http://grbio.org/cool/r9n2-w98n" name="Bayerische Staatssammlung fuer Palaeontologie und Geologie" pageId="5" pageNumber="416">BSPG</collectionCode>
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1997 XIII 499, 500); 1 cloacal vertebra (
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<collectionCode id="24DDAE5EFFF0FFA5466CF9EDFDD3ECE5" box="[527,589,1649,1673]" country="0" httpUri="http://grbio.org/cool/r9n2-w98n" name="Bayerische Staatssammlung fuer Palaeontologie und Geologie" pageId="5" pageNumber="416">BSPG</collectionCode>
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1997 XIII 501).
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</paragraph>
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<paragraph id="4273369BFFF0FFA546C6FF44FC78EA83" blockId="5.[677,998,215,239]" box="[677,998,215,239]" pageId="5" pageNumber="416">LOCALITY. — Griesbeckerzell 1a.</paragraph>
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</subSubSection>
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<subSubSection id="0AD66510FFF0FFA646C6FE84FEE8ECC8" lastPageId="6" lastPageNumber="417" pageId="5" pageNumber="416" type="description">
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<paragraph id="4273369BFFF0FFA546C6FE84FCABEB5C" blockId="5.[674,1212,280,1700]" box="[677,821,280,306]" pageId="5" pageNumber="416">DESCRIPTION</paragraph>
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<paragraph id="4273369BFFF0FFA546C6FEABFC2EEB3D" blockId="5.[674,1212,280,1700]" box="[677,944,311,337]" pageId="5" pageNumber="416">
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<emphasis id="70B8EA89FFF0FFA546C6FEABFC2EEB3D" box="[677,944,311,337]" italics="true" pageId="5" pageNumber="416">
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Trunk vertebrae (
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<figureCitation id="DAF72A1EFFF0FFA5473BFEABFC36EB3D" box="[856,936,311,337]" captionStart="FIG" captionStartId="7.[113,124,1127,1144]" captionTargetBox="[121,1202,217,1099]" captionTargetId="figure-228@7.[117,1205,210,1105]" captionTargetPageId="7" captionText="FIG. 4. — Bavarioboa aff.hermi from the Middle Miocene (MN 6, base) of Griesbeckerzell 1a in lateral (l), dorsal (d), ventral (v), cranial (cr), and caudal (ca) views:A, middle trunk vertebra (BSPG 1997 XIII 499);B, cloacal vertebra (BSPG 1997 XIII 501).Abbreviations:cd, condyle; ct, cotyle; hae, haemapophysis; hk, haemal keel; lf, lateral foramen; na, neural arch; nc, neural canal; ns, neural spine; pr, prezygapophysis; prf, prezygapophyseal articular facet; prp, prezygapophyseal process; po, postzygapophysis; pof, postzygapophyseal articular facet; scf, subcentral foramen; scr, subcentral ridge; syn, synapophysis; zy, zygosphene; zyg, zygantrum. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5378177" httpUri="https://zenodo.org/record/5378177/files/figure.png" pageId="5" pageNumber="416">Fig. 4A</figureCitation>
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)
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</emphasis>
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</paragraph>
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<paragraph id="4273369BFFF0FFA546C6FECBFB1DE921" blockId="5.[674,1212,280,1700]" pageId="5" pageNumber="416">In lateral view, the more complete vertebra (BSPG 1997 XIII 499) is slightly higher than long. The neural spine is low and short, occupying about one third of the neural arch length. It was not possible to observe the typical dorsal thickening of the neural spine because the dorsal tip is not preserved, but we assume that at least in one vertebra the dorsal thickening was not developed. The lateral foramina are indistinct. The short interzygapophyseal ridges are sharp. The paradiapophyses are damaged; the subdivision into para- and diapophysis is indistinct. The diapophysis originally was much more smaller than the parapophysis. The relatively straight or slightly dorsally arched subcentral ridges are strongly developed and reach the condylar base. The ventral margin of the haemal keel is arched dorsally.</paragraph>
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<paragraph id="4273369BFFF0FFA546D8FCCDFBF8EEE6" blockId="5.[674,1212,280,1700]" pageId="5" pageNumber="416">In dorsal view, the vertebrae are wider than long. The prezygapophyseal articular facets are subtriangular and elongated in lateral direction. Prezygapophyseal processes are not visible from above.The cranial margin of the zygosphene is almost straight; sometimes two small lateral lobes are developed.The base of the neural spine rises at the level of the posterior margin of the prezygapophyseal articular facets.The posterior end of the neural spine is widened. The interzygapophyseal constriction is moderately deep.</paragraph>
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<paragraph id="4273369BFFF0FFA546D8FB12FCCBEF0B" blockId="5.[674,1212,280,1700]" pageId="5" pageNumber="416">In ventral view, the centrum is slightly wider than long. The prominent haemal keel is limited by the deep subcentral grooves. Subcentral foramina are small and situated at the base of the anterior part of the haemal keel. The subtriangular to oval postzygapophyseal articular facets are strongly laterally elongated.</paragraph>
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<paragraph id="4273369BFFF0FFA546D8FAF0FC09EC6A" blockId="5.[674,1212,280,1700]" pageId="5" pageNumber="416">In cranial view, the zygosphene is straight and is either as wide as the cotyle or markedly wider. In one vertebra the zygosphenal lip is thickened. The cotylar rim is circular; paracotylar foramina in deep depressions are absent.</paragraph>
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<paragraph id="4273369BFFF0FFA546D8F996FD43ECC8" blockId="5.[674,1212,280,1700]" pageId="5" pageNumber="416">
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In caudal view, the neural arch is weakly vaulted, and the condyle is rounded and situated on a short neck. Measurements of the larger vertebra are as follows: cl =
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<quantity id="85349B7EFFF0FFA5474DF9F5FC0AECE8" box="[814,916,1641,1668]" metricMagnitude="-3" metricUnit="m" metricValue="5.37" pageId="5" pageNumber="416" unit="mm" value="5.37">5.37 mm</quantity>
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; naw =
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<quantity id="85349B7EFFF0FFA54784F9F5FBCEECE8" box="[999,1104,1641,1668]" metricMagnitude="-3" metricUnit="m" metricValue="6.32" pageId="5" pageNumber="416" unit="mm" value="6.32">6.32 mm</quantity>
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.; cl/naw = 0.85.
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</paragraph>
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<caption id="16B36613FFF3FFA64412FA91FBBDEFFC" ID-DOI="http://doi.org/10.5281/zenodo.5378175" ID-Zenodo-Dep="5378175" httpUri="https://zenodo.org/record/5378175/files/figure.png" pageId="6" pageNumber="417" startId="6.[113,124,1293,1310]" targetBox="[123,1206,214,1251]" targetPageId="6">
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<paragraph id="4273369BFFF3FFA64412FA91FBBDEFFC" blockId="6.[112,1211,1293,1424]" pageId="6" pageNumber="417">
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FIG. 3. — Synoptical chart of the chronology for the Early to Middle Miocene lithostratigraphic units in the Bavarian part of the NAFB (modified from
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<bibRefCitation id="265D4B6AFFF3FFA64491FAB8FE33EF59" author="ABDUL AZIZ H. & BOHME M. & ROCHOLL A. & PRIETO J. & WIJBRANS J. & BACHTADSE V." box="[242,429,1316,1333]" pageId="6" pageNumber="417" pagination="1859 - 1886" refId="ref20091" refString="ABDUL AZIZ H., BOHME M., ROCHOLL A., PRIETO J., WIJBRANS J. & BACHTADSE V. 2010. - Integrated stratigraphy and 40 Ar / 39 Ar chronology of the Early to Middle Miocene Upper Freshwater Molasse in western Bavaria (Germany). International Journal of Earth Sciences, 99 (8): 1859 - 1886." type="journal article" year="2010">
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Abdul Aziz
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<emphasis id="70B8EA89FFF3FFA64530FAB8FEE9EF59" box="[339,375,1316,1333]" italics="true" pageId="6" pageNumber="417">et al</emphasis>
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. 2010
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</bibRefCitation>
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) and stratigraphic position of the Griesbeckerzell localities (
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<emphasis id="70B8EA89FFF3FFA647C9FAB8FC25EF5A" bold="true" box="[938,955,1316,1334]" pageId="6" pageNumber="417">*</emphasis>
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):
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<emphasis id="70B8EA89FFF3FFA647A8FAB8FC4BEF59" bold="true" box="[971,981,1316,1333]" pageId="6" pageNumber="417">1</emphasis>
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, Marine Molasse;
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<emphasis id="70B8EA89FFF3FFA64010FAB8FBE3EF59" bold="true" box="[1139,1149,1316,1333]" pageId="6" pageNumber="417">2</emphasis>
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, Grimmelfingen beds;
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<emphasis id="70B8EA89FFF3FFA6449EFAA7FE99EF20" bold="true" box="[253,263,1339,1356]" pageId="6" pageNumber="417">3</emphasis>
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, Albstein;
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<emphasis id="70B8EA89FFF3FFA64503FAA7FEF4EF20" bold="true" box="[352,362,1339,1356]" pageId="6" pageNumber="417">4</emphasis>
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, Kirchberg Formation;
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<emphasis id="70B8EA89FFF3FFA64648FAA7FDABEF20" bold="true" box="[555,565,1339,1356]" pageId="6" pageNumber="417">5</emphasis>
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, Sand-Kalkmergel-Serie and untere Bunte Mergel Serie;
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<emphasis id="70B8EA89FFF3FFA64071FAA7FB82EF20" bold="true" box="[1042,1052,1339,1356]" pageId="6" pageNumber="417">6</emphasis>
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, Limnische Untere Serie;
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<emphasis id="70B8EA89FFF3FFA644C6FACDFF31EF0E" bold="true" box="[165,175,1361,1378]" pageId="6" pageNumber="417">7</emphasis>
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, NÖrdlicher Vollschotter,lower part;
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<emphasis id="70B8EA89FFF3FFA645B9FACDFE7AEF0E" bold="true" box="[474,484,1361,1378]" pageId="6" pageNumber="417">8</emphasis>
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, Fluviatile Untere Serie;
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<emphasis id="70B8EA89FFF3FFA646C8FACDFD2BEF0E" bold="true" box="[683,693,1361,1378]" pageId="6" pageNumber="417">9</emphasis>
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, NÖrdlicher Vollschotter, upper part;
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<emphasis id="70B8EA89FFF3FFA64781FACDFC69EF0E" bold="true" box="[994,1015,1361,1378]" pageId="6" pageNumber="417">10</emphasis>
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, Fluviatile Untere Serie;
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<emphasis id="70B8EA89FFF3FFA64412FAF4FF18EF15" bold="true" box="[113,134,1384,1401]" pageId="6" pageNumber="417">11</emphasis>
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, Zwischenmergel;
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<emphasis id="70B8EA89FFF3FFA6457DFAF4FEACEF15" bold="true" box="[286,306,1384,1401]" pageId="6" pageNumber="417">12</emphasis>
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, NÖrdlicher Vollschotter,upper part;
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<emphasis id="70B8EA89FFF3FFA6463AFAF4FDF3EF15" bold="true" box="[601,621,1384,1401]" pageId="6" pageNumber="417">13</emphasis>
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, GerÖllsand Serie;
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<emphasis id="70B8EA89FFF3FFA64765FAF4FC84EF15" bold="true" box="[774,794,1384,1401]" pageId="6" pageNumber="417">14</emphasis>
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, Brock-horizon;
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<emphasis id="70B8EA89FFF3FFA647C2FAF4FC2BEF15" bold="true" box="[929,949,1384,1401]" pageId="6" pageNumber="417">15</emphasis>
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, Sand-Mergel-Decke;
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<emphasis id="70B8EA89FFF3FFA6400EFAF4FB1FEF15" bold="true" box="[1133,1153,1384,1401]" pageId="6" pageNumber="417">16</emphasis>
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, dated volcanic ash;
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<emphasis id="70B8EA89FFF3FFA64480FAE3FF66EFFC" bold="true" box="[227,248,1407,1424]" pageId="6" pageNumber="417">17</emphasis>
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, undated volcanic ash;
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<emphasis id="70B8EA89FFF3FFA645DCFAE3FE4AEFFC" bold="true" box="[447,468,1407,1424]" pageId="6" pageNumber="417">18</emphasis>
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, Lower Laimering Series, Ubergangsschichten;
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<emphasis id="70B8EA89FFF3FFA64701FAE3FCE9EFFC" bold="true" box="[866,887,1407,1424]" pageId="6" pageNumber="417">19</emphasis>
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, Steinbalmensande.
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</paragraph>
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</caption>
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<paragraph id="4273369BFFF3FFA64412FA57FEE1EF89" blockId="6.[111,648,1483,1700]" box="[113,383,1483,1509]" pageId="6" pageNumber="417">
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<emphasis id="70B8EA89FFF3FFA64412FA57FEE1EF89" box="[113,383,1483,1509]" italics="true" pageId="6" pageNumber="417">
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Cloacal vertebra (
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<figureCitation id="DAF72A1EFFF3FFA6454BFA57FEE9EF89" box="[296,375,1483,1509]" captionStart="FIG" captionStartId="7.[113,124,1127,1144]" captionTargetBox="[121,1202,217,1099]" captionTargetId="figure-228@7.[117,1205,210,1105]" captionTargetPageId="7" captionText="FIG. 4. — Bavarioboa aff.hermi from the Middle Miocene (MN 6, base) of Griesbeckerzell 1a in lateral (l), dorsal (d), ventral (v), cranial (cr), and caudal (ca) views:A, middle trunk vertebra (BSPG 1997 XIII 499);B, cloacal vertebra (BSPG 1997 XIII 501).Abbreviations:cd, condyle; ct, cotyle; hae, haemapophysis; hk, haemal keel; lf, lateral foramen; na, neural arch; nc, neural canal; ns, neural spine; pr, prezygapophysis; prf, prezygapophyseal articular facet; prp, prezygapophyseal process; po, postzygapophysis; pof, postzygapophyseal articular facet; scf, subcentral foramen; scr, subcentral ridge; syn, synapophysis; zy, zygosphene; zyg, zygantrum. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5378177" httpUri="https://zenodo.org/record/5378177/files/figure.png" pageId="6" pageNumber="417">Fig. 4B</figureCitation>
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)
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</emphasis>
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</paragraph>
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<paragraph id="4273369BFFF3FFA64412FA77FEE8ECC8" blockId="6.[111,648,1483,1700]" pageId="6" pageNumber="417">A single fragmentary vertebra is preserved with broken off prezygapophyses. Bases of the incipient haemapophyses occupy unusually anterior position. They are developed at the ventral margin of the haemal keel, which is typical for the cloacal region of the vertebral column.</paragraph>
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</subSubSection>
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<subSubSection id="0AD66510FFF3FFA846C6FA50FEE8EEE6" lastPageId="8" lastPageNumber="419" pageId="6" pageNumber="417" type="discussion">
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<paragraph id="4273369BFFF3FFA646C6FA50FCB8EF88" blockId="6.[674,1211,1484,1700]" box="[677,806,1484,1510]" pageId="6" pageNumber="417">DISCUSSION</paragraph>
|
|
<paragraph id="4273369BFFF3FFA746C6FA77FB24ECC8" blockId="6.[674,1211,1484,1700]" lastBlockId="7.[674,1212,1293,1700]" lastPageId="7" lastPageNumber="418" pageId="6" pageNumber="417">
|
|
Although paracotylar foramina are absent, the vertebrae can be assigned to the Boinae based on the relatively small dimensions (compared with Miocene
|
|
<taxonomicName id="85CC4D18FFF3FFA646C1F9D6FC59EC09" authority="Fitzinger, 1826" authorityName="Fitzinger" authorityYear="1826" box="[674,967,1610,1637]" class="Reptilia" family="Colubridae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="417" phylum="Chordata" rank="subFamily" subFamily="Pythoninae">Pythoninae Fitzinger, 1826</taxonomicName>
|
|
, e.g.,
|
|
<taxonomicName id="85CC4D18FFF3FFA6406AF9D6FC0FECE8" authority="Szyndlar & Rage, 2003" authorityName="Szyndlar & Rage" authorityYear="2003" class="Reptilia" family="Pythonidae" genus="Python" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="417" phylum="Chordata" rank="species" species="europaeus">
|
|
<emphasis id="70B8EA89FFF3FFA6406AF9D6FB24EC08" box="[1033,1210,1610,1636]" italics="true" pageId="6" pageNumber="417">Python europaeus</emphasis>
|
|
<bibRefCitation id="265D4B6AFFF3FFA646C6F9F5FC0FECE8" author="SZYNDLAR Z. & RAGE J. - C." box="[677,913,1641,1668]" pageId="6" pageNumber="417" refId="ref23455" refString="SZYNDLAR Z. & RAGE J. - C. 2003. - Non-Erycine Booidea from the Oligocene and Miocene of Europe. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, 109 p." type="book" year="2003">Szyndlar & Rage, 2003</bibRefCitation>
|
|
</taxonomicName>
|
|
[cf.
|
|
<bibRefCitation id="265D4B6AFFF3FFA647D8F9F5FBA3ECE8" author="IVANOV M." box="[955,1085,1641,1668]" pageId="6" pageNumber="417" pagination="559 - 588" refId="ref21638" refString="IVANOV M. 2000. - Snakes of the lower / middle Miocene transition at Vieux Collonges (Rhone; France), with comments on the colonisation of western Europe by colubroids. Geodiversitas 22 (4): 559 - 588." type="journal article" year="2000">Ivanov 2000</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="265D4B6AFFF3FFA64026F9F5FC8AECC8" author="SZYNDLAR Z. & RAGE J. - C." pageId="6" pageNumber="417" refId="ref23455" refString="SZYNDLAR Z. & RAGE J. - C. 2003. - Non-Erycine Booidea from the Oligocene and Miocene of Europe. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, 109 p." type="book" year="2003">Szyndlar & Rage 2003</bibRefCitation>
|
|
]) and several typical features of the genus
|
|
<taxonomicName id="85CC4D18FFF2FFA74412FA91FF7DEF4B" authorityName="Szyndlar & Schleich" authorityYear="1993" box="[113,227,1293,1319]" class="Reptilia" family="Boidae" genus="Bavarioboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="418" phylum="Chordata" rank="genus">
|
|
<emphasis id="70B8EA89FFF2FFA74412FA91FF7DEF4B" box="[113,227,1293,1319]" italics="true" pageId="7" pageNumber="418">Bavarioboa</emphasis>
|
|
</taxonomicName>
|
|
. The general morphology concurs with that of
|
|
<taxonomicName id="85CC4D18FFF2FFA744A3FAB1FEAFEF2B" authorityName="Szyndlar & Schleich" authorityYear="1993" box="[192,305,1325,1351]" class="Reptilia" family="Boidae" genus="Bavarioboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="418" phylum="Chordata" rank="genus">
|
|
<emphasis id="70B8EA89FFF2FFA744A3FAB1FEAFEF2B" box="[192,305,1325,1351]" italics="true" pageId="7" pageNumber="418">Bavarioboa</emphasis>
|
|
</taxonomicName>
|
|
, including the absence of dorsal thickening of the neural spine in the middle trunk vertebrae,the straight cranial margin of the zygosphene in cranial view, and the weakly vaulted neural arch (cf.
|
|
<bibRefCitation id="265D4B6AFFF2FFA744FAFA37FE1BEFAA" author="SZYNDLAR Z. & RAGE J. - C." box="[153,389,1451,1478]" pageId="7" pageNumber="418" refId="ref23455" refString="SZYNDLAR Z. & RAGE J. - C. 2003. - Non-Erycine Booidea from the Oligocene and Miocene of Europe. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, 109 p." type="book" year="2003">Szyndlar & Rage 2003</bibRefCitation>
|
|
). Vertebrae of
|
|
<taxonomicName id="85CC4D18FFF2FFA74676FA37FF4AEF89" authorityName="Szyndlar & Schleich" authorityYear="1993" class="Reptilia" family="Boidae" genus="Bavarioboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="418" phylum="Chordata" rank="species" species="hermi">
|
|
<emphasis id="70B8EA89FFF2FFA74676FA37FD19EFA9" box="[533,647,1451,1477]" italics="true" pageId="7" pageNumber="418">Bavarioboa</emphasis>
|
|
aff.
|
|
<emphasis id="70B8EA89FFF2FFA744FAFA57FF4AEF89" box="[153,212,1483,1509]" italics="true" pageId="7" pageNumber="418">hermi</emphasis>
|
|
</taxonomicName>
|
|
resemble those of
|
|
<taxonomicName id="85CC4D18FFF2FFA745FBFA57FDD2EF89" authorityName="Szyndlar & Schleich" authorityYear="1993" box="[408,588,1483,1509]" class="Reptilia" family="Boidae" genus="Bavarioboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="418" phylum="Chordata" rank="species" species="hermi">
|
|
<emphasis id="70B8EA89FFF2FFA745FBFA57FDD2EF89" box="[408,588,1483,1509]" italics="true" pageId="7" pageNumber="418">Bavarioboa hermi</emphasis>
|
|
</taxonomicName>
|
|
from the
|
|
<typeStatus id="9D778839FFF2FFA744FAFA77FF58EC6A" box="[153,198,1515,1542]" pageId="7" pageNumber="418">type</typeStatus>
|
|
locality Petersbuch 2,
|
|
<collectingCountry id="3ADB760BFFF2FFA745D3FA77FD8CEC6A" box="[432,530,1515,1542]" name="Germany" pageId="7" pageNumber="418">Germany</collectingCountry>
|
|
(Early Miocene, late Eggenburgian) in the following features: 1) the cranial margin of the zygosphene is straight and small lateral lobes may occur; 2) the haemal keel is widened posteriorly, especially in
|
|
<specimenCount id="54CAFD12FFF2FFA74591F9F5FD1BECE8" box="[498,645,1641,1668]" pageId="7" pageNumber="418" type="generic">one specimen</specimenCount>
|
|
of a posterior precaudal vertebra; and 3) the neural spine of trunk vertebrae is not thickened. However, the trunk vertebrae are somewhat lower compared to those of
|
|
<taxonomicName id="85CC4D18FFF2FFA7469AFAD0FC37EF0A" authorityName="Szyndlar & Schleich" authorityYear="1993" box="[761,937,1356,1382]" class="Reptilia" family="Boidae" genus="Bavarioboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="418" phylum="Chordata" rank="species" species="hermi">
|
|
<emphasis id="70B8EA89FFF2FFA7469AFAD0FC37EF0A" box="[761,937,1356,1382]" italics="true" pageId="7" pageNumber="418">Bavarioboa hermi</emphasis>
|
|
</taxonomicName>
|
|
, but the neural spine is not as low as the neural spine of the late Early Miocene (Karpatian)
|
|
<taxonomicName id="85CC4D18FFF2FFA7477FFA10FB24EFCB" authority="Szyndlar & Rage, 2003" authorityName="Szyndlar & Rage" authorityYear="2003" box="[796,1210,1420,1447]" class="Reptilia" family="Boidae" genus="Bavarioboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="418" phylum="Chordata" rank="species" species="ultima">
|
|
<emphasis id="70B8EA89FFF2FFA7477FFA10FC51EFCA" box="[796,975,1420,1446]" italics="true" pageId="7" pageNumber="418">Bavarioboa ultima</emphasis>
|
|
<bibRefCitation id="265D4B6AFFF2FFA747B0FA10FB24EFCB" author="SZYNDLAR Z. & RAGE J. - C." box="[979,1210,1420,1447]" pageId="7" pageNumber="418" refId="ref23455" refString="SZYNDLAR Z. & RAGE J. - C. 2003. - Non-Erycine Booidea from the Oligocene and Miocene of Europe. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, 109 p." type="book" year="2003">Szyndlar &Rage,2003</bibRefCitation>
|
|
</taxonomicName>
|
|
from Rothenstein 13,
|
|
<collectingCountry id="3ADB760BFFF2FFA747E3FA37FC7CEFAA" box="[896,994,1451,1478]" name="Germany" pageId="7" pageNumber="418">Germany</collectingCountry>
|
|
(cf.
|
|
<bibRefCitation id="265D4B6AFFF2FFA7406FFA37FD42EF8A" author="SZYNDLAR Z. & RAGE J. - C." pageId="7" pageNumber="418" refId="ref23455" refString="SZYNDLAR Z. & RAGE J. - C. 2003. - Non-Erycine Booidea from the Oligocene and Miocene of Europe. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, 109 p." type="book" year="2003">Szyndlar & Rage 2003</bibRefCitation>
|
|
). Morphologically, vertebrae of
|
|
<taxonomicName id="85CC4D18FFF2FFA74041FA57FD7FEC69" authorityName="Szyndlar & Schleich" authorityYear="1993" class="Reptilia" family="Boidae" genus="Bavarioboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="418" phylum="Chordata" rank="species" species="hermi">
|
|
<emphasis id="70B8EA89FFF2FFA74041FA57FB0AEF89" box="[1058,1172,1483,1509]" italics="true" pageId="7" pageNumber="418">Bavarioboa</emphasis>
|
|
aff.
|
|
<emphasis id="70B8EA89FFF2FFA746C6FA77FD7FEC69" box="[677,737,1515,1541]" italics="true" pageId="7" pageNumber="418">hermi</emphasis>
|
|
</taxonomicName>
|
|
from Griesbeckerzell 1a seem to occupy an intermediate position between the Early Miocene
|
|
<taxonomicName id="85CC4D18FFF2FFA746C5F9B6FCFBEC28" authorityName="Szyndlar & Schleich" authorityYear="1993" box="[678,869,1578,1604]" class="Reptilia" family="Boidae" genus="Bavarioboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="418" phylum="Chordata" rank="species" species="hermi">
|
|
<emphasis id="70B8EA89FFF2FFA746C5F9B6FCFBEC28" box="[678,869,1578,1604]" italics="true" pageId="7" pageNumber="418">Bavarioboa hermi</emphasis>
|
|
</taxonomicName>
|
|
from Petersbuch 2 (
|
|
<collectingCountry id="3ADB760BFFF2FFA7402BF9B6FB2CEC29" box="[1096,1202,1578,1605]" name="Germany" pageId="7" pageNumber="418">Germany</collectingCountry>
|
|
) and Dolnice (
|
|
<collectingCountry id="3ADB760BFFF2FFA74757F9D6FC45EC09" box="[820,987,1610,1637]" name="Czech Republic" pageId="7" pageNumber="418">Czech Republic</collectingCountry>
|
|
; Ottnangian) and the late Early Miocene
|
|
<taxonomicName id="85CC4D18FFF2FFA74708F9F5FBBBECEF" authorityName="Szyndlar & Rage" authorityYear="2003" box="[875,1061,1641,1667]" class="Reptilia" family="Boidae" genus="Bavarioboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="418" phylum="Chordata" rank="species" species="ultima">
|
|
<emphasis id="70B8EA89FFF2FFA74708F9F5FBBBECEF" box="[875,1061,1641,1667]" italics="true" pageId="7" pageNumber="418">Bavarioboa ultima</emphasis>
|
|
</taxonomicName>
|
|
from Rothenstein 13 (
|
|
<collectingCountry id="3ADB760BFFF2FFA74760F915FCF4ECC8" box="[771,874,1673,1700]" name="Germany" pageId="7" pageNumber="418">Germany</collectingCountry>
|
|
), which has been regarded as the
|
|
</paragraph>
|
|
<caption id="16B36613FFF2FFA74412FBFBFBFBEEBF" ID-DOI="http://doi.org/10.5281/zenodo.5378177" ID-Zenodo-Dep="5378177" httpUri="https://zenodo.org/record/5378177/files/figure.png" pageId="7" pageNumber="418" startId="7.[113,124,1127,1144]" targetBox="[121,1202,217,1099]" targetPageId="7">
|
|
<paragraph id="4273369BFFF2FFA74412FBFBFBFBEEBF" blockId="7.[112,1211,1127,1235]" pageId="7" pageNumber="418">
|
|
FIG. 4. —
|
|
<taxonomicName id="85CC4D18FFF2FFA744DEFBFBFEF7EE14" authorityName="Szyndlar & Schleich" authorityYear="1993" box="[189,361,1127,1144]" class="Reptilia" family="Boidae" genus="Bavarioboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="418" phylum="Chordata" rank="species" species="hermi">
|
|
<emphasis id="70B8EA89FFF2FFA744DEFBFBFE87EE14" box="[189,281,1127,1144]" italics="true" pageId="7" pageNumber="418">Bavarioboa</emphasis>
|
|
aff.
|
|
<emphasis id="70B8EA89FFF2FFA74558FBFBFEF7EE14" box="[315,361,1127,1144]" italics="true" pageId="7" pageNumber="418">hermi</emphasis>
|
|
</taxonomicName>
|
|
from the Middle Miocene (MN 6, base) of Griesbeckerzell 1a in lateral (
|
|
<emphasis id="70B8EA89FFF2FFA747C2FBFBFC38EE14" bold="true" box="[929,934,1127,1144]" pageId="7" pageNumber="418">l</emphasis>
|
|
), dorsal (
|
|
<emphasis id="70B8EA89FFF2FFA7478CFBFBFC64EE14" bold="true" box="[1007,1018,1127,1144]" pageId="7" pageNumber="418">d</emphasis>
|
|
), ventral (
|
|
<emphasis id="70B8EA89FFF2FFA7402BFBFBFBCCEE14" bold="true" box="[1096,1106,1127,1144]" pageId="7" pageNumber="418">v</emphasis>
|
|
), cranial (
|
|
<emphasis id="70B8EA89FFF2FFA740FDFBFBFB2EEE14" bold="true" box="[1182,1200,1127,1144]" pageId="7" pageNumber="418">cr</emphasis>
|
|
), and caudal (
|
|
<emphasis id="70B8EA89FFF2FFA744B1FBE2FF79EEE3" bold="true" box="[210,231,1150,1167]" pageId="7" pageNumber="418">ca</emphasis>
|
|
) views:
|
|
<emphasis id="70B8EA89FFF2FFA74545FBE2FEADEEE3" bold="true" box="[294,307,1150,1167]" pageId="7" pageNumber="418">A</emphasis>
|
|
, middle trunk vertebra (BSPG 1997 XIII 499);
|
|
<emphasis id="70B8EA89FFF2FFA746F4FBE2FD3AEEE3" bold="true" box="[663,676,1150,1167]" pageId="7" pageNumber="418">B</emphasis>
|
|
, cloacal vertebra (BSPG 1997 XIII 501).Abbreviations:
|
|
<emphasis id="70B8EA89FFF2FFA74035FBE2FBF2EEE3" bold="true" box="[1110,1132,1150,1167]" pageId="7" pageNumber="418">cd</emphasis>
|
|
, condyle;
|
|
<emphasis id="70B8EA89FFF2FFA74413FB08FF1FEEC9" bold="true" box="[112,129,1172,1189]" pageId="7" pageNumber="418">ct</emphasis>
|
|
, cotyle;
|
|
<emphasis id="70B8EA89FFF2FFA744ABFB08FF76EEC9" bold="true" box="[200,232,1172,1189]" pageId="7" pageNumber="418">hae</emphasis>
|
|
, haemapophysis;
|
|
<emphasis id="70B8EA89FFF2FFA7451EFB08FE0CEEC9" bold="true" box="[381,402,1172,1189]" pageId="7" pageNumber="418">hk</emphasis>
|
|
, haemal keel;
|
|
<emphasis id="70B8EA89FFF2FFA7466AFB08FD8AEEC9" bold="true" box="[521,532,1172,1189]" pageId="7" pageNumber="418">lf</emphasis>
|
|
, lateral foramen;
|
|
<emphasis id="70B8EA89FFF2FFA746C1FB08FD29EEC9" bold="true" box="[674,695,1172,1189]" pageId="7" pageNumber="418">na</emphasis>
|
|
, neural arch;
|
|
<emphasis id="70B8EA89FFF2FFA74745FB08FCA5EEC9" bold="true" box="[806,827,1172,1189]" pageId="7" pageNumber="418">nc</emphasis>
|
|
, neural canal;
|
|
<emphasis id="70B8EA89FFF2FFA747D0FB08FC56EEC9" bold="true" box="[947,968,1172,1189]" pageId="7" pageNumber="418">ns</emphasis>
|
|
, neural spine;
|
|
<emphasis id="70B8EA89FFF2FFA74023FB08FBCCEEC9" bold="true" box="[1088,1106,1172,1189]" pageId="7" pageNumber="418">pr</emphasis>
|
|
, prezygapophysis;
|
|
<emphasis id="70B8EA89FFF2FFA744D3FB37FF56EED0" bold="true" box="[176,200,1195,1212]" pageId="7" pageNumber="418">prf</emphasis>
|
|
, prezygapophyseal articular facet;
|
|
<emphasis id="70B8EA89FFF2FFA74587FB37FD9CEED0" bold="true" box="[484,514,1195,1212]" pageId="7" pageNumber="418">prp</emphasis>
|
|
, prezygapophyseal process;
|
|
<emphasis id="70B8EA89FFF2FFA74693FB37FC98EED0" bold="true" box="[752,774,1195,1212]" pageId="7" pageNumber="418">po</emphasis>
|
|
, postzygapophysis;
|
|
<emphasis id="70B8EA89FFF2FFA747CDFB37FC54EED0" bold="true" box="[942,970,1195,1212]" pageId="7" pageNumber="418">pof</emphasis>
|
|
, postzygapophyseal articular facet;
|
|
<emphasis id="70B8EA89FFF2FFA744C7FB5EFF20EEBF" bold="true" box="[164,190,1218,1235]" pageId="7" pageNumber="418">scf</emphasis>
|
|
, subcentral foramen;
|
|
<emphasis id="70B8EA89FFF2FFA74513FB5EFE15EEBF" bold="true" box="[368,395,1218,1235]" pageId="7" pageNumber="418">scr</emphasis>
|
|
, subcentral ridge;
|
|
<emphasis id="70B8EA89FFF2FFA74647FB5EFDDCEEBF" bold="true" box="[548,578,1218,1235]" pageId="7" pageNumber="418">syn</emphasis>
|
|
, synapophysis;
|
|
<emphasis id="70B8EA89FFF2FFA746A5FB5EFD47EEBF" bold="true" box="[710,729,1218,1235]" pageId="7" pageNumber="418">zy</emphasis>
|
|
, zygosphene;
|
|
<emphasis id="70B8EA89FFF2FFA74733FB5EFCF0EEBF" bold="true" box="[848,878,1218,1235]" pageId="7" pageNumber="418">zyg</emphasis>
|
|
, zygantrum. Scale bar: 2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="4273369BFFFDFFA8460BFC34FDE4E9D0" blockId="8.[616,634,936,956]" box="[616,634,936,956]" pageId="8" pageNumber="419">ct</paragraph>
|
|
<caption id="16B36613FFFDFFA84412FC7DFC5EEE65" ID-DOI="http://doi.org/10.5281/zenodo.5378179" ID-Zenodo-Dep="5378179" httpUri="https://zenodo.org/record/5378179/files/figure.png" pageId="8" pageNumber="419" startId="8.[113,124,993,1010]" targetBox="[147,1199,215,930]" targetPageId="8">
|
|
<paragraph id="4273369BFFFDFFA84412FC7DFC5EEE65" blockId="8.[113,1209,993,1033]" pageId="8" pageNumber="419">
|
|
FIG. 5. — cf.
|
|
<taxonomicName id="85CC4D18FFFDFFA844BBFC7DFECBE99E" box="[216,341,993,1010]" class="Reptilia" family="Boidae" genus="Bavarioboa" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="419" phylum="Chordata" rank="species" species="undetermined">
|
|
<emphasis id="70B8EA89FFFDFFA844BBFC7DFEA8E99E" box="[216,310,993,1010]" italics="true" pageId="8" pageNumber="419">Bavarioboa</emphasis>
|
|
sp.
|
|
</taxonomicName>
|
|
from the Middle Miocene (MN 6, base) of Griesbeckerzell 1a. Anterior trunk vertebra (BSPG 1997 XIII 502) in lateral (
|
|
<emphasis id="70B8EA89FFFDFFA844A1FC64FF59EE65" bold="true" box="[194,199,1016,1033]" pageId="8" pageNumber="419">l</emphasis>
|
|
), dorsal (
|
|
<emphasis id="70B8EA89FFFDFFA84570FC64FE80EE65" bold="true" box="[275,286,1016,1033]" pageId="8" pageNumber="419">d</emphasis>
|
|
), ventral (
|
|
<emphasis id="70B8EA89FFFDFFA84513FC64FEE4EE65" bold="true" box="[368,378,1016,1033]" pageId="8" pageNumber="419">v</emphasis>
|
|
), and cranial (
|
|
<emphasis id="70B8EA89FFFDFFA8458DFC64FD9EEE65" bold="true" box="[494,512,1016,1033]" pageId="8" pageNumber="419">cr</emphasis>
|
|
) views. Abbreviations: see Figure 4. Scale bar: 2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="4273369BFFFDFFA84412FBD3FEE8EEE6" blockId="8.[113,647,1103,1162]" pageId="8" pageNumber="419">
|
|
last representative of the genus
|
|
<taxonomicName id="85CC4D18FFFDFFA845C9FBD3FD82EE05" authorityName="Szyndlar & Schleich" authorityYear="1993" box="[426,540,1103,1129]" class="Reptilia" family="Boidae" genus="Bavarioboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="419" phylum="Chordata" rank="genus">
|
|
<emphasis id="70B8EA89FFFDFFA845C9FBD3FD82EE05" box="[426,540,1103,1129]" italics="true" pageId="8" pageNumber="419">Bavarioboa</emphasis>
|
|
</taxonomicName>
|
|
in Europe (
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|
<bibRefCitation id="265D4B6AFFFDFFA8441BFBF3FEF4EEE6" author="SZYNDLAR Z. & RAGE J. - C." box="[120,362,1135,1162]" pageId="8" pageNumber="419" refId="ref23455" refString="SZYNDLAR Z. & RAGE J. - C. 2003. - Non-Erycine Booidea from the Oligocene and Miocene of Europe. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, 109 p." type="book" year="2003">Szyndlar & Rage 2003</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |